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<mods:title id="541B8349EBCBF2D565F3310E9F071A42">A taxonomic review of the newly recorded genus Cyanotiphia Cameron, 1907 (Hymenoptera, Tiphiidae, Tiphiinae) from China, with a key to all known species</mods:title>
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<mods:namePart id="6128B96DAA993D05F2C572B72AB7BDB3">Feng, Zi-Yan</mods:namePart>
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<mods:namePart id="AEA295CF8EA7EFB8A62A4A1DDCC585AF">Chen, Bin</mods:namePart>
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<mods:affiliation id="DDB87A312983636919956BFE8C31ACBE">Chongqing Key Laboratory of Vector Control and Utilization; Institute of Entomology and Molecular Biology, Chongqing Normal University, Chongqing 401331, China</mods:affiliation>
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<mods:affiliation id="6D9522493EF4E21D54A3982B1D741687">Chongqing Key Laboratory of Vector Control and Utilization; Institute of Entomology and Molecular Biology, Chongqing Normal University, Chongqing 401331, China</mods:affiliation>
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. 2022: 88
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species
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.
<taxonomicName id="4C021C9CA02EFFDCFEF593E6B330FC4B" authority="Cameron, 1907: 287" authorityName="Cameron" authorityPageNumber="287" authorityYear="1907" box="[313,807,932,958]" class="Insecta" family="Vespidae" genus="Cyanotiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="species" species="ruficauda">
<emphasis id="B976BB0DA02EFFDCFEF593E6B22FFC4B" box="[313,568,932,958]" italics="true" pageId="1" pageNumber="593">Cyanotiphia ruficauda</emphasis>
<bibRefCitation id="EF931AEEA02EFFDCFDF393E6B330FC4B" author="Cameron, P." box="[575,807,932,958]" pageId="1" pageNumber="593" pagination="287 - 288" refId="ref4392" refString="Cameron, P. (1907) On the bornean Tiphiidae, including a new genus. The Entomologist, 40, 287-288." type="journal article" year="1907">Cameron, 1907: 287</bibRefCitation>
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, 288, by original designation and monotypy.
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<emphasis id="B976BB0DA02EFFDCFF0B938AB12DFC17" bold="true" box="[199,314,968,994]" pageId="1" pageNumber="593">Diagnosis</emphasis>
. Dorsal and lateral pronotum with dense punctures; disc of propodeum laterally with at least one complete, longitudinal carina running from anterior to posterior margin of this area and posteriorly connected with transverse carina (
<figureCitation id="13397B9AA02EFFDCFE939452B1B0FBDF" box="[351,423,1040,1066]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 4</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFE7D9452B1C7FBDF" box="[433,464,1040,1066]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">12</figureCitation>
); posterior surface of propodeum with radial wrinkles (
<figureCitation id="13397B9AA02EFFDCFBE19452B463FBDF" box="[1069,1140,1040,1066]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 4</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFBB39452B489FBDF" box="[1151,1182,1040,1066]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">12</figureCitation>
); vein
<emphasis id="B976BB0DA02EFFDCFB2A9452B504FBDF" box="[1254,1299,1040,1066]" italics="true" pageId="1" pageNumber="593">2RS</emphasis>
incomplete, not reaching
<emphasis id="B976BB0DA02EFFDCFEE79476B163FBBB" box="[299,372,1076,1102]" italics="true" pageId="1" pageNumber="593">RS+M</emphasis>
(
<figureCitation id="13397B9AA02EFFDCFE489476B1DBFBBB" box="[388,460,1076,1102]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 1</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFE179476B1FCFBBB" box="[475,491,1076,1102]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">9</figureCitation>
); tegula distinctly large, slightly shorter than mesonotum (
<figureCitation id="13397B9AA02EFFDCFB4C9476B4DFFBBB" box="[1152,1224,1076,1102]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 1</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFB1A9476B4F1FBBB" box="[1238,1254,1076,1102]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">9</figureCitation>
); mid tibia with two spurs apically; anterior T1 separated by distinct transverse carina followed by short longitudinal wrinkles (
<figureCitation id="13397B9AA02EFFDCFAA6941AB0B2FB63" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 1</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFF79943EB0D2FB63" box="[181,197,1148,1174]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">9</figureCitation>
); T1 densely punctate, with posterior margin impunctate (
<figureCitation id="13397B9AA02EFFDCFC93943EB3BFFB63" box="[863,936,1148,1174]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 1</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFC74943EB3DFFB63" box="[952,968,1148,1174]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">9</figureCitation>
); S1 laterally with strongly raised carina (
<figureCitation id="13397B9AA02EFFDCFF5394E2B0FDFB4E" box="[159,234,1184,1211]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 5</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFF3494E2B100FB4F" box="[248,279,1184,1210]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">14</figureCitation>
); in female, mid and hind femora cylindrical, not flat (
<bibRefCitation id="EF931AEEA02EFFDCFC4194E2B407FB4F" author="Allen, H. W." box="[909,1040,1184,1210]" pageId="1" pageNumber="593" pagination="353 - 438" refId="ref4228" refString="Allen, H. W. (1969) Redescriptions of types of Tiphiinae from Asia, Africa, Oceania in the British Museum (NH) and at Oxford University. Transactions of the American Entomological Society, 95, 353-438." type="journal article" year="1969">Allen 1969</bibRefCitation>
;
<bibRefCitation id="EF931AEEA02EFFDCFBD294E2B4DEFB4F" author="Cameron, P." box="[1054,1225,1184,1210]" pageId="1" pageNumber="593" pagination="287 - 288" refId="ref4392" refString="Cameron, P. (1907) On the bornean Tiphiidae, including a new genus. The Entomologist, 40, 287-288." type="journal article" year="1907">Cameron 1907</bibRefCitation>
;
<bibRefCitation id="EF931AEEA02EFFDCFB1B94E2B58FFB4F" author="Saini, J. &amp; Gupta, D. &amp; Gupta, K. C. S. K." box="[1239,1432,1184,1210]" pageId="1" pageNumber="593" pagination="87 - 92" refId="ref4546" refString="Saini, J., Gupta, D. &amp; Gupta, K. C. S. K. (2022) Taxonomic notes on the little-known genus Cyanotiphia Cameron, 1907 (Hymenoptera: Tiphidae: Tiphinae). Zootaxa, 5154 (1), 87-92. https://doi.org/10.11646/zootaxa.5154.1.6" type="journal article" year="2022">
Saini
<emphasis id="B976BB0DA02EFFDCFAD594E3B543FB4F" box="[1305,1364,1184,1210]" italics="true" pageId="1" pageNumber="593">et al.</emphasis>
2022
</bibRefCitation>
;
<bibRefCitation id="EF931AEEA02EFFDCFF5B9486B125FB2B" author="Tsuneki, K." box="[151,306,1220,1246]" pageId="1" pageNumber="593" pagination="1 - 8" refId="ref4671" refString="Tsuneki, K. (1986) A contribution to the knowledge of the Taiwanese Tiphiidae (Hymenoptera, Tiphiidae). Special Publications of the Japan Hymenopterists Association, 33, 1-8." type="journal article" year="1986">Tsuneki 1986</bibRefCitation>
).
</paragraph>
<paragraph id="8BBD671FA02EFFDCFF0B94AAB3EBFAD3" blockId="1.[151,1437,932,1786]" pageId="1" pageNumber="593">
<emphasis id="B976BB0DA02EFFDCFF0B94AAB186FAF7" bold="true" box="[199,401,1256,1282]" pageId="1" pageNumber="593">Species included</emphasis>
. There are three species in this genus:
<taxonomicName id="4C021C9CA02EFFDCFCA694AAB516FAF7" authority="Feng, Chen &amp; Li" authorityName="Feng, Chen &amp; Li" authorityYear="2025" box="[874,1281,1256,1283]" class="Insecta" family="Vespidae" genus="Cyanotiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="species" species="breviclypeola" status="sp. nov.">
<emphasis id="B976BB0DA02EFFDCFCA694AAB431FAF7" box="[874,1062,1256,1282]" italics="true" pageId="1" pageNumber="593">C. breviclypeola</emphasis>
Feng, Chen &amp; Li
</taxonomicName>
,
<emphasis id="B976BB0DA02EFFDCFADD94AAB567FAF7" bold="true" box="[1297,1392,1256,1282]" pageId="1" pageNumber="593">
<taxonomicNameLabel id="A2450676A02EFFDCFADD94AAB567FAF7" box="[1297,1392,1256,1282]" pageId="1" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C021C9CA02EFFDCFA4F94AAB1FDFAD3" authority="(Tsuneki, 1986)" authorityName="Feng &amp; Chen &amp; Li" authorityYear="2025" baseAuthorityName="Tsuneki" baseAuthorityYear="1986" class="Insecta" family="Tiphiidae" genus="Tiphia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="species" species="fenchihuensis" status="comb. nov.">
<emphasis id="B976BB0DA02EFFDCFA4F94AAB138FAD3" italics="true" pageId="1" pageNumber="593">C. fenchihuensis</emphasis>
(
<bibRefCitation id="EF931AEEA02EFFDCFEF2954EB1F5FAD3" author="Tsuneki, K." box="[318,482,1292,1318]" pageId="1" pageNumber="593" pagination="1 - 8" refId="ref4671" refString="Tsuneki, K. (1986) A contribution to the knowledge of the Taiwanese Tiphiidae (Hymenoptera, Tiphiidae). Special Publications of the Japan Hymenopterists Association, 33, 1-8." type="journal article" year="1986">Tsuneki, 1986</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B976BB0DA02EFFDCFE39954EB261FAD3" bold="true" box="[501,630,1292,1318]" pageId="1" pageNumber="593">
<taxonomicNameLabel id="A2450676A02EFFDCFE39954EB261FAD3" box="[501,630,1292,1318]" pageId="1" pageNumber="593" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C021C9CA02EFFDCFD7F954EB3E0FAD3" authority="Cameron, 1907" authorityName="Cameron" authorityYear="1907" box="[691,1015,1292,1318]" class="Insecta" family="Vespidae" genus="Cyanotiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="species" species="ruficauda">
<emphasis id="B976BB0DA02EFFDCFD7F954EB356FAD3" box="[691,833,1292,1318]" italics="true" pageId="1" pageNumber="593">C. ruficauda</emphasis>
<bibRefCitation id="EF931AEEA02EFFDCFC84954EB3E0FAD3" author="Cameron, P." box="[840,1015,1292,1318]" pageId="1" pageNumber="593" pagination="287 - 288" refId="ref4392" refString="Cameron, P. (1907) On the bornean Tiphiidae, including a new genus. The Entomologist, 40, 287-288." type="journal article" year="1907">Cameron, 1907</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3183494A02EFFDCFF0B9572B173FA9B" pageId="1" pageNumber="593" type="distribution">
<paragraph id="8BBD671FA02EFFDCFF0B9572B173FA9B" blockId="1.[151,1437,932,1786]" pageId="1" pageNumber="593">
<emphasis id="B976BB0DA02EFFDCFF0B9572B14EFABF" bold="true" box="[199,345,1328,1354]" pageId="1" pageNumber="593">Distribution</emphasis>
.
<collectingCountry id="F315278FA02EFFDCFEA09572B1B8FABF" box="[364,431,1328,1354]" name="China" pageId="1" pageNumber="593">China</collectingCountry>
(
<collectingCountry id="F315278FA02EFFDCFE0D9572B20FFABF" box="[449,536,1328,1354]" name="Taiwan" pageId="1" pageNumber="593">Taiwan</collectingCountry>
, *Guangdong, *Chongqing),
<collectingCountry id="F315278FA02EFFDCFCBC9572B3CAFABF" box="[880,989,1328,1354]" name="Indonesia" pageId="1" pageNumber="593">Indonesia</collectingCountry>
(
<collectingRegion id="49C6A9FDA02EFFDCFC239572B489FABF" box="[1007,1182,1328,1354]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="593">North Sumatra</collectingRegion>
,
<collectingRegion id="49C6A9FDA02EFFDCFB619573B509FABF" box="[1197,1310,1328,1355]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="593">East Java</collectingRegion>
),
<collectingCountry id="F315278FA02EFFDCFAFA9572B58AFABF" box="[1334,1437,1328,1354]" name="Malaysia" pageId="1" pageNumber="593">Malaysia</collectingCountry>
(
<collectingRegion id="49C6A9FDA02EFFDCFF539516B0FFFA9B" box="[159,232,1364,1390]" country="Malaysia" name="Sabah" pageId="1" pageNumber="593">Sabah</collectingRegion>
,
<collectingCountry id="F315278FA02EFFDCFF3F9516B14EFA9B" box="[243,345,1364,1390]" name="Malaysia" pageId="1" pageNumber="593">Sarawak</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3183494A02EFFDCFF0B953AB1FCF90F" pageId="1" pageNumber="593" type="discussion">
<paragraph id="8BBD671FA02EFFDCFF0B953AB1FCF90F" blockId="1.[151,1437,932,1786]" pageId="1" pageNumber="593">
<emphasis id="B976BB0DA02EFFDCFF0B953AB123FA67" bold="true" box="[199,308,1400,1426]" pageId="1" pageNumber="593">Remarks</emphasis>
. The three genera,
<taxonomicName id="4C021C9CA02EFFDCFDC7953AB35BFA67" authority="Cameron, 1907" authorityName="Cameron" authorityYear="1907" box="[523,844,1400,1426]" class="Insecta" family="Vespidae" genus="Cyanotiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B976BB0DA02EFFDCFDC7953AB282FA67" box="[523,661,1400,1426]" italics="true" pageId="1" pageNumber="593">Cyanotiphia</emphasis>
<bibRefCitation id="EF931AEEA02EFFDCFD50953AB35BFA67" author="Cameron, P." box="[668,844,1400,1426]" pageId="1" pageNumber="593" pagination="287 - 288" refId="ref4392" refString="Cameron, P. (1907) On the bornean Tiphiidae, including a new genus. The Entomologist, 40, 287-288." type="journal article" year="1907">Cameron, 1907</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C021C9CA02EFFDCFC9B953AB441FA66" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[855,1110,1400,1427]" class="Insecta" family="Tiphiidae" genus="Tiphia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B976BB0DA02EFFDCFC9B953AB388FA67" box="[855,927,1400,1426]" italics="true" pageId="1" pageNumber="593">Tiphia</emphasis>
<bibRefCitation id="EF931AEEA02EFFDCFC69953AB441FA66" author="Fabricius, J. C." box="[933,1110,1400,1427]" pageId="1" pageNumber="593" refId="ref4418" refString="Fabricius, J. C. (1775) Systema Entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Officina Libraria Kortii, Flensburgi et Lipsiae, [32] + 832 pp. https://doi.org/10.5962/bhl.title.36510" type="book" year="1775">Fabricius, 1775</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C021C9CA02EFFDCFB5C953AB58BFA67" authority="Sichel, 1864" authorityName="Sichel" authorityYear="1864" box="[1168,1436,1400,1426]" class="Insecta" family="Tiphiidae" genus="Paratiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B976BB0DA02EFFDCFB5C953AB51EFA67" box="[1168,1289,1400,1426]" italics="true" pageId="1" pageNumber="593">Paratiphia</emphasis>
Sichel, 1864
</taxonomicName>
are more or less similar in morphology.
<taxonomicName id="4C021C9CA02EFFDCFD8A95DEB32BFA43" authority="Cameron" authorityName="Cameron" authorityYear="1907" box="[582,828,1436,1462]" class="Insecta" family="Vespidae" genus="Cyanotiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B976BB0DA02EFFDCFD8A95DEB2C7FA43" box="[582,720,1436,1462]" italics="true" pageId="1" pageNumber="593">Cyanotiphia</emphasis>
Cameron
</taxonomicName>
differs from
<taxonomicName id="4C021C9CA02EFFDCFC0495DEB46AFA43" authority="Fabricius" authorityName="Fabricius" authorityYear="1775" box="[968,1149,1436,1462]" class="Insecta" family="Tiphiidae" genus="Tiphia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B976BB0DA02EFFDCFC0495DEB407FA43" box="[968,1040,1436,1462]" italics="true" pageId="1" pageNumber="593">Tiphia</emphasis>
Fabricius
</taxonomicName>
in these characters: dorsal and lateral pronotum with dense punctures; disc of propodeum laterally with at least one complete, longitudinal carina running from anterior to posterior margin of this area and posteriorly connected with transverse carina (
<figureCitation id="13397B9AA02EFFDCFAA695A6B0B0F9D7" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 4</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFF7A964AB0C2F9D7" box="[182,213,1544,1570]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">12</figureCitation>
); posterior surface of propodeum with radial wrinkles (
<figureCitation id="13397B9AA02EFFDCFC99964AB3B7F9D7" box="[853,928,1544,1570]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 4</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFC63964AB3D9F9D7" box="[943,974,1544,1570]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">12</figureCitation>
); mid tibia with two spurs apically; and anterior T1 separated by distinct transverse carina followed by short longitudinal wrinkles (
<figureCitation id="13397B9AA02EFFDCFB5B966EB4C9F9B3" box="[1175,1246,1580,1606]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 1</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFB20966EB4EBF9B3" box="[1260,1276,1580,1606]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">9</figureCitation>
).
<taxonomicName id="4C021C9CA02EFFDCFADE966EB0E9F99F" authority="Cameron" authorityName="Cameron" authorityYear="1907" class="Insecta" family="Vespidae" genus="Cyanotiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B976BB0DA02EFFDCFADE966EB58BF9B3" box="[1298,1436,1580,1606]" italics="true" pageId="1" pageNumber="593">Cyanotiphia</emphasis>
Cameron
</taxonomicName>
differs from
<taxonomicName id="4C021C9CA02EFFDCFE419612B245F99F" authority="Sichel" authorityName="Sichel" authorityYear="1864" box="[397,594,1616,1642]" class="Insecta" family="Tiphiidae" genus="Paratiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B976BB0DA02EFFDCFE419612B211F99F" box="[397,518,1616,1642]" italics="true" pageId="1" pageNumber="593">Paratiphia</emphasis>
Sichel
</taxonomicName>
by the following: lateral pronotum in the male without a prominent, forward directed process; ventral aspect of hind femur without a carina in female (
<bibRefCitation id="EF931AEEA02EFFDCFC1E9636B487F97B" author="Saini, J. &amp; Gupta, D. &amp; Gupta, K. C. S. K." box="[978,1168,1652,1678]" pageId="1" pageNumber="593" pagination="87 - 92" refId="ref4546" refString="Saini, J., Gupta, D. &amp; Gupta, K. C. S. K. (2022) Taxonomic notes on the little-known genus Cyanotiphia Cameron, 1907 (Hymenoptera: Tiphidae: Tiphinae). Zootaxa, 5154 (1), 87-92. https://doi.org/10.11646/zootaxa.5154.1.6" type="journal article" year="2022">
Saini
<emphasis id="B976BB0DA02EFFDCFBD89637B45BF97B" box="[1044,1100,1652,1678]" italics="true" pageId="1" pageNumber="593">et al.</emphasis>
2022
</bibRefCitation>
); tegula distinctly large, slightly shorter than mesonotum (
<figureCitation id="13397B9AA02EFFDCFDDB96DAB248F947" box="[535,607,1688,1714]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 1</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFDA396DAB268F947" box="[623,639,1688,1714]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">9</figureCitation>
); propodeum posteriorly with only one transverse carina (
<figureCitation id="13397B9AA02EFFDCFADB96DAB576F947" box="[1303,1377,1688,1714]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 4</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFAA396DAB599F947" box="[1391,1422,1688,1714]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">12</figureCitation>
); lateral T1 without an abruptly depressed, finely setiferous plaque (
<bibRefCitation id="EF931AEEA02EFFDCFCB196FEB417F923" author="Allen, H. W." box="[893,1024,1724,1750]" pageId="1" pageNumber="593" pagination="25 - 109" refId="ref4198" refString="Allen, H. W. (1968) A monographic study of the genus Paratiphia. Transactions of the American Entomological Society, 94, 25-109." type="journal article" year="1968">Allen 1968</bibRefCitation>
); abdominal segments with posterior margin impunctate (
<figureCitation id="13397B9AA02EFFDCFEB496A2B1A8F90F" box="[376,447,1760,1786]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 1</figureCitation>
,
<figureCitation id="13397B9AA02EFFDCFE0196A2B1CAF90F" box="[461,477,1760,1786]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1309,1333]" captionTargetBox="[151,1436,187,1285]" captionTargetId="figure-18@4.[151,1436,181,1285]" captionTargetPageId="4" captionText="FIGURES 9–16. Cyanotiphia fenchihuensis (Tsuneki, 1986), comb. nov. (♂). 9. Habitus in dorsal view. 10. Head in frontal view (the arrow shows medial longitudinal carina). 11. Head in dorsal view. 12. Metanotum and dorsal face of propodeum. 13. Lateral face of propodeum. 14. S1 in ventral view. 15. Metasoma in ventral view. 16. T7 in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14853718" httpUri="https://zenodo.org/record/14853718/files/figure.png" pageId="1" pageNumber="593">9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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