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<document id="7235B0A1FEEDD4D6FD12B99717D2AE12" ID-DOI="10.26879/756" ID-ISSN="1094-8074" ID-Zenodo-Dep="11062937" ID-ZooBank="5EE56527-5F98-4724-8120-18F7471BB497" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1714012330871" checkinUser="felipe" docAuthor="López-Torres, Sergi, Silcox, Mary T. &amp; Holroyd, Patricia A." docDate="2018" docId="038D87899233486721ACFD72F959B5DB" docLanguage="en" docName="PalaeontolElectron.21.3.37A.pdf" docOrigin="Palaeontologia Electronica (37 A) 21 (3)" docSource="http://dx.doi.org/10.26879/756" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Walshina esmaraldensis López-Torres &amp; Silcox &amp; Holroyd 2018, gen. et sp. nov." docType="treatment" docVersion="3" lastPageNumber="7" masterDocId="FFB4FFF192354860222AFFE4F808B326" masterDocTitle="New omomyoids (Euprimates, Mammalia) from the late Uintan of southern California, USA, and the question of the extinction of the Paromomyidae (Plesiadapiformes, Primates)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="7" updateTime="1715625529881" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="7B7746C6EB0EE7D095BCC5B79CFB2B91">New omomyoids (Euprimates, Mammalia) from the late Uintan of southern California, USA, and the question of the extinction of the Paromomyidae (Plesiadapiformes, Primates)</mods:title>
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<mods:namePart id="94E22349F8C0FD67FE587CDB1FFB5C1C">López-Torres, Sergi</mods:namePart>
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<mods:namePart id="74F35F027308F29F60D9BC165E740CA1">Silcox, Mary T.</mods:namePart>
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<mods:namePart id="925CA4211144190E8322E0CD69DDA96A">Holroyd, Patricia A.</mods:namePart>
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<mods:title id="5681C059D1433A2C16145097A1B54872">Palaeontologia Electronica</mods:title>
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<mods:date id="88966C784C403C34CE6B90A8944B4A0B">2018</mods:date>
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<mods:title id="87CEDEC596F81B71D4A560E16EB50F7A">37 A</mods:title>
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<mods:number id="071A6B8212D69448D721CAE14AF03AD0">2018-09-30</mods:number>
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<treatment id="038D87899233486721ACFD72F959B5DB" LSID="urn:lsid:plazi:treatment:038D87899233486721ACFD72F959B5DB" httpUri="http://treatment.plazi.org/id/038D87899233486721ACFD72F959B5DB" lastPageId="7" lastPageNumber="7" pageId="6" pageNumber="7">
<subSubSection id="C33E65149233486621ACFD72FD6BB189" box="[902,1379,662,687]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8B9B369F9233486621ACFD72FD6BB189" blockId="6.[902,1379,662,718]" box="[902,1379,662,687]" pageId="6" pageNumber="7">
<heading id="D0D381F39233486621ACFD72FD6BB189" box="[902,1379,662,687]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="7" reason="7">
<taxonomicName id="4C244D1C9233486621ACFD72FCAFB189" authority="López-Torres &amp; Silcox &amp; Holroyd, 2018" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" box="[902,1191,662,687]" class="Mammalia" family="Omomyidae" genus="Walshina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="esmaraldensis" status="gen. et sp. nov.">
<emphasis id="B950EA8D9233486621ACFD72FCAFB189" box="[902,1191,662,687]" italics="true" pageId="6" pageNumber="7">Walshina esmaraldensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6923348662684FD72FD6BB189" box="[1198,1379,662,687]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33E6514923348662184FD51FB83B019" pageId="6" pageNumber="7" type="description">
<paragraph id="8B9B369F923348662184FD51FD32B1E8" blockId="6.[902,1379,662,718]" box="[942,1338,693,718]" pageId="6" pageNumber="7">
<figureCitation id="131F2A1A923348662184FD51FC6DB1E8" box="[942,1125,693,718]" captionStart="FIGURE 4" captionStartId="6.[196,287,856,879]" captionTargetBox="[241,760,175,815]" captionTargetId="figure-613@6.[201,788,162,832]" captionTargetPageId="6" captionText="FIGURE 4. Environmental SEM images of four teeth of Walshina esmaraldensis gen. et sp. nov. 1: left M3, SDSNH 76276; 2: right M1, SDSNH 76337; 3: right M2, SDSNH 76338; 4: right M3, SDSNH 72583. All teeth are in occlusal view. Arrow indicates the location of the fovea." figureDoi="http://doi.org/10.5281/zenodo.11062945" httpUri="https://zenodo.org/record/11062945/files/figure.png" pageId="6" pageNumber="7">Figures 4.1-4.4</figureCitation>
,
<figureCitation id="131F2A1A92334866265AFD51FCC5B1E8" box="[1136,1229,693,718]" captionStart="FIGURE 5" captionStartId="7.[188,278,1450,1473]" captionTargetBox="[366,1296,178,1422]" captionTargetId="figure-75@7.[318,1311,168,1422]" captionTargetPageId="7" captionText="FIGURE 5. Photographs taken with digital camera (1, 4-11) and Micro-CT scan reconstructions generated using Avizo 7 (2, 3). Walshina shifrae comb. nov. (1, 4, 7, 10) 1: right M1, CM 15797 (holotype; mirrored), in occlusal view; 4: left M2, CM 15103, in occlusal view; 7: left M, CM 21637, in occlusal view; 10: left M; CM 15726, in occlusal view. 2 3 Walshina esmaraldensis gen. et sp. nov. (2, 5) 2: left M1, LACM 40198 (holotype), in occlusal view; 5: left M2, SDSNH 62850, in occlusal view. Walshina mcgrewi comb. nov. (3, 6, 8, 9, 11) 3: left M1, CM 15635 (holotype), in occlusal view; 6: left M2, CM15794, in occlusal view; 8, 9, 11: left mandibular fragment with M, CM 29005, in occlusal 2 (8), buccal (9), and lingual (11) views." figureDoi="http://doi.org/10.5281/zenodo.11062947" httpUri="https://zenodo.org/record/11062947/files/figure.png" pageId="6" pageNumber="7">5.2, 5.5</figureCitation>
,
<figureCitation id="131F2A1A9233486626F2FD51FD32B1E8" box="[1240,1338,693,718]" captionStart="FIGURE 6" captionStartId="8.[198,289,1210,1233]" captionTargetBox="[467,1188,193,1157]" captionTargetId="figure-245@8.[412,1221,164,1192]" captionTargetPageId="8" captionText="FIGURE 6. Micro-CT scan reconstructions of specimens of Walshina esmaraldensis gen. et sp. nov. generated using Avizo 7: 1: left M1, LACM 40198 (holotype), in occlusal view; 2: left M2, SDSNH 87336, in occlusal view; 3: lingual fragment of a left M2, SDSNH 87337, in occlusal view; 4: lingual fragment of a right M2, SDSNH 42268, in occlusal view; 5, 6, 9, 10: left M2, SDSNH 87332, in occlusal (5), buccal (6), mesial (9) and lingual (10) views; 7, 8, 11, 12: mesial fragment of a left M1, SDSNH 87331, in occlusal (7), buccal (8), mesial (11) and lingual (12) views; 13, 14, 17, 18: left M3, SDSNH 87334, in buccal (13), occlusal (14), lingual (17) and mesial (18) views; 15, 16, 19: distal fragment of a right M3, SDSNH 87335, in buccal (15), occlusal (16) and lingual (19) views." figureDoi="http://doi.org/10.5281/zenodo.11062949" httpUri="https://zenodo.org/record/11062949/files/figure.png" pageId="6" pageNumber="7">6.1-6.19</figureCitation>
</paragraph>
<paragraph id="8B9B369F923348662162FD04FD87B1D2" blockId="6.[840,1423,736,756]" box="[840,1423,736,756]" pageId="6" pageNumber="7">
<emphasis id="B950EA8D923348662162FD04FD87B1D2" bold="true" box="[840,1423,736,756]" pageId="6" pageNumber="7">
zoobank.org/
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</emphasis>
</paragraph>
<paragraph id="8B9B369F923348662162FCE2FB83B019" blockId="6.[840,1441,773,1877]" pageId="6" pageNumber="7">
<taxonomicName id="4C244D1C923348662162FCE2FD33B039" authority="Mason, 1990" authorityName="Mason" authorityYear="1990" box="[840,1339,773,799]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="shifrae">
<emphasis id="B950EA8D923348662162FCE2FBFCB039" box="[840,1012,774,799]" italics="true" pageId="6" pageNumber="7">Phenacolemur</emphasis>
cf.
<emphasis id="B950EA8D923348662618FCE1FC8AB038" box="[1074,1154,773,798]" italics="true" pageId="6" pageNumber="7">shifrae</emphasis>
Mason, 1990
</taxonomicName>
; Walsh, 1996.
</paragraph>
</subSubSection>
<subSubSection id="C33E6514923348662162FCADFB81B774" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8B9B369F923348662162FCADFCCFB044" blockId="6.[840,1441,773,1877]" box="[840,1223,838,866]" pageId="6" pageNumber="7">
<materialsCitation id="3B4C3CC2923348662162FCADFCCFB044" box="[840,1223,838,866]" collectionCode="LACM" pageId="6" pageNumber="7" specimenCode="LACM 40198" specimenCount="1" typeStatus="holotype">
<emphasis id="B950EA8D923348662162FCADFBC9B044" bold="true" box="[840,961,841,866]" pageId="6" pageNumber="7">
<typeStatus id="549F883D923348662162FCADFBB5B044" box="[840,957,841,866]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DB829EE49233486621E3FCACFC60B044" box="[969,1128,840,866]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="6" pageNumber="7" type="Museum">LACM 40198</specimenCode>
, left M
<superScript id="7C519BD792334866269EFCA2FCC8B07C" attach="left" box="[1204,1216,838,858]" fontSize="8" pageId="6" pageNumber="7">1</superScript>
.
</materialsCitation>
</paragraph>
<caption id="DF5B66179233486622EEFCBCF90EB728" ID-DOI="http://doi.org/10.5281/zenodo.11062945" ID-Zenodo-Dep="11062945" httpUri="https://zenodo.org/record/11062945/files/figure.png" pageId="6" pageNumber="7" startId="6.[196,287,856,879]" targetBox="[241,760,175,815]" targetPageId="6" targetType="figure">
<paragraph id="8B9B369F9233486622EEFCBCF90EB728" blockId="6.[196,790,856,1038]" pageId="6" pageNumber="7">
<emphasis id="B950EA8D9233486622EEFCBCF933B049" bold="true" box="[196,315,856,879]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
Environmental SEM images of four teeth of
<taxonomicName id="4C244D1C9233486622EEFC9DF9C7B0B6" authority="López-Torres &amp; Silcox &amp; Holroyd, 2018" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" box="[196,463,889,912]" class="Mammalia" family="Omomyidae" genus="Walshina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="esmaraldensis" status="gen. et sp. nov.">
<emphasis id="B950EA8D9233486622EEFC9DF9C7B0B6" box="[196,463,889,912]" italics="true" pageId="6" pageNumber="7">Walshina esmaraldensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F69233486623F6FC9DFA86B0B6" box="[476,654,889,912]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
1: left M3, SDSNH 76276; 2: right M
<subScript id="17A034DA9233486623F2FC44F9EBB094" attach="left" box="[472,483,928,946]" fontSize="7" pageId="6" pageNumber="7">1</subScript>
, SDSNH 76337; 3: right M
<subScript id="17A034DA92334866212FFC44FB18B094" attach="left" box="[773,784,928,946]" fontSize="7" pageId="6" pageNumber="7">2</subScript>
, SDSNH 76338; 4: right M
<subScript id="17A034DA9233486623F9FC26F9D6B0F2" attach="left" box="[467,478,962,980]" fontSize="7" pageId="6" pageNumber="7">3</subScript>
, SDSNH 72583. All teeth are in occlusal view. Arrow indicates the location of the fovea.
</paragraph>
</caption>
<paragraph id="8B9B369F923348662162FC88FD6BB0D4" blockId="6.[840,1441,773,1877]" pageId="6" pageNumber="7">
<emphasis id="B950EA8D923348662162FC88FBDAB0A3" bold="true" box="[840,978,876,901]" pageId="6" pageNumber="7">Hypodigm.</emphasis>
M
<superScript id="7C519BD79233486621D2FC8DFC0CB05B" attach="left" box="[1016,1028,873,893]" fontSize="8" pageId="6" pageNumber="7">2</superScript>
:
<collectionCode id="ED35AE5A923348662636FC8FFC71B0A3" box="[1052,1145,875,901]" pageId="6" pageNumber="7">SDSNH</collectionCode>
42268, 62850, 87336, 87337; M
<superScript id="7C519BD7923348662194FC68FBC2B086" attach="left" box="[958,970,908,928]" fontSize="8" pageId="6" pageNumber="7">3</superScript>
:
<collectionCode id="ED35AE5A9233486621CAFC6BFC35B08F" box="[992,1085,911,937]" pageId="6" pageNumber="7">SDSNH</collectionCode>
76267; M
<subScript id="17A034DA9233486626EBFC7DFCC5B08B" attach="left" box="[1217,1229,921,941]" fontSize="8" pageId="6" pageNumber="7">1</subScript>
:
<collectionCode id="ED35AE5A9233486626C9FC6BFD48B08F" box="[1251,1344,911,937]" pageId="6" pageNumber="7">SDSNH</collectionCode>
76337, 87331; M
<subScript id="17A034DA9233486621EAFC5AFBC4B0F4" attach="left" box="[960,972,958,978]" fontSize="8" pageId="6" pageNumber="7">2</subScript>
:
<collectionCode id="ED35AE5A9233486621CEFC57FC49B0EB" box="[996,1089,947,973]" pageId="6" pageNumber="7">SDSNH</collectionCode>
76338, 87332, 87333; M
<subScript id="17A034DA9233486627A7FC5AFD91B0F4" attach="left" box="[1421,1433,958,978]" fontSize="8" pageId="6" pageNumber="7">3</subScript>
:
<collectionCode id="ED35AE5A923348662162FC3CFBADB0D4" box="[840,933,984,1010]" pageId="6" pageNumber="7">SDSNH</collectionCode>
72583, 76266, 76339, 87334, 87335.
</paragraph>
<paragraph id="8B9B369F923348662162FC1DFB81B774" blockId="6.[840,1441,773,1877]" pageId="6" pageNumber="7">
<emphasis id="B950EA8D923348662162FC1DFBE7B734" bold="true" box="[840,1007,1017,1042]" pageId="6" pageNumber="7">Type locality.</emphasis>
<materialsCitation id="3B4C3CC29233486621D2FC1CFB8DB774" collectionCode="LACM" country="United States of America" county="Ventura County" location="Simi Valley" municipality="Late Uintan" pageId="6" pageNumber="7" specimenCount="1" stateProvince="California">
<collectionCode id="ED35AE5A9233486621D2FC1CFC48B734" box="[1016,1088,1016,1042]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="6" pageNumber="7" type="Museum">LACM</collectionCode>
(CIT) 180, north of
<location id="8EFB604492334866271FFC1CFB61B714" LSID="urn:lsid:plazi:treatment:038D87899233486721ACFD72F959B5DB:8EFB604492334866271FFC1CFB61B714" country="United States of America" county="Ventura County" municipality="Late Uintan" name="Simi Valley" pageId="6" pageNumber="7" stateProvince="California">Simi Valley</location>
,
<collectingCounty id="62FA4E13923348662151FBFDFC3DB714" box="[891,1077,1048,1074]" pageId="6" pageNumber="7">Ventura County</collectingCounty>
,
<collectingRegion id="49E0F87D92334866266DFBFCFCB0B714" box="[1095,1208,1048,1074]" country="United States of America" name="California" pageId="6" pageNumber="7">California</collectingRegion>
,
<collectingCountry id="F333760F9233486626E1FBFCFD0AB714" box="[1227,1282,1048,1074]" name="United States of America" pageId="6" pageNumber="7">USA</collectingCountry>
.
<collectingMunicipality id="6BFFACE592334866273EFBFDFDA8B714" box="[1300,1440,1049,1074]" pageId="6" pageNumber="7">Late Uintan</collectingMunicipality>
(Ui3)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E6514923348662162FBBDFCF7B7B4" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8B9B369F923348662162FBBDFCF7B7B4" blockId="6.[840,1441,773,1877]" pageId="6" pageNumber="7">
<emphasis id="B950EA8D923348662162FBBDFBDEB754" bold="true" box="[840,982,1113,1138]" pageId="6" pageNumber="7">Etymology.</emphasis>
In reference to the localities of Emerald Ridge (Vulgar Latin:
<emphasis id="B950EA8D92334866264FFB9DFCE7B7B4" box="[1125,1263,1145,1170]" italics="true" pageId="6" pageNumber="7">esmaraldus</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E6514923348672162FB7DF959B5DB" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8B9B369F923348672162FB7DF959B5DB" blockId="6.[840,1441,773,1877]" lastBlockId="7.[192,792,1732,1789]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B950EA8D923348662162FB7DFBC7B794" bold="true" box="[840,975,1177,1202]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
Intermediate in size between
<taxonomicName id="4C244D1C9233486627AAFB7DFBA3B7F4" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="2018" class="Mammalia" family="Paromomyidae" genus="Ignacius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="mcgrewi" status="comb. nov.">
<emphasis id="B950EA8D9233486627AAFB7DFBA3B7F4" italics="true" pageId="6" pageNumber="7">W. mcgrewi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6923348662199FB5DFC3CB7F4" box="[947,1076,1209,1234]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C244D1C923348662645FB5DFCEFB7F4" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="1978" box="[1135,1255,1209,1234]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="shifrae" status="comb. nov.">
<emphasis id="B950EA8D923348662645FB5DFCEFB7F4" box="[1135,1255,1209,1234]" italics="true" pageId="6" pageNumber="7">W. shifrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F69233486626C4FB5DFD67B7F4" box="[1262,1391,1209,1234]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
Differs from
<taxonomicName id="4C244D1C923348662193FB3DFC4CB7D4" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="2018" box="[953,1092,1241,1266]" class="Mammalia" family="Paromomyidae" genus="Ignacius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="mcgrewi" status="comb. nov.">
<emphasis id="B950EA8D923348662193FB3DFC4CB7D4" box="[953,1092,1241,1266]" italics="true" pageId="6" pageNumber="7">W. mcgrewi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6923348662666FB3DFCC5B7D4" box="[1100,1229,1241,1266]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
in having a hypoconulid closer to the hypoconid than to the entoconid on M
<subScript id="17A034DA92334866262EFAC7FC18B611" attach="left" box="[1028,1040,1315,1335]" fontSize="8" pageId="6" pageNumber="7">2</subScript>
, similar to
<taxonomicName id="4C244D1C9233486626B3FAFDFD1BB614" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="1978" box="[1177,1299,1305,1330]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="shifrae" status="comb. nov.">
<emphasis id="B950EA8D9233486626B3FAFDFD1BB614" box="[1177,1299,1305,1330]" italics="true" pageId="6" pageNumber="7">W. shifrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6923348662737FAFDFDA9B614" box="[1309,1441,1305,1330]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
Very large hypoconulids on M
<subScript id="17A034DA92334866269DFAACFCCBB67A" attach="left" box="[1207,1219,1352,1372]" fontSize="8" pageId="6" pageNumber="7">1</subScript>
and M
<subScript id="17A034DA923348662736FAACFD20B67A" attach="left" box="[1308,1320,1352,1372]" fontSize="8" pageId="6" pageNumber="7">2</subScript>
, whereas they are weakly developed in
<taxonomicName id="4C244D1C923348662697FA86FD43B65D" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="2018" box="[1213,1355,1378,1403]" class="Mammalia" family="Paromomyidae" genus="Ignacius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="mcgrewi" status="comb. nov.">
<emphasis id="B950EA8D923348662697FA86FD43B65D" box="[1213,1355,1378,1403]" italics="true" pageId="6" pageNumber="7">W. mcgrewi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6923348662772FA86FB71B6BD" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C244D1C92334866219FFA66FC24B6BD" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="1978" box="[949,1068,1410,1435]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="shifrae" status="comb. nov.">
<emphasis id="B950EA8D92334866219FFA66FC24B6BD" box="[949,1068,1410,1435]" italics="true" pageId="6" pageNumber="7">W. shifrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F692334866261EFA66FCBEB6BD" box="[1076,1206,1410,1435]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
M
<subScript id="17A034DA9233486626FEFA69FCE8B687" attach="left" box="[1236,1248,1421,1441]" fontSize="8" pageId="6" pageNumber="7">2</subScript>
trigonid and talonid of similar width, whereas
<taxonomicName id="4C244D1C923348662695FA43FD44B6E6" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="2018" box="[1215,1356,1447,1472]" class="Mammalia" family="Paromomyidae" genus="Ignacius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="mcgrewi" status="comb. nov.">
<emphasis id="B950EA8D923348662695FA43FD44B6E6" box="[1215,1356,1447,1472]" italics="true" pageId="6" pageNumber="7">W. mcgrewi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6923348662772FA43FB71B6C6" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C244D1C923348662196FA23FC3EB6C6" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="1978" box="[956,1078,1479,1504]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="shifrae" status="comb. nov.">
<emphasis id="B950EA8D923348662196FA23FC3EB6C6" box="[956,1078,1479,1504]" italics="true" pageId="6" pageNumber="7">W. shifrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6923348662668FA23FCCFB6C6" box="[1090,1223,1479,1504]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
have significantly wider M
<subScript id="17A034DA92334866218FFA16FBB9B520" attach="left" box="[933,945,1522,1542]" fontSize="8" pageId="6" pageNumber="7">2</subScript>
talonid bases relative to the trigonid base. M
<subScript id="17A034DA923348662174F9F3FB62B50D" attach="left" box="[862,874,1559,1579]" fontSize="8" pageId="6" pageNumber="7">3</subScript>
with larger paraconid and taller metaconid relative to the protoconid than in
<taxonomicName id="4C244D1C92334866268AF9D5FD10B56C" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="1978" box="[1184,1304,1585,1610]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="shifrae" status="comb. nov.">
<emphasis id="B950EA8D92334866268AF9D5FD10B56C" box="[1184,1304,1585,1610]" italics="true" pageId="6" pageNumber="7">W. shifrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6923348662735F9D5FDA9B56C" box="[1311,1441,1585,1610]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
Hypoconulid lobe of M
<subScript id="17A034DA923348662676F9B8FC60B556" attach="left" box="[1116,1128,1628,1648]" fontSize="8" pageId="6" pageNumber="7">3</subScript>
not expanded as far distally from the apex of the hypoconulid as in
<taxonomicName id="4C244D1C9233486627AAF992FB90B589" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="1978" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="shifrae" status="comb. nov.">
<emphasis id="B950EA8D9233486627AAF992FB90B589" italics="true" pageId="6" pageNumber="7">W. shifrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F692334866218BF972FC2DB589" box="[929,1061,1686,1711]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
Differs from
<taxonomicName id="4C244D1C9233486626E8F972FD46B589" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="2018" box="[1218,1358,1686,1711]" class="Mammalia" family="Paromomyidae" genus="Ignacius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="mcgrewi" status="comb. nov.">
<emphasis id="B950EA8D9233486626E8F972FD46B589" box="[1218,1358,1686,1711]" italics="true" pageId="6" pageNumber="7">W. mcgrewi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6923348662772F972FB71B5E9" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
(but not
<taxonomicName id="4C244D1C9233486621D1F952FC73B5E9" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="1978" box="[1019,1147,1718,1743]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="shifrae" status="comb. nov.">
<emphasis id="B950EA8D9233486621D1F952FC73B5E9" box="[1019,1147,1718,1743]" italics="true" pageId="6" pageNumber="7">W. shifrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F69233486626A1F952FD1FB5E9" box="[1163,1303,1718,1743]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
) in having upper molars with smaller distolingual basins. In mesial view, M
<superScript id="7C519BD79233486621D3F912FC0DB42C" attach="left" box="[1017,1029,1782,1802]" fontSize="8" pageId="6" pageNumber="7">1</superScript>
with a less clearly delineated preprotocrista, like
<taxonomicName id="4C244D1C923348662622F8F8FC9CB413" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="2018" box="[1032,1172,1820,1845]" class="Mammalia" family="Paromomyidae" genus="Ignacius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="mcgrewi" status="comb. nov.">
<emphasis id="B950EA8D923348662622F8F8FC9CB413" box="[1032,1172,1820,1845]" italics="true" pageId="6" pageNumber="7">W. mcgrewi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F69233486626B5F8F8FD28B413" box="[1183,1312,1820,1845]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
M
<superScript id="7C519BD7923348662768F8FEFD46B408" attach="left" box="[1346,1358,1818,1838]" fontSize="8" pageId="6" pageNumber="7">2</superScript>
with a much more lingually expanded protocone lobe than in
<taxonomicName id="4C244D1C9232486722CBF920F955B5FB" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="1978" box="[225,349,1732,1757]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="shifrae" status="comb. nov.">
<emphasis id="B950EA8D9232486722CBF920F955B5FB" box="[225,349,1732,1757]" italics="true" pageId="7" pageNumber="8">W. shifrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6923248672340F920F9E3B5FB" box="[362,491,1732,1757]" pageId="7" pageNumber="8" rank="species">comb. nov.</taxonomicNameLabel>
(but not in
<taxonomicName id="4C244D1C9232486720A3F920FB10B5FB" authorityName="López-Torres &amp; Silcox &amp; Holroyd" authorityYear="2018" baseAuthorityName="López-Torres &amp; Silcox &amp; Holroyd" baseAuthorityYear="2018" box="[649,792,1732,1757]" class="Mammalia" family="Paromomyidae" genus="Ignacius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="mcgrewi" status="comb. nov.">
<emphasis id="B950EA8D9232486720A3F920FB10B5FB" box="[649,792,1732,1757]" italics="true" pageId="7" pageNumber="8">W. mcgrewi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F69232486722EAF900F94EB5DB" box="[192,326,1764,1789]" pageId="7" pageNumber="8" rank="species">comb. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
</treatment>
</document>