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<document id="3CB68F7171265A5FDA64CDCF512EB453" ID-DOI="10.11646/zootaxa.5569.3.7" ID-GBIF-Dataset="8b2f7d82-7f71-4a1e-b35e-b2fe914a33c2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734390" ID-ZooBank="B5C3024E-24CA-48D2-87A4-1ACAE8C849C0" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1737735340294" checkinUser="plazi" docAuthor="Thacker, Dimple, Myers, Alan A. &amp; Trivedi, Jigneshkumar N." docDate="2025" docId="038D87BCFFACFFEE5A9AFABB8C90F8DE" docLanguage="en" docName="zootaxa.5569.3.7.pdf" docOrigin="Zootaxa 5569 (3)" docSource="https://doi.org/10.11646/zootaxa.5569.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Audulla sindhu Thacker, Myers &amp; Trivedi, 2025, sp. nov." docType="treatment" docVersion="5" lastPageNumber="526" masterDocId="FFB4FFC4FFADFFEB5A0DFF808D11FF93" masterDocTitle="One new species and one new record of family Photidae Boeck, 1871 (Amphipoda: Senticaudata) from Gujarat, India" masterLastPageNumber="530" masterPageNumber="521" pageNumber="522" updateTime="1738075966129" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="F859A485810B9D87B3A8992ABBCB148F">One new species and one new record of family Photidae Boeck, 1871 (Amphipoda: Senticaudata) from Gujarat, India</mods:title>
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<mods:namePart id="7664F0069556C10D1799439563CFA8BD">Thacker, Dimple</mods:namePart>
<mods:nameIdentifier id="957EDDCF677BE127224E31584D4724F7" type="ORCID">0000-0003-3101-0028</mods:nameIdentifier>
<mods:affiliation id="4AECDA8534EFA439591FD79797C23A42">Department of Life sciences, Hemchandracharya North Gujarat University, Patan- 384265, Gujarat, India.</mods:affiliation>
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<mods:namePart id="79276F79313FAB5310CD19245A37A758">Myers, Alan A.</mods:namePart>
<mods:nameIdentifier id="BE1A9E858478258F3C15CE68AC3B8A91" type="ORCID">0000-0003-3256-2123</mods:nameIdentifier>
<mods:affiliation id="50B0CAFC212CB216157AD95E4198F365">School of Biological, Earth and Environmental Sciences, University College Cork, Cork Enterprise Centre, Distillery Fields, North Mall, Cork, Ireland.</mods:affiliation>
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<mods:namePart id="BDE60BB42B1F009745C9CAE4E605DFA0">Trivedi, Jigneshkumar N.</mods:namePart>
<mods:affiliation id="5AF17F73FA2F625A270475720FE84C8B">Department of Life sciences, Hemchandracharya North Gujarat University, Patan- 384265, Gujarat, India.</mods:affiliation>
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<treatment id="038D87BCFFACFFEE5A9AFABB8C90F8DE" ID-DOI="http://doi.org/10.5281/zenodo.14734408" ID-GBIF-Taxon="252879530" ID-Zenodo-Dep="14734408" LSID="urn:lsid:plazi:treatment:038D87BCFFACFFEE5A9AFABB8C90F8DE" httpUri="http://treatment.plazi.org/id/038D87BCFFACFFEE5A9AFABB8C90F8DE" lastPageId="5" lastPageNumber="526" pageId="1" pageNumber="522" scope_family="Photidae" scope_order="Amphipoda" scope_subOrder="Senticaudata" scope_superFamily="Photoidea">
<subSubSection id="C33E6521FFACFFEA5A9AFABB8CA9FAC6" box="[151,440,1339,1366]" pageId="1" pageNumber="522" type="nomenclature">
<paragraph id="8B9B36AAFFACFFEA5A9AFABB8CA9FAC6" blockId="1.[151,440,1339,1401]" box="[151,440,1339,1366]" pageId="1" pageNumber="522">
<heading id="D0D381C6FFACFFEA5A9AFABB8CA9FAC6" bold="true" box="[151,440,1339,1366]" fontSize="11" level="1" pageId="1" pageNumber="522" reason="1">
<emphasis id="B950EAB8FFACFFEA5A9AFABB8CA9FAC6" bold="true" box="[151,440,1339,1366]" pageId="1" pageNumber="522">
<taxonomicName id="4C244D29FFACFFEA5A9AFABB8C42FAC5" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2025" box="[151,339,1339,1366]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="sindhu" status="sp. nov.">
<emphasis id="B950EAB8FFACFFEA5A9AFABB8C42FAC5" bold="true" box="[151,339,1339,1366]" italics="true" pageId="1" pageNumber="522">Audulla sindhu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357C3FFACFFEA5B56FABB8CA9FAC6" box="[347,440,1339,1365]" pageId="1" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C33E6521FFACFFEA5A9AFADF8C53FAEA" box="[151,322,1375,1401]" pageId="1" pageNumber="522" type="description">
<paragraph id="8B9B36AAFFACFFEA5A9AFADF8C53FAEA" blockId="1.[151,440,1339,1401]" box="[151,322,1375,1401]" pageId="1" pageNumber="522">
(
<figureCitation id="131F2A2FFFACFFEA5A92FADF8C12FAEA" box="[159,259,1375,1401]" captionStart="FIGURE 1" captionStartId="2.[152,255,1874,1898]" captionTargetBox="[151,1435,381,1850]" captionTargetId="figure-125@2.[151,1435,381,1850]" captionTargetPageId="2" captionText="FIGURE 1. (A) Audulla sindhu sp. nov., male paratype (LFSC.ZRC-222): 5.0 mm; (B) Latigammaropsis photisimilis (Ruffo, 1969), male (LFSC.ZRC-224): 4.0 mm, Veraval (20°54'35&quot;N 70°21'09&quot;E) Gujarat, India. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14734392" httpUri="https://zenodo.org/record/14734392/files/figure.png" pageId="1" pageNumber="522">Figs. 1A</figureCitation>
,
<figureCitation id="131F2A2FFFACFFEA5B02FADF8C2BFAEA" box="[271,314,1375,1401]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[152,255,1943,1967]" captionStartId-1="4.[152,255,1943,1967]" captionTargetBox-0="[171,1413,185,1915]" captionTargetBox-1="[171,1417,185,1916]" captionTargetId-0="figure-18@3.[171,1417,181,1919]" captionTargetId-1="figure-22@4.[167,1420,181,1919]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 2. Audulla sindhu sp. nov., male holotype (LFSC.ZRC-221): 5.0 mm, Veraval (20°54'35&quot;N 70°21'09&quot;E), Gujarat, India. Scale bars = 0.25 mm. The first article of A2 peduncle is not drawn as it was damaged during dissection." captionText-1="FIGURE 3. Audulla sindhu sp. nov., male holotype (LFSC.ZRC-221): 5.0 mm, Veraval (20°54'35&quot;N 70°21'09&quot;E), Gujarat, India. Scale bars = 0.25 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14734394" figureDoi-1="http://doi.org/10.5281/zenodo.14734400" httpUri-0="https://zenodo.org/record/14734394/files/figure.png" httpUri-1="https://zenodo.org/record/14734400/files/figure.png" pageId="1" pageNumber="522">23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33E6521FFACFFEA5A9AFA248979FA2F" box="[151,1128,1443,1468]" pageId="1" pageNumber="522" type="reference_group">
<paragraph id="8B9B36AAFFACFFEA5A9AFA248979FA2F" blockId="1.[151,1128,1443,1468]" box="[151,1128,1443,1468]" pageId="1" pageNumber="522">
<treatmentCitation id="0A8510BBFFACFFEA5A9AFA248F95FA28" author="Sivaprakasam, T. E." box="[151,644,1443,1468]" page="566" pageId="1" pageNumber="522" year="1969">
<taxonomicName id="4C244D29FFACFFEA5A9AFA248F95FA28" ID-CoL="7RBB4" authority="Sivaprakasam, 1969: 564 - 566" authorityName="Sivaprakasam" authorityPageNumber="564 - 566" authorityYear="1969" box="[151,644,1443,1468]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFACFFEA5A9AFA248C59FA2F" box="[151,328,1444,1468]" italics="true" pageId="1" pageNumber="522">Audulla chelifera</emphasis>
<bibRefCitation id="EFB54B5BFFACFFEA5B43FA248F95FA28" author="Sivaprakasam, T. E." box="[334,644,1443,1468]" pageId="1" pageNumber="522" pagination="560 - 575" refId="ref4470" refString="Sivaprakasam, T. E. (1969) Amphipoda from the East Coast of India Part 2 Gammaridea and Caprellidea. Journal of Bombay Natural History Society, 66 (3), 560 - 575." type="journal article" year="1969">Sivaprakasam, 1969: 564566</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 7. (not
<taxonomicName id="4C244D29FFACFFEA590DFA24894DFA2F" ID-CoL="7RBB4" authority="Chevreux, 1901" authorityName="Chevreux" authorityYear="1901" box="[768,1116,1444,1468]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFACFFEA590DFA248EA1FA2F" box="[768,944,1444,1468]" italics="true" pageId="1" pageNumber="522">Audulla chelifera</emphasis>
<bibRefCitation id="EFB54B5BFFACFFEA59BBFA24894DFA2F" author="Chevreux, E." box="[950,1116,1444,1468]" pageId="1" pageNumber="522" pagination="388 - 438" refId="ref3611" refString="Chevreux, E. (1901) Crustaces Amphipodes. Mission scientifique de M. Ch. Alluaud aux Iles Sechelles (Mars, Avril, Mai 1892). Memoires de La Societe zoologique de France, 14, 388 - 438. [in French]" type="journal article" year="1901">Chevreux, 1901</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E6521FFACFFEA5A9AFA6A89C8F9BB" pageId="1" pageNumber="522" type="materials_examined">
<paragraph id="8B9B36AAFFACFFEA5A9AFA6A89C8F9BB" blockId="1.[151,1437,1513,2008]" pageId="1" pageNumber="522">
<emphasis id="B950EAB8FFACFFEA5A9AFA6A8C56F990" bold="true" box="[151,327,1513,1540]" pageId="1" pageNumber="522">Type material.</emphasis>
<materialsCitation id="3B4C3CF7FFACFFEA5B5CFA6A8C78F9BB" ID-GBIF-Occurrence="5051887302" collectingDate="2024-01-16" collectionCode="LFSC, R" collectorName="D. R. Thacker" latitude="20.909721" location="Veraval" longLatPrecision="21" longitude="70.3525" municipality="Veraval" pageId="1" pageNumber="522" specimenCode="LFSC.ZRC-221" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549F8808FFACFFEA5B5CFA6A8CADF997" box="[337,444,1514,1540]" pageId="1" pageNumber="522" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D22FD23FFACFFEA5BC4FA6A8F13F997" box="[457,514,1514,1540]" pageId="1" pageNumber="522" type="male">male</specimenCount>
, 5.0 mm (
<specimenCode id="DB829ED1FFACFFEA5874FA6A8E22F997" box="[633,819,1514,1540]" pageId="1" pageNumber="522">LFSC.ZRC-221</specimenCode>
),
<collectingMunicipality id="6BFFACD0FFACFFEA594AFA6A8E8CF997" box="[839,925,1514,1540]" pageId="1" pageNumber="522">Veraval</collectingMunicipality>
(
<geoCoordinate id="EE10506DFFACFFEA59A2FA6A8925F996" box="[943,1076,1514,1541]" degrees="20" direction="north" minutes="54" orientation="latitude" pageId="1" pageNumber="522" precision="15" seconds="35" value="20.909721">20°54'35&quot;N</geoCoordinate>
<geoCoordinate id="EE10506DFFACFFEA5E30FA6A89D1F997" box="[1085,1216,1514,1540]" degrees="70" direction="east" minutes="21" orientation="longitude" pageId="1" pageNumber="522" precision="15" seconds="09" value="70.3525">70°21'09&quot;E</geoCoordinate>
),
<date id="FF9A106AFFACFFEA5EDAFA6A8889F997" box="[1239,1432,1514,1540]" pageId="1" pageNumber="522" value="2024-01-16">
<collectingDate id="EFDEE982FFACFFEA5EDAFA6A8889F997" box="[1239,1432,1514,1540]" pageId="1" pageNumber="522" value="2024-01-16">16 January 2024</collectingDate>
</date>
, coll.
<collectorName id="26D1537CFFACFFEA5AC2F98E8C78F9BB" box="[207,361,1550,1576]" pageId="1" pageNumber="522">D.R. Thacker</collectorName>
</materialsCitation>
.
<materialsCitation id="3B4C3CF7FFACFFEA5B7EF98E89C4F9BB" ID-GBIF-Occurrence="5051887301" box="[371,1237,1550,1576]" collectingDate="2024-01-16" collectionCode="LFSC" collectorName="D. R. Thacker" latitude="20.909721" location="Veraval" longLatPrecision="21" longitude="70.3525" municipality="Veraval" pageId="1" pageNumber="522" specimenCode="LFSC.ZRC-222" specimenCount="8" specimenCount-male="8" typeStatus="paratype">
<typeStatus id="549F8808FFACFFEA5B7EF98E8CF2F9BB" box="[371,483,1550,1576]" pageId="1" pageNumber="522" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D22FD23FFACFFEA5BE3F98E8F56F9BB" box="[494,583,1550,1576]" count="8" pageId="1" pageNumber="522" type="male">8 males</specimenCount>
,
<quantity id="4CDC9B4FFFACFFEA585FF98E8FBEF9BB" box="[594,687,1550,1576]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="1" pageNumber="522" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
(
<specimenCode id="DB829ED1FFACFFEA58B0F98E8E66F9BB" box="[701,887,1550,1576]" pageId="1" pageNumber="522">LFSC.ZRC-222</specimenCode>
), data same as for the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E6521FFACFFEA5ACAF9B28DFAF9E3" pageId="1" pageNumber="522" type="etymology">
<paragraph id="8B9B36AAFFACFFEA5ACAF9B28DFAF9E3" blockId="1.[151,1437,1513,2008]" pageId="1" pageNumber="522">
<emphasis id="B950EAB8FFACFFEA5ACAF9B28C5FF9DF" bold="true" box="[199,334,1586,1612]" pageId="1" pageNumber="522">Etymology.</emphasis>
The specific epithet sindhu is derived from the historical name of Arabian Sea the “Sindhu Sagar”.
</paragraph>
</subSubSection>
<subSubSection id="C33E6521FFACFFEA5ACAF9F98E00F94F" pageId="1" pageNumber="522" type="distribution">
<paragraph id="8B9B36AAFFACFFEA5ACAF9F98E00F94F" blockId="1.[151,1437,1513,2008]" pageId="1" pageNumber="522">
<emphasis id="B950EAB8FFACFFEA5ACAF9F98CE2F900" bold="true" box="[199,499,1657,1684]" pageId="1" pageNumber="522">Distribution and habitat.</emphasis>
<taxonomicName id="4C244D29FFACFFEA5BF6F9FA8FB6F907" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2025" box="[507,679,1658,1684]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="sindhu" status="sp. nov.">
<emphasis id="B950EAB8FFACFFEA5BF6F9FA8FB6F907" box="[507,679,1658,1684]" italics="true" pageId="1" pageNumber="522">Audulla sindhu</emphasis>
</taxonomicName>
<emphasis id="B950EAB8FFACFFEA58A2F9FA8E18F907" bold="true" box="[687,777,1658,1684]" pageId="1" pageNumber="522">
<taxonomicNameLabel id="A26357C3FFACFFEA58A2F9FA8E18F907" box="[687,777,1658,1684]" pageId="1" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is reported from Cape Comorin and Pamban,
<collectingRegion id="49E0F848FFACFFEA5F1AF9FA8DC3F92B" country="India" name="Tamil Nadu" pageId="1" pageNumber="522">Tamil Nadu State</collectingRegion>
,
<collectingCountry id="F333763AFFACFFEA5AD0F91E8C06F92B" box="[221,279,1694,1720]" name="India" pageId="1" pageNumber="522">India</collectingCountry>
<bibRefCitation id="EFB54B5BFFACFFEA5B10F91E8F1CF92B" author="Sivaprakasam, T. E." box="[285,525,1694,1720]" pageId="1" pageNumber="522" pagination="560 - 575" refId="ref4470" refString="Sivaprakasam, T. E. (1969) Amphipoda from the East Coast of India Part 2 Gammaridea and Caprellidea. Journal of Bombay Natural History Society, 66 (3), 560 - 575." type="journal article" year="1969">Sivaprakasam (1969)</bibRefCitation>
, Veraval coast of
<collectingRegion id="49E0F848FFACFFEA58DCF91E8E75F92B" box="[721,868,1694,1720]" country="India" name="Gujarat" pageId="1" pageNumber="522">Gujarat State</collectingRegion>
(present study). In Veraval it was found among the algal beds spread along the intertidal region of the coast.
</paragraph>
</subSubSection>
<subSubSection id="C33E6521FFACFFEE5ACAF9658C64FB56" lastPageId="5" lastPageNumber="526" pageId="1" pageNumber="522" type="description">
<paragraph id="8B9B36AAFFACFFEA5ACAF9658F61F893" blockId="1.[151,1437,1513,2008]" box="[199,624,1765,1793]" pageId="1" pageNumber="522">
<emphasis id="B950EAB8FFACFFEA5ACAF9658C48F96C" bold="true" box="[199,345,1765,1791]" pageId="1" pageNumber="522">Description.</emphasis>
Based on male, 5.0 mm.
</paragraph>
<paragraph id="8B9B36AAFFACFFE95ACAF88A8E28FEF6" blockId="1.[151,1437,1513,2008]" lastBlockId="2.[151,1437,150,357]" lastPageId="2" lastPageNumber="523" pageId="1" pageNumber="522">
<emphasis id="B950EAB8FFACFFEA5ACAF88A8C1FF8B7" bold="true" box="[199,270,1802,1828]" pageId="1" pageNumber="522">Head.</emphasis>
<emphasis id="B950EAB8FFACFFEA5B1BF88B8C5AF8B7" box="[278,331,1803,1828]" italics="true" pageId="1" pageNumber="522">Eyes</emphasis>
medium.
<emphasis id="B950EAB8FFACFFEA5BB1F88A8F3EF8B7" box="[444,559,1802,1828]" italics="true" pageId="1" pageNumber="522">Antenna 1</emphasis>
subequal to antenna 2. Article 1 0.63 x as long as article 2; article 2 1.41 x as long as article 3; accessory flagellum 6 articulated.
<emphasis id="B950EAB8FFACFFEA58D1F8AE8E5EF8DB" box="[732,847,1838,1864]" italics="true" pageId="1" pageNumber="522">Antenna 2</emphasis>
peduncular article 4 subequal to article 5; flagellum paddle-shaped, 1.25 x as long as peduncular article 5.
<emphasis id="B950EAB8FFACFFEA58F6F8D28E45F8FF" box="[763,852,1874,1900]" italics="true" pageId="1" pageNumber="522">Labrum</emphasis>
broader at distal end and narrower at proximal end, distal margin with minute setae.
<emphasis id="B950EAB8FFACFFEA5803F8F68F75F803" box="[526,612,1910,1936]" italics="true" pageId="1" pageNumber="522">Labium</emphasis>
inner lobes well developed, oval in shape; outer lobes apically rounded covered with minute setae.
<emphasis id="B950EAB8FFACFFEA5BDDF81A8F6FF827" box="[464,638,1946,1972]" italics="true" pageId="1" pageNumber="522">Right mandible</emphasis>
with 5 dentate incisors; 4 dentate lacinia mobilis; accessory setal row with 8 setae, palp 2 articulate, article 1 upper middle part slightly swollen, article 2 apically setose;
<emphasis id="B950EAB8FFACFFEA5F0CF83E888DF84B" box="[1281,1436,1982,2008]" italics="true" pageId="1" pageNumber="522">Left mandible</emphasis>
with 5 dentate incisors, 3 dentate lacinia mobilis, accessory setal row with 8 setae.
<emphasis id="B950EAB8FFAFFFE95E55FF1789D6FF22" box="[1112,1223,151,177]" italics="true" pageId="2" pageNumber="523">Maxilla 1</emphasis>
inner plate with 6 long plumose setae and 3 small simple setae; outer plate with apical 9 dentate setae; palp 2 articulate, article 2 with several apex setae.
<emphasis id="B950EAB8FFAFFFE95B78FF5F8CF2FF6A" box="[373,483,223,249]" italics="true" pageId="2" pageNumber="523">Maxilla 2</emphasis>
inner plate broadened apically, with distinct oblique row of setae.
<emphasis id="B950EAB8FFAFFFE95EECFF5F884BFF6A" box="[1249,1370,223,249]" italics="true" pageId="2" pageNumber="523">Maxilliped</emphasis>
inner plate well developed, with several apical and marginal plumose setae; outer plate with 4 apical long, thick plumose setae, 9 marginal dentate spines of gradually decreasing size, several setae on the upper face; palp 4 articulated second article longest; article 4 smallest, apical claw absent.
</paragraph>
<caption id="DF5B6622FFAFFFE95A95F8D289EFF81E" ID-DOI="http://doi.org/10.5281/zenodo.14734392" ID-Zenodo-Dep="14734392" httpUri="https://zenodo.org/record/14734392/files/figure.png" pageId="2" pageNumber="523" startId="2.[152,255,1874,1898]" targetBox="[151,1435,381,1850]" targetPageId="2" targetType="figure">
<paragraph id="8B9B36AAFFAFFFE95A95F8D289EFF81E" blockId="2.[151,1437,1874,1934]" pageId="2" pageNumber="523">
<emphasis id="B950EAB8FFAFFFE95A95F8D28C36F8F9" bold="true" box="[152,295,1874,1898]" pageId="2" pageNumber="523">FIGURE 1. (</emphasis>
A)
<taxonomicName id="4C244D29FFAFFFE95B45F8D28CF2F8F9" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2025" box="[328,483,1874,1898]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="sindhu" status="sp. nov.">
<emphasis id="B950EAB8FFAFFFE95B45F8D28CF2F8F9" box="[328,483,1874,1898]" italics="true" pageId="2" pageNumber="523">Audulla sindhu</emphasis>
</taxonomicName>
<emphasis id="B950EAB8FFAFFFE95BE4F8D28F51F8F9" bold="true" box="[489,576,1874,1898]" pageId="2" pageNumber="523">
<taxonomicNameLabel id="A26357C3FFAFFFE95BE4F8D28F2DF8F9" box="[489,572,1874,1898]" pageId="2" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male paratype (LFSC.ZRC-222): 5.0 mm; (B)
<taxonomicName id="4C244D29FFAFFFE95E12F8D38DC5F81D" authority="(Ruffo, 1969)" baseAuthorityName="Ruffo" baseAuthorityYear="1969" class="Malacostraca" family="Photidae" genus="Latigammaropsis" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="photisimilis">
<emphasis id="B950EAB8FFAFFFE95E12F8D3885DF8F9" box="[1055,1356,1874,1898]" italics="true" pageId="2" pageNumber="523">Latigammaropsis photisimilis</emphasis>
(
<bibRefCitation id="EFB54B5BFFAFFFE95F57F8D28DDCF81D" author="Ruffo, S." pageId="2" pageNumber="523" pagination="1 - 77" refId="ref4392" refString="Ruffo, S. (1969) Studi sui Crostacei Anfipodi LXVII terzo contributo alla conoscenza degli Anfipodi del Mar Rosso. Memorie del Museo civico di storia naturale di Verona, 17, 1 - 77, 24 figs. [in Italian]" type="journal article" year="1969">Ruffo, 1969</bibRefCitation>
)
</taxonomicName>
, male (LFSC.ZRC-224): 4.0 mm, Veraval (20°54'35&quot;N 70°21'09&quot;E) Gujarat, India. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF5B6622FFAEFFE85A95F81789EDF847" ID-DOI="http://doi.org/10.5281/zenodo.14734394" ID-Zenodo-Dep="14734394" httpUri="https://zenodo.org/record/14734394/files/figure.png" pageId="3" pageNumber="524" startId="3.[152,255,1943,1967]" targetBox="[171,1413,185,1915]" targetPageId="3" targetType="figure">
<paragraph id="8B9B36AAFFAEFFE85A95F81789EDF847" blockId="3.[151,1437,1943,2004]" pageId="3" pageNumber="524">
<emphasis id="B950EAB8FFAEFFE85A95F8178C0BF83C" bold="true" box="[152,282,1943,1967]" pageId="3" pageNumber="524">FIGURE 2.</emphasis>
<taxonomicName id="4C244D29FFAEFFE85B2FF8188CAEF823" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2025" box="[290,447,1944,1968]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="sindhu" status="sp. nov.">
<emphasis id="B950EAB8FFAEFFE85B2FF8188CAEF823" box="[290,447,1944,1968]" italics="true" pageId="3" pageNumber="524">Audulla sindhu</emphasis>
</taxonomicName>
<emphasis id="B950EAB8FFAEFFE85BCBF8188F0EF823" bold="true" box="[454,543,1944,1968]" pageId="3" pageNumber="524">
<taxonomicNameLabel id="A26357C3FFAEFFE85BCBF8188F0AF823" box="[454,539,1944,1968]" pageId="3" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male holotype (LFSC.ZRC-221): 5.0 mm, Veraval (20°54'35&quot;N 70°21'09&quot;E), Gujarat, India. Scale bars = 0.25 mm. The first article of A2 peduncle is not drawn as it was damaged during dissection.
</paragraph>
</caption>
<caption id="DF5B6622FFA9FFEF5A95F8178CADF840" ID-DOI="http://doi.org/10.5281/zenodo.14734400" ID-Zenodo-Dep="14734400" httpUri="https://zenodo.org/record/14734400/files/figure.png" pageId="4" pageNumber="525" startId="4.[152,255,1943,1967]" targetBox="[171,1417,185,1916]" targetPageId="4" targetType="figure">
<paragraph id="8B9B36AAFFA9FFEF5A95F8178CADF840" blockId="4.[151,1437,1943,2004]" pageId="4" pageNumber="525">
<emphasis id="B950EAB8FFA9FFEF5A95F8178C0BF83C" bold="true" box="[152,282,1943,1967]" pageId="4" pageNumber="525">FIGURE 3.</emphasis>
<taxonomicName id="4C244D29FFA9FFEF5B2FF8188CAEF823" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2025" box="[290,447,1944,1968]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="sindhu" status="sp. nov.">
<emphasis id="B950EAB8FFA9FFEF5B2FF8188CAEF823" box="[290,447,1944,1968]" italics="true" pageId="4" pageNumber="525">Audulla sindhu</emphasis>
</taxonomicName>
<emphasis id="B950EAB8FFA9FFEF5BCBF8188F0EF823" bold="true" box="[454,543,1944,1968]" pageId="4" pageNumber="525">
<taxonomicNameLabel id="A26357C3FFA9FFEF5BCBF8188F0AF823" box="[454,539,1944,1968]" pageId="4" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male holotype (LFSC.ZRC-221): 5.0 mm, Veraval (20°54'35&quot;N 70°21'09&quot;E), Gujarat, India. Scale bars = 0.25 mm.
</paragraph>
</caption>
<paragraph id="8B9B36AAFFA8FFEE5ACAFF178E87FCAA" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="526">
<emphasis id="B950EAB8FFA8FFEE5ACAFF178C33FF22" bold="true" box="[199,290,151,177]" pageId="5" pageNumber="526">Pereon.</emphasis>
<emphasis id="B950EAB8FFA8FFEE5B26FF178CD1FF22" box="[299,448,151,177]" italics="true" pageId="5" pageNumber="526">Gnathopod 1</emphasis>
coxa subrectangular, ventral margin with small setae; basis 1.18 x as long as anterior margin of carpus; carpus with 5 tufts of setae on posterior margin and 3 tufts of setae on medial surface; propodus oval, subequal to anterior margin of carpus; palm subchelate, lined with unequal sized setae; dactylus subequal to palm.
<emphasis id="B950EAB8FFA8FFEE5AEDFE838C67FE8E" box="[224,374,259,285]" italics="true" pageId="5" pageNumber="526">Gnathopod 2</emphasis>
coxa subrectangular, 1.48 x as long as broad; basis both margins moderately setose; carpus triangular, anterodistal corner with 2 setae, posterodistal corner with a tuft of several setae. Propodus subrectangular with posterodistal corner produced, 1.26 x as long as basis, anterior margin with 3 tufts of setae on distal end, entire posterior margin with several tufts of setae, palm with a small protuberance at the base of the dactylus; dactylus longer than palm, slightly bulged above the base, posterior margin dentate.
<emphasis id="B950EAB8FFA8FFEE59F8FE138966FE3E" box="[1013,1143,403,429]" italics="true" pageId="5" pageNumber="526">Pereopod 3</emphasis>
coxa subrectangular with several small setae on ventral margin; basis rectangular with 4 small setae on posterior margin, anterior margin with several small and 2 medium sized setae; carpus subequal to propodus; dactylus 0.57 x as long as propodus.
<emphasis id="B950EAB8FFA8FFEE5A9AFE7F8C0AFD8A" box="[151,283,511,537]" italics="true" pageId="5" pageNumber="526">Pereopod 4</emphasis>
coxa subrectangular, ventral margin with 1 seta; basis rectangular with 2 small and 1 long seta on anterior margin, posterior margin with 3 medium sized setae, posterodistal corner with 3 medium sized setae; carpus subequal to propodus; dactylus 0.40 x as long as propodus.
<emphasis id="B950EAB8FFA8FFEE593AFDC78EA9FDF2" box="[823,952,583,609]" italics="true" pageId="5" pageNumber="526">Pereopod 5</emphasis>
coxa bilobed, basis narrower at distal half, anterior margin with 4 small robust setae, anterodistal corner with 1 long and one short setae; merus 1.30 x as long as carpus; propodus 2.39 x as long as carpus; dactylus 0.27 x as long as propodus.
<emphasis id="B950EAB8FFA8FFEE5E48FD0F89D7FD3B" box="[1093,1222,654,681]" italics="true" pageId="5" pageNumber="526">Pereopod 6</emphasis>
coxa ellipse; basis subrectangular with 4 small setae on anterior margin and 1 long robust seta on posterior margin, anterior margin with 4 small robust setae, anterodistal corner with 2 robust setae, posterior margin with few small setae, merus 1.27 x as long as carpus; propodus twice as long as carpus; dactylus small, 1/4 of the length of the propodus.
<emphasis id="B950EAB8FFA8FFEE5F11FD7B888DFC86" box="[1308,1436,763,789]" italics="true" pageId="5" pageNumber="526">Pereopod 7</emphasis>
similar to pereopod 6 except, last 2 articles are in reversed direction.
</paragraph>
<paragraph id="8B9B36AAFFA8FFEE5ACAFCC38C64FB56" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="526">
<emphasis id="B950EAB8FFA8FFEE5ACAFCC38C00FCCE" bold="true" box="[199,273,835,861]" pageId="5" pageNumber="526">Pleon.</emphasis>
<emphasis id="B950EAB8FFA8FFEE5B16FCC38CB1FCCE" box="[283,416,835,861]" italics="true" pageId="5" pageNumber="526">Epimeron 1</emphasis>
with almost straight ventral margin, posteroventral corner developed into a small notch, bearing 1 small setae.
<emphasis id="B950EAB8FFA8FFEE5B80FCE78F1FFC12" box="[397,526,871,897]" italics="true" pageId="5" pageNumber="526">Epimeron 2</emphasis>
ventral margin rounded, posterodistal corner produced into a small notch, bearing 1 small setae.
<emphasis id="B950EAB8FFA8FFEE5B4DFC0B8CD7FC36" box="[320,454,907,933]" italics="true" pageId="5" pageNumber="526">Epimeron 3</emphasis>
ventral margin rounded, posterodistal corner produced into a small notch, bearing 1 small setae, posterior margin with 1 medium sized setae located on the middle.
<emphasis id="B950EAB8FFA8FFEE5E25FC2F8984FC5A" box="[1064,1173,943,969]" italics="true" pageId="5" pageNumber="526">Uropod 1</emphasis>
peduncle inner margin with 4 small robust setae with 1 long robust seta on distal corner; outer margin with 3 small robust setae and with 1 medium robust seta on distal corner, inter-ramus spur reaches 1/3 of the endopod; endopod as long as peduncle, outer margin with 4 small robust setae, inner margin bare; exopod 0.87 x as long as endopod.
<emphasis id="B950EAB8FFA8FFEE5ECEFB9B883EFBA6" box="[1219,1327,1051,1077]" italics="true" pageId="5" pageNumber="526">Uropod 2</emphasis>
peduncle subequal to exopod, inner margin with 3 small robust setae and 1 long robust seta at distal corner, outer margin with 3 small robust setae; exopod 1.2 x as long as endopod.
<emphasis id="B950EAB8FFA8FFEE5909FBE38E7EFBEE" box="[772,879,1123,1149]" italics="true" pageId="5" pageNumber="526">Uropod 3</emphasis>
peduncle 1.14 x as long as subequal rami.
<taxonomicName id="4C244D29FFA8FFEE5F59FBE3888DFBEE" authorityName="Pearse" authorityYear="1952" box="[1364,1436,1123,1149]" class="Copepoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="5" pageNumber="526" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAB8FFA8FFEE5F59FBE3888DFBEE" box="[1364,1436,1123,1149]" italics="true" pageId="5" pageNumber="526">Telson</emphasis>
</taxonomicName>
as long as broad, with a triangular notch on the distal margin, each arm with 1 small robust seta, 2 hair line setae on each lateral margin.
</paragraph>
</subSubSection>
<subSubSection id="C33E6521FFA8FFEE5ACAFB4F8C90F8DE" pageId="5" pageNumber="526" type="discussion">
<paragraph id="8B9B36AAFFA8FFEE5ACAFB4F8F7EF9BE" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="526">
<emphasis id="B950EAB8FFA8FFEE5ACAFB4F8C2AFB7A" bold="true" box="[199,315,1231,1257]" pageId="5" pageNumber="526">Remarks.</emphasis>
<taxonomicName id="4C244D29FFA8FFEE5B49FB4F8CE1FB7A" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2025" box="[324,496,1231,1257]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="sindhu" status="sp. nov.">
<emphasis id="B950EAB8FFA8FFEE5B49FB4F8CE1FB7A" box="[324,496,1231,1257]" italics="true" pageId="5" pageNumber="526">Audulla sindhu</emphasis>
</taxonomicName>
<emphasis id="B950EAB8FFA8FFEE5BF5FB4F8F42FB7A" bold="true" box="[504,595,1231,1257]" pageId="5" pageNumber="526">
<taxonomicNameLabel id="A26357C3FFA8FFEE5BF5FB4F8F42FB7A" box="[504,595,1231,1257]" pageId="5" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is classified under a genus
<taxonomicName id="4C244D29FFA8FFEE599DFB4F89B7FB7A" authority="Chevreux, 1901" authorityName="Chevreux" authorityYear="1901" box="[912,1190,1231,1257]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAB8FFA8FFEE599DFB4F8EF8FB7A" box="[912,1001,1231,1257]" italics="true" pageId="5" pageNumber="526">Audulla</emphasis>
<bibRefCitation id="EFB54B5BFFA8FFEE59FCFB4F89B7FB7A" author="Chevreux, E." box="[1009,1190,1231,1257]" pageId="5" pageNumber="526" pagination="388 - 438" refId="ref3611" refString="Chevreux, E. (1901) Crustaces Amphipodes. Mission scientifique de M. Ch. Alluaud aux Iles Sechelles (Mars, Avril, Mai 1892). Memoires de La Societe zoologique de France, 14, 388 - 438. [in French]" type="journal article" year="1901">Chevreux, 1901</bibRefCitation>
</taxonomicName>
as it is having robust and elongated peduncles of both pairs of antennae, antenna 2 flagellum paddle-shaped, mandible palp robust and maxilla 1 inner plate obliquely truncate. But
<taxonomicName id="4C244D29FFA8FFEE5885FA978FE3FAA2" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2025" box="[648,754,1303,1329]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="sindhu" status="sp. nov.">
<emphasis id="B950EAB8FFA8FFEE5885FA978FE3FAA2" box="[648,754,1303,1329]" italics="true" pageId="5" pageNumber="526">A. sindhu</emphasis>
</taxonomicName>
<emphasis id="B950EAB8FFA8FFEE58F5FA978E40FAA2" bold="true" box="[760,849,1303,1329]" pageId="5" pageNumber="526">
<taxonomicNameLabel id="A26357C3FFA8FFEE58F5FA978E40FAA2" box="[760,849,1303,1329]" pageId="5" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from its only congener
<taxonomicName id="4C244D29FFA8FFEE5F49FA978CAEFAC6" authority="Chevreux (1901)" authorityName="Chevreux" authorityYear="1901" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFA8FFEE5F49FA978DEAFAC6" italics="true" pageId="5" pageNumber="526">Audulla chelifera</emphasis>
<bibRefCitation id="EFB54B5BFFA8FFEE5B0CFABB8CAEFAC6" author="Chevreux, E." box="[257,447,1339,1365]" pageId="5" pageNumber="526" pagination="388 - 438" refId="ref3611" refString="Chevreux, E. (1901) Crustaces Amphipodes. Mission scientifique de M. Ch. Alluaud aux Iles Sechelles (Mars, Avril, Mai 1892). Memoires de La Societe zoologique de France, 14, 388 - 438. [in French]" type="journal article" year="1901">Chevreux (1901)</bibRefCitation>
</taxonomicName>
in the following characters: antenna 2 flagellum has 12 articles (fig. 2, A2) (vs. 8 articles in
<taxonomicName id="4C244D29FFA8FFEE5AB8FADF8CE7FAEA" authority="Chevreux, 1901" authorityName="Chevreux" authorityYear="1901" box="[181,502,1375,1401]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFA8FFEE5AB8FADF8C28FAEA" box="[181,313,1375,1401]" italics="true" pageId="5" pageNumber="526">A. chelifera</emphasis>
<bibRefCitation id="EFB54B5BFFA8FFEE5B4DFADF8CE7FAEA" author="Chevreux, E." box="[320,502,1375,1401]" pageId="5" pageNumber="526" pagination="388 - 438" refId="ref3611" refString="Chevreux, E. (1901) Crustaces Amphipodes. Mission scientifique de M. Ch. Alluaud aux Iles Sechelles (Mars, Avril, Mai 1892). Memoires de La Societe zoologique de France, 14, 388 - 438. [in French]" type="journal article" year="1901">Chevreux, 1901</bibRefCitation>
</taxonomicName>
, fig. 57), maxilla 1 inner plate has 6 long setae and 3 short setae (fig. 2, Mx1) (vs. 8 long setae in
<taxonomicName id="4C244D29FFA8FFEE5B43FA038F86FA0E" authority="Chevreux, 1901" authorityName="Chevreux" authorityYear="1901" box="[334,663,1411,1437]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFA8FFEE5B43FA038CC4FA0E" box="[334,469,1411,1437]" italics="true" pageId="5" pageNumber="526">A. chelifera</emphasis>
<bibRefCitation id="EFB54B5BFFA8FFEE5BD2FA038F86FA0E" author="Chevreux, E." box="[479,663,1411,1437]" pageId="5" pageNumber="526" pagination="388 - 438" refId="ref3611" refString="Chevreux, E. (1901) Crustaces Amphipodes. Mission scientifique de M. Ch. Alluaud aux Iles Sechelles (Mars, Avril, Mai 1892). Memoires de La Societe zoologique de France, 14, 388 - 438. [in French]" type="journal article" year="1901">Chevreux, 1901</bibRefCitation>
</taxonomicName>
, fig. 59), maxilliped outer plate with 9 marginal dentate setae and several setae on upper face and (vs. outer plate with 8 marginal dentate setae and upper face bare in in
<taxonomicName id="4C244D29FFA8FFEE5F14FA278C5DFA76" authority="Chevreux, 1901" authorityName="Chevreux" authorityYear="1901" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFA8FFEE5F14FA27888CFA52" box="[1305,1437,1447,1473]" italics="true" pageId="5" pageNumber="526">A. chelifera</emphasis>
<bibRefCitation id="EFB54B5BFFA8FFEE5A9AFA4B8C5DFA76" author="Chevreux, E." box="[151,332,1483,1509]" pageId="5" pageNumber="526" pagination="388 - 438" refId="ref3611" refString="Chevreux, E. (1901) Crustaces Amphipodes. Mission scientifique de M. Ch. Alluaud aux Iles Sechelles (Mars, Avril, Mai 1892). Memoires de La Societe zoologique de France, 14, 388 - 438. [in French]" type="journal article" year="1901">Chevreux, 1901</bibRefCitation>
</taxonomicName>
, fig. 60); gnathopod 2 propodus, sparsely setose and 1.8 x as long as broad (fig. 3, G2) (vs. densely setose and 1.5 x as long as broad in
<taxonomicName id="4C244D29FFA8FFEE5827FA6F8E78F99A" authority="Chevreux, 1901" authorityName="Chevreux" authorityYear="1901" box="[554,873,1519,1545]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFA8FFEE5827FA6F8FBCF99A" box="[554,685,1519,1545]" italics="true" pageId="5" pageNumber="526">A. chelifera</emphasis>
<bibRefCitation id="EFB54B5BFFA8FFEE58B9FA6F8E78F99A" author="Chevreux, E." box="[692,873,1519,1545]" pageId="5" pageNumber="526" pagination="388 - 438" refId="ref3611" refString="Chevreux, E. (1901) Crustaces Amphipodes. Mission scientifique de M. Ch. Alluaud aux Iles Sechelles (Mars, Avril, Mai 1892). Memoires de La Societe zoologique de France, 14, 388 - 438. [in French]" type="journal article" year="1901">Chevreux, 1901</bibRefCitation>
</taxonomicName>
, fig. 62), uropod 1 inter-ramus spur reaches 1/3 of the endopod (vs. inter-ramus spur absent).
</paragraph>
<paragraph id="8B9B36AAFFA8FFEE5ACAF9B78C90F8DE" blockId="5.[151,1437,151,1869]" pageId="5" pageNumber="526">
<bibRefCitation id="EFB54B5BFFA8FFEE5ACAF9B78CAFF9C2" author="Sivaprakasam, T. E." box="[199,446,1591,1617]" pageId="5" pageNumber="526" pagination="560 - 575" refId="ref4470" refString="Sivaprakasam, T. E. (1969) Amphipoda from the East Coast of India Part 2 Gammaridea and Caprellidea. Journal of Bombay Natural History Society, 66 (3), 560 - 575." type="journal article" year="1969">Sivaprakasam (1969)</bibRefCitation>
recorded
<taxonomicName id="4C244D29FFA8FFEE5835F9B78FD0F9C2" authorityName="Chevreux" authorityYear="1901" box="[568,705,1591,1617]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFA8FFEE5835F9B78FD0F9C2" box="[568,705,1591,1617]" italics="true" pageId="5" pageNumber="526">A. chelifera</emphasis>
</taxonomicName>
on the basis of specimens collected from Cape Comorin and Pamban located in the
<collectingRegion id="49E0F848FFA8FFEE5B9CF9DB8F42F9E6" box="[401,595,1627,1653]" country="India" name="Tamil Nadu" pageId="5" pageNumber="526">Tamil Nadu State</collectingRegion>
of
<collectingCountry id="F333763AFFA8FFEE5878F9DB8FA5F9E6" box="[629,692,1627,1653]" name="India" pageId="5" pageNumber="526">India</collectingCountry>
.The description and figures of
<taxonomicName id="4C244D29FFA8FFEE5E02F9DB8983F9E6" authorityName="Chevreux" authorityYear="1901" box="[1039,1170,1627,1653]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFA8FFEE5E02F9DB8983F9E6" box="[1039,1170,1627,1653]" italics="true" pageId="5" pageNumber="526">A. chelifera</emphasis>
</taxonomicName>
given by
<bibRefCitation id="EFB54B5BFFA8FFEE5EF1F9DB8DF2F90A" author="Sivaprakasam, T. E." pageId="5" pageNumber="526" pagination="560 - 575" refId="ref4470" refString="Sivaprakasam, T. E. (1969) Amphipoda from the East Coast of India Part 2 Gammaridea and Caprellidea. Journal of Bombay Natural History Society, 66 (3), 560 - 575." type="journal article" year="1969">Sivaprakasam (1969)</bibRefCitation>
shows similarities with
<taxonomicName id="4C244D29FFA8FFEE5BE2F9FF8F49F90A" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2025" box="[495,600,1663,1689]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="sindhu" status="sp. nov.">
<emphasis id="B950EAB8FFA8FFEE5BE2F9FF8F49F90A" box="[495,600,1663,1689]" italics="true" pageId="5" pageNumber="526">A. sindhu</emphasis>
</taxonomicName>
<emphasis id="B950EAB8FFA8FFEE5850F9FF8FA7F90A" bold="true" box="[605,694,1663,1689]" pageId="5" pageNumber="526">
<taxonomicNameLabel id="A26357C3FFA8FFEE5850F9FF8FA7F90A" box="[605,694,1663,1689]" pageId="5" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having characters such as: gnathopod 2 propodus comparatively less setose and 1.7 x as long as broad, gnathopod 1 dactylus posterior margin serrated (vs. propodus densely setose and 1.5 x as long as broad; gnathopod 1 dactylus not serrated in
<taxonomicName id="4C244D29FFA8FFEE5981F9478903F972" authorityName="Chevreux" authorityYear="1901" box="[908,1042,1735,1761]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFA8FFEE5981F9478903F972" box="[908,1042,1735,1761]" italics="true" pageId="5" pageNumber="526">A. chelifera</emphasis>
</taxonomicName>
). These characters suggest that
<taxonomicName id="4C244D29FFA8FFEE5F89F9478DEAF896" authorityName="Chevreux" authorityYear="1901" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFA8FFEE5F89F9478DEAF896" italics="true" pageId="5" pageNumber="526">A. chelifera</emphasis>
</taxonomicName>
recorded by
<bibRefCitation id="EFB54B5BFFA8FFEE5B9CF96B8F90F896" author="Sivaprakasam, T. E." box="[401,641,1771,1797]" pageId="5" pageNumber="526" pagination="560 - 575" refId="ref4470" refString="Sivaprakasam, T. E. (1969) Amphipoda from the East Coast of India Part 2 Gammaridea and Caprellidea. Journal of Bombay Natural History Society, 66 (3), 560 - 575." type="journal article" year="1969">Sivaprakasam (1969)</bibRefCitation>
from
<collectingRegion id="49E0F848FFA8FFEE58CAF96B8E5CF896" box="[711,845,1771,1797]" country="India" name="Tamil Nadu" pageId="5" pageNumber="526">Tamil Nadu</collectingRegion>
is showing more affinities to
<taxonomicName id="4C244D29FFA8FFEE5EAFF96B881CF896" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2025" box="[1186,1293,1771,1797]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="sindhu" status="sp. nov.">
<emphasis id="B950EAB8FFA8FFEE5EAFF96B881CF896" box="[1186,1293,1771,1797]" italics="true" pageId="5" pageNumber="526">A. sindhu</emphasis>
</taxonomicName>
<emphasis id="B950EAB8FFA8FFEE5F18F96B8861F896" bold="true" box="[1301,1392,1771,1797]" pageId="5" pageNumber="526">
<taxonomicNameLabel id="A26357C3FFA8FFEE5F18F96B8861F896" box="[1301,1392,1771,1797]" pageId="5" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
So, we herewith confirm that the records of
<taxonomicName id="4C244D29FFA8FFEE585AF88F8FCBF8BA" authorityName="Chevreux" authorityYear="1901" box="[599,730,1807,1833]" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="chelifera">
<emphasis id="B950EAB8FFA8FFEE585AF88F8FCBF8BA" box="[599,730,1807,1833]" italics="true" pageId="5" pageNumber="526">A. chelifera</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB54B5BFFA8FFEE590EF88F8EE0F8BA" author="Sivaprakasam, T. E." box="[771,1009,1807,1833]" pageId="5" pageNumber="526" pagination="560 - 575" refId="ref4470" refString="Sivaprakasam, T. E. (1969) Amphipoda from the East Coast of India Part 2 Gammaridea and Caprellidea. Journal of Bombay Natural History Society, 66 (3), 560 - 575." type="journal article" year="1969">Sivaprakasam (1969)</bibRefCitation>
from
<collectingRegion id="49E0F848FFA8FFEE5E38F88F89EAF8BA" box="[1077,1275,1807,1833]" country="India" name="Tamil Nadu" pageId="5" pageNumber="526">Tamil Nadu State</collectingRegion>
,
<collectingCountry id="F333763AFFA8FFEE5F08F88F8850F8BA" box="[1285,1345,1807,1833]" name="India" pageId="5" pageNumber="526">India</collectingCountry>
, is of
<taxonomicName id="4C244D29FFA8FFEE5F89F88F8DF3F8DE" authorityName="Thacker &amp; Myers &amp; Trivedi" authorityYear="2025" class="Malacostraca" family="Photidae" genus="Audulla" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="sindhu" status="sp. nov.">
<emphasis id="B950EAB8FFA8FFEE5F89F88F8DF3F8DE" italics="true" pageId="5" pageNumber="526">A. sindhu</emphasis>
</taxonomicName>
<emphasis id="B950EAB8FFA8FFEE5AE4F8B38C53F8DE" bold="true" box="[233,322,1843,1869]" pageId="5" pageNumber="526">
<taxonomicNameLabel id="A26357C3FFA8FFEE5AE4F8B38C53F8DE" box="[233,322,1843,1869]" pageId="5" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
only.
</paragraph>
</subSubSection>
</treatment>
</document>