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<document id="5A78A0ABF0C7A98BEC1C99ADA667F68F" ID-DOI="10.1111/j.1096-3642.2005.00189.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5435260" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630434122956" checkinUser="diego" docAuthor="Daugeron, Christophe &amp; Grootaert, Patrick" docDate="2005" docId="039D7838FFCDFFD93CA9DAE7FECFFACF" docLanguage="en" docName="j.1096-3642.2005.00189.x.pdf" docOrigin="Zoological Journal of the Linnean Society 145 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2005.00189.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Empis (Coptophlebia) atratata Daugeron &amp; Grootaert 2003" docType="treatment" docVersion="4" lastPageNumber="360" masterDocId="FFA40040FFD8FFCF3F28DF3EFFC4FFD9" masterDocTitle="Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae)" masterLastPageNumber="391" masterPageNumber="339" pageNumber="360" updateTime="1699269823662" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="F0159365D773A4A9C8A702AE23745F78">Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae)</mods:title>
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<mods:namePart id="94A458DAE1A7DE22960D70F4A3CCAD71">Daugeron, Christophe</mods:namePart>
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<mods:namePart id="540077E2DF6EB771D7551E962C455840">Grootaert, Patrick</mods:namePart>
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<mods:date id="4366809A821EE4102AB2D4A05C62C340">2005</mods:date>
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<treatment id="039D7838FFCDFFD93CA9DAE7FECFFACF" ID-DOI="http://doi.org/10.5281/zenodo.5490722" ID-Zenodo-Dep="5490722" LSID="urn:lsid:plazi:treatment:039D7838FFCDFFD93CA9DAE7FECFFACF" httpUri="http://treatment.plazi.org/id/039D7838FFCDFFD93CA9DAE7FECFFACF" lastPageId="22" lastPageNumber="360" pageId="21" pageNumber="360">
<subSubSection id="C32E9AA5FFCDFFDA3CA9DAE7FC7FF943" pageId="21" pageNumber="360" type="nomenclature">
<paragraph id="8B8BC92EFFCDFFDA3CA9DAE7FA9FFA36" blockId="21.[897,1371,1497,1521]" box="[897,1371,1497,1521]" pageId="21" pageNumber="360">
<heading id="D0C37E42FFCDFFDA3CA9DAE7FA9FFA36" box="[897,1371,1497,1521]" centered="true" fontSize="9" level="2" pageId="21" pageNumber="360" reason="2">
<taxonomicName id="4C34B2ADFFCDFFDA3CA9DAE7FACEFA36" ID-CoL="8BZ3L" authorityName="Daugeron &amp; Grootaert" authorityYear="2003" box="[897,1290,1497,1521]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="363" phylum="Arthropoda" rank="species" species="atratata" subGenus="Coptophlebia">
<emphasis id="B940153CFFCDFFDA3CA9DAE7FC0EFA36" box="[897,970,1497,1520]" italics="true" pageId="21" pageNumber="360">EMPIS</emphasis>
(
<emphasis id="B940153CFFCDFFDA3CF3DAE7FB43FA36" box="[987,1159,1497,1521]" italics="true" pageId="21" pageNumber="360">COPTOPHLEBIA</emphasis>
)
<emphasis id="B940153CFFCDFFDA3BB0DAE2FACEFA36" box="[1176,1290,1500,1519]" italics="true" pageId="21" pageNumber="360">ATRATATA</emphasis>
</taxonomicName>
CLADE
</heading>
</paragraph>
<paragraph id="8B8BC92EFFCDFFDA3C12D937FC7FF943" blockId="21.[826,1442,1545,1690]" pageId="21" pageNumber="360">
<emphasis id="B940153CFFCDFFDA3C12D937FC7AF9C7" box="[826,958,1545,1566]" italics="true" pageId="21" pageNumber="360">Monophyly:</emphasis>
The monophyly of this clade is supported by two characters: epandrium with a dorsal double brush of spine-like bristles [character 30(1)], and epandrium with at least 1 very strong, long bristle at tip [32 (1)].
</paragraph>
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<subSubSection id="C32E9AA5FFCDFFD93C12D98CFECFFACF" lastPageId="22" lastPageNumber="361" pageId="21" pageNumber="360" type="distribution">
<paragraph id="8B8BC92EFFCDFFDA3C12D98CFC18F8DC" blockId="21.[826,1441,1714,1797]" pageId="21" pageNumber="360">
<emphasis id="B940153CFFCDFFDA3C12D98CFBF0F91E" box="[826,1076,1714,1735]" italics="true" pageId="21" pageNumber="360">
Distribution (
<figureCitation id="130FD5ABFFCDFFDA3CFFD98CFBE3F91E" box="[983,1063,1714,1735]" captionStart="Figure 16" captionStartId="24.[142,220,1274,1293]" captionTargetBox="[377,1185,198,1239]" captionTargetId="figure-285@24.[368,1200,191,1250]" captionTargetPageId="24" captionText="Figure 16. Distribution of the Empis (Coptophlebia) atratata clade: E. (C.) atratata Daugeron &amp; Grootaert = 2; E. (C.) kosametensis Daugeron &amp; Grootaert = 2; E. (C.) velutinella Frey = 1." figureDoi="http://doi.org/10.5281/zenodo.5435296" httpUri="https://zenodo.org/record/5435296/files/figure.png" pageId="21" pageNumber="360">Fig. 16</figureCitation>
):
</emphasis>
This group of three species is typically South-east Asian, recorded from
<collectingCountry id="F32389BEFFCDFFDA3A19D9EFFA65F93F" box="[1329,1441,1745,1766]" name="Myanmar" pageId="21" pageNumber="360">Myanmar</collectingCountry>
and
<collectingCountry id="F32389BEFFCDFFDA3C44D9D1FC13F8DC" box="[876,983,1775,1797]" name="Thailand" pageId="21" pageNumber="360">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="8B8BC92EFFCDFFD93C12D823FECFFACF" blockId="21.[826,1442,1821,1905]" lastBlockId="22.[142,758,1065,1302]" lastPageId="22" lastPageNumber="361" pageId="21" pageNumber="360">
<emphasis id="B940153CFFCDFFDA3C12D823FC62F8EB" box="[826,934,1821,1842]" italics="true" pageId="21" pageNumber="360">Remarks:</emphasis>
Although
<taxonomicName id="4C34B2ADFFCDFFDA3B19D820FA85F8EA" authority="Frey" authorityName="Frey" box="[1073,1345,1821,1843]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="360" phylum="Arthropoda" rank="species" species="velutinella" subGenus="Coptophlebia">
<emphasis id="B940153CFFCDFFDA3B19D820FB8EF8EA" box="[1073,1098,1822,1843]" italics="true" pageId="21" pageNumber="360">E.</emphasis>
(
<emphasis id="B940153CFFCDFFDA3B71D820FBB5F8EA" box="[1113,1137,1822,1843]" italics="true" pageId="21" pageNumber="360">C.</emphasis>
)
<emphasis id="B940153CFFCDFFDA3B57D823FB3CF8EB" box="[1151,1272,1821,1842]" italics="true" pageId="21" pageNumber="360">velutinella</emphasis>
Frey
</taxonomicName>
is not included in the phylogenetic analysis and not reviewed taxonomically, a preliminary morphological study allows inclusion of this species in the
<taxonomicName id="4C34B2ADFFCEFFD93FA6DB76FEFEFB84" authorityName="Daugeron &amp; Grootaert" authorityYear="2003" box="[142,314,1096,1117]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="361" phylum="Arthropoda" rank="species" species="atratata" subGenus="Coptophlebia">
<emphasis id="B940153CFFCEFFD93FA6DB76FF63FB84" box="[142,167,1096,1117]" italics="true" pageId="22" pageNumber="361">E.</emphasis>
(
<emphasis id="B940153CFFCEFFD93F9EDB76FF0AFB84" box="[182,206,1096,1117]" italics="true" pageId="22" pageNumber="361">C.</emphasis>
)
<emphasis id="B940153CFFCEFFD93FF4DB76FEFEFB84" box="[220,314,1096,1117]" italics="true" pageId="22" pageNumber="361">atratata</emphasis>
</taxonomicName>
clade as well as in the key given below. A fourth species from
<collectingCountry id="F32389BEFFCEFFD93ECDDB58FDEEFBA5" box="[485,554,1126,1148]" name="China" pageId="22" pageNumber="361">China</collectingCountry>
(
<collectingRegion id="49F007CCFFCEFFD93D14DB58FD53FBA5" box="[572,663,1126,1148]" country="China" name="Yunnan" pageId="22" pageNumber="361">Yunnan</collectingRegion>
) with 2 very strong, long bristles at the tip of the epandrium seems to also belong to this clade. It is known from
<specimenCount id="9D3202A7FFCEFFD93DC0DB9AFF02FB01" pageId="22" pageNumber="361" type="male">1 male</specimenCount>
and
<specimenCount id="9D3202A7FFCEFFD93E0ADBFCFE65FB01" box="[290,417,1218,1240]" pageId="22" pageNumber="361" type="female">2 females</specimenCount>
(KBINIRScNB) but not described here because of the poor condition of the specimens.
</paragraph>
</subSubSection>
</treatment>
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