treatments-xml/data/03/90/32/03903223FD6EFFC0FD0FD39E8842FE68.xml

<document id="1529787A8DB4501DFC54E9FA55F74DE2" ID-CLB-Dataset="8827" ID-DOI="10.5852/ejt.2021.731.1205" ID-GBIF-Dataset="d48c7b51-c9b4-49e7-8abf-1ecee8db36a1" ID-ISSN="2118-9773" ID-Zenodo-Dep="4446916" ID-ZooBank="CBF3D6E0-9896-4852-AA9F-6DD49C950795" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610975383020" checkinUser="plazi" docAuthor="Verhaegen, Gerlien &amp; Haase, Martin" docDate="2021" docId="03903223FD6EFFC0FD0FD39E8842FE68" docLanguage="en" docName="ejt-731_71-96.pdf" docOrigin="European Journal of Taxonomy 731" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Catapyrgus jami Verhaegen &amp; Haase 2021, sp. nov." docType="treatment" docUuid="98955097-7D04-4822-8408-30E37B79766B" docUuidSource="ZooBank" docVersion="7" lastPageNumber="82" masterDocId="FFA94A5BFD66FFCBFF8AD12E8D70FFB7" masterDocTitle="All-inclusive descriptions of new freshwater snail taxa of the hyperdiverse family Tateidae (Gastropoda, Caenogastropoda) from the South Island of New Zealand" masterLastPageNumber="96" masterPageNumber="71" pageNumber="79" updateTime="1698924542573" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="53324322DB8576B0BFEEA2B87B67812A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="44A82A84C1345E8B5CE627E15273E81E">
<mods:title id="AE6BCCE0C01A1796BB80D5DFECF8BA9B">All-inclusive descriptions of new freshwater snail taxa of the hyperdiverse family Tateidae (Gastropoda, Caenogastropoda) from the South Island of New Zealand</mods:title>
</mods:titleInfo>
<mods:name id="8FC692DE0C667A18633696C0B2266828" type="personal">
<mods:role id="F0939FB2BBD4748138671CDC13314AF1">
<mods:roleTerm id="8EC464FA07F16233C42441A3BD1ECF5C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F84C9C66A48BBAEE4F6AE3BD999D3CC">Verhaegen, Gerlien</mods:namePart>
<mods:nameIdentifier id="D792829C0A0A68AEFACD45C5760A402C" type="LSID">218491F7-1BE3-438C-8CAE-F11733A06D5C</mods:nameIdentifier>
<mods:affiliation id="D5866E6F8C1EBD07FB89237315795941">AG Vogelwarte, Zoological Institute and Museum, University of Greifswald, Soldmannstrasse 23, Greifswald, Germany; Present address: Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Institute for Extra-cutting-edge Science and Technology Avant-garde Research (X-star), Advanced Science-Technology Research (ASTER) Program, 2 - 15, Natsushimacho, Yokosuka, Kanagawa 237 - 0061, Japan.</mods:affiliation>
<mods:nameIdentifier id="581B8AD0D797B15060BC60D6C6DB1FE9" type="email">gerlienverhaegen@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="81CC57D5CA0738BB8A9442F7D6E520C9" type="personal">
<mods:role id="0A365286282DAD9E071CDF29597C750B">
<mods:roleTerm id="C52A82BC50F92E8A282EC2B6545E592A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D657F7CD59D86DF675D8AABF499CAAFC">Haase, Martin</mods:namePart>
<mods:nameIdentifier id="0FF6757344CAE6E0D609E3FC3E03D465" type="LSID">8CE4B347-14C5-464F-99C4-DE2263FC55D5</mods:nameIdentifier>
<mods:affiliation id="B9EB80B9F1BCA0E4B5B47C3C1C89246E">Vogelwarte, Zoological Institute and Museum, University of Greifswald, Soldmannstrasse 23, Greifswald, Germany.</mods:affiliation>
<mods:nameIdentifier id="662972EE4618AEAC1AE409BA855B5CAB" type="email">martin.haase@uni-greifswald.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="867136E5C68F59BF4A4F6EC0AC25452C">text</mods:typeOfResource>
<mods:relatedItem id="2EA5DAF9888C7F03966DAA27E6AF6D56" type="host">
<mods:titleInfo id="CE4030A2F1DCF78B0126779DF1EADC77">
<mods:title id="B2C048963668097C94983CD6C0FE58E2">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="D405B8BEE522C51BBA8582FEDF0518E4">
<mods:date id="C2F43F7B1E70E25D656D0622097924EE">2021</mods:date>
<mods:detail id="91ED85FD3FC42B7E91693660AE826EB9" type="pubDate">
<mods:number id="35CC79B9E925C1837E42E10567A4A53F">2021-01-18</mods:number>
</mods:detail>
<mods:detail id="2D94F32C42D18FA7075D9A025E8F19D6" type="volume">
<mods:number id="3602268352531310A9756850C6FB9EC8">731</mods:number>
</mods:detail>
<mods:extent id="6D94A8DAFF99345E19442A2B3DD42421" unit="page">
<mods:start id="94D0AB1923F550DDC98C2048EB33FBB5">71</mods:start>
<mods:end id="05D569B89E1AB978621E115BB9D7196E">96</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9579B6DBEF6BDDB4F4D781A7D413597B">journal article</mods:classification>
<mods:identifier id="CF8EF29B5844114EB0B98C5DDB7FD1F3" type="CLB-Dataset">8827</mods:identifier>
<mods:identifier id="E701BB77C2055B47EE7C0704FDD379ED" type="DOI">10.5852/ejt.2021.731.1205</mods:identifier>
<mods:identifier id="41F81F21070B0576B55EB93F1EA6FC3F" type="GBIF-Dataset">d48c7b51-c9b4-49e7-8abf-1ecee8db36a1</mods:identifier>
<mods:identifier id="C11D1A4851E2B3EE41FA9D97CB1F6050" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="C0B7C3FC5E52A3C9AC6E522C91BA4FB2" type="Zenodo-Dep">4446916</mods:identifier>
<mods:identifier id="595297CE6021512E86221A6AA5A62379" type="ZooBank">CBF3D6E0-9896-4852-AA9F-6DD49C950795</mods:identifier>
</mods:mods>
<treatment id="03903223FD6EFFC0FD0FD39E8842FE68" ID-DOI="http://doi.org/10.5281/zenodo.4446918" ID-GBIF-Taxon="177006642" ID-Zenodo-Dep="4446918" LSID="urn:lsid:zoobank.org:act:98955097-7D04-4822-8408-30E37B79766B" httpUri="http://treatment.plazi.org/id/03903223FD6EFFC0FD0FD39E8842FE68" lastPageId="11" lastPageNumber="82" pageId="8" pageNumber="79">
<subSubSection id="C323D0BEFD6EFFC3FD0FD39E8EDFFD7C" box="[645,943,688,715]" pageId="8" pageNumber="79" type="nomenclature">
<paragraph id="8B868335FD6EFFC3FD0FD39E8EDFFD7C" blockId="8.[383,1204,688,785]" box="[645,943,688,715]" pageId="8" pageNumber="79">
<heading id="D0CE3459FD6EFFC3FD0FD39E8EDFFD7C" box="[645,943,688,715]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="79" reason="2">
<taxonomicName id="4C39F8B6FD6EFFC3FD0FD39E8E3DFD7D" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[645,845,688,714]" class="Gastropoda" family="Hydrobiidae" genus="Catapyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="79" phylum="Mollusca" rank="species" species="jami" status="sp. nov.">
<emphasis id="B94D5F27FD6EFFC3FD0FD39E8E3DFD7D" bold="true" box="[645,845,688,714]" italics="true" pageId="8" pageNumber="79">Catapyrgus jami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6EFFC3FCDED39F8EDFFD7C" box="[852,943,689,715]" pageId="8" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C323D0BEFD6EFFC3FEF5D3FD897AFCA6" pageId="8" pageNumber="79" type="description">
<paragraph id="8B868335FD6EFFC3FEF5D3FD89C4FD59" blockId="8.[383,1204,688,785]" box="[383,1204,723,750]" pageId="8" pageNumber="79">
<uri id="FFA88F37FD6EFFC3FEF5D3FD89C4FD59" box="[383,1204,723,750]" pageId="8" pageNumber="79">
urn:lsid:zoobank.org:act: 
<uuid id="FF9FB9E0FD6EFFC3FD16D3FD89C4FD59" box="[668,1204,723,750]" pageId="8" pageNumber="79">98955097-7D04-4822-8408-30E37B79766B</uuid>
</uri>
</paragraph>
<paragraph id="8B868335FD6EFFC3FDA3D3D9897AFCA6" blockId="8.[383,1204,688,785]" box="[553,1034,758,785]" pageId="8" pageNumber="79">
<figureCitation id="13029FB0FD6EFFC3FDA3D3D98F02FCA6" box="[553,626,759,785]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[189,1398,1043,1871]" captionTargetId="figure-163@9.[189,1400,1043,1871]" captionTargetPageId="9" captionText="Fig. 4. Types. A–B. Catapyrgus jami sp. nov. C–D. Opacuincola lisannea sp. nov. (D from northern locality). E–F. Op. gretathunbergae sp. nov. G–H. Op. mete kahurangi subsp. nov. I–J. Obtusopyrgus farri sp. nov. A, C, E, G, I, holotypes, rest paratypes." figureDoi="http://doi.org/10.5281/zenodo.4446930" httpUri="https://zenodo.org/record/4446930/files/figure.png" pageId="8" pageNumber="79">Figs 4</figureCitation>
A–B, 
<figureCitation id="13029FB0FD6EFFC3FD32D3D88FAEFCA6" box="[696,734,758,785]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,962,1876]" captionTargetId="figure-214@10.[189,1399,962,1876]" captionTargetPageId="10" captionText="Fig. 5. Shells. SEM photographs of paratypes. A. Catapyrgus jami sp. nov. B–C. Opacuincola lisannea sp. nov. (B, topotype; C, northern locality).D–E. Op. gretathunbergae sp. nov.F–G. Op. mete kahurangi subsp. nov. H–I. Obtusopyrgus farri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446932" httpUri="https://zenodo.org/record/4446932/files/figure.png" pageId="8" pageNumber="79">5A</figureCitation>
, 
<figureCitation id="13029FB0FD6EFFC3FD61D3D98F8AFCA6" box="[747,762,759,785]" captionStart="Fig" captionStartId="12.[189,232,1017,1043]" captionTargetBox="[190,1399,266,946]" captionTargetId="figure-15@12.[190,1399,266,946]" captionTargetPageId="12" captionText="Fig. 6. Protoconch. SEM photographs of paratypes. A. Catapyrgus jami sp. nov. B–C. Opacuincola lisannea sp. nov. (B, topotype; C, northern locality). D. Op. gretathunbergae sp. nov. E. Op. mete kahurangi subsp. nov. F. Obtusopyrgus farri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446934" httpUri="https://zenodo.org/record/4446934/files/figure.png" pageId="8" pageNumber="79">6</figureCitation>
A, 
<figureCitation id="13029FB0FD6EFFC3FC97D3D98E5EFCA6" box="[797,814,759,785]" captionStart="Fig" captionStartId="12.[189,232,1998,2024]" captionTargetBox="[264,1332,1160,1938]" captionTargetId="figure-17@12.[264,1332,1160,1938]" captionTargetPageId="12" captionText="Fig. 7. Operculum of Catapyrgus jami sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446938" httpUri="https://zenodo.org/record/4446938/files/figure.png" pageId="8" pageNumber="79">7</figureCitation>
, 
<figureCitation id="13029FB0FD6EFFC3FCB0D3D98E38FCA6" box="[826,840,759,785]" captionStart="Fig" captionStartId="13.[189,232,1964,1990]" captionTargetBox="[189,1398,1079,1914]" captionTargetId="figure-275@13.[189,1398,1079,1914]" captionTargetPageId="13" captionText="Fig. 8. Radula. SEM photographs. A. Catapyrgus jami sp. nov. B. Opacuincola lisannea sp. nov. (from type locality). C. Op. gretathunbergae sp. nov. D. Obtusopyrgus farri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446940" httpUri="https://zenodo.org/record/4446940/files/figure.png" pageId="8" pageNumber="79">8</figureCitation>
A, 
<figureCitation id="13029FB0FD6EFFC3FCE7D3D98E0EFCA6" box="[877,894,759,785]" captionStart="Fig" captionStartId="14.[190,233,1178,1204]" captionTargetBox="[262,1268,567,1095]" captionTargetId="figure-124@14.[187,1397,541,1114]" captionTargetPageId="14" captionText="Fig. 9. Genitalia of Catapyrgus jami sp. nov. µCT scans. A. ♀. B. ³. Abbreviations: ac = anterior capsule gland; ag = albumen gland; bc = bursa copulatrix; od = oviduct; ov = ovary; pc = posterior capsule gland; pr = prostate; pv = pallial vas deferens; rs = receptaculum seminis; te = testis; vc = ventral channel; vd = vas deferens; vs = vesicula seminalis. Scale bars = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4446942" httpUri="https://zenodo.org/record/4446942/files/figure.png" pageId="8" pageNumber="79">9</figureCitation>
; Tables 
<tableCitation id="C6BBB68EFD6EFFC3FC54D3D98E98FCA6" box="[990,1000,759,785]" captionStart-0="Table 1" captionStart-1="Table 1" captionStartId-0="5.[189,258,265,291]" captionStartId-1="6.[189,258,265,291]" captionTargetBox-0="[200,1288,480,1976]" captionTargetBox-1="[200,1288,362,975]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Table 1 (continued on next page). Morphometrics of new taxa. A–E following the names refer to the localities in Figure 1. Abbreviations: AH = aperture height; AW = aperture width; BWW = bodywhorl width; CV = coefficient of variation corrected for unequal sample sizes; max = maximum; min = minimum; SD = standard deviation; SH = shell height; SW = shell width; W = number of whorls. Measurements in mm." captionText-1="Table 1. (continued)." httpUri-0="http://table.plazi.org/id/DF46D3BDFD63FFCEFF37D0278CB5FE07" httpUri-1="http://table.plazi.org/id/DF46D3BDFD60FFCDFF37D0278CC6FE93" pageId="8" pageNumber="79" tableUuid-0="DF46D3BDFD63FFCEFF37D0278CB5FE07" tableUuid-1="DF46D3BDFD60FFCDFF37D0278CC6FE93">1</tableCitation>
– 
<tableCitation id="C6BBB68EFD6EFFC3FC70D3D9897AFCA6" box="[1018,1034,759,785]" captionStart="Table 2" captionStartId="7.[189,258,265,291]" captionTargetBox="[200,1382,478,1976]" captionTargetPageId="7" captionText="Table 2. Diagnostic alignment positions. All characters are of type 1, i.e., they are fixed in a particular taxon. Positions refer to the alignments of COI and 16S rRNA in Electronic Supplements 1 and 2, respectively. Position 382 in bold is the only diagnostic position for Opacuincola gretathunbergae across all samples of the genus. Therefore, it was also compared to its sister species Op. ngatapuna. In Ob. farri, sequencing of COI failed." httpUri="http://table.plazi.org/id/DF46D3BDFD61FFCCFF37D0278F14FE07" pageId="8" pageNumber="79" tableUuid="DF46D3BDFD61FFCCFF37D0278F14FE07">2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C323D0BEFD6EFFC3FF37D2168FB9FC70" pageId="8" pageNumber="79" type="diagnosis">
<paragraph id="8B868335FD6EFFC3FF37D2168C46FCE5" blockId="8.[189,310,824,850]" box="[189,310,824,850]" pageId="8" pageNumber="79">
<heading id="D0CE3459FD6EFFC3FF37D2168C46FCE5" bold="true" box="[189,310,824,850]" fontSize="11" level="3" pageId="8" pageNumber="79" reason="3">
<emphasis id="B94D5F27FD6EFFC3FF37D2168C46FCE5" bold="true" box="[189,310,824,850]" pageId="8" pageNumber="79">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B868335FD6EFFC3FF37D2498FB9FC70" blockId="8.[189,1398,870,967]" pageId="8" pageNumber="79">
<taxonomicName id="4C39F8B6FD6EFFC3FF37D2498CF3FC37" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[189,387,871,897]" class="Gastropoda" family="Hydrobiidae" genus="Catapyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="79" phylum="Mollusca" rank="species" species="jami" status="sp. nov.">
<emphasis id="B94D5F27FD6EFFC3FF37D2498CF3FC37" box="[189,387,871,897]" italics="true" pageId="8" pageNumber="79">Catapyrgus jami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6EFFC3FE07D2498C9AFC36" box="[397,490,871,897]" pageId="8" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to 
<taxonomicName id="4C39F8B6FD6EFFC3FD5BD2498E0AFC37" authorityName="Haase" authorityYear="2008" box="[721,890,871,897]" class="Gastropoda" family="Hydrobiidae" genus="Catapyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="79" phylum="Mollusca" rank="species" species="matapango">
<emphasis id="B94D5F27FD6EFFC3FD5BD2498E0AFC37" box="[721,890,871,897]" italics="true" pageId="8" pageNumber="79">C. matapango</emphasis>
</taxonomicName>
, however but is, genetically distinct at 59 alignment positions. Furthermore, it differs from its congeners in the shape of the bursa copulatrix, which is globular rather than kidney-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C323D0BEFD6EFFC3FF37D2C18EACFBCA" pageId="8" pageNumber="79" type="etymology">
<paragraph id="8B868335FD6EFFC3FF37D2C18C34FBBE" blockId="8.[189,324,1007,1033]" box="[189,324,1007,1033]" pageId="8" pageNumber="79">
<heading id="D0CE3459FD6EFFC3FF37D2C18C34FBBE" bold="true" box="[189,324,1007,1033]" fontSize="11" level="3" pageId="8" pageNumber="79" reason="3">
<emphasis id="B94D5F27FD6EFFC3FF37D2C18C34FBBE" bold="true" box="[189,324,1007,1033]" pageId="8" pageNumber="79">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B868335FD6EFFC3FF37D5338EACFBCA" blockId="8.[189,1398,1053,1150]" pageId="8" pageNumber="79">
<taxonomicName id="4C39F8B6FD6EFFC3FF37D5338CF5FB80" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[189,389,1053,1079]" class="Gastropoda" family="Hydrobiidae" genus="Catapyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="79" phylum="Mollusca" rank="species" species="jami" status="sp. nov.">
<emphasis id="B94D5F27FD6EFFC3FF37D5338CF5FB80" box="[189,389,1053,1079]" italics="true" pageId="8" pageNumber="79">Catapyrgus jami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6EFFC3FE18D5308C82FB8F" box="[402,498,1054,1080]" pageId="8" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
is named after Jochen A. Modess, musician and composer who until his retirement from the university in 2019 has significantly shaped the cultural life of the city of Greifswald for over 25 years. The name is based on the initials of the dedicatee.
</paragraph>
</subSubSection>
<caption id="DF46D3BDFD6EFFC3FF34D6E38810F850" ID-DOI="http://doi.org/10.5281/zenodo.4446928" ID-Zenodo-Dep="4446928" box="[190,1376,1997,2024]" httpUri="https://zenodo.org/record/4446928/files/figure.png" pageId="8" pageNumber="79" startId="8.[190,233,1997,2023]" targetBox="[218,1368,1343,1904]" targetPageId="8">
<paragraph id="8B868335FD6EFFC3FF34D6E38810F850" blockId="8.[190,1376,1997,2024]" box="[190,1376,1997,2024]" pageId="8" pageNumber="79">
<emphasis id="B94D5F27FD6EFFC3FF34D6E38C7BF850" bold="true" box="[190,267,1997,2023]" pageId="8" pageNumber="79">Fig. 3.</emphasis>
Shell morphology. Principal component analysis based on five shell measurements of Table 1.
</paragraph>
</caption>
<subSubSection id="C323D0BEFD6FFFC2FF37D02689D4FDA8" pageId="9" pageNumber="80" type="materials_examined">
<paragraph id="8B868335FD6FFFC2FF37D0268CDAFE95" blockId="9.[189,426,264,290]" box="[189,426,264,290]" pageId="9" pageNumber="80">
<heading id="D0CE3459FD6FFFC2FF37D0268CDAFE95" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="9" pageNumber="80" reason="3">
<emphasis id="B94D5F27FD6FFFC2FF37D0268CDAFE95" bold="true" box="[189,426,264,290]" pageId="9" pageNumber="80">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B868335FD6FFFC2FF60D0198F4CFE0C" blockId="9.[189,1399,311,443]" pageId="9" pageNumber="80">
<materialsCitation id="3B518968FD6FFFC2FF60D0198F4CFE0C" ID-GBIF-Occurrence="3019435303" collectedFrom="on leaves, stones, woody debris in small spring" collectingDate="2016-02-28" collectionCode="NMNZ" collectorName="G. Verhaegen &amp; M. Haase" country="New Zealand" county="Kaituna Track" latitude="-40.710083" location="Kahurangi National Park" longLatPrecision="1" longitude="172.56009" municipality="Collingwood" pageId="9" pageNumber="80" specimenCode="NMNZ.M.330187" specimenCount="1" typeStatus="holotype">
<emphasis id="B94D5F27FD6FFFC2FF60D0198C2CFEE6" bold="true" box="[234,348,311,337]" pageId="9" pageNumber="80">
<typeStatus id="54823D97FD6FFFC2FF60D0198C2CFEE6" box="[234,348,311,337]" pageId="9" pageNumber="80" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="13029FB0FD6FFFC2FEE6D0198CB8FEE6" box="[364,456,311,337]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[189,1398,1043,1871]" captionTargetId="figure-163@9.[189,1400,1043,1871]" captionTargetPageId="9" captionText="Fig. 4. Types. A–B. Catapyrgus jami sp. nov. C–D. Opacuincola lisannea sp. nov. (D from northern locality). E–F. Op. gretathunbergae sp. nov. G–H. Op. mete kahurangi subsp. nov. I–J. Obtusopyrgus farri sp. nov. A, C, E, G, I, holotypes, rest paratypes." figureDoi="http://doi.org/10.5281/zenodo.4446930" httpUri="https://zenodo.org/record/4446930/files/figure.png" pageId="9" pageNumber="80">Fig. 4A</figureCitation>
) 
<collectingCountry id="F32EC3A5FD6FFFC2FF37D0758CE6FEC3" box="[189,406,346,373]" name="New Zealand" pageId="9" pageNumber="80">NEW ZEALAND</collectingCountry>
• 
<location id="8EE6D5EEFD6FFFC2FE33D0748F9CFEC3" LSID="urn:lsid:plazi:treatment:03903223FD6EFFC0FD0FD39E8842FE68:8EE6D5EEFD6FFFC2FE33D0748F9CFEC3" box="[441,748,346,372]" country="New Zealand" county="Kaituna Track" latitude="-40.710083" longLatPrecision="1" longitude="172.56009" municipality="Collingwood" name="Kahurangi National Park" pageId="9" pageNumber="80">Kahurangi National Park</location>
, 
<locationDeviation id="9A349A17FD6FFFC2FD77D07489C6FEC3" box="[765,1206,346,372]" pageId="9" pageNumber="80">SW of Collingwood, Kaituna Track</locationDeviation>
; 
<geoCoordinate id="EE0DE5F2FD6FFFC2FB4DD0778801FEC3" box="[1223,1393,345,373]" degrees="40" direction="south" minutes="42" orientation="latitude" pageId="9" pageNumber="80" precision="1" seconds="36.3" value="-40.710083">40°42ʹ36.3ʺ S</geoCoordinate>
, 
<geoCoordinate id="EE0DE5F2FD6FFFC2FF37D0528C06FE2F" box="[189,374,380,408]" degrees="172" direction="east" minutes="33" orientation="longitude" pageId="9" pageNumber="80" precision="1" seconds="36.3" value="172.56009">172°33ʹ36.3ʺ E</geoCoordinate>
; 
<date id="FF87A5F5FD6FFFC2FE0FD0508F59FE20" box="[389,553,381,408]" pageId="9" pageNumber="80" value="2016-02-28">
<collectingDate id="EFC35C1DFD6FFFC2FE0FD0508F59FE20" box="[389,553,381,408]" pageId="9" pageNumber="80" value="2016-02-28">28 Feb. 2016</collectingDate>
</date>
; 
<collectorName id="26CCE6E3FD6FFFC2FDB2D0508FABFE20" box="[568,731,381,408]" pageId="9" pageNumber="80">G. Verhaegen</collectorName>
and 
<collectorName id="26CCE6E3FD6FFFC2FC90D0508EFDFE2F" box="[794,909,382,408]" pageId="9" pageNumber="80">M. Haase</collectorName>
leg.; 
<collectedFrom id="3825C0C8FD6FFFC2FC5FD0508C21FE0C" pageId="9" pageNumber="80">on leaves, stones, woody debris in small spring</collectedFrom>
; 
<specimenCode id="DB9F2B4EFD6FFFC2FED7D08F8F47FE0C" box="[349,567,417,443]" pageId="9" pageNumber="80">NMNZ.M.330187</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B868335FD6FFFC2FF60D0CC8CABFE4B" blockId="9.[189,1188,482,544]" box="[234,475,482,508]" pageId="9" pageNumber="80">
<emphasis id="B94D5F27FD6FFFC2FF60D0CC8C17FE4B" bold="true" box="[234,359,482,508]" pageId="9" pageNumber="80">
<typeStatus id="54823D97FD6FFFC2FF60D0CC8C17FE4B" box="[234,359,482,508]" pageId="9" pageNumber="80" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<figureCitation id="13029FB0FD6FFFC2FEF2D0CC8CA2FE4B" box="[376,466,482,508]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[189,1398,1043,1871]" captionTargetId="figure-163@9.[189,1400,1043,1871]" captionTargetPageId="9" captionText="Fig. 4. Types. A–B. Catapyrgus jami sp. nov. C–D. Opacuincola lisannea sp. nov. (D from northern locality). E–F. Op. gretathunbergae sp. nov. G–H. Op. mete kahurangi subsp. nov. I–J. Obtusopyrgus farri sp. nov. A, C, E, G, I, holotypes, rest paratypes." figureDoi="http://doi.org/10.5281/zenodo.4446930" httpUri="https://zenodo.org/record/4446930/files/figure.png" pageId="9" pageNumber="80">Fig. 4B</figureCitation>
)
</paragraph>
<paragraph id="8B868335FD6FFFC2FF37D32889D4FDA8" blockId="9.[189,1188,482,544]" box="[189,1188,517,544]" pageId="9" pageNumber="80">
<materialsCitation id="3B518968FD6FFFC2FF37D32889D0FDA8" ID-GBIF-Occurrence="3019435306" box="[189,1184,517,544]" collectingDate="2016-02-28" collectionCode="NMNZ" collectorName="G. Verhaegen &amp; M. Haase" country="New Zealand" county="Kaituna Track" latitude="-40.710083" location="NEW ZEALAND" longLatPrecision="1" longitude="172.56009" municipality="Collingwood" pageId="9" pageNumber="80" specimenCode="NMNZ.M.330188" specimenCount="9" typeStatus="paratype">
<collectingCountry id="F32EC3A5FD6FFFC2FF37D3288CE1FDA8" box="[189,401,517,544]" name="New Zealand" pageId="9" pageNumber="80">NEW ZEALAND</collectingCountry>
• 
<specimenCount id="9D3F48BCFD6FFFC2FE20D32B8CC9FDA8" box="[426,441,517,543]" pageId="9" pageNumber="80">9</specimenCount>
specs; same collection data as for holotype; 
<specimenCode id="DB9F2B4EFD6FFFC2FC4FD32B89D0FDA8" box="[965,1184,517,543]" pageId="9" pageNumber="80">NMNZ.M.330188</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C323D0BEFD6FFFC1FF37D3688ECDFCB6" lastPageId="10" lastPageNumber="81" pageId="9" pageNumber="80" type="description">
<paragraph id="8B868335FD6FFFC2FF37D3688C3EFDD7" blockId="9.[189,334,582,608]" box="[189,334,582,608]" pageId="9" pageNumber="80">
<heading id="D0CE3459FD6FFFC2FF37D3688C3EFDD7" bold="true" box="[189,334,582,608]" fontSize="11" level="3" pageId="9" pageNumber="80" reason="3">
<emphasis id="B94D5F27FD6FFFC2FF37D3688C3EFDD7" bold="true" box="[189,334,582,608]" pageId="9" pageNumber="80">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B868335FD6FFFC2FF37D35B8F92FD4E" blockId="9.[189,1400,629,761]" pageId="9" pageNumber="80">
SHELL (
<figureCitation id="13029FB0FD6FFFC2FE9ED35B8C2FFD38" box="[276,351,629,655]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[189,1398,1043,1871]" captionTargetId="figure-163@9.[189,1400,1043,1871]" captionTargetPageId="9" captionText="Fig. 4. Types. A–B. Catapyrgus jami sp. nov. C–D. Opacuincola lisannea sp. nov. (D from northern locality). E–F. Op. gretathunbergae sp. nov. G–H. Op. mete kahurangi subsp. nov. I–J. Obtusopyrgus farri sp. nov. A, C, E, G, I, holotypes, rest paratypes." figureDoi="http://doi.org/10.5281/zenodo.4446930" httpUri="https://zenodo.org/record/4446930/files/figure.png" pageId="9" pageNumber="80">Figs 4</figureCitation>
A–B, 5A). Short turriform, about 1.7 times as high as than wide, white-translucent with light brown periostracum; protoconch with fine pits comprising 0.75 whorls (
<figureCitation id="13029FB0FD6FFFC2FBCDD3B689D6FD04" box="[1095,1190,664,691]" captionStart="Fig" captionStartId="12.[189,232,1017,1043]" captionTargetBox="[190,1399,266,946]" captionTargetId="figure-15@12.[190,1399,266,946]" captionTargetPageId="12" captionText="Fig. 6. Protoconch. SEM photographs of paratypes. A. Catapyrgus jami sp. nov. B–C. Opacuincola lisannea sp. nov. (B, topotype; C, northern locality). D. Op. gretathunbergae sp. nov. E. Op. mete kahurangi subsp. nov. F. Obtusopyrgus farri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446934" httpUri="https://zenodo.org/record/4446934/files/figure.png" pageId="9" pageNumber="80">Fig. 6A</figureCitation>
); entire shell with 3.875 to 4.25 whorls without structure apart from growth lines; umbilicus narrow; aperture orthocline, almost circular, only slightly higher than wide.
</paragraph>
<paragraph id="8B868335FD6FFFC2FF37D20E8C6CFCEA" blockId="9.[187,1398,800,861]" pageId="9" pageNumber="80">
OPERCULUM (
<figureCitation id="13029FB0FD6FFFC2FEDBD20E8CE5FC8C" box="[337,405,800,827]" captionStart="Fig" captionStartId="12.[189,232,1998,2024]" captionTargetBox="[264,1332,1160,1938]" captionTargetId="figure-17@12.[264,1332,1160,1938]" captionTargetPageId="12" captionText="Fig. 7. Operculum of Catapyrgus jami sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446938" httpUri="https://zenodo.org/record/4446938/files/figure.png" pageId="9" pageNumber="80">Fig. 7</figureCitation>
). Yellowish to light orange, paucispiral; with non-calcareous white peg in submarginal nucleus.
</paragraph>
<paragraph id="8B868335FD6FFFC2FF37D2AB8F70FC75" blockId="9.[189,1399,901,962]" pageId="9" pageNumber="80">EXTERNAL FEATURES. Epidermis without pigment; eyes small with bean-shaped pigment spot; tentacles without particular ciliation.</paragraph>
<caption id="DF46D3BDFD6FFFC2FF37D6A68E58F85E" ID-DOI="http://doi.org/10.5281/zenodo.4446930" ID-Zenodo-Dep="4446930" httpUri="https://zenodo.org/record/4446930/files/figure.png" pageId="9" pageNumber="80" startId="9.[189,232,1928,1954]" targetBox="[189,1398,1043,1871]" targetPageId="9">
<paragraph id="8B868335FD6FFFC2FF37D6A68E58F85E" blockId="9.[189,1399,1928,2025]" pageId="9" pageNumber="80">
<emphasis id="B94D5F27FD6FFFC2FF37D6A68C7DF815" bold="true" box="[189,269,1928,1954]" pageId="9" pageNumber="80">Fig. 4.</emphasis>
Types. 
<emphasis id="B94D5F27FD6FFFC2FEE4D6A68CD4F815" bold="true" box="[366,420,1928,1954]" pageId="9" pageNumber="80">A–B</emphasis>
. 
<taxonomicName id="4C39F8B6FD6FFFC2FE3FD6A68F0AF815" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[437,634,1928,1954]" class="Gastropoda" family="Hydrobiidae" genus="Catapyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="80" phylum="Mollusca" rank="species" species="jami" status="sp. nov.">
<emphasis id="B94D5F27FD6FFFC2FE3FD6A68F0AF815" box="[437,634,1928,1954]" italics="true" pageId="9" pageNumber="80">Catapyrgus jami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6FFFC2FD0ED6A68F91F815" box="[644,737,1928,1954]" pageId="9" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B94D5F27FD6FFFC2FD66D6A68E53F815" bold="true" box="[748,803,1928,1954]" pageId="9" pageNumber="80">C–D</emphasis>
. 
<taxonomicName id="4C39F8B6FD6FFFC2FCBED6A68948F815" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[820,1080,1928,1954]" class="Gastropoda" family="Hydrobiidae" genus="Opacuincola" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="80" phylum="Mollusca" rank="species" species="lisannea" status="sp. nov.">
<emphasis id="B94D5F27FD6FFFC2FCBED6A68948F815" box="[820,1080,1928,1954]" italics="true" pageId="9" pageNumber="80">Opacuincola lisannea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6FFFC2FBC8D6A689EFF815" box="[1090,1183,1928,1954]" pageId="9" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
(D from northern locality). 
<emphasis id="B94D5F27FD6FFFC2FEA4D6858C12F872" bold="true" box="[302,354,1963,1990]" pageId="9" pageNumber="80">E–F</emphasis>
. 
<taxonomicName id="4C39F8B6FD6FFFC2FEFBD6828F16F872" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[369,614,1963,1990]" class="Gastropoda" family="Hydrobiidae" genus="Opacuincola" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="80" phylum="Mollusca" rank="species" species="gretathunbergae" status="sp. nov.">
<emphasis id="B94D5F27FD6FFFC2FEFBD6828F16F872" box="[369,614,1963,1990]" italics="true" pageId="9" pageNumber="80">Op. gretathunbergae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6FFFC2FDE4D6828FB9F871" box="[622,713,1964,1990]" pageId="9" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B94D5F27FD6FFFC2FD5BD6858E7DF872" bold="true" box="[721,781,1963,1990]" pageId="9" pageNumber="80">G–H</emphasis>
. 
<taxonomicName id="4C39F8B6FD6FFFC2FC96D6828978F872" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[796,1032,1963,1990]" class="Gastropoda" family="Hydrobiidae" genus="Opacuincola" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="80" phylum="Mollusca" rank="subSpecies" species="mete" subSpecies="kahurangi">
<emphasis id="B94D5F27FD6FFFC2FC96D6828978F872" box="[796,1032,1963,1990]" italics="true" pageId="9" pageNumber="80">Op. mete kahurangi</emphasis>
</taxonomicName>
subsp. nov. 
<emphasis id="B94D5F27FD6FFFC2FB16D68589B4F872" bold="true" box="[1180,1220,1963,1990]" pageId="9" pageNumber="80">I–J</emphasis>
. 
<taxonomicName id="4C39F8B6FD6FFFC2FB59D6858D83F85F" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" class="Gastropoda" family="Tateidae" genus="Obtusopyrgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="80" phylum="Mollusca" rank="species" species="farri" status="sp. nov.">
<emphasis id="B94D5F27FD6FFFC2FB59D6858D83F85F" italics="true" pageId="9" pageNumber="80">Obtusopyrgus farri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6FFFC2FF70D6E18C25F85E" box="[250,341,1999,2025]" pageId="9" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
A, C, E, G, I, holotypes, rest paratypes.
</paragraph>
</caption>
<paragraph id="8B868335FD6CFFC1FF37D0278EA4FE94" blockId="10.[189,980,265,291]" box="[189,980,265,291]" pageId="10" pageNumber="81">MANTLE CAVITY. No ctenidial filaments; osphradium ovate-elongate.</paragraph>
<paragraph id="8B868335FD6CFFC1FF37D06589E7FE3F" blockId="10.[189,1398,330,392]" pageId="10" pageNumber="81">
DIGESTIVE SYSTEM. Radula has formula R 5-6 1 5-6/2 2, L 4-5 1 7, M1 25-29, M2 22-24 (
<figureCitation id="13029FB0FD6CFFC1FA8DD0648818FED2" box="[1287,1384,330,357]" captionStart="Fig" captionStartId="13.[189,232,1964,1990]" captionTargetBox="[189,1398,1079,1914]" captionTargetId="figure-275@13.[189,1398,1079,1914]" captionTargetPageId="13" captionText="Fig. 8. Radula. SEM photographs. A. Catapyrgus jami sp. nov. B. Opacuincola lisannea sp. nov. (from type locality). C. Op. gretathunbergae sp. nov. D. Obtusopyrgus farri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446940" httpUri="https://zenodo.org/record/4446940/files/figure.png" pageId="10" pageNumber="81">Fig. 8A</figureCitation>
); stomach without caecum; intestine makes narrow, long coil in roof of mantle cavity.
</paragraph>
<paragraph id="8B868335FD6CFFC1FF37D09E8E30FDE1" blockId="10.[188,1400,431,598]" pageId="10" pageNumber="81">
FEMALE GENITALIA (n = 3; 
<figureCitation id="13029FB0FD6CFFC1FE7FD0818F3EFE7E" box="[501,590,431,457]" captionStart="Fig" captionStartId="14.[190,233,1178,1204]" captionTargetBox="[262,1268,567,1095]" captionTargetId="figure-124@14.[187,1397,541,1114]" captionTargetPageId="14" captionText="Fig. 9. Genitalia of Catapyrgus jami sp. nov. µCT scans. A. ♀. B. ³. Abbreviations: ac = anterior capsule gland; ag = albumen gland; bc = bursa copulatrix; od = oviduct; ov = ovary; pc = posterior capsule gland; pr = prostate; pv = pallial vas deferens; rs = receptaculum seminis; te = testis; vc = ventral channel; vd = vas deferens; vs = vesicula seminalis. Scale bars = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4446942" httpUri="https://zenodo.org/record/4446942/files/figure.png" pageId="10" pageNumber="81">Fig 9A</figureCitation>
). Ovary short, simple sac not reaching stomach; renal oviduct first coiling 180° clockwise, then 270° counter-clockwise; one small, distal receptaculum seminis; bursa copulatrix globular, behind albumen gland, bursal duct entering ventrally; ovoviviparous, brooding at least one embryo in pallial oviduct, pallial oviduct as brood pouch with albumen gland as well as capsule gland with a larger posterior and short anterior section.
</paragraph>
<paragraph id="8B868335FD6CFFC1FF37D3508ECDFCB6" blockId="10.[189,1400,637,769]" pageId="10" pageNumber="81">
MALE GENITALIA (n = 1; 
<figureCitation id="13029FB0FD6CFFC1FE5ED3538F40FD20" box="[468,560,637,664]" captionStart="Fig" captionStartId="14.[190,233,1178,1204]" captionTargetBox="[262,1268,567,1095]" captionTargetId="figure-124@14.[187,1397,541,1114]" captionTargetPageId="14" captionText="Fig. 9. Genitalia of Catapyrgus jami sp. nov. µCT scans. A. ♀. B. ³. Abbreviations: ac = anterior capsule gland; ag = albumen gland; bc = bursa copulatrix; od = oviduct; ov = ovary; pc = posterior capsule gland; pr = prostate; pv = pallial vas deferens; rs = receptaculum seminis; te = testis; vc = ventral channel; vd = vas deferens; vs = vesicula seminalis. Scale bars = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4446942" httpUri="https://zenodo.org/record/4446942/files/figure.png" pageId="10" pageNumber="81">Fig. 9B</figureCitation>
). Testis sac with short lobes, starting more than one whorl below apex, comprises ca 0.5 whorl, anteriorly just reaching stomach; vesicula seminalis coils along anterior half of testis; proximal and distal vasa deferentia insert closely in about middle of flat pear-shaped prostate; penis slender, unclear from CT-scans whether it has a lobe or not.
</paragraph>
</subSubSection>
<caption id="DF46D3BDFD6CFFC1FF37D6A68FB9F85E" ID-DOI="http://doi.org/10.5281/zenodo.4446932" ID-Zenodo-Dep="4446932" httpUri="https://zenodo.org/record/4446932/files/figure.png" pageId="10" pageNumber="81" startId="10.[189,232,1928,1954]" targetBox="[189,1398,962,1876]" targetPageId="10">
<paragraph id="8B868335FD6CFFC1FF37D6A68FB9F85E" blockId="10.[189,1399,1928,2025]" pageId="10" pageNumber="81">
<emphasis id="B94D5F27FD6CFFC1FF37D6A68C7AF815" bold="true" box="[189,266,1928,1954]" pageId="10" pageNumber="81">Fig. 5.</emphasis>
Shells. SEM photographs of paratypes. 
<emphasis id="B94D5F27FD6CFFC1FD6AD6A68F85F815" bold="true" box="[736,757,1928,1954]" pageId="10" pageNumber="81">A</emphasis>
. 
<taxonomicName id="4C39F8B6FD6CFFC1FC8ED6A68EB7F815" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[772,967,1928,1954]" class="Gastropoda" family="Hydrobiidae" genus="Catapyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="81" phylum="Mollusca" rank="species" species="jami" status="sp. nov.">
<emphasis id="B94D5F27FD6CFFC1FC8ED6A68EB7F815" box="[772,967,1928,1954]" italics="true" pageId="10" pageNumber="81">Catapyrgus jami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6CFFC1FC44D6A68958F815" box="[974,1064,1928,1954]" pageId="10" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B94D5F27FD6CFFC1FBA4D6A68916F815" bold="true" box="[1070,1126,1928,1954]" pageId="10" pageNumber="81">B–C</emphasis>
. 
<taxonomicName id="4C39F8B6FD6CFFC1FBFFD6A68806F815" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[1141,1398,1928,1954]" class="Gastropoda" family="Hydrobiidae" genus="Opacuincola" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="81" phylum="Mollusca" rank="species" species="lisannea" status="sp. nov.">
<emphasis id="B94D5F27FD6CFFC1FBFFD6A68806F815" box="[1141,1398,1928,1954]" italics="true" pageId="10" pageNumber="81">Opacuincola lisannea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6CFFC1FF37D6828C66F871" box="[189,278,1964,1990]" pageId="10" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
(B, topotype; C, northern locality). 
<emphasis id="B94D5F27FD6CFFC1FD3AD6858F9AF872" bold="true" box="[688,746,1963,1990]" pageId="10" pageNumber="81">D–E</emphasis>
. 
<taxonomicName id="4C39F8B6FD6CFFC1FD7CD6828E98F872" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[758,1000,1963,1990]" class="Gastropoda" family="Hydrobiidae" genus="Opacuincola" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="81" phylum="Mollusca" rank="species" species="gretathunbergae" status="sp. nov.">
<emphasis id="B94D5F27FD6CFFC1FD7CD6828E98F872" box="[758,1000,1963,1990]" italics="true" pageId="10" pageNumber="81">Op. gretathunbergae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6CFFC1FC64D6828937F871" box="[1006,1095,1964,1990]" pageId="10" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B94D5F27FD6CFFC1FBC0D68589F3F872" bold="true" box="[1098,1155,1963,1990]" pageId="10" pageNumber="81">F–G</emphasis>
. 
<taxonomicName id="4C39F8B6FD6CFFC1FB1AD6828807F872" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[1168,1399,1963,1990]" class="Gastropoda" family="Hydrobiidae" genus="Opacuincola" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="81" phylum="Mollusca" rank="subSpecies" species="mete" subSpecies="kahurangi">
<emphasis id="B94D5F27FD6CFFC1FB1AD6828807F872" box="[1168,1399,1963,1990]" italics="true" pageId="10" pageNumber="81">Op. mete kahurangi</emphasis>
</taxonomicName>
subsp. nov. 
<emphasis id="B94D5F27FD6CFFC1FECDD6E08C08F85F" bold="true" box="[327,376,1998,2025]" pageId="10" pageNumber="81">H–I</emphasis>
. 
<taxonomicName id="4C39F8B6FD6CFFC1FE0DD6E08F18F85F" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[391,616,1998,2024]" class="Gastropoda" family="Tateidae" genus="Obtusopyrgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="81" phylum="Mollusca" rank="species" species="farri" status="sp. nov.">
<emphasis id="B94D5F27FD6CFFC1FE0DD6E08F18F85F" box="[391,616,1998,2024]" italics="true" pageId="10" pageNumber="81">Obtusopyrgus farri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6CFFC1FDE5D6E18FB9F85E" box="[623,713,1999,2025]" pageId="10" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="C323D0BEFD6DFFC0FF37D0278842FE68" pageId="11" pageNumber="82" type="discussion">
<paragraph id="8B868335FD6DFFC0FF37D0278C5FFE94" blockId="11.[189,303,265,291]" box="[189,303,265,291]" pageId="11" pageNumber="82">
<heading id="D0CE3459FD6DFFC0FF37D0278C5FFE94" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="11" pageNumber="82" reason="3">
<emphasis id="B94D5F27FD6DFFC0FF37D0278C5FFE94" bold="true" box="[189,303,265,291]" pageId="11" pageNumber="82">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B868335FD6DFFC0FF37D0198842FE68" blockId="11.[187,1398,311,479]" pageId="11" pageNumber="82">
<taxonomicName id="4C39F8B6FD6DFFC0FF37D0198CF3FEE6" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[189,387,311,337]" class="Gastropoda" family="Hydrobiidae" genus="Catapyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="82" phylum="Mollusca" rank="species" species="jami" status="sp. nov.">
<emphasis id="B94D5F27FD6DFFC0FF37D0198CF3FEE6" box="[189,387,311,337]" italics="true" pageId="11" pageNumber="82">Catapyrgus jami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6DFFC0FE00D0168C95FEE5" box="[394,485,312,338]" pageId="11" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to 
<taxonomicName id="4C39F8B6FD6DFFC0FD36D0198E14FEE6" authorityName="Haase" authorityYear="2008" box="[700,868,311,337]" class="Gastropoda" family="Hydrobiidae" genus="Catapyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="82" phylum="Mollusca" rank="species" species="matapango">
<emphasis id="B94D5F27FD6DFFC0FD36D0198E14FEE6" box="[700,868,311,337]" italics="true" pageId="11" pageNumber="82">C. matapango</emphasis>
</taxonomicName>
in terms of shell size and shape (
<figureCitation id="13029FB0FD6DFFC0FB61D0198843FEE6" box="[1259,1331,311,337]" captionStart="Fig" captionStartId="8.[190,233,1997,2023]" captionTargetBox="[218,1368,1343,1904]" captionTargetId="figure-199@8.[189,1399,1305,1953]" captionTargetPageId="8" captionText="Fig. 3. Shell morphology. Principal component analysis based on five shell measurements of Table 1." figureDoi="http://doi.org/10.5281/zenodo.4446928" httpUri="https://zenodo.org/record/4446928/files/figure.png" pageId="11" pageNumber="82">Fig. 3</figureCitation>
). The latter is the only other species of the genus for which sequence data exist. The divergence of the species was fairly large (
<figureCitation id="13029FB0FD6DFFC0FE08D0538CB9FE2F" box="[386,457,381,408]" captionStart="Fig" captionStartId="4.[189,232,1927,1953]" captionTargetBox="[223,1363,820,1877]" captionTargetId="figure-161@4.[186,1401,746,1891]" captionTargetPageId="4" captionText="Fig. 2. Phylogenetic analysis. Maximum likelihood tree with bootstrap support&gt; 50 / posterior probabilities from Bayesian analysis&gt; 0.50 for nodes. New taxa in bold. Letters A-E after new taxa refer to localities in Figure 1." figureDoi="http://doi.org/10.5281/zenodo.4446924" httpUri="https://zenodo.org/record/4446924/files/figure.png" pageId="11" pageNumber="82">Fig. 2</figureCitation>
) with a COI p-distance of 0.07, and 45 and 14 diagnostic characters in COI and 16S, respectively (
<tableCitation id="C6BBB68EFD6DFFC0FE13D08F8C86FE0C" box="[409,502,417,443]" captionStart="Table 2" captionStartId="7.[189,258,265,291]" captionTargetPageId="7" captionText="Table 2. Diagnostic alignment positions. All characters are of type 1, i.e., they are fixed in a particular taxon. Positions refer to the alignments of COI and 16S rRNA in Electronic Supplements 1 and 2, respectively. Position 382 in bold is the only diagnostic position for Opacuincola gretathunbergae across all samples of the genus. Therefore, it was also compared to its sister species Op. ngatapuna. In Ob. farri, sequencing of COI failed." httpUri="http://table.plazi.org/id/DF46D3BDFD61FFCCFF37D0278F14FE07" pageId="11" pageNumber="82" tableUuid="DF46D3BDFD61FFCCFF37D0278F14FE07">Table 2</tableCitation>
). The reduced eyes suggest that this species dwells in the transition zone of epigean and ground waters. The new species occurred sympatrically with 
<taxonomicName id="4C39F8B6FD6DFFC0FBBFD0EA89BEFE69" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[1077,1230,452,478]" class="Gastropoda" family="Hydrobiidae" genus="Opacuincola" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="82" phylum="Mollusca" rank="species" species="lisannea" status="sp. nov.">
<emphasis id="B94D5F27FD6DFFC0FBBFD0EA89BEFE69" box="[1077,1230,452,478]" italics="true" pageId="11" pageNumber="82">Op. lisannea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD6DFFC0FB5CD0EB8842FE68" box="[1238,1330,453,479]" pageId="11" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>