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<document ID-DOI="http://doi.org/10.5281/zenodo.4760318" ID-GBIF-Dataset="00423bac-ed5d-426e-9c93-c78cfa131dae" ID-GBIF-Taxon="182075239" ID-ISSN="1854-0392" ID-Zenodo-Dep="4760318" ID-ZooBank="EF081AA7-D54B-4F49-AE22-92B10A2CD652" LSID="urn:lsid:zoobank.org:act:4E107EF2-A371-4269-BDDF-8238BC95A182" checkinTime="1620964352749" checkinUser="felipe" docAuthor="Vinçon, Gilles, Dia, Aref, Kovács, Tibor &amp; Murányi, Dávid" docDate="2013" docId="039E87B7F9622C02FF60F9BD3102FA4B" docLanguage="en" docName="Illiesia.9.03.18-27.pdf" docOrigin="Illiesia 9 (3)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Perlodes thomasi Vincon, Dia &amp; Kovacs 2013, sp. n." docType="treatment" docUuid="4E107EF2-A371-4269-BDDF-8238BC95A182" docUuidSource="ZooBank" docVersion="7" lastPageId="5" lastPageNumber="23" masterDocId="FFA7FFCFF9602C07FFBAFFD33431FFFB" masterDocTitle="A New Stonefly From Lebanon, Perlodes Thomasi Sp. N. (Plecoptera: Perlodidae)" masterLastPageNumber="27" masterPageNumber="18" pageId="2" pageNumber="20" updateTime="1643531936469" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A New Stonefly From Lebanon, Perlodes Thomasi Sp. N. (Plecoptera: Perlodidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vinçon, Gilles</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dia, Aref</mods:namePart>
<mods:affiliation>Faculty of Sciences 1. Lebanese University and National Council for Scientific Research, Beirut, Lebanon E-mail: Arefdia @ yahoo. com</mods:affiliation>
<mods:nameIdentifier type="email">Arefdia@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kovács, Tibor</mods:namePart>
<mods:affiliation>Mátra Museum, Kossuth Lajos u. 40, H- 3200 Gyöngyös, Hungary E-mail: koati @ t-online. hu</mods:affiliation>
<mods:nameIdentifier type="email">koati@t-online.hu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Murányi, Dávid</mods:namePart>
<mods:affiliation>Department of Zoology, Hungarian Natural History Museum, Baross u. 13, H- 1088 Budapest, Hungary E-mail: muranyi @ zool. nhmus. hu</mods:affiliation>
<mods:nameIdentifier type="email">muranyi@zool.nhmus.hu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>9</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>18</mods:start>
<mods:end>27</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760840</mods:identifier>
<mods:identifier type="GBIF-Dataset">00423bac-ed5d-426e-9c93-c78cfa131dae</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4760840</mods:identifier>
<mods:identifier type="ZooBank">EF081AA7-D54B-4F49-AE22-92B10A2CD652</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4760318" ID-GBIF-Taxon="182075239" ID-Zenodo-Dep="4760318" LSID="urn:lsid:zoobank.org:act:4E107EF2-A371-4269-BDDF-8238BC95A182" httpUri="http://treatment.plazi.org/id/039E87B7F9622C02FF60F9BD3102FA4B" lastPageId="5" lastPageNumber="23" pageId="2" pageNumber="20">
<subSubSection box="[218,771,1645,1670]" pageId="2" pageNumber="20" type="nomenclature">
<paragraph blockId="2.[218,771,1645,1670]" box="[218,771,1645,1670]" pageId="2" pageNumber="20">
<heading bold="true" box="[218,771,1645,1670]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="20" reason="2">
<emphasis bold="true" box="[218,771,1645,1670]" pageId="2" pageNumber="20">
<taxonomicName authority="Vincon, Dia &amp; Kovacs" authorityName="Vincon, Dia &amp; Kovacs" authorityYear="2013" box="[218,700,1645,1670]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="thomasi" status="sp. n.">
<emphasis bold="true" box="[218,423,1646,1670]" italics="true" pageId="2" pageNumber="20">Perlodes thomasi</emphasis>
Vinçon, Dia &amp; Kovács
</taxonomicName>
<taxonomicNameLabel box="[707,771,1646,1670]" pageId="2" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[320,669,1682,1706]" pageId="2" pageNumber="20" type="description">
<paragraph blockId="2.[320,669,1682,1706]" box="[320,669,1682,1706]" pageId="2" pageNumber="20">
(
<figureCitation box="[328,405,1682,1706]" captionStart="Fig" captionStartId="2.[189,230,1113,1137]" captionTargetBox="[286,1353,245,1051]" captionTargetId="figure-50@2.[274,1358,235,1058]" captionTargetPageId="2" captionText="Fig. 2. P. thomasi sp. n. a: male head (2 mm), b: female subgenital plate (2 mm), c: male paraproct in ¾ view (0.5 mm), d: male paraproct in side view (same scale), e: male everted penis (2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760844" httpUri="https://zenodo.org/record/4760844/files/figure.png" pageId="2" pageNumber="20">Figs. 2</figureCitation>
a-e, 3a-d, 4a-d, 5 right)
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="20" type="materials_examined">
<paragraph blockId="2.[189,800,1753,1884]" lastBlockId="2.[832,1444,1255,1883]" pageId="2" pageNumber="20">
<emphasis bold="true" box="[189,444,1753,1777]" pageId="2" pageNumber="20">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111752302" collectingDate="1996-05-18" collectingDateMax="1996-05-18" collectingDateMin="1996-04-24" collectionCode="ZML" country="Lebanon" county="Zoology Museum of Lausanne" elevation="1500" location="Ouadi Qadicha torrent" municipality="Abou Aali" pageId="2" pageNumber="20" specimenCount="1" typeStatus="holotype">
<typeStatus box="[470,582,1753,1777]" pageId="2" pageNumber="20">Holotype</typeStatus>
<specimenCount box="[607,667,1753,1777]" pageId="2" pageNumber="20" type="male">male</specimenCount>
: northern
<collectingCountry box="[189,293,1789,1813]" name="Lebanon" pageId="2" pageNumber="20">Lebanon</collectingCountry>
, western slope of Lebanon Mount,
<location LSID="urn:lsid:plazi:treatment:039E87B7F9622C02FF60F9BD3102FA4B:8EE8607AF9622C05FD6CF92E3548F8C3" country="Lebanon" county="Zoology Museum of Lausanne" municipality="Abou Aali" name="Ouadi Qadicha torrent" pageId="2" pageNumber="20">Ouâdi Qâdîcha torrent</location>
,
<elevation box="[387,529,1824,1848]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="20" unit="m" value="1500.0">
<quantity box="[387,470,1824,1848]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="20" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
,
<locationDeviation pageId="2" pageNumber="20">
about
<quantity box="[613,682,1824,1848]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="20" unit="m" value="500.0">500 m</quantity>
above the village of Bcherré, Abou Aali water basin
</locationDeviation>
,
<date box="[676,796,1860,1884]" pageId="2" pageNumber="20" value="1996-05-18">
<collectingDate box="[676,796,1860,1884]" pageId="2" pageNumber="20" value="1996-05-18">18.05.1996</collectingDate>
</date>
, deposited in the Zoology Museum of Lausanne (
<collectionCode box="[839,901,1291,1315]" country="Russia" httpUri="http://grbio.org/cool/nhq2-n6y1" name="St Petersburg State University" pageId="2" pageNumber="20">ZML</collectionCode>
) (
<figureCitation box="[922,985,1290,1315]" captionStart="Fig" captionStartId="1.[189,230,1219,1243]" captionTargetBox="[493,1138,250,1155]" captionTargetId="figure-50@1.[488,1144,241,1158]" captionTargetPageId="1" captionText="Fig. 1. Map of Lebanon, with the two collecting sites for Perlodes thomasi sp. n." figureDoi="http://doi.org/10.5281/zenodo.4760842" httpUri="https://zenodo.org/record/4760842/files/figure.png" pageId="2" pageNumber="20">Fig. 1</figureCitation>
, station 1b).
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111752303" collectingDate="1996-04-24" collectingDateMax="1996-05-18" collectingDateMin="1996-04-24" collectionCode="ZML" country="Lebanon" county="Zoology Museum of Lausanne" elevation="1500" location="Mount" municipality="Abou Aali" pageId="2" pageNumber="20" specimenCount="2" typeStatus="holotype">
One female
<typeStatus box="[1269,1374,1291,1315]" pageId="2" pageNumber="20">paratype</typeStatus>
, same location and date, in the
<collectionCode box="[1117,1180,1327,1350]" country="Russia" httpUri="http://grbio.org/cool/nhq2-n6y1" name="St Petersburg State University" pageId="2" pageNumber="20">ZML</collectionCode>
. One larva,
<date box="[1315,1438,1326,1350]" pageId="2" pageNumber="20" value="1996-04-24">
<collectingDate box="[1315,1438,1326,1350]" pageId="2" pageNumber="20" value="1996-04-24">24.04.1996</collectingDate>
</date>
, same location, also in the
<collectionCode box="[1134,1197,1362,1385]" country="Russia" httpUri="http://grbio.org/cool/nhq2-n6y1" name="St Petersburg State University" pageId="2" pageNumber="20">ZML</collectionCode>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111752306" pageId="2" pageNumber="20">
Other material from the same locality:
<specimenCount box="[1080,1114,1396,1421]" pageId="2" pageNumber="20" type="male">1♂</specimenCount>
,
<specimenCount box="[1139,1173,1396,1421]" pageId="2" pageNumber="20" type="female">1♀</specimenCount>
,
<date box="[1198,1321,1397,1421]" pageId="2" pageNumber="20" value="1995-05-30">30.05.1995</date>
,
<specimenCount box="[1344,1438,1397,1421]" pageId="2" pageNumber="20" type="larva">1 larva</specimenCount>
,
<date box="[832,953,1432,1456]" pageId="2" pageNumber="20" value="1996-04-24">24.04.1996</date>
(in the collection of G. Vinçon);
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111752304" pageId="2" pageNumber="20">
<specimenCount box="[1331,1439,1432,1457]" pageId="2" pageNumber="20" type="larva">25 larvae</specimenCount>
,
<date box="[832,942,1468,1492]" pageId="2" pageNumber="20" value="1996-04-24">24.4.1996</date>
;
<specimenCount box="[951,998,1467,1492]" pageId="2" pageNumber="20" type="male">18♂</specimenCount>
,
<specimenCount box="[1010,1043,1467,1492]" pageId="2" pageNumber="20" type="female">7♀</specimenCount>
,
<specimenCount box="[1055,1148,1468,1492]" pageId="2" pageNumber="20" type="larva">8 larvae</specimenCount>
, 2 exuviae,
<date box="[1275,1395,1468,1492]" pageId="2" pageNumber="20" value="1996-05-18">18.05.1996</date>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111752301" pageId="2" pageNumber="20">
<specimenCount box="[1404,1437,1467,1492]" pageId="2" pageNumber="20" type="male">2♂</specimenCount>
,
<specimenCount box="[832,865,1502,1528]" pageId="2" pageNumber="20" type="female">6♀</specimenCount>
,
<specimenCount box="[878,986,1503,1528]" pageId="2" pageNumber="20" type="larva">15 larvae</specimenCount>
,
<date box="[997,1107,1503,1527]" pageId="2" pageNumber="20" value="1999-05-06">6.05.1999</date>
;
<specimenCount box="[1117,1151,1502,1527]" pageId="2" pageNumber="20" type="male">7♂</specimenCount>
,
<specimenCount box="[1164,1198,1502,1527]" pageId="2" pageNumber="20" type="female">5♀</specimenCount>
,
<date box="[1211,1335,1503,1527]" pageId="2" pageNumber="20" value="1999-05-30">30.05.1999</date>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111752305" pageId="2" pageNumber="20">
Qâdîcha spring, resurgence of the Abou Aali stream,
<quantity box="[1358,1442,1539,1563]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="2" pageNumber="20" unit="m" value="1760.0">1760 m</quantity>
a.s.l. (
<figureCitation box="[913,991,1575,1599]" captionStart="Fig" captionStartId="1.[189,230,1219,1243]" captionTargetBox="[493,1138,250,1155]" captionTargetId="figure-50@1.[488,1144,241,1158]" captionTargetPageId="1" captionText="Fig. 1. Map of Lebanon, with the two collecting sites for Perlodes thomasi sp. n." figureDoi="http://doi.org/10.5281/zenodo.4760842" httpUri="https://zenodo.org/record/4760842/files/figure.png" pageId="2" pageNumber="20">Fig. 1</figureCitation>
, station 1a),
<specimenCount box="[1182,1215,1573,1599]" pageId="2" pageNumber="20" type="female">8♀</specimenCount>
,
<date box="[1242,1365,1574,1598]" pageId="2" pageNumber="20" value="1995-05-30">30.05.1995</date>
;
<specimenCount box="[1389,1437,1573,1599]" pageId="2" pageNumber="20" type="female">10♀</specimenCount>
,
<date box="[832,955,1610,1634]" pageId="2" pageNumber="20" value="1996-06-15">15.06.1996</date>
;
<specimenCount box="[966,1000,1609,1634]" pageId="2" pageNumber="20" type="female">4♀</specimenCount>
,
<date box="[1013,1123,1610,1634]" pageId="2" pageNumber="20" value="1995-07-09">9.07.1995</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="23" pageId="2" pageNumber="20" type="description">
<paragraph blockId="2.[832,1444,1255,1883]" lastBlockId="3.[189,801,935,1883]" lastPageId="3" lastPageNumber="21" pageId="2" pageNumber="20">
<emphasis bold="true" box="[832,975,1646,1670]" pageId="2" pageNumber="20">Description</emphasis>
. Medium sized
<taxonomicName box="[1164,1347,1646,1670]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[1164,1254,1646,1670]" italics="true" pageId="2" pageNumber="20">Perlodes</emphasis>
species
</taxonomicName>
:
<specimenCount box="[1357,1377,1645,1670]" pageId="2" pageNumber="20" type="male">♂</specimenCount>
body length
<quantity box="[913,1032,1681,1706]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="2" pageNumber="20" unit="cm" value="1.55" valueMax="1.6" valueMin="1.5">1.5-1.6 cm</quantity>
,
<specimenCount box="[1043,1063,1680,1705]" pageId="2" pageNumber="20" type="female">♀</specimenCount>
2.0-
<quantity box="[1112,1189,1682,1706]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="20" unit="cm" value="2.2">2.2 cm</quantity>
.
<specimenCount box="[1199,1219,1680,1705]" pageId="2" pageNumber="20" type="male">♂</specimenCount>
micropterous with relative wing length (RWL) between 1.1 and 1.3 (RWL = wing length / head width at the position of eyes).
<specimenCount box="[910,930,1787,1812]" pageId="2" pageNumber="20" type="female">♀</specimenCount>
normal winged (wing length:
<quantity box="[1306,1431,1788,1812]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="2" pageNumber="20" unit="cm" value="2.5" valueMax="2.6" valueMin="2.4">2.4-2.6 cm</quantity>
). General colour brown. Head dark brown except yellowish occiput and oval yellow spot in the ocellar triangle (
<figureCitation box="[296,377,935,959]" captionStart="Fig" captionStartId="2.[189,230,1113,1137]" captionTargetBox="[286,1353,245,1051]" captionTargetId="figure-50@2.[274,1358,235,1058]" captionTargetPageId="2" captionText="Fig. 2. P. thomasi sp. n. a: male head (2 mm), b: female subgenital plate (2 mm), c: male paraproct in ¾ view (0.5 mm), d: male paraproct in side view (same scale), e: male everted penis (2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760844" httpUri="https://zenodo.org/record/4760844/files/figure.png" pageId="3" pageNumber="21">Fig. 2a</figureCitation>
); two dark spots on each side of the occiput, behind the compound eyes. Antennae dark brown. Median part of pro- and mesonotum with lateral dark brown patterns and contrasting median yellowish strip. Legs brownish.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4760846" ID-Zenodo-Dep="4760846" box="[189,1189,829,853]" httpUri="https://zenodo.org/record/4760846/files/figure.png" pageId="3" pageNumber="21" startId="3.[189,230,829,853]" targetBox="[269,1362,235,761]" targetPageId="3">
<paragraph blockId="3.[189,1189,829,853]" box="[189,1189,829,853]" pageId="3" pageNumber="21">
Fig. 3.
<taxonomicName authority="Vinçon &amp; Dia &amp; Kovács &amp; Murányi, 2013" authorityName="Vinçon &amp; Dia &amp; Kovács &amp; Murányi" authorityYear="2013" box="[267,381,829,853]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="thomasi" status="sp. n.">
<emphasis box="[267,381,829,853]" italics="true" pageId="3" pageNumber="21">P. thomasi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[388,450,829,852]" pageId="3" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
larva. a: head (5 mm), b: maxilla (1 mm), c: paraprocts (1 mm).
</paragraph>
</caption>
<paragraph blockId="3.[189,801,935,1883]" pageId="3" pageNumber="21">
<emphasis bold="true" box="[189,364,1113,1137]" pageId="3" pageNumber="21">Male genitalia</emphasis>
. The paraprocts are nearly pyramidal, mostly membranous on their inner side and protected by a wide outer sclerite that narrows progressively towards the tip of the paraproct. This expansion of the sclerite ends in a thin blade, extending around the tip of the paraproct; it is more visible in ¾ view (
<figureCitation box="[410,489,1326,1350]" captionStart="Fig" captionStartId="2.[189,230,1113,1137]" captionTargetBox="[286,1353,245,1051]" captionTargetId="figure-50@2.[274,1358,235,1058]" captionTargetPageId="2" captionText="Fig. 2. P. thomasi sp. n. a: male head (2 mm), b: female subgenital plate (2 mm), c: male paraproct in ¾ view (0.5 mm), d: male paraproct in side view (same scale), e: male everted penis (2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760844" httpUri="https://zenodo.org/record/4760844/files/figure.png" pageId="3" pageNumber="21">Fig. 2c</figureCitation>
) than in lateral view (
<figureCitation captionStart="Fig" captionStartId="2.[189,230,1113,1137]" captionTargetBox="[286,1353,245,1051]" captionTargetId="figure-50@2.[274,1358,235,1058]" captionTargetPageId="2" captionText="Fig. 2. P. thomasi sp. n. a: male head (2 mm), b: female subgenital plate (2 mm), c: male paraproct in ¾ view (0.5 mm), d: male paraproct in side view (same scale), e: male everted penis (2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760844" httpUri="https://zenodo.org/record/4760844/files/figure.png" pageId="3" pageNumber="21">Fig. 2d</figureCitation>
). The membranous part of the paraprocts is of very variable size, from hardly turgescent (
<figureCitation box="[683,755,1397,1421]" captionStart="Fig" captionStartId="2.[189,230,1113,1137]" captionTargetBox="[286,1353,245,1051]" captionTargetId="figure-50@2.[274,1358,235,1058]" captionTargetPageId="2" captionText="Fig. 2. P. thomasi sp. n. a: male head (2 mm), b: female subgenital plate (2 mm), c: male paraproct in ¾ view (0.5 mm), d: male paraproct in side view (same scale), e: male everted penis (2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760844" httpUri="https://zenodo.org/record/4760844/files/figure.png" pageId="3" pageNumber="21">Figs. 2</figureCitation>
c-d) to strongly turgescent (
<figureCitation box="[476,558,1433,1457]" captionStart="Fig" captionStartId="2.[189,230,1113,1137]" captionTargetBox="[286,1353,245,1051]" captionTargetId="figure-50@2.[274,1358,235,1058]" captionTargetPageId="2" captionText="Fig. 2. P. thomasi sp. n. a: male head (2 mm), b: female subgenital plate (2 mm), c: male paraproct in ¾ view (0.5 mm), d: male paraproct in side view (same scale), e: male everted penis (2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760844" httpUri="https://zenodo.org/record/4760844/files/figure.png" pageId="3" pageNumber="21">Fig. 2e</figureCitation>
: male with everted penis). The penis is often not visible since it is retracted in the abdomen; in some specimens it could be reverted (perhaps during mating or due to finger pressure of the collector); it is fully membranous and of complicated structure comprising two wide lateral lobes extending on each side of the abdomen tip, two upper lobes with a median depression and two less developed posterior lobes (
<figureCitation box="[510,591,1717,1741]" captionStart="Fig" captionStartId="2.[189,230,1113,1137]" captionTargetBox="[286,1353,245,1051]" captionTargetId="figure-50@2.[274,1358,235,1058]" captionTargetPageId="2" captionText="Fig. 2. P. thomasi sp. n. a: male head (2 mm), b: female subgenital plate (2 mm), c: male paraproct in ¾ view (0.5 mm), d: male paraproct in side view (same scale), e: male everted penis (2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760844" httpUri="https://zenodo.org/record/4760844/files/figure.png" pageId="3" pageNumber="21">Fig. 2e</figureCitation>
).
</paragraph>
<paragraph blockId="3.[189,801,935,1883]" lastBlockId="3.[832,1443,935,1883]" pageId="3" pageNumber="21">
<emphasis bold="true" box="[189,403,1752,1777]" pageId="3" pageNumber="21">Female genitalia</emphasis>
. Subgenital plate wide, semicircular, with rounded edges and shallow median depression on the outer edge. The plate sides nearly reach the lateral edges of the abdomen. Two lateral dark spots occur near the anterior edge of the plate (
<figureCitation box="[840,923,971,995]" captionStart="Fig" captionStartId="2.[189,230,1113,1137]" captionTargetBox="[286,1353,245,1051]" captionTargetId="figure-50@2.[274,1358,235,1058]" captionTargetPageId="2" captionText="Fig. 2. P. thomasi sp. n. a: male head (2 mm), b: female subgenital plate (2 mm), c: male paraproct in ¾ view (0.5 mm), d: male paraproct in side view (same scale), e: male everted penis (2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760844" httpUri="https://zenodo.org/record/4760844/files/figure.png" pageId="3" pageNumber="21">Fig. 2b</figureCitation>
).
</paragraph>
<paragraph blockId="3.[832,1443,935,1883]" pageId="3" pageNumber="21">
<emphasis bold="true" box="[832,900,1006,1030]" pageId="3" pageNumber="21">Larva</emphasis>
(
<figureCitation box="[923,1008,1006,1030]" captionStart="Fig" captionStartId="3.[189,230,829,853]" captionTargetBox="[269,1362,235,761]" captionTargetId="figure-50@3.[261,1370,226,764]" captionTargetPageId="3" captionText="Fig. 3. P. thomasi sp. n. larva. a: head (5 mm), b: maxilla (1 mm), c: paraprocts (1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760846" httpUri="https://zenodo.org/record/4760846/files/figure.png" pageId="3" pageNumber="21">Figs. 3</figureCitation>
a-c). Head brownish, with a wide yellow marking covering the occiput and an oval median yellow spot between the three ocelli (
<figureCitation box="[1353,1429,1078,1102]" captionStart="Fig" captionStartId="3.[189,230,829,853]" captionTargetBox="[269,1362,235,761]" captionTargetId="figure-50@3.[261,1370,226,764]" captionTargetPageId="3" captionText="Fig. 3. P. thomasi sp. n. larva. a: head (5 mm), b: maxilla (1 mm), c: paraprocts (1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760846" httpUri="https://zenodo.org/record/4760846/files/figure.png" pageId="3" pageNumber="21">Fig. 3a</figureCitation>
). M-line yellowish, sinuous and poorly visible. The lacinia is typical of the genus
<taxonomicName authorityName="THOMASI" authorityYear="2013" box="[1198,1288,1149,1173]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis box="[1198,1288,1149,1173]" italics="true" pageId="3" pageNumber="21">Perlodes</emphasis>
</taxonomicName>
, with a very strong outer tooth, gently curved over its length, and one inner tooth which is about twice as long as the smaller. Two strong bristles are visible, one on the inner edge of the lacinia, proximally to the smaller tooth, and the other between the two teeth (
<figureCitation box="[1348,1430,1326,1350]" captionStart="Fig" captionStartId="3.[189,230,829,853]" captionTargetBox="[269,1362,235,761]" captionTargetId="figure-50@3.[261,1370,226,764]" captionTargetPageId="3" captionText="Fig. 3. P. thomasi sp. n. larva. a: head (5 mm), b: maxilla (1 mm), c: paraprocts (1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760846" httpUri="https://zenodo.org/record/4760846/files/figure.png" pageId="3" pageNumber="21">Fig. 3b</figureCitation>
). Paraprocts triangular, well sclerotized, and covered with only few scattered short spine-like bristles (
<figureCitation captionStart="Fig" captionStartId="3.[189,230,829,853]" captionTargetBox="[269,1362,235,761]" captionTargetId="figure-50@3.[261,1370,226,764]" captionTargetPageId="3" captionText="Fig. 3. P. thomasi sp. n. larva. a: head (5 mm), b: maxilla (1 mm), c: paraprocts (1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760846" httpUri="https://zenodo.org/record/4760846/files/figure.png" pageId="3" pageNumber="21">Fig. 3c</figureCitation>
).
</paragraph>
<paragraph blockId="3.[832,1443,935,1883]" lastBlockId="5.[189,801,971,1919]" lastPageId="5" lastPageNumber="23" pageId="3" pageNumber="21">
<emphasis bold="true" box="[832,877,1469,1493]" pageId="3" pageNumber="21">Egg</emphasis>
(
<figureCitation box="[894,973,1468,1492]" captionStart="Fig" captionStartId="4.[189,230,1752,1776]" captionTargetBox="[287,1345,251,1645]" captionTargetId="figure-50@4.[287,1345,251,1648]" captionTargetPageId="4" captionText="Fig. 4. Comparison of the eggs of P. thomasi sp. n. (a-d) and P. floridus (e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale." figureDoi="http://doi.org/10.5281/zenodo.4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="3" pageNumber="21">Figs. 4</figureCitation>
a-d). Exceptionally large (about
<quantity box="[1355,1443,1468,1492]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="21" unit="mm" value="0.5">0.5 mm</quantity>
length and
<quantity box="[963,1049,1504,1528]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="21" unit="mm" value="0.4">0.4 mm</quantity>
width), bullet-shaped. The ridges of the egg are clearly visible, rather thick and bulged (not flat). The hatching line is strong, bulged, well delimiting the operculum. The attachment disc is flanked by several concentric rows of peripheral cells similar to the petals of a flower. These cells are placed on the anterior surface of the attachment disc (surface of the disc connected to the base of the egg). Therefore, the cells are more visible in lateral view (
<figureCitation box="[840,919,1824,1848]" captionStart="Fig" captionStartId="4.[189,230,1752,1776]" captionTargetBox="[287,1345,251,1645]" captionTargetId="figure-50@4.[287,1345,251,1648]" captionTargetPageId="4" captionText="Fig. 4. Comparison of the eggs of P. thomasi sp. n. (a-d) and P. floridus (e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale." figureDoi="http://doi.org/10.5281/zenodo.4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="3" pageNumber="21">Fig. 4a</figureCitation>
) than in basal view (
<figureCitation box="[1164,1247,1824,1848]" captionStart="Fig" captionStartId="4.[189,230,1752,1776]" captionTargetBox="[287,1345,251,1645]" captionTargetId="figure-50@4.[287,1345,251,1648]" captionTargetPageId="4" captionText="Fig. 4. Comparison of the eggs of P. thomasi sp. n. (a-d) and P. floridus (e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale." figureDoi="http://doi.org/10.5281/zenodo.4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="3" pageNumber="21">Fig. 4b</figureCitation>
). The cells of the inner rows are rounded and smaller than those of the outer rows that are more oval or even quadrangular (
<figureCitation box="[197,294,1006,1030]" captionStart="Fig" captionStartId="4.[189,230,1752,1776]" captionTargetBox="[287,1345,251,1645]" captionTargetId="figure-50@4.[287,1345,251,1648]" captionTargetPageId="4" captionText="Fig. 4. Comparison of the eggs of P. thomasi sp. n. (a-d) and P. floridus (e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale." figureDoi="http://doi.org/10.5281/zenodo.4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="5" pageNumber="23">Figs. 4c</figureCitation>
). Numerous mushroom-like globules are gathered on the surface of the attachment disc near its outer edge (
<figureCitation box="[350,425,1077,1101]" captionStart="Fig" captionStartId="4.[189,230,1752,1776]" captionTargetBox="[287,1345,251,1645]" captionTargetId="figure-50@4.[287,1345,251,1648]" captionTargetPageId="4" captionText="Fig. 4. Comparison of the eggs of P. thomasi sp. n. (a-d) and P. floridus (e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale." figureDoi="http://doi.org/10.5281/zenodo.4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="5" pageNumber="23">Fig. 4</figureCitation>
b-c). The chorionic surface is covered by a conspicuous polygonal network of follicular cell impressions (FCIs) (
<figureCitation box="[602,731,1148,1172]" captionStart="Fig" captionStartId="4.[189,230,1752,1776]" captionTargetBox="[287,1345,251,1645]" captionTargetId="figure-50@4.[287,1345,251,1648]" captionTargetPageId="4" captionText="Fig. 4. Comparison of the eggs of P. thomasi sp. n. (a-d) and P. floridus (e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale." figureDoi="http://doi.org/10.5281/zenodo.4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="5" pageNumber="23">Figs. 4a, d</figureCitation>
). The surface of each polygon is covered by several rounded granulations (
<figureCitation box="[462,545,1219,1243]" captionStart="Fig" captionStartId="4.[189,230,1752,1776]" captionTargetBox="[287,1345,251,1645]" captionTargetId="figure-50@4.[287,1345,251,1648]" captionTargetPageId="4" captionText="Fig. 4. Comparison of the eggs of P. thomasi sp. n. (a-d) and P. floridus (e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale." figureDoi="http://doi.org/10.5281/zenodo.4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="5" pageNumber="23">Fig. 4d</figureCitation>
).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4760848" ID-Zenodo-Dep="4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="4" pageNumber="22" startId="4.[189,230,1752,1776]" targetBox="[287,1345,251,1645]" targetPageId="4">
<paragraph blockId="4.[189,1443,1752,1812]" pageId="4" pageNumber="22">
Fig. 4. Comparison of the eggs of
<taxonomicName authority="Vinçon &amp; Dia &amp; Kovács &amp; Murányi, 2013" authorityName="Vinçon &amp; Dia &amp; Kovács &amp; Murányi" authorityYear="2013" box="[604,721,1752,1776]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="22" phylum="Arthropoda" rank="species" species="thomasi" status="sp. n.">
<emphasis box="[604,721,1752,1776]" italics="true" pageId="4" pageNumber="22">P. thomasi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[731,796,1753,1776]" pageId="4" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
(a-d) and
<taxonomicName authorityName="Kovacs, Tibor &amp; Vincon" authorityYear="2012" box="[927,1042,1752,1776]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="22" phylum="Arthropoda" rank="species" species="floridus">
<emphasis box="[927,1042,1752,1776]" italics="true" pageId="4" pageNumber="22">P. floridus</emphasis>
</taxonomicName>
(e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4760850" ID-Zenodo-Dep="4760850" box="[189,1390,829,853]" httpUri="https://zenodo.org/record/4760850/files/figure.png" pageId="5" pageNumber="23" startId="5.[189,230,829,853]" targetBox="[249,1383,251,750]" targetPageId="5">
<paragraph blockId="5.[189,1390,829,853]" box="[189,1390,829,853]" pageId="5" pageNumber="23">
Fig. 5.
<taxonomicName authorityName="THOMASI" authorityYear="2013" box="[267,357,829,853]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis box="[267,357,829,853]" italics="true" pageId="5" pageNumber="23">Perlodes</emphasis>
</taxonomicName>
eggs, from left to right:
<taxonomicName authority="floridus" authorityName="floridus Kovacs &amp; Vincon" authorityYear="2012" box="[641,843,829,853]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="floridus" subSpecies="floridus">
<emphasis box="[641,843,829,853]" italics="true" pageId="5" pageNumber="23">P. floridus floridus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Kovacs &amp; Vincon" authorityYear="2012" box="[856,1148,829,853]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="floridus" subSpecies="peloponnesiacus">
<emphasis box="[856,1148,829,853]" italics="true" pageId="5" pageNumber="23">P. floridus peloponnesiacus</emphasis>
</taxonomicName>
and
<taxonomicName authority="Vinçon &amp; Dia &amp; Kovács &amp; Murányi, 2013" authorityName="Vinçon &amp; Dia &amp; Kovács &amp; Murányi" authorityYear="2013" box="[1206,1320,829,853]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="thomasi" status="sp. n.">
<emphasis box="[1206,1320,829,853]" italics="true" pageId="5" pageNumber="23">P. thomasi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1327,1390,829,852]" pageId="5" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection pageId="5" pageNumber="23" type="discussion">
<paragraph blockId="5.[189,801,971,1919]" lastBlockId="5.[832,1444,971,1457]" pageId="5" pageNumber="23">
<emphasis bold="true" box="[189,307,1255,1279]" pageId="5" pageNumber="23">Affinities</emphasis>
. In the genus
<taxonomicName authorityName="THOMASI" authorityYear="2013" box="[493,583,1255,1279]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis box="[493,583,1255,1279]" italics="true" pageId="5" pageNumber="23">Perlodes</emphasis>
</taxonomicName>
,
<taxonomicName authority="Vinçon &amp; Dia &amp; Kovács &amp; Murányi, 2013" authorityName="Vinçon &amp; Dia &amp; Kovács &amp; Murányi" authorityYear="2013" box="[602,721,1255,1279]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="thomasi" status="sp. n.">
<emphasis box="[602,721,1255,1279]" italics="true" pageId="5" pageNumber="23">P. thomasi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[733,800,1255,1278]" pageId="5" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
shares a very distinct position together with
<taxonomicName authority="floridus" authorityName="floridus Kovacs &amp; Vincon" authorityYear="2012" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="floridus" subSpecies="floridus">
<emphasis italics="true" pageId="5" pageNumber="23">P. floridus floridus</emphasis>
</taxonomicName>
from the Balkan Peninsula and
<taxonomicName authorityName="Kovacs &amp; Vincon" authorityYear="2012" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="floridus" subSpecies="peloponnesiacus">
<emphasis italics="true" pageId="5" pageNumber="23">P. floridus peloponnesiacus</emphasis>
</taxonomicName>
from the Greek
<collectingRegion box="[657,794,1362,1386]" country="Greece" name="Peloponnisos" pageId="5" pageNumber="23">Peloponnes</collectingRegion>
. Both species have bullet-shaped eggs with a distinctive flower-like attachment disc (
<figureCitation box="[649,724,1433,1457]" captionStart="Fig" captionStartId="4.[189,230,1752,1776]" captionTargetBox="[287,1345,251,1645]" captionTargetId="figure-50@4.[287,1345,251,1648]" captionTargetPageId="4" captionText="Fig. 4. Comparison of the eggs of P. thomasi sp. n. (a-d) and P. floridus (e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale." figureDoi="http://doi.org/10.5281/zenodo.4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="5" pageNumber="23">Figs. 4</figureCitation>
b-c, 4fg). Nevertheless, the eggs of
<taxonomicName authority="Vinçon &amp; Dia &amp; Kovács &amp; Murányi, 2013" authorityName="Vinçon &amp; Dia &amp; Kovács &amp; Murányi" authorityYear="2013" box="[553,672,1468,1492]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="thomasi" status="sp. n.">
<emphasis box="[553,672,1468,1492]" italics="true" pageId="5" pageNumber="23">P. thomasi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[684,752,1469,1492]" pageId="5" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
are about one third wider and longer than those of
<taxonomicName authority="floridus" authorityName="floridus Kovacs &amp; Vincon" authorityYear="2012" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="floridus" subSpecies="floridus">
<emphasis italics="true" pageId="5" pageNumber="23">P. floridus floridus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Kovacs &amp; Vincon" authorityYear="2012" box="[414,702,1539,1563]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="floridus" subSpecies="peloponnesiacus">
<emphasis box="[414,702,1539,1563]" italics="true" pageId="5" pageNumber="23">P. floridus peloponnesiacus</emphasis>
</taxonomicName>
(
<figureCitation captionStart="Fig" captionStartId="4.[189,230,1752,1776]" captionTargetBox="[287,1345,251,1645]" captionTargetId="figure-50@4.[287,1345,251,1648]" captionTargetPageId="4" captionText="Fig. 4. Comparison of the eggs of P. thomasi sp. n. (a-d) and P. floridus (e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale." figureDoi="http://doi.org/10.5281/zenodo.4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="5" pageNumber="23">Figs 4a, 4e</figureCitation>
,
<figureCitation box="[237,252,1575,1599]" captionStart="Fig" captionStartId="5.[189,230,829,853]" captionTargetBox="[249,1383,251,750]" captionTargetId="figure-50@5.[249,1383,251,750]" captionTargetPageId="5" captionText="Fig. 5. Perlodes eggs, from left to right: P. floridus floridus, P. floridus peloponnesiacus and P. thomasi sp. n." figureDoi="http://doi.org/10.5281/zenodo.4760850" httpUri="https://zenodo.org/record/4760850/files/figure.png" pageId="5" pageNumber="23">5</figureCitation>
). The chorion is covered by a network of conspicuous convex polygonal cells FCIs (
<figureCitation box="[700,786,1610,1634]" captionStart="Fig" captionStartId="4.[189,230,1752,1776]" captionTargetBox="[287,1345,251,1645]" captionTargetId="figure-50@4.[287,1345,251,1648]" captionTargetPageId="4" captionText="Fig. 4. Comparison of the eggs of P. thomasi sp. n. (a-d) and P. floridus (e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale." figureDoi="http://doi.org/10.5281/zenodo.4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="5" pageNumber="23">Fig. 4d</figureCitation>
), while that of
<taxonomicName authority="floridus" authorityName="floridus Kovacs &amp; Vincon" authorityYear="2012" box="[360,571,1646,1670]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="floridus" subSpecies="floridus">
<emphasis box="[360,571,1646,1670]" italics="true" pageId="5" pageNumber="23">P. floridus floridus</emphasis>
</taxonomicName>
just shows hardly visible depressions and is densely covered with numerous rounded globules (
<figureCitation box="[546,629,1717,1741]" captionStart="Fig" captionStartId="4.[189,230,1752,1776]" captionTargetBox="[287,1345,251,1645]" captionTargetId="figure-50@4.[287,1345,251,1648]" captionTargetPageId="4" captionText="Fig. 4. Comparison of the eggs of P. thomasi sp. n. (a-d) and P. floridus (e-h). a, e: egg in side view; b, f: attachment disc; c, g: peripheral cells of the attachment disc; d, h: chorion ornamentation; a, b, f, e same scale." figureDoi="http://doi.org/10.5281/zenodo.4760848" httpUri="https://zenodo.org/record/4760848/files/figure.png" pageId="5" pageNumber="23">Fig. 4h</figureCitation>
). In
<taxonomicName authorityName="Kovacs &amp; Vincon" authorityYear="2012" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="floridus" subSpecies="peloponnesiacus">
<emphasis italics="true" pageId="5" pageNumber="23">P. floridus peloponnesiacus</emphasis>
</taxonomicName>
the polygonal cells of the chorion are concave instead of convex (
<bibRefCitation author="Kovacs, T. &amp; G. Vincon &amp; D. Muranyi &amp; I. Sivec" box="[505,707,1788,1812]" pageId="5" pageNumber="23" pagination="182 - 192" refId="ref3776" refString="Kovacs, T., G. Vincon, D. Muranyi &amp; I. Sivec. 2012. A new Perlodes species and its subspecies from the Balkan Peninsula (Plecoptera: Perlodidae). Illiesia, 8 (20): 182 - 192." type="journal article" year="2012">Kovács et al. 2012</bibRefCitation>
, Figs 12, 13, 15). The male paraproctal sclerite is also different: in
<taxonomicName authorityName="Vinçon &amp; Dia &amp; Kovács &amp; Murányi" authorityYear="2013" box="[235,364,1859,1883]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="thomasi">
<emphasis box="[235,364,1859,1883]" italics="true" pageId="5" pageNumber="23">P. thomasi</emphasis>
</taxonomicName>
the sclerite expansion is short, narrowing quickly toward the tip (
<figureCitation box="[588,664,1895,1919]" captionStart="Fig" captionStartId="2.[189,230,1113,1137]" captionTargetBox="[286,1353,245,1051]" captionTargetId="figure-50@2.[274,1358,235,1058]" captionTargetPageId="2" captionText="Fig. 2. P. thomasi sp. n. a: male head (2 mm), b: female subgenital plate (2 mm), c: male paraproct in ¾ view (0.5 mm), d: male paraproct in side view (same scale), e: male everted penis (2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760844" httpUri="https://zenodo.org/record/4760844/files/figure.png" pageId="5" pageNumber="23">Fig. 2c</figureCitation>
), while in
<taxonomicName authorityName="Kovacs, Tibor &amp; Vincon" authorityYear="2012" box="[778,915,971,1918]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="floridus">
<emphasis box="[778,915,971,1918]" italics="true" pageId="5" pageNumber="23">P. floridus</emphasis>
</taxonomicName>
it is more elongated and slender (
<bibRefCitation author="Kovacs, T. &amp; G. Vincon &amp; D. Muranyi &amp; I. Sivec" pageId="5" pageNumber="23" pagination="182 - 192" refId="ref3776" refString="Kovacs, T., G. Vincon, D. Muranyi &amp; I. Sivec. 2012. A new Perlodes species and its subspecies from the Balkan Peninsula (Plecoptera: Perlodidae). Illiesia, 8 (20): 182 - 192." type="journal article" year="2012">Kovács et al. 2012</bibRefCitation>
,
<figureCitation box="[899,979,1006,1030]" captionStart="Fig" captionStartId="3.[189,230,829,853]" captionTargetBox="[269,1362,235,761]" captionTargetId="figure-50@3.[261,1370,226,764]" captionTargetPageId="3" captionText="Fig. 3. P. thomasi sp. n. larva. a: head (5 mm), b: maxilla (1 mm), c: paraprocts (1 mm)." figureDoi="http://doi.org/10.5281/zenodo.4760846" httpUri="https://zenodo.org/record/4760846/files/figure.png" pageId="5" pageNumber="23">Fig. 3a</figureCitation>
). In both species the apex of the sclerite of the paraproct turns around the tip of the paraproct, just as in
<taxonomicName authorityName="Aubert" authorityYear="1946" box="[951,1091,1078,1101]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="jurassicus">
<emphasis box="[951,1091,1078,1101]" italics="true" pageId="5" pageNumber="23">P. jurassicus</emphasis>
</taxonomicName>
and to a lesser degree for
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="intricatus">
<emphasis italics="true" pageId="5" pageNumber="23">P. intricatus</emphasis>
</taxonomicName>
(
<bibRefCitation author="Lubini, V. &amp; Knispel, S. &amp; G. Vincon" box="[958,1183,1113,1137]" pageId="5" pageNumber="23" refId="ref3823" refString="Lubini, V., Knispel, S. &amp; G. Vincon. 2012. Les plecopteres de Suisse: identification et distribution. Fauna Helvetica 27, CSCF &amp; SEG," type="book" year="2012">Lubini et al. 2012</bibRefCitation>
, Vinçon, Murányi &amp; Kovács in prep.). In
<taxonomicName baseAuthorityName="Zwick" baseAuthorityYear="2011" box="[1063,1244,1148,1172]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="microcephalus">
<emphasis box="[1063,1244,1148,1172]" italics="true" pageId="5" pageNumber="23">P. microcephalus</emphasis>
</taxonomicName>
the sclerite of the paraproct ends in a rather long thin expansion that is just slightly curved at the tip, and in
<taxonomicName baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[1296,1396,1219,1243]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="dispar">
<emphasis box="[1296,1396,1219,1243]" italics="true" pageId="5" pageNumber="23">P. dispar</emphasis>
</taxonomicName>
the sclerite of the paraproct is very short with a rounded apex (
<bibRefCitation author="Marten, M." box="[902,1051,1291,1315]" pageId="5" pageNumber="23" pagination="387 - 401" refId="ref3858" refString="Marten, M. 1991. The effect of temperature on the egg incubation period of Perlodes microcephalus, Perlodes dispar (Plecoptera: Perlodidae) and Perla burmeisteriana (Plecoptera: Perlidae). In: Overview and Strategies of Ephemeroptera and Plecoptera (eds J. Alba-Tercedor and A. Sanchez- Ortega), Sandhill Crane Press, Gainesville. 387 - 401." type="book chapter" year="1991">Marten 1991</bibRefCitation>
,
<bibRefCitation author="Lubini, V. &amp; Knispel, S. &amp; G. Vincon" box="[1062,1268,1290,1315]" pageId="5" pageNumber="23" refId="ref3823" refString="Lubini, V., Knispel, S. &amp; G. Vincon. 2012. Les plecopteres de Suisse: identification et distribution. Fauna Helvetica 27, CSCF &amp; SEG," type="book" year="2012">Lubini et al. 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="23" type="etymology">
<paragraph blockId="5.[832,1444,971,1457]" pageId="5" pageNumber="23">
<emphasis bold="true" box="[832,963,1326,1350]" pageId="5" pageNumber="23">Etymology</emphasis>
. This species is named in honor of Prof. Alain Thomas (Toulouse) for his important contributions to the knowledge of Mediterranean aquatic Insects, especially Ephemeroptera.
</paragraph>
</subSubSection>
</treatment>
</document>
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