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<document id="B52EE2C69CD6B8539525B27BD6155F7C" ID-CLB-Dataset="279456" ID-DOI="10.11646/zootaxa.5374.4.1" ID-GBIF-Dataset="6f75d877-d9e0-49de-8ce6-b739b532136a" ID-ISSN="1175-5326" ID-Zenodo-Dep="10158722" ID-ZooBank="EEFF13BD-664A-4FD8-9FED-4E4B21A475E4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1700469135046" checkinUser="plazi" docAuthor="Ahrens, Dirk, Lukic, Daniel, Pham, Phu, Li, Wei &amp; Liu, Wangang" docDate="2023" docId="03B73169BE7E2743D9E4083AFDABFA55" docLanguage="en" docName="zootaxa.5374.4.1.pdf" docOrigin="Zootaxa 5374 (4)" docSource="https://mapress.com/zt/article/download/zootaxa.5374.4.1/52301" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tetraserica gialaiensis Pham &amp; Ahrens 2023, new species" docType="treatment" docVersion="7" lastPageNumber="465" masterDocId="FF8E4911BE73274DD9730A19FFFAFFD1" masterDocTitle="Tetraserica Ahrens, 2004 of continental Southeast Asia: new records, new species and an updated key to species (Coleoptera: Scarabaeidae: Sericinae: Sericini)" masterLastPageNumber="486" masterPageNumber="451" pageNumber="464" updateTime="1732584714017" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7E3DA614380F440EF770F0C5B2389C5B">Tetraserica Ahrens, 2004 of continental Southeast Asia: new records, new species and an updated key to species (Coleoptera: Scarabaeidae: Sericinae: Sericini)</mods:title>
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<mods:namePart id="F19532973EC907CB941D2C304FA10DAD">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="1EA917D3CC84A87A656CBA9019320742" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:affiliation id="2C537005E4C26859F703C1F1DDC546B6">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; ahrens. dirk _ col @ gmx. de; https: // orcid. org / 0000 - 0003 - 3524 - 7153</mods:affiliation>
<mods:nameIdentifier id="AF5FF409368D3764989DE92F4D102180" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:name id="FC9CC979C6CFD36A66C942C3B57F28BD" type="personal">
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<mods:namePart id="62E17751222B4A79A786E34CA8545BD6">Lukic, Daniel</mods:namePart>
<mods:affiliation id="D857E9E1DE00B0FD3C3602619FF88451">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; University of Salzburg, Salzburg, Austria</mods:affiliation>
</mods:name>
<mods:name id="B1814E34BF93536752CFE5423D867F53" type="personal">
<mods:role id="9A1FD06E2DE0147BEDD71124CB1DA4A7">
<mods:roleTerm id="39C137483ED35B5D5212F9D0722C88A4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B4A66A32DD9C5953246AC20D7FD6BCA2">Pham, Phu</mods:namePart>
<mods:affiliation id="310A7CF376DB04AE21CEA898E49C7179">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:name id="D1E9C3D2F1F7EBF2BEFE8B6FA899F6C3" type="personal">
<mods:role id="045A0A96A50B8D8EEEADB64617FFEEDB">
<mods:roleTerm id="B0F547D85F4751D1047F24547DAA54D3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DEFE0D8A9BA59DDE2BB878E55BFB1ED8">Li, Wei</mods:namePart>
<mods:nameIdentifier id="9B3D86A26C5CB7CD30B23560364F5CBE" type="ORCID">0009-0001-0195-7889</mods:nameIdentifier>
<mods:affiliation id="80C0094BB5675D4B6C5D1E9B27BB2FBA">Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97 #, Yanta District, Xi’an 710061, P. R. China. &amp; liweiv @ stumail. nwu. edu. cn; https: // orcid. org / 0009 - 0001 - 0195 - 7889</mods:affiliation>
<mods:nameIdentifier id="15EF2411DB0CFF4D797556B8B5EEDD70" type="email">liweiv@stumail.nwu.edu.cn</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="5031EDB46E5E2DE88C084E1B2E25DFE1">Liu, Wangang</mods:namePart>
<mods:nameIdentifier id="8B1CF5F18F9B4DA87B0D205FACAB8279" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:affiliation id="3D1C91EA4D94C88C1B11C7F6FA8EEDBB">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97 #, Yanta District, Xi’an 710061, P. R. China. &amp; liuwangang @ ieecas. cn; https: // orcid. org / 0000 - 0003 - 4788 - 7967</mods:affiliation>
<mods:nameIdentifier id="83960C6A6CA52ECC68835C7FD99BE455" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:title id="A1D4FFD020C69143652B874C49372726">Zootaxa</mods:title>
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<mods:date id="9E2B30E8DB61391F5908FEACD35F4742">2023</mods:date>
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<mods:classification id="91A0A0D68E57062A4A4C86B474F4AFEF">journal article</mods:classification>
<mods:identifier id="C9B83E76479F91856ACEA0D65EAD66CC" type="CLB-Dataset">279456</mods:identifier>
<mods:identifier id="BB4E1644CB899AA7D695AF83CF1130A8" type="DOI">10.11646/zootaxa.5374.4.1</mods:identifier>
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<treatment id="03B73169BE7E2743D9E4083AFDABFA55" ID-DOI="http://doi.org/10.5281/zenodo.10169410" ID-GBIF-Taxon="219263341" ID-Zenodo-Dep="10169410" LSID="urn:lsid:plazi:treatment:03B73169BE7E2743D9E4083AFDABFA55" httpUri="http://treatment.plazi.org/id/03B73169BE7E2743D9E4083AFDABFA55" lastPageId="14" lastPageNumber="465" pageId="13" pageNumber="464" scope_family="Scarabaeidae" scope_order="Coleoptera" scope_tribe="Sericini">
<subSubSection id="C304D3F4BE7E2740D9E4083AFCEFFDED" box="[151,789,546,573]" pageId="13" pageNumber="464" type="nomenclature">
<paragraph id="8BA1807FBE7E2740D9E4083AFCEFFDED" blockId="13.[151,789,546,609]" box="[151,789,546,573]" pageId="13" pageNumber="464">
<heading id="D0E93713BE7E2740D9E4083AFCEFFDED" bold="true" box="[151,789,546,573]" fontSize="11" level="1" pageId="13" pageNumber="464" reason="1">
<emphasis id="B96A5C6DBE7E2740D9E4083AFCEFFDED" bold="true" box="[151,789,546,573]" pageId="13" pageNumber="464">
<taxonomicName id="4C1EFBFCBE7E2740D9E4083AFD82FDED" ID-CoL="BW5GQ" authority="Pham &amp; Ahrens" authorityName="Pham &amp; Ahrens" authorityYear="2023" box="[151,632,546,573]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="gialaiensis" status="sp. nov.">
<emphasis id="B96A5C6DBE7E2740D9E4083AFE5CFDEC" bold="true" box="[151,422,546,573]" italics="true" pageId="13" pageNumber="464">Tetraserica gialaiensis</emphasis>
Pham &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE7E2740DBF6083AFCEFFDED" box="[645,789,546,573]" pageId="13" pageNumber="464" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<figureCitation id="13259CFABE7E2740D9E4085EFEF5FDB0" box="[151,271,582,609]" captionStart="FIGURE 5" captionStartId="14.[152,255,883,907]" captionTargetBox="[162,1425,193,848]" captionTargetId="figure-23@14.[151,1436,181,859]" captionTargetPageId="14" captionText="FIGURE 5. A–D: Tetraserica gialaiensis Pham &amp; Ahrens, new species (holotype). A: aedeagus, left side lateral view; B: parameres, dorsal view; C: aedeagus, right side lateral view; D: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158734" httpUri="https://zenodo.org/record/10158734/files/figure.png" pageId="13" pageNumber="464">Fig. 5A–D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE7E2740D9E40896FEF7FCC4" pageId="13" pageNumber="464" type="materials_examined">
<paragraph id="8BA1807FBE7E2740D9E40896FEF7FCC4" blockId="13.[151,1437,654,2013]" pageId="13" pageNumber="464">
<emphasis id="B96A5C6DBE7E2740D9E40896FE46FD79" bold="true" box="[151,444,654,681]" pageId="13" pageNumber="464">Type material examined.</emphasis>
<materialsCitation id="3B768A22BE7E2740D8B20896FD39FD1C" ID-GBIF-Occurrence="4442319368" collectingDate="2022-04-28" collectionCode="IEBR" collectorName="P. V. Pham" country="Vietnam" elevation="578" latitude="14.29675" location="Gia Lai" longLatPrecision="1" longitude="108.44949" pageId="13" pageNumber="464" specimenCount="1" stateProvince="Gia Lai" typeStatus="holotype">
<typeStatus id="54A53EDDBE7E2740D8B20896FDD4FD78" box="[449,558,655,681]" pageId="13" pageNumber="464" type="holotype">Holotype</typeStatus>
: ♂ “[
<collectingCountry id="F309C0EFBE7E2740DB120896FD32FD78" box="[609,712,655,681]" name="Vietnam" pageId="13" pageNumber="464">Vietnam</collectingCountry>
]
<collectingRegion id="49DA4E9DBE7E2740DBA00896FCDCFD78" box="[723,806,655,681]" country="Vietnam" name="Gia Lai" pageId="13" pageNumber="464">Gia Lai</collectingRegion>
Pro., Kon Ka Kinh NP,
<date id="FFA0A6BFBE7E2740DD580896FB5EFD78" box="[1067,1188,655,681]" pageId="13" pageNumber="464" value="2022-04-28">
<collectingDate id="EFE45F57BE7E2740DD580896FB5EFD78" box="[1067,1188,655,681]" pageId="13" pageNumber="464" value="2022-04-28">28.iv.2022</collectingDate>
</date>
;
<collectingMethod id="525FF868BE7E2740DDDD0896FAEFFD78" box="[1198,1301,655,681]" pageId="13" pageNumber="464">light trap</collectingMethod>
;
<geoCoordinate id="EE2AE6B8BE7E2740DC6C0897FA66FD78" box="[1311,1436,654,681]" degrees="14,29675" direction="north" orientation="latitude" pageId="13" pageNumber="464" precision="1" value="14.29675">14,29675N</geoCoordinate>
<geoCoordinate id="EE2AE6B8BE7E2740D9E408AAFED8FD1C" box="[151,290,691,717]" degrees="108,44949" direction="east" orientation="longitude" pageId="13" pageNumber="464" precision="1" value="108.44949">108,44949E</geoCoordinate>
;
<elevation id="0033674CBE7E2740D85E08ABFE55FD1C" box="[301,431,690,717]" metricMagnitude="2" metricUnit="m" metricValue="5.779999999999999" pageId="13" pageNumber="464" unit="m" value="578.0">
<quantity id="4CE62D9ABE7E2740D85E08ABFE89FD1C" box="[301,371,690,717]" metricMagnitude="2" metricUnit="m" metricValue="5.779999999999999" pageId="13" pageNumber="464" unit="m" value="578.0">578 m</quantity>
a.s.l.
</elevation>
;
<collectorName id="26EBE5A9BE7E2740D8C808AAFDD6FD1C" box="[443,556,691,717]" pageId="13" pageNumber="464">P.V. Pham</collectorName>
leg.” (
<collectionCode id="ED0F18BABE7E2740DB0608AAFD41FD1C" box="[629,699,691,717]" pageId="13" pageNumber="464">IEBR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B768A22BE7E2740DBBE08AAFEF3FCC4" ID-GBIF-Occurrence="4442319366" collectingDate="2003-05-13" collectionCode="NHMW" collectorName="Boukal" country="Myanmar" elevation="350" latitude="22.31855" location="Division" longLatPrecision="1" longitude="94.4753" pageId="13" pageNumber="464" specimenCount="1" stateProvince="Sagaing" typeStatus="paratype">
<typeStatus id="54A53EDDBE7E2740DBBE08AAFCC9FD1C" box="[717,819,691,717]" pageId="13" pageNumber="464" type="paratype">Paratype</typeStatus>
: 1 ♂ “
<collectingCountry id="F309C0EFBE7E2740DA0508AAFC10FD1C" box="[886,1002,691,717]" name="Myanmar" pageId="13" pageNumber="464">Myanmar</collectingCountry>
:
<collectingRegion id="49DA4E9DBE7E2740DA8008AAFBB7FD1C" box="[1011,1101,691,717]" country="Myanmar" name="Sagaing" pageId="13" pageNumber="464">Sagaing</collectingRegion>
<location id="8EC1D6A4BE7E2740DD2008AAFB49FD1C" LSID="urn:lsid:plazi:treatment:03B73169BE7E2743D9E4083AFDABFA55:8EC1D6A4BE7E2740DD2008AAFB49FD1C" box="[1107,1203,691,717]" country="Myanmar" latitude="22.31855" longLatPrecision="1" longitude="94.4753" name="Division" pageId="13" pageNumber="464" stateProvince="Sagaing">Division</location>
<taxonomicName id="4C1EFBFCBE7E2740DDCB08AAFF40FD20" authority="Kathapa NP" authorityName="Kathapa NP" pageId="13" pageNumber="464" phylum="Alaungdaw" rank="phylum">Alaungdaw Kathapa NP</taxonomicName>
<geoCoordinate id="EE2AE6B8BE7E2740D9B308CEFEABFD20" box="[192,337,727,753]" degrees="22" direction="north" minutes="19.113" orientation="latitude" pageId="13" pageNumber="464" precision="1" value="22.31855">22°19.113’N</geoCoordinate>
<geoCoordinate id="EE2AE6B8BE7E2740D82408CFFE1CFD20" box="[343,486,726,753]" degrees="94" direction="east" minutes="28.518" orientation="longitude" pageId="13" pageNumber="464" precision="1" value="94.4753">94°28.518’E</geoCoordinate>
3.-
<date id="FFA0A6BFBE7E2740DB7708CFFD87FD20" box="[516,637,726,753]" pageId="13" pageNumber="464" value="2003-05-13">
<collectingDate id="EFE45F57BE7E2740DB7708CFFD87FD20" box="[516,637,726,753]" pageId="13" pageNumber="464" value="2003-05-13">13.5.2003</collectingDate>
</date>
, ca.
<quantity id="4CE62D9ABE7E2740DBDD08CFFD0BFD20" box="[686,753,726,753]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="464" unit="m" value="350.0">
<elevation id="0033674CBE7E2740DBDD08CFFD0BFD20" box="[686,753,726,753]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="464" unit="m" value="350.0">350m</elevation>
</quantity>
, light leg.
<collectorName id="26EBE5A9BE7E2740DA1608CEFC4CFD20" box="[869,950,727,753]" pageId="13" pageNumber="464">Boukal</collectorName>
&amp; Schillhammer/ 1319 Asia Sericini spec.” (
<collectionCode id="ED0F18BABE7E2740D9EC08E2FEF8FCC4" box="[159,258,763,789]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="464" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE7E2740D9B40907FCA1F80C" pageId="13" pageNumber="464" type="description">
<paragraph id="8BA1807FBE7E2740D9B40907FCF0FC8C" blockId="13.[151,1437,654,2013]" pageId="13" pageNumber="464">
<emphasis id="B96A5C6DBE7E2740D9B40907FDF7FCE8" bold="true" box="[199,525,798,825]" pageId="13" pageNumber="464">
Description of the
<typeStatus id="54A53EDDBE7E2740D8D30906FDF3FCE8" box="[416,521,799,825]" pageId="13" pageNumber="464" type="holotype">holotype</typeStatus>
.
</emphasis>
Length:
<quantity id="4CE62D9ABE7E2740DB030906FD3DFCE8" box="[624,711,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="13" pageNumber="464" unit="mm" value="8.4">8.4 mm</quantity>
, length of elytra:
<quantity id="4CE62D9ABE7E2740DAF40906FC24FCE8" box="[903,990,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="13" pageNumber="464" unit="mm" value="6.8">6.8 mm</quantity>
, width:
<quantity id="4CE62D9ABE7E2740DD450907FB77FCE8" box="[1078,1165,798,825]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="13" pageNumber="464" unit="mm" value="5.1">5.1 mm</quantity>
. Body oval, dark brown, labroclypeus and dorsal surface glabrous, without setae.
</paragraph>
<paragraph id="8BA1807FBE7E2740D9B4097EFC0EFB70" blockId="13.[151,1437,654,2013]" pageId="13" pageNumber="464">
Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins weakly convex, anterior margin shallowly emarginate medially, anterior angles strongly convex, lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed; surface moderately convex medially, coarsely and moderately densely punctate, without setae; frontoclypeal suture moderately impressed and moderately angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus moderate and triangular (1/2.5 of ocular diameter), weakly punctate, with one terminal seta. Frons weakly shiny and punctate, bearing only a few erect setae beside eyes. Eyes moderately small, ratio of diameter/ interocular width: 0.52. Antenna yellowish brown, with ten antennomeres, club in male with four antennomeres, as long as remaining antennomeres combined, first joint of club slightly shorter than club.
<collectingRegion id="49DA4E9DBE7E2740D87E0E9EFE96FB70" box="[269,364,1159,1185]" country="Myanmar" name="Chin" pageId="13" pageNumber="464">Mentum</collectingRegion>
anteriorly weakly elevated and weakly convex in middle.
</paragraph>
<paragraph id="8BA1807FBE7E2740D9B40EB2FA9BFAE0" blockId="13.[151,1437,654,2013]" pageId="13" pageNumber="464">Pronotum transverse, widest just before base, lateral margins moderately convex and evenly convergent anteriorly; anterior angles distinctly produced, posterior angles strongly convex. Anterior margin almost straight medially, with a fine, complete marginal line. Surface densely and finely punctate, with minute setae in punctures; lateral margins sparsely setose. Hypomeron not carinate. Scutellum triangular, with fine and dense punctures.</paragraph>
<paragraph id="8BA1807FBE7E2740D9B40F22FDA3FA10" blockId="13.[151,1437,654,2013]" pageId="13" pageNumber="464">Elytra oblong, widest at posterior third, striae moderately impressed, finely and densely punctate; intervals moderately convex, with fine, dense punctures, glabrous, except minute setae in punctures; epipleural edge robust, ending at convex external apical angle of elytra, epipleura robustly sparsely setose, apical border membraneous, with a fine rim of short microtrichomes.</paragraph>
<paragraph id="8BA1807FBE7E2740D9B40FD2FC63F9A4" blockId="13.[151,1437,654,2013]" pageId="13" pageNumber="464">Ventral surface dull, reddish brown, thorax moderate and dense punctures, sparsely setose, metasternum sparsely covered with fine, short, or minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures each bearing a robust seta. Mesosternum between mesocoxae wider than mesofemur. Ratio of length of metepisternum/ metacoxa: 1/2.0. Pygidium moderately convex and dull, coarsely densely punctate, without smooth midline.</paragraph>
<paragraph id="8BA1807FBE7E2740D9B40C66FBFEF868" blockId="13.[151,1437,654,2013]" pageId="13" pageNumber="464">Legs moderately wide; metafemur moderately shiny, anterior margin acute, lacking an adjacent serrated line; ventral posterior margin nearly straight and smooth, slightly widened in apical half; dorsal posterior margin smooth, without setae basally. Metatibia moderately short, widest at middle, ratio width/ length: 1/ 3.3; dorsal margin longitudinally convex, with two groups of spines, basal one at about first third, apical one at nearly two thirds of metatibial length; lateral face smooth, sparsely finely punctate; ventral margin finely serrate, with four robust setae; medial face smooth; apex interiorly near tarsal articulation slightly concave. Tarsomeres in distal half with fine, dense setae ventrally, dorsally smooth; metatarsomeres ventrally with a strong serrated ridge and glabrous, first metatarsomere shorter than following two tarsomeres combined and longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws sharp at apex.</paragraph>
<paragraph id="8BA1807FBE7E2740D9B40DDAFCA1F80C" blockId="13.[151,1437,654,2013]" box="[199,859,1986,2013]" pageId="13" pageNumber="464">
Aedeagus.
<figureCitation id="13259CFABE7E2740D8370DDAFE47F80C" box="[324,445,1986,2013]" captionStart="FIGURE 5" captionStartId="14.[152,255,883,907]" captionTargetBox="[162,1425,193,848]" captionTargetId="figure-23@14.[151,1436,181,859]" captionTargetPageId="14" captionText="FIGURE 5. A–D: Tetraserica gialaiensis Pham &amp; Ahrens, new species (holotype). A: aedeagus, left side lateral view; B: parameres, dorsal view; C: aedeagus, right side lateral view; D: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158734" httpUri="https://zenodo.org/record/10158734/files/figure.png" pageId="13" pageNumber="464">Fig. 5A–C</figureCitation>
. Habitus:
<figureCitation id="13259CFABE7E2740DB5D0DDAFD7CF80C" box="[558,646,1986,2013]" captionStart="FIGURE 5" captionStartId="14.[152,255,883,907]" captionTargetBox="[162,1425,193,848]" captionTargetId="figure-23@14.[151,1436,181,859]" captionTargetPageId="14" captionText="FIGURE 5. A–D: Tetraserica gialaiensis Pham &amp; Ahrens, new species (holotype). A: aedeagus, left side lateral view; B: parameres, dorsal view; C: aedeagus, right side lateral view; D: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158734" httpUri="https://zenodo.org/record/10158734/files/figure.png" pageId="13" pageNumber="464">Fig. 5D</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<caption id="DF61D0F7BE7D2743D9EB096AFB12FC7E" ID-DOI="http://doi.org/10.5281/zenodo.10158734" ID-Zenodo-Dep="10158734" httpUri="https://zenodo.org/record/10158734/files/figure.png" pageId="14" pageNumber="465" startId="14.[152,255,883,907]" targetBox="[162,1425,193,848]" targetPageId="14" targetType="figure">
<paragraph id="8BA1807FBE7D2743D9EB096AFB12FC7E" blockId="14.[151,1437,883,943]" pageId="14" pageNumber="465">
<emphasis id="B96A5C6DBE7D2743D9EB096AFEA7FC5A" bold="true" box="[152,349,883,907]" pageId="14" pageNumber="465">FIGURE 5. A–D:</emphasis>
<taxonomicName id="4C1EFBFCBE7D2743D815096DFCF8FC5A" authority="Pham &amp; Ahrens" authorityName="Pham &amp; Ahrens" authorityYear="2023" box="[358,770,883,907]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="gialaiensis" status="sp. nov.">
<emphasis id="B96A5C6DBE7D2743D815096DFDB5FC5A" box="[358,591,883,907]" italics="true" pageId="14" pageNumber="465">Tetraserica gialaiensis</emphasis>
Pham &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE7D2743DA7C096DFC71FC5A" box="[783,907,883,907]" pageId="14" pageNumber="465" rank="species">new species</taxonomicNameLabel>
(holotype).
<emphasis id="B96A5C6DBE7D2743DD7F096AFBDDFC5A" bold="true" box="[1036,1063,883,907]" pageId="14" pageNumber="465">A:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B96A5C6DBE7D2743DCF0096AFA67FC5A" bold="true" box="[1411,1437,883,907]" pageId="14" pageNumber="465">B:</emphasis>
parameres, dorsal view;
<emphasis id="B96A5C6DBE7D2743D8FC098EFE50FC7E" bold="true" box="[399,426,919,943]" pageId="14" pageNumber="465">C:</emphasis>
aedeagus, right side lateral view;
<emphasis id="B96A5C6DBE7D2743DA70098EFCE7FC7E" bold="true" box="[771,797,919,943]" pageId="14" pageNumber="465">D:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<subSubSection id="C304D3F4BE7D2743D9B40E18FE38FAC6" pageId="14" pageNumber="465" type="diagnosis">
<paragraph id="8BA1807FBE7D2743D9B40E18FE38FAC6" blockId="14.[151,1437,1025,1412]" pageId="14" pageNumber="465">
<emphasis id="B96A5C6DBE7D2743D9B40E18FEB8FBCA" bold="true" box="[199,322,1025,1051]" pageId="14" pageNumber="465">Diagnosis.</emphasis>
<taxonomicName id="4C1EFBFCBE7D2743D83E0E1BFCE0FBCA" authority="Pham &amp; Ahrens" authorityName="Pham &amp; Ahrens" authorityYear="2023" box="[333,794,1025,1052]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="gialaiensis" status="sp. nov.">
<emphasis id="B96A5C6DBE7D2743D83E0E1BFDB5FBCA" box="[333,591,1025,1051]" italics="true" pageId="14" pageNumber="465">Tetraserica gialaiensis</emphasis>
Pham &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE7D2743DA5A0E1BFC4EFBCA" box="[809,948,1025,1052]" pageId="14" pageNumber="465" rank="species">new species</taxonomicNameLabel>
differs from the similar
<taxonomicName id="4C1EFBFCBE7D2743DDA80E1BFDE0FB91" authority="Liu, Fabrizi, Bai, Yang &amp; Ahrens, 2014" authorityName="Liu, Fabrizi, Bai, Yang &amp; Ahrens" authorityYear="2014" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="ruiliensis">
<emphasis id="B96A5C6DBE7D2743DDA80E1BFA99FBCA" box="[1243,1379,1025,1051]" italics="true" pageId="14" pageNumber="465">T. ruiliensis</emphasis>
<bibRefCitation id="EF8FFD8EBE7D2743DC1D0E18FDE0FB91" author="Liu, W. &amp; Fabrizi, S. &amp; Bai, M. &amp; Yang, X. - K. &amp; Ahrens, D." pageId="14" pageNumber="465" pagination="83 - 121" refId="ref44587" refString="Liu, W., Fabrizi, S., Bai, M., Yang, X. - K. &amp; Ahrens, D. (2014) A taxonomic review of the Tetraserica Ahrens, 2004 species of China (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 448, 83 - 121. https: // doi. org / 10.3897 / zookeys. 443.8429" type="journal article" year="2014">Liu, Fabrizi, Bai, Yang &amp; Ahrens, 2014</bibRefCitation>
</taxonomicName>
by the distinctly shorter (compared to the length of phallobase) and more robust left paramere (lateral view), which is also at apex curved more strongly, and by the longer apical portion (distal to the strong bent at middle) which is subequal in length to the basal portion (shorter than the basal portion in
<taxonomicName id="4C1EFBFCBE7D2743D9E40E8BFEE7FB7A" authorityName="Liu, Fabrizi, Bai, Yang &amp; Ahrens" authorityYear="2014" box="[151,285,1169,1195]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="ruiliensis">
<emphasis id="B96A5C6DBE7D2743D9E40E8BFEE7FB7A" box="[151,285,1169,1195]" italics="true" pageId="14" pageNumber="465">T. ruiliensis</emphasis>
</taxonomicName>
). From
<taxonomicName id="4C1EFBFCBE7D2743D80D0E8BFBD0FB7D" authority="Fabrizi, Dalstein &amp; Ahrens, 2019" authorityName="Fabrizi, Dalstein &amp; Ahrens" authorityYear="2019" box="[382,1066,1169,1196]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="semiruiliensis">
<emphasis id="B96A5C6DBE7D2743D80D0E8BFD58FB7A" box="[382,674,1169,1195]" italics="true" pageId="14" pageNumber="465">Tetraserica semiruiliensis</emphasis>
<bibRefCitation id="EF8FFD8EBE7D2743DBDF0E88FBD0FB7D" author="Fabrizi, S. &amp; Dalstein, V. &amp; Ahrens, D." box="[684,1066,1169,1196]" pageId="14" pageNumber="465" pagination="1 - 155" refId="ref44465" refString="Fabrizi, S., Dalstein, V. &amp; Ahrens, D. (2019) A monograph on the genus Tetraserica from the Indochinese region (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 837, 1 - 155. https: // doi. org / 10.3897 / zookeys. 837.32057" type="journal article" year="2019">Fabrizi, Dalstein &amp; Ahrens, 2019</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1EFBFCBE7D2743DD150E8BFE4EFB01" authority="Fabrizi, Dalstein &amp; Ahrens, 2019" authorityName="Fabrizi, Dalstein &amp; Ahrens" authorityYear="2019" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="pseudoruiliensis">
<emphasis id="B96A5C6DBE7D2743DD150E8BFAC7FB7A" box="[1126,1341,1169,1195]" italics="true" pageId="14" pageNumber="465">T. pseudoruiliensis</emphasis>
<bibRefCitation id="EF8FFD8EBE7D2743DC350E88FE4EFB01" author="Fabrizi, S. &amp; Dalstein, V. &amp; Ahrens, D." pageId="14" pageNumber="465" pagination="1 - 155" refId="ref44465" refString="Fabrizi, S., Dalstein, V. &amp; Ahrens, D. (2019) A monograph on the genus Tetraserica from the Indochinese region (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 837, 1 - 155. https: // doi. org / 10.3897 / zookeys. 837.32057" type="journal article" year="2019">Fabrizi, Dalstein &amp; Ahrens, 2019</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1EFBFCBE7D2743D8B20EAFFCE2FB1E" authority="Pham &amp; Ahrens" authorityName="Pham &amp; Ahrens" authorityYear="2023" box="[449,792,1205,1232]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="gialaiensis" status="sp. nov.">
<emphasis id="B96A5C6DBE7D2743D8B20EAFFDACFB1E" box="[449,598,1205,1231]" italics="true" pageId="14" pageNumber="465">T. gialaiensis</emphasis>
Pham &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE7D2743DA570EAFFC56FB1E" box="[804,940,1205,1232]" pageId="14" pageNumber="465" rank="species">new species</taxonomicNameLabel>
differs by the strongly bent right paramere, whose apical portion (distal to the strong bent at middle) is subequal in length to the basal portion (basal to the strong bent) (lateral view).
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE7D2743D9B40F38FB9EFAED" box="[199,1124,1313,1340]" pageId="14" pageNumber="465" type="description">
<paragraph id="8BA1807FBE7D2743D9B40F38FB9EFAED" blockId="14.[151,1437,1025,1412]" box="[199,1124,1313,1340]" pageId="14" pageNumber="465">
<emphasis id="B96A5C6DBE7D2743D9B40F38FEC5FAEA" bold="true" box="[199,319,1313,1339]" pageId="14" pageNumber="465">Variation.</emphasis>
Length:
<quantity id="4CE62D9ABE7D2743D8D70F38FDD7FAED" box="[420,557,1313,1340]" metricMagnitude="-3" metricUnit="m" metricValue="8.45" metricValueMax="8.5" metricValueMin="8.4" pageId="14" pageNumber="465" unit="mm" value="8.45" valueMax="8.5" valueMin="8.4">8.4–8.5 mm</quantity>
, length of elytra:
<quantity id="4CE62D9ABE7D2743DB810F38FC81FAED" box="[754,891,1313,1340]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="6.9" metricValueMin="6.8" pageId="14" pageNumber="465" unit="mm" value="6.85" valueMax="6.9" valueMin="6.8">6.8–6.9 mm</quantity>
, width:
<quantity id="4CE62D9ABE7D2743DAA60F38FBA5FAED" box="[981,1119,1313,1340]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.2" metricValueMin="5.1" pageId="14" pageNumber="465" unit="mm" value="5.15" valueMax="5.2" valueMin="5.1">5.1–5.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE7D2743D9B40F5FFDABFA55" pageId="14" pageNumber="465" type="etymology">
<paragraph id="8BA1807FBE7D2743D9B40F5FFDABFA55" blockId="14.[151,1437,1025,1412]" pageId="14" pageNumber="465">
<emphasis id="B96A5C6DBE7D2743D9B40F5FFEB4FAB1" bold="true" box="[199,334,1350,1376]" pageId="14" pageNumber="465">Etymology.</emphasis>
This species is named after its occurrence in the
<collectingRegion id="49DA4E9DBE7D2743DA1B0F5CFBDDFA8E" box="[872,1063,1349,1375]" country="Vietnam" name="Gia Lai" pageId="14" pageNumber="465">Gia Lai Province</collectingRegion>
, where the
<typeStatus id="54A53EDDBE7D2743DDD00F5FFB29FAB1" box="[1187,1235,1350,1376]" pageId="14" pageNumber="465">type</typeStatus>
locality is located (adjective in nominative singular case).
</paragraph>
</subSubSection>
</treatment>
</document>
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