989 lines
72 KiB
XML
989 lines
72 KiB
XML
<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3793.3.1" ID-GBIF-Dataset="f2f9eed1-b32c-40ab-8f6c-1a7ef8f59f52" ID-ISSN="1175-5326" ID-Zenodo-Dep="229629" ID-ZooBank="874ED359-D4A4-4EF0-BB3B-C0A70563472E" checkinTime="1460413274041" checkinUser="plazi" docAuthor="Koeda, Keita, Yoshino, Tetsuo, Imai, Hideyuki & Tachihara, Katsunori" docDate="2014" docId="03869D6CFFE44123B9AAF991CAEDFDA6" docLanguage="en" docName="zt03793p330.pdf" docOrigin="Zootaxa 3793 (3)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@324da50a" docStyleName="zootaxa.2013.journal_article" docTitle="Pempheris tominagai Koeda, Yoshino & Tachihara, sp. nov." docType="treatment" docUuid="E2DA30A0-1E27-4677-A29D-CBE9BB791ED9" docUuidSource="ZooBank" docVersion="10" lastPageNumber="321" masterDocId="FFBFE514FFF44137B93DFFE5C914FF84" masterDocTitle="A review of the genus Pempheri s (Perciformes, Pempheridae) of the Red Sea, with description of a new species" masterLastPageNumber="330" masterPageNumber="301" pageNumber="317" updateTime="1677849786491" updateUser="juliana">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the genus Pempheri s (Perciformes, Pempheridae) of the Red Sea, with description of a new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Koeda, Keita</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yoshino, Tetsuo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Imai, Hideyuki</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tachihara, Katsunori</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>3793</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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<mods:extent unit="page">
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<mods:start>301</mods:start>
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<mods:end>330</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3793.3.1</mods:identifier>
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<mods:identifier type="GBIF-Dataset">f2f9eed1-b32c-40ab-8f6c-1a7ef8f59f52</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">229629</mods:identifier>
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<mods:identifier type="ZooBank">874ED359-D4A4-4EF0-BB3B-C0A70563472E</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6124657" ID-GBIF-Taxon="119424149" ID-Zenodo-Dep="6124657" LSID="urn:lsid:zoobank.org:act:E2DA30A0-1E27-4677-A29D-CBE9BB791ED9" httpUri="http://treatment.plazi.org/id/03869D6CFFE44123B9AAF991CAEDFDA6" lastPageId="20" lastPageNumber="321" pageId="16" pageNumber="317">
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<subSubSection pageId="16" pageNumber="317" type="nomenclature">
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<paragraph blockId="16.[151,879,1652,1711]" box="[151,879,1652,1678]" pageId="16" pageNumber="317">
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<heading bold="true" box="[151,879,1652,1678]" fontSize="11" level="1" pageId="16" pageNumber="317" reason="1">
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<emphasis bold="true" box="[151,879,1652,1678]" pageId="16" pageNumber="317">
|
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<taxonomicName ID-CoL="76G44" ID-ENA="1159865" authority="Koeda, Yoshino & Tachihara" authorityName="Koeda, Yoshino & Tachihara" box="[151,779,1652,1678]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="tominagai" status="sp. nov.">
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<emphasis bold="true" box="[151,409,1652,1678]" italics="true" pageId="16" pageNumber="317">Pempheris tominagai</emphasis>
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Koeda, Yoshino & Tachihara
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</taxonomicName>
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<taxonomicNameLabel box="[785,879,1652,1678]" pageId="16" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="16.[151,879,1652,1711]" box="[151,436,1686,1711]" pageId="16" pageNumber="317">
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<figureCitation box="[151,218,1686,1711]" captionStart="FIGURE 7 a" captionStartId="17.[151,250,1144,1166]" captionTargetBox="[340,1246,199,1132]" captionTargetId="figure@17.[340,1247,187,1133]" captionTargetPageId="17" captionText="FIGURE 7 a. Holotype of Pempheris tominagai (NSMT-P 106826), 95.1 mm SL, Sharm el Moya Bay, Egypt. b: Underwater photograph of P. tominagai, Tulear, Madagascar, taken by G. R. Allen." httpUri="https://zenodo.org/record/229636/files/figure.png" pageId="16" pageNumber="317">
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Fig.
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<number box="[202,218,1687,1710]" pageId="16" pageNumber="317" value="7.0">7</number>
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||
</figureCitation>
|
||
;
|
||
<tableCitation box="[231,354,1686,1711]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="5.[151,239,544,567]" captionStartId-1="5.[151,239,1002,1025]" captionStartId-2="5.[151,239,1470,1493]" captionText-0="TABLE 1. Counts of anal soft rays of four Pempheris species distributed in the Red Sea, and P. adusta from Andaman Sea and Pacific Ocean" captionText-1="TABLE 2. Counts of pored lateral line scales of four Pempheris species distributed in the Red Sea, and P. adusta from Andaman Sea and Pacific Ocean" captionText-2="TABLE 3. Counts of scale rows above lateral line of four Pempheris species distributed in the Red Sea, and P. adusta from Andaman Sea and Pacific Ocean." pageId="16" pageNumber="317">
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Tables
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<number box="[311,322,1687,1711]" pageId="16" pageNumber="317" value="1.0">1</number>
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–3
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</tableCitation>
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||
,
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<tableCitation box="[367,396,1687,1711]" captionStart="TABLE 11" captionStartId="18.[151,239,152,175]" captionText="TABLE 11. Counts of Pempheris tominagai, holotype, paratypes and non-types, syntypes of P. schwenkii, and the other specimens." pageId="16" pageNumber="317">11</tableCitation>
|
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,
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<tableCitation box="[408,436,1687,1711]" captionStart="TABLE 12" captionStartId="19.[151,239,152,175]" captionText="TABLE 12. Measurements of Pempheris tominagai, holotype, paratypes and non-types, syntypes of P. schwenkii, and the other specimens." pageId="16" pageNumber="317">
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<number box="[408,436,1687,1711]" pageId="16" pageNumber="317" value="12.0">12</number>
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</tableCitation>
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||
</paragraph>
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||
<paragraph blockId="16.[151,1436,1752,1896]" box="[151,767,1752,1774]" pageId="16" pageNumber="317">
|
||
<treatmentCitationGroup box="[151,767,1752,1774]" pageId="16" pageNumber="317">
|
||
<taxonomicName ID-CoL="76G58" box="[151,351,1752,1774]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="molucca">
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<emphasis box="[151,351,1752,1774]" italics="true" pageId="16" pageNumber="317">Pempheris molucca</emphasis>
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||
</taxonomicName>
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(not of Cuvier):
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<treatmentCitation author="Mouneimne" box="[523,763,1752,1774]" page="105" pageId="16" pageNumber="317" year="1979">
|
||
<bibRefCitation author="Mouneimne" box="[523,712,1752,1774]" pageId="16" pageNumber="317" refString="Mouneimne, N. (1979) Poissons nouveaux pour les cotes Libanaises. Cybium, 6, 105 - 110." type="journal article" year="1979">
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||
Mouneimne,
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||
<number box="[658,712,1752,1774]" pageId="16" pageNumber="317" value="1979.0">1979</number>
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||
</bibRefCitation>
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||
:
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<number box="[722,763,1752,1774]" pageId="16" pageNumber="317" value="105.0">105</number>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
|
||
<paragraph blockId="16.[151,1436,1752,1896]" box="[151,1205,1782,1804]" pageId="16" pageNumber="317">
|
||
<treatmentCitationGroup box="[151,1205,1782,1804]" pageId="16" pageNumber="317">
|
||
<taxonomicName ID-CoL="76FTG" box="[151,361,1782,1804]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="oualensis">
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<emphasis box="[151,361,1782,1804]" italics="true" pageId="16" pageNumber="317">Pempheris oualensis</emphasis>
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</taxonomicName>
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(not of Cuvier, in Cuvier & Valenciennes):
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||
<treatmentCitation author="Allen" box="[805,1064,1782,1804]" page="55" pageId="16" pageNumber="317" year="1987">
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<bibRefCitation author="Allen" box="[805,1027,1782,1804]" pageId="16" pageNumber="317" refString="Allen, G. R. & Steene, R. C. (1987) Reef Fishes of the Indian Ocean. TFH Publications, Neptune City, New Jersey, 240 pp." type="book" year="1987">
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||
Allen & Steene,
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<number box="[973,1027,1782,1804]" pageId="16" pageNumber="317" value="1987.0">1987</number>
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||
</bibRefCitation>
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||
:
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<number box="[1037,1064,1782,1804]" pageId="16" pageNumber="317" value="55.0">55</number>
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</treatmentCitation>
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, pl.
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<number box="[1107,1135,1782,1804]" pageId="16" pageNumber="317" value="36.0">36</number>
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, fig.
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<number box="[1186,1201,1783,1804]" pageId="16" pageNumber="317" value="7.0">7</number>
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.
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="317" type="reference_group">
|
||
<paragraph blockId="16.[151,1436,1752,1896]" pageId="16" pageNumber="317">
|
||
<treatmentCitationGroup pageId="16" pageNumber="317">
|
||
<taxonomicName ID-CoL="76FRP" box="[151,365,1813,1835]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="schwenkii">
|
||
<emphasis box="[151,365,1813,1835]" italics="true" pageId="16" pageNumber="317">Pempheris schwenkii</emphasis>
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||
</taxonomicName>
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||
(not of Bleeker):
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||
<treatmentCitation author="Smith" box="[546,852,1813,1835]" page="669" pageId="16" pageNumber="317" year="1986">
|
||
<bibRefCitation author="Smith" box="[546,802,1813,1835]" pageId="16" pageNumber="317" refString="Smith, M. M. & Heemstra, P. C. (1986) Smith's sea fishes. Springer-Verlag, New York. 1047 pp." type="book" year="1986">
|
||
Smith & Heemstra,
|
||
<number box="[748,802,1813,1835]" pageId="16" pageNumber="317" value="1986.0">1986</number>
|
||
</bibRefCitation>
|
||
:
|
||
<number box="[811,852,1813,1835]" pageId="16" pageNumber="317" value="669.0">669</number>
|
||
</treatmentCitation>
|
||
, pl.
|
||
<number box="[894,922,1813,1835]" pageId="16" pageNumber="317" value="84.0">84</number>
|
||
;
|
||
<treatmentCitation author="Smith" box="[933,1111,1813,1835]" page="247" pageId="16" pageNumber="317" year="1965">
|
||
<bibRefCitation author="Smith" box="[933,1060,1813,1835]" pageId="16" pageNumber="317" refString="Smith, J. L. B. (1965) The Sea Fishes of Southern Africa, 5 th Ed. Central News Agency L. T. D., Cape Town, 580 pp." type="book" year="1965">
|
||
Smith,
|
||
<number box="[1006,1060,1813,1835]" pageId="16" pageNumber="317" value="1965.0">1965</number>
|
||
</bibRefCitation>
|
||
:
|
||
<number box="[1070,1111,1813,1835]" pageId="16" pageNumber="317" value="247.0">247</number>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation author="Fricke" box="[1121,1303,1813,1835]" page="322" pageId="16" pageNumber="317" year="1999">
|
||
<bibRefCitation author="Fricke" box="[1121,1252,1813,1835]" pageId="16" pageNumber="317" refString="Fricke, R. (1999) Fishes of the Mascarene Islands (Reunion, Mauritius, Rodriguez). An annotated checklist, with descriptions of new species. Koeltz Scientific Books, Koenigstein, Germany, viii + 759 pp." type="book" year="1999">
|
||
Fricke,
|
||
<number box="[1198,1252,1813,1835]" pageId="16" pageNumber="317" value="1999.0">1999</number>
|
||
</bibRefCitation>
|
||
:
|
||
<number box="[1262,1303,1813,1835]" pageId="16" pageNumber="317" value="322.0">322</number>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation author="Heemstra" page="327" pageId="16" pageNumber="317" year="2004">
|
||
<bibRefCitation author="Heemstra" pageId="16" pageNumber="317" refString="Heemstra, P. & Heemstra, E. (2004) Coastal fishes of Southern Africa. NISC and SAIAB, Grahamstown (South Africa), 488 pp." type="book" year="2004">
|
||
Heemstra & Heemstra,
|
||
<number box="[308,363,1844,1866]" pageId="16" pageNumber="317" value="2004.0">2004</number>
|
||
</bibRefCitation>
|
||
:
|
||
<number box="[372,413,1844,1866]" pageId="16" pageNumber="317" value="327.0">327</number>
|
||
</treatmentCitation>
|
||
;
|
||
<bibRefCitation author="Laith" box="[424,642,1844,1866]" pageId="16" pageNumber="317" refString="Laith, J. & Koeda, K. (2013) Records of Pempheris schwenkii Bleeker (1855) and Pempheris mangula Cuvier (1829) from the Omani waters. Journal of Applied Ichthyology, 29 (2013), 1378 - 1379. http: // dx. doi. org / 10.1111 / jai. 12240" type="journal article" year="2013">
|
||
Laith & Koeda,
|
||
<number box="[588,642,1844,1866]" pageId="16" pageNumber="317" value="2013.0">2013</number>
|
||
</bibRefCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="16.[151,1436,1752,1896]" box="[151,1360,1874,1896]" pageId="16" pageNumber="317">
|
||
<treatmentCitationGroup box="[151,1360,1874,1896]" pageId="16" pageNumber="317">
|
||
<taxonomicName ID-CoL="76G3V" box="[151,390,1874,1896]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="317" phylum="Chordata" rank="species" species="vanicolensis">
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||
<emphasis box="[151,390,1874,1896]" italics="true" pageId="16" pageNumber="317">Pempheris vanicolensis</emphasis>
|
||
</taxonomicName>
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||
(not of Cuvier, in Cuvier & Valenciennes):
|
||
<treatmentCitation author="Randall" box="[834,1031,1874,1896]" page="244" pageId="16" pageNumber="317" year="1995">
|
||
<bibRefCitation author="Randall" box="[834,981,1874,1896]" pageId="16" pageNumber="317" refString="Randall, J. E. (1995) Coastal fishes of Oman. University of Hawaii Press, Honolulu, 439 pp." type="book" year="1995">
|
||
Randall,
|
||
<number box="[926,981,1874,1896]" pageId="16" pageNumber="317" value="1995.0">1995</number>
|
||
</bibRefCitation>
|
||
:
|
||
<number box="[990,1031,1874,1896]" pageId="16" pageNumber="317" value="244.0">244</number>
|
||
</treatmentCitation>
|
||
, fig.
|
||
<number box="[1081,1123,1874,1896]" pageId="16" pageNumber="317" value="638.0">638</number>
|
||
:
|
||
<treatmentCitation author="Siliotti" box="[1132,1305,1874,1896]" page="31" pageId="16" pageNumber="317" year="2009">
|
||
<bibRefCitation author="Siliotti" box="[1132,1268,1874,1896]" pageId="16" pageNumber="317" refString="Siliotti, A. (2009) Das Rote Meer: Ein Korallengarten. Delius Klasing, 142 pp." type="book" year="2009">
|
||
Siliotti,
|
||
<number box="[1214,1268,1874,1896]" pageId="16" pageNumber="317" value="2009.0">2009</number>
|
||
</bibRefCitation>
|
||
:
|
||
<number box="[1278,1305,1874,1896]" pageId="16" pageNumber="317" value="31112.0">31</number>
|
||
</treatmentCitation>
|
||
, 112.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317" type="materials_examined">
|
||
<paragraph blockId="16.[151,1188,1935,1960]" box="[151,1188,1935,1960]" pageId="16" pageNumber="317">
|
||
<emphasis bold="true" box="[151,267,1935,1960]" pageId="16" pageNumber="317">
|
||
<typeStatus box="[151,262,1935,1960]" pageId="16" pageNumber="317">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
NSMT-P 106826, 95 mm, Sharm el-Sheikh (
|
||
<geoCoordinate box="[774,926,1935,1960]" direction="east" orientation="longitude" pageId="16" pageNumber="317" precision="15" value="27.858055">
|
||
<number box="[774,804,1936,1960]" pageId="16" pageNumber="317" value="27.0">27</number>
|
||
°
|
||
<number box="[821,851,1935,1960]" pageId="16" pageNumber="317" value="51.0">51</number>
|
||
'
|
||
<number box="[861,890,1936,1960]" pageId="16" pageNumber="317" value="29.0">29</number>
|
||
'' E
|
||
</geoCoordinate>
|
||
;
|
||
<geoCoordinate box="[938,1093,1936,1960]" direction="north" orientation="latitude" pageId="16" pageNumber="317" precision="15" value="34.294724">
|
||
<number box="[938,968,1936,1960]" pageId="16" pageNumber="317" value="34.0">34</number>
|
||
°
|
||
<number box="[984,1014,1936,1960]" pageId="16" pageNumber="317" value="17.0">17</number>
|
||
'
|
||
<number box="[1024,1053,1936,1960]" pageId="16" pageNumber="317" value="41.0">41</number>
|
||
'' N
|
||
</geoCoordinate>
|
||
),
|
||
<collectingCountry box="[1113,1184,1936,1960]" name="Egypt" pageId="16" pageNumber="317">Egypt</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<caption httpUri="https://zenodo.org/record/229636/files/figure.png" pageId="17" pageNumber="318" targetBox="[340,1246,199,1132]" targetPageId="17">
|
||
<paragraph blockId="17.[151,1435,1144,1197]" pageId="17" pageNumber="318">
|
||
<emphasis bold="true" box="[151,289,1144,1166]" pageId="17" pageNumber="318">FIGURE 7a.</emphasis>
|
||
Holotype of
|
||
<taxonomicName box="[427,645,1144,1166]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="318" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[427,645,1144,1166]" italics="true" pageId="17" pageNumber="318">Pempheris tominagai</emphasis>
|
||
</taxonomicName>
|
||
(NSMT-P106826), 95.1 mm SL, Sharm el Moya Bay, Egypt.
|
||
<emphasis bold="true" box="[1286,1309,1144,1166]" pageId="17" pageNumber="318">b:</emphasis>
|
||
Underwater photograph of
|
||
<taxonomicName authority="Tulear" authorityName="Tulear" box="[300,507,1174,1197]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="318" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[300,427,1176,1197]" italics="true" pageId="17" pageNumber="318">
|
||
P.
|
||
<taxonomicName authority="Tulear" authorityName="Tulear" box="[325,427,1176,1197]" pageId="17" pageNumber="318" rank="species" species="tominagai">tominagai</taxonomicName>
|
||
</emphasis>
|
||
, Tulear
|
||
</taxonomicName>
|
||
, Madagascar, taken by G.R. Allen.
|
||
</paragraph>
|
||
</caption>
|
||
<caption box="[151,1095,1637,1659]" httpUri="https://zenodo.org/record/229637/files/figure.png" pageId="17" pageNumber="318" targetBox="[390,1196,1283,1621]" targetPageId="17">
|
||
<paragraph blockId="17.[151,1095,1637,1659]" box="[151,1095,1637,1659]" pageId="17" pageNumber="318">
|
||
<emphasis bold="true" box="[151,274,1637,1659]" pageId="17" pageNumber="318">FIGURE 8.</emphasis>
|
||
Illustrations of opened posterior nostril (a), and compressed posterior nostril (b).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="17.[151,1437,1698,2035]" pageId="17" pageNumber="318">
|
||
<emphasis bold="true" box="[199,326,1698,1723]" pageId="17" pageNumber="318">
|
||
<typeStatus box="[199,321,1698,1723]" pageId="17" pageNumber="318">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
NSMT-P 106827, 106828 (
|
||
<number box="[646,662,1699,1723]" pageId="17" pageNumber="318" value="2.0">2</number>
|
||
), 82, 99 mm, Sharm el-Sheikh,
|
||
<collectingCountry box="[1031,1102,1699,1723]" name="Egypt" pageId="17" pageNumber="318">Egypt</collectingCountry>
|
||
;
|
||
<collectionCode LSID="urn:lsid:biocol.org:col:34984" box="[1115,1199,1699,1723]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="17" pageNumber="318">SAIAB</collectionCode>
|
||
77935, 92 mm, Cap Matoopa,
|
||
<collectingCountry box="[266,390,1733,1758]" name="Seychelles" pageId="17" pageNumber="318">Seychelles</collectingCountry>
|
||
;
|
||
<collectionCode LSID="urn:lsid:biocol.org:col:34984" box="[402,486,1734,1758]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="17" pageNumber="318">SAIAB</collectionCode>
|
||
<number box="[495,565,1733,1758]" pageId="17" pageNumber="318" value="97406.0">97406</number>
|
||
(
|
||
<number box="[583,599,1734,1758]" pageId="17" pageNumber="318" value="2.0">2</number>
|
||
),
|
||
<quantity box="[622,774,1733,1758]" metricMagnitude="2" metricUnit="m" metricValue="1.16117" pageId="17" pageNumber="318" unit="mm" value="116117.0">116, 117 mm</quantity>
|
||
, Libanona Beach,
|
||
<collectingCountry box="[988,1127,1733,1758]" name="Madagascar" pageId="17" pageNumber="318">Madagascar</collectingCountry>
|
||
;
|
||
<collectionCode LSID="urn:lsid:biocol.org:col:34984" box="[1139,1223,1734,1758]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="17" pageNumber="318">SAIAB</collectionCode>
|
||
62146, 75871 (
|
||
<number box="[1406,1421,1733,1758]" pageId="17" pageNumber="318" value="5.0">5</number>
|
||
),
|
||
<quantity box="[151,289,1767,1792]" metricMagnitude="-2" metricUnit="m" metricValue="8.85" metricValueMax="11.7" metricValueMin="6.0" pageId="17" pageNumber="318" unit="mm" value="88.5" valueMax="117.0" valueMin="60.0">60–117 mm</quantity>
|
||
, KwaZulu-Natal,
|
||
<collectingCountry box="[494,643,1767,1792]" name="South Africa" pageId="17" pageNumber="318">South Africa</collectingCountry>
|
||
; URM-P
|
||
<quantity box="[754,937,1767,1792]" metricMagnitude="4" metricUnit="m" metricValue="4.63361" pageId="17" pageNumber="318" unit="mm" value="4.63361E7">46336, 100 mm</quantity>
|
||
,
|
||
<collectingCountry box="[951,1036,1768,1791]" name="Oman" pageId="17" pageNumber="318">Muscat</collectingCountry>
|
||
,
|
||
<collectingCountry box="[1049,1121,1768,1792]" name="Oman" pageId="17" pageNumber="318">Oman</collectingCountry>
|
||
; URM-P 46567, 46568 (
|
||
<number box="[1406,1422,1768,1792]" pageId="17" pageNumber="318" value="2.0">2</number>
|
||
),
|
||
<quantity box="[151,287,1803,1827]" metricMagnitude="1" metricUnit="m" metricValue="9.309999999999999" pageId="17" pageNumber="318" unit="mm" value="93100.0">93, 100 mm</quantity>
|
||
, Sharm el-Sheikh,
|
||
<collectingCountry box="[499,570,1803,1827]" name="Egypt" pageId="17" pageNumber="318">Egypt</collectingCountry>
|
||
;
|
||
<collectionCode box="[580,664,1803,1826]" pageId="17" pageNumber="318">BMNH</collectionCode>
|
||
<number box="[671,753,1803,1827]" pageId="17" pageNumber="318" value="1982.9">1982.9</number>
|
||
.
|
||
<number box="[757,792,1803,1827]" pageId="17" pageNumber="318" value="9.1">9.1</number>
|
||
,
|
||
<quantity box="[804,884,1802,1827]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="318" unit="mm" value="60.0">60 mm</quantity>
|
||
,
|
||
<collectingCountry box="[896,1009,1802,1827]" name="Kenya" pageId="17" pageNumber="318">Mombasa</collectingCountry>
|
||
,
|
||
<collectingCountry box="[1020,1097,1803,1827]" name="Kenya" pageId="17" pageNumber="318">Kenya</collectingCountry>
|
||
;
|
||
<collectionCode LSID="urn:lsid:biocol.org:col:34840" box="[1107,1183,1803,1827]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="17" pageNumber="318">SMNS</collectionCode>
|
||
<quantity box="[1190,1367,1803,1827]" metricMagnitude="4" metricUnit="m" metricValue="2.1172107" pageId="17" pageNumber="318" unit="mm" value="2.1172107E7">21172, 107 mm</quantity>
|
||
, Saint Paul (
|
||
<geoCoordinate box="[216,365,1838,1862]" direction="south" orientation="latitude" pageId="17" pageNumber="318" precision="15" value="-21.078056">
|
||
<number box="[216,246,1838,1862]" pageId="17" pageNumber="318" value="21.0">21</number>
|
||
° 04'
|
||
<number box="[303,332,1838,1862]" pageId="17" pageNumber="318" value="41.0">41</number>
|
||
'' S
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[378,529,1837,1862]" direction="east" orientation="longitude" pageId="17" pageNumber="318" precision="15" value="55.2175">
|
||
<number box="[378,408,1837,1862]" pageId="17" pageNumber="318" value="55.0">55</number>
|
||
°
|
||
<number box="[424,454,1838,1862]" pageId="17" pageNumber="318" value="13.0">13</number>
|
||
' 03'' E
|
||
</geoCoordinate>
|
||
),
|
||
<collectingCountry box="[549,662,1837,1862]" name="Mauritius" pageId="17" pageNumber="318">Mauritius</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="19" lastPageNumber="320" pageId="17" pageNumber="318" type="diagnosis">
|
||
<paragraph blockId="17.[151,1437,1698,2035]" pageId="17" pageNumber="318">
|
||
<emphasis bold="true" box="[199,321,1871,1896]" pageId="17" pageNumber="318">Diagnosis.</emphasis>
|
||
Pored lateral-line scales
|
||
<number box="[612,686,1871,1896]" pageId="17" pageNumber="318" value="49.5" valueMax="52.0" valueMin="47.0">47–52</number>
|
||
; scale rows above lateral line
|
||
<number box="[1036,1050,1872,1896]" pageId="17" pageNumber="318" value="4.0">4</number>
|
||
<number box="[1059,1067,1873,1895]" pageId="17" pageNumber="318" value="1.0">1</number>
|
||
/
|
||
<number box="[1076,1089,1873,1895]" pageId="17" pageNumber="318" value="2.0">2</number>
|
||
; scale rows below lateral line
|
||
<number box="[151,223,1907,1931]" pageId="17" pageNumber="318" value="10.5" valueMax="11.0" valueMin="10.0">10–11</number>
|
||
; predorsal scales
|
||
<number box="[434,507,1906,1931]" pageId="17" pageNumber="318" value="27.5" valueMax="30.0" valueMin="25.0">25–30</number>
|
||
; circumpeduncular scales
|
||
<number box="[816,847,1907,1931]" pageId="17" pageNumber="318" value="12.0">12</number>
|
||
; body depth 40.0–
|
||
<number box="[1070,1119,1907,1931]" pageId="17" pageNumber="318" value="47.7">47.7</number>
|
||
% SL; no blackish spot on pectoral fin base; anterior edge of dorsal fin usually blackish, the rest faint orange; anal fin pale with base zonal black, and margin very faintly black; chest keeled, rarely weakly so; posterior (outer) nostril (closest to eye) usually open, not compressed (
|
||
<figureCitation box="[497,566,2010,2035]" captionStart="FIGURE 8" captionStartId="17.[151,250,1637,1659]" captionTargetBox="[390,1196,1283,1621]" captionTargetId="figure@17.[390,1196,1283,1622]" captionTargetPageId="17" captionText="FIGURE 8. Illustrations of opened posterior nostril (a), and compressed posterior nostril (b)." httpUri="https://zenodo.org/record/229637/files/figure.png" pageId="17" pageNumber="318">
|
||
Fig.
|
||
<number box="[550,566,2011,2035]" pageId="17" pageNumber="318" value="8.0">8</number>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption pageId="18" pageNumber="319">
|
||
<paragraph blockId="18.[151,1436,151,210]" pageId="18" pageNumber="319">
|
||
<emphasis bold="true" box="[151,276,152,175]" pageId="18" pageNumber="319">TABLE 11.</emphasis>
|
||
Counts of
|
||
<taxonomicName box="[402,631,151,175]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[402,631,151,175]" italics="true" pageId="18" pageNumber="319">Pempheris tominagai</emphasis>
|
||
</taxonomicName>
|
||
, holotype, paratypes and non-types, syntypes of
|
||
<taxonomicName box="[1152,1283,151,175]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="schwenkii">
|
||
<emphasis box="[1152,1283,151,175]" italics="true" pageId="18" pageNumber="319">P. schwenkii</emphasis>
|
||
</taxonomicName>
|
||
, and the other specimens.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="18.[519,1324,238,291]" lastBlockId="18.[159,1312,352,415]" pageId="18" pageNumber="319">
|
||
<emphasis box="[519,818,240,261]" italics="true" pageId="18" pageNumber="319">
|
||
<taxonomicName box="[519,647,240,261]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="tominagai">P. tominagai</taxonomicName>
|
||
<taxonomicName box="[690,818,240,261]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="tominagai">P. tominagai</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicName box="[969,1096,238,260]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="schwenkii">P. schwenkii</taxonomicName>
|
||
<taxonomicName box="[1198,1324,238,260]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="schwenkii">P. schwenkii</taxonomicName>
|
||
<typeStatus box="[519,613,269,291]" pageId="18" pageNumber="319">Holotype</typeStatus>
|
||
<typeStatus box="[690,789,270,291]" pageId="18" pageNumber="319">Paratypes</typeStatus>
|
||
and non-types
|
||
<typeStatus box="[969,1062,269,291]" pageId="18" pageNumber="319">Syntypes</typeStatus>
|
||
non-types Number of Individuals
|
||
<number box="[519,532,352,374]" pageId="18" pageNumber="319" value="1.0">1</number>
|
||
<number box="[690,728,352,374]" pageId="18" pageNumber="319" value="124.0">124</number>
|
||
<number box="[969,982,352,374]" pageId="18" pageNumber="319" value="2.0">2</number>
|
||
<number box="[1198,1236,352,374]" pageId="18" pageNumber="319" value="176.0">176</number>
|
||
</paragraph>
|
||
<paragraph blockId="18.[159,1312,352,415]" lastBlockId="18.[159,1431,476,611]" pageId="18" pageNumber="319">
|
||
Standard length (mm)
|
||
<number box="[519,563,393,415]" pageId="18" pageNumber="319" value="95.1">95.1</number>
|
||
<number box="[690,804,393,415]" pageId="18" pageNumber="319" value="80.15" valueMax="106.9" valueMin="53.4">53.4‒106.9</number>
|
||
<number box="[969,1016,393,415]" pageId="18" pageNumber="319" value="87.2">87.2</number>
|
||
,
|
||
<number box="[1026,1070,393,415]" pageId="18" pageNumber="319" value="89.7">89.7</number>
|
||
<number box="[1198,1312,393,415]" pageId="18" pageNumber="319" value="80.65" valueMax="125.9" valueMin="35.4">35.4‒125.9</number>
|
||
Counts
|
||
</paragraph>
|
||
<paragraph blockId="18.[159,1431,476,611]" box="[159,1431,517,539]" pageId="18" pageNumber="319">
|
||
Dorsal-fin rays VI,
|
||
<date box="[558,719,517,539]" pageId="18" pageNumber="319">
|
||
<number box="[558,571,517,539]" pageId="18" pageNumber="319" value="9.0">9</number>
|
||
VI
|
||
</date>
|
||
,
|
||
<date box="[729,998,517,539]" pageId="18" pageNumber="319">
|
||
<number box="[729,742,517,539]" pageId="18" pageNumber="319" value="9.0">9</number>
|
||
VI
|
||
</date>
|
||
,
|
||
<date box="[1009,1227,517,539]" pageId="18" pageNumber="319">
|
||
<number box="[1009,1022,517,539]" pageId="18" pageNumber="319" value="9.0">9</number>
|
||
VI
|
||
</date>
|
||
, VII rarely VII,
|
||
</paragraph>
|
||
<paragraph blockId="18.[159,1431,476,611]" lastBlockId="18.[151,1437,1175,1946]" pageId="18" pageNumber="319">
|
||
<number box="[1198,1249,548,570]" pageId="18" pageNumber="319" value="9.5" valueMax="10.0" valueMin="9.0">9‒10</number>
|
||
rarely
|
||
<number box="[1321,1346,548,570]" pageId="18" pageNumber="319" value="10.0">10</number>
|
||
Anal-fin rays III,
|
||
<date box="[557,718,589,611]" pageId="18" pageNumber="319" value="1938-03">
|
||
<number box="[557,582,589,611]" pageId="18" pageNumber="319" value="38.0">38</number>
|
||
III
|
||
</date>
|
||
,
|
||
<number box="[728,791,589,611]" pageId="18" pageNumber="319" value="37.5" valueMax="40.0" valueMin="35.0">35‒40</number>
|
||
rarely 35,
|
||
<date box="[900,997,589,611]" pageId="18" pageNumber="319" value="1936-03">36 III</date>
|
||
, 35,
|
||
<date box="[1045,1226,589,611]" pageId="18" pageNumber="319" value="1936-03">36 III</date>
|
||
,
|
||
<number box="[1236,1300,589,611]" pageId="18" pageNumber="319" value="38.5" valueMax="42.0" valueMin="35.0">35‒42</number>
|
||
<emphasis bold="true" box="[199,345,1175,1200]" pageId="18" pageNumber="319">Description.</emphasis>
|
||
Body strongly compressed, body width
|
||
<quantity box="[793,906,1175,1200]" metricMagnitude="-1" metricUnit="m" metricValue="1.8415" metricValueMax="2.0827999999999998" metricValueMin="1.6001999999999998" pageId="18" pageNumber="319" unit="in" value="7.25" valueMax="8.2" valueMin="6.3">6.3–8.2 in</quantity>
|
||
SL; body deep, body depth
|
||
<quantity box="[1218,1330,1175,1200]" metricMagnitude="-2" metricUnit="m" metricValue="5.715" metricValueMax="5.842" metricValueMin="5.588" pageId="18" pageNumber="319" unit="in" value="2.25" valueMax="2.3" valueMin="2.2">2.2–2.3 in</quantity>
|
||
SL; head length
|
||
<quantity box="[229,343,1209,1234]" metricMagnitude="-2" metricUnit="m" metricValue="8.382" metricValueMax="8.636" metricValueMin="8.128" pageId="18" pageNumber="319" unit="in" value="3.3" valueMax="3.4" valueMin="3.2">3.2–3.4 in</quantity>
|
||
SL; snout very short, rounded, snout length
|
||
<quantity box="[851,965,1209,1234]" metricMagnitude="-1" metricUnit="m" metricValue="1.0795" metricValueMax="1.2192" metricValueMin="0.9398" pageId="18" pageNumber="319" unit="in" value="4.25" valueMax="4.8" valueMin="3.7">3.7–4.8 in</quantity>
|
||
HL; eye large
|
||
<quantity box="[1137,1251,1209,1234]" metricMagnitude="-2" metricUnit="m" metricValue="5.969" metricValueMax="6.35" metricValueMin="5.588" pageId="18" pageNumber="319" unit="in" value="2.35" valueMax="2.5" valueMin="2.2">2.2–2.5 in</quantity>
|
||
HL; interorbital very narrow 3.0–
|
||
<quantity box="[354,423,1242,1267]" metricMagnitude="-2" metricUnit="m" metricValue="9.398" pageId="18" pageNumber="319" unit="in" value="3.7">3.7 in</quantity>
|
||
HL; dorsal outline convex to dorsal fin base, extending somewhat linearly to caudal peduncle; ventral outline convex to anal fin base, extending somewhat linearly to caudal peduncle.
|
||
</paragraph>
|
||
<paragraph pageId="18" pageNumber="319">
|
||
<table box="[159,1431,630,1107]" gridcols="4" gridrows="5" pageId="18" pageNumber="319">
|
||
<tr box="[159,1431,630,652]" gridrow="0" pageId="18" pageNumber="319">
|
||
<th box="[159,584,630,652]" gridcol="0" gridrow="0" pageId="18" pageNumber="319">Pectoral-fin rays I, 17</th>
|
||
<th box="[689,805,630,652]" gridcol="1" gridrow="0" pageId="18" pageNumber="319">I, 16‒17</th>
|
||
<th box="[969,1060,630,652]" gridcol="2" gridrow="0" pageId="18" pageNumber="319">I, 17</th>
|
||
<th box="[1198,1431,630,652]" gridcol="3" gridrow="0" pageId="18" pageNumber="319">I, 15‒17</th>
|
||
</tr>
|
||
<tr box="[159,1431,672,776]" gridrow="1" pageId="18" pageNumber="319">
|
||
<th box="[159,584,672,776]" gridcol="0" gridrow="1" pageId="18" pageNumber="319">Pelvic-fin rays I, 5 Caudal-fin rays 9+8 Proccurent caudal rays 8+5</th>
|
||
<td box="[689,805,672,776]" gridcol="1" gridrow="1" pageId="18" pageNumber="319">I, 5 9+8 6‒8+4‒6</td>
|
||
<td box="[969,1060,672,776]" gridcol="2" gridrow="1" pageId="18" pageNumber="319">I, 5 9+8 N/A</td>
|
||
<td box="[1198,1431,672,776]" gridcol="3" gridrow="1" pageId="18" pageNumber="319">I, 5 9+8 6‒8+4‒6</td>
|
||
</tr>
|
||
<tr box="[159,1431,796,900]" gridrow="2" pageId="18" pageNumber="319">
|
||
<th box="[159,584,796,900]" gridcol="0" gridrow="2" pageId="18" pageNumber="319">Left pored lateral-line scales 51 Right pored lateral-line scales 49 Scale rows above lateral line 4 1/2</th>
|
||
<td box="[689,805,796,900]" gridcol="1" gridrow="2" pageId="18" pageNumber="319">47‒52 47‒52 4 1/2</td>
|
||
<td box="[969,1060,796,900]" gridcol="2" gridrow="2" pageId="18" pageNumber="319">48 48, 49 3 1/2</td>
|
||
<td box="[1198,1431,796,900]" gridcol="3" gridrow="2" pageId="18" pageNumber="319">44‒53 45‒53 3 1/2</td>
|
||
</tr>
|
||
<tr box="[159,1431,920,1024]" gridrow="3" pageId="18" pageNumber="319">
|
||
<th box="[159,584,920,1024]" gridcol="0" gridrow="3" pageId="18" pageNumber="319">Scale rows below lateral line 11 Predorsal scales 30 Circumpeduncular scales 12</th>
|
||
<td box="[689,805,920,1024]" gridcol="1" gridrow="3" pageId="18" pageNumber="319">10‒11 25‒30 12</td>
|
||
<td box="[969,1060,920,1024]" gridcol="2" gridrow="3" pageId="18" pageNumber="319">10 damaged 12</td>
|
||
<td box="[1198,1431,920,1024]" gridcol="3" gridrow="3" pageId="18" pageNumber="319">10‒12 23‒30 10‒12</td>
|
||
</tr>
|
||
<tr box="[159,1431,1044,1107]" gridrow="4" pageId="18" pageNumber="319">
|
||
<th box="[159,584,1044,1107]" gridcol="0" gridrow="4" pageId="18" pageNumber="319">Gill rakers 9+19 Vertebrae 10+15</th>
|
||
<td box="[689,805,1044,1107]" gridcol="1" gridrow="4" pageId="18" pageNumber="319">7‒9+19‒22 10+15</td>
|
||
<td box="[969,1060,1044,1107]" gridcol="2" gridrow="4" pageId="18" pageNumber="319">7+18‒19 N/A</td>
|
||
<td box="[1198,1431,1044,1107]" gridcol="3" gridrow="4" pageId="18" pageNumber="319">6‒9+18‒22 10+15</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="18.[151,1437,1175,1946]" pageId="18" pageNumber="319">
|
||
Mouth large and strongly oblique, upper jaw length
|
||
<quantity box="[793,908,1309,1334]" metricMagnitude="-2" metricUnit="m" metricValue="4.699" metricValueMax="4.826" metricValueMin="4.572" pageId="18" pageNumber="319" unit="in" value="1.85" valueMax="1.9" valueMin="1.8">1.8–1.9 in</quantity>
|
||
HL; the maxilla extending to middle of pupil; maxilla broadly expanded posteriorly, its surface with level; lower jaw projecting beyond upper jaw; narrow band of villiform teeth in jaws.
|
||
</paragraph>
|
||
<paragraph blockId="18.[151,1437,1175,1946]" pageId="18" pageNumber="319">Scales ctenoid, thin, semicircular in shape, far wider than long, and very deciduous; concealed small scales under surface scales; scales of ventral area larger than those of dorsal area; almost all parts of head scaled, except for just anterior to orbit; about one-third of basal part of anal fin covered with small scales; scales above lateral line more deciduous than body side scales; pored lateral-line scales adherent, running parallel to dorsal outline to middle of caudal fin.</paragraph>
|
||
<paragraph blockId="18.[151,1437,1175,1946]" pageId="18" pageNumber="319">
|
||
<emphasis box="[199,458,1597,1621]" italics="true" pageId="18" pageNumber="319">Coloration when fresh.</emphasis>
|
||
Body scales gold in daylight (
|
||
<bibRefCitation author="Allen" box="[795,1042,1597,1622]" pageId="18" pageNumber="319" refString="Allen, G. R. & Steene, R. C. (1987) Reef Fishes of the Indian Ocean. TFH Publications, Neptune City, New Jersey, 240 pp." type="book" year="1987">
|
||
Allen & Steene,
|
||
<number box="[982,1042,1598,1622]" pageId="18" pageNumber="319" value="1987.0">1987</number>
|
||
</bibRefCitation>
|
||
); paired fins pink to orange; caudal fin usually yellowish; body coloration silvery, and coloration of fins pale at night.
|
||
</paragraph>
|
||
<paragraph blockId="18.[151,1437,1175,1946]" pageId="18" pageNumber="319">
|
||
<emphasis box="[199,515,1669,1693]" italics="true" pageId="18" pageNumber="319">Coloration when preserved.</emphasis>
|
||
Body scales light brown; areas missing scales pale brown with irregular vertical dark brown lines marking scale pockets; black pigmentation on dorsal and anal fin usually persistent; blackish pigmentation on anal fin margin sometimes not discernable; posterior edge of caudal fin dusky; paired fins hyaline.
|
||
</paragraph>
|
||
<paragraph blockId="18.[151,1437,1175,1946]" pageId="18" pageNumber="319">
|
||
<emphasis bold="true" box="[199,309,1777,1802]" pageId="18" pageNumber="319">Genetics.</emphasis>
|
||
On the mitochondrial
|
||
<number box="[570,599,1777,1802]" pageId="18" pageNumber="319" value="16.0">16</number>
|
||
S ribosomal
|
||
<collectionCode LSID="urn:lsid:biocol.org:col:15536" box="[743,807,1778,1802]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="18" pageNumber="319">DNA</collectionCode>
|
||
, two haplotypes occurred in
|
||
<taxonomicName box="[1139,1282,1778,1801]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[1139,1282,1778,1801]" italics="true" pageId="18" pageNumber="319">P. tominagai</emphasis>
|
||
</taxonomicName>
|
||
, and a single specimen from the Indian Ocean (
|
||
<collectingCountry box="[542,665,1813,1838]" name="Seychelles" pageId="18" pageNumber="319">Seychelles</collectingCountry>
|
||
) had slightly different sequence from the other Red Sea specimens (
|
||
<figureCitation box="[159,233,1849,1874]" captionStart="FIGURE 2" captionStartId="7.[151,250,1821,1843]" captionTargetBox="[162,1423,204,1793]" captionTargetId="figure@7.[151,1436,194,1800]" captionTargetPageId="7" captionText="FIGURE 2. Phylogenetic tree of the 15 Pempheris species recovered from mitochondria 16 S, estimated by the maximum likelihood (ML) method, using as out group Parapriacanthus ransonetii (Pempheridae). Only bootstrap values based on 1000 replications higher than 50 % by ML (left) and neighbor-joining (right) are displayed, respectively." httpUri="https://zenodo.org/record/229632/files/figure.png" pageId="18" pageNumber="319">
|
||
Fig.
|
||
<number box="[216,233,1850,1874]" pageId="18" pageNumber="319" value="2.0">2</number>
|
||
</figureCitation>
|
||
). The molecular analysis shows that
|
||
<taxonomicName box="[671,816,1850,1873]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[671,816,1850,1873]" italics="true" pageId="18" pageNumber="319">P. tominagai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[879,1021,1849,1873]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="schwenkii">
|
||
<emphasis box="[879,1021,1849,1873]" italics="true" pageId="18" pageNumber="319">P. schwenkii</emphasis>
|
||
</taxonomicName>
|
||
, which is distributed in the Pacific Ocean, are sister species. However, these two species can clearly be separated by a minimum of
|
||
<number box="[1266,1302,1886,1910]" pageId="18" pageNumber="319" value="2.1">2.1</number>
|
||
% sequence divergence, and comprised different monophyletic groups.
|
||
</paragraph>
|
||
<caption pageId="19" pageNumber="320">
|
||
<paragraph blockId="19.[151,1435,151,214]" pageId="19" pageNumber="320">
|
||
<emphasis bold="true" box="[151,280,152,175]" pageId="19" pageNumber="320">TABLE 12.</emphasis>
|
||
Measurements of
|
||
<taxonomicName box="[490,720,151,175]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[490,720,151,175]" italics="true" pageId="19" pageNumber="320">Pempheris tominagai</emphasis>
|
||
</taxonomicName>
|
||
, holotype, paratypes and non-types, syntypes of
|
||
<taxonomicName box="[1250,1383,151,175]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="schwenkii">
|
||
<emphasis box="[1250,1383,151,175]" italics="true" pageId="19" pageNumber="320">P. schwenkii</emphasis>
|
||
</taxonomicName>
|
||
, and the other specimens.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="19.[159,1367,243,1524]" pageId="19" pageNumber="320">
|
||
<emphasis box="[478,771,244,265]" italics="true" pageId="19" pageNumber="320">
|
||
<taxonomicName box="[478,605,244,265]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="tominagai">P. tominagai</taxonomicName>
|
||
<taxonomicName box="[644,771,244,265]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="tominagai">P. tominagai</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicName box="[956,1082,243,265]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="schwenkii">P. schwenkii</taxonomicName>
|
||
<taxonomicName box="[1198,1324,243,265]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="schwenkii">P. schwenkii</taxonomicName>
|
||
<typeStatus box="[478,572,273,295]" pageId="19" pageNumber="320">Holotype</typeStatus>
|
||
<typeStatus box="[644,743,274,295]" pageId="19" pageNumber="320">Paratypes</typeStatus>
|
||
and non-types
|
||
<typeStatus box="[956,1049,273,295]" pageId="19" pageNumber="320">Syntypes</typeStatus>
|
||
non-types Number of Individuals
|
||
<date box="[478,969,356,378]" pageId="19" pageNumber="320" value="1919-01-02">
|
||
<number box="[478,490,356,378]" pageId="19" pageNumber="320" value="1.0">1</number>
|
||
<number box="[644,670,356,378]" pageId="19" pageNumber="320" value="19.0">19</number>
|
||
<number box="[956,969,356,378]" pageId="19" pageNumber="320" value="2.0">2</number>
|
||
</date>
|
||
<number box="[1198,1223,356,378]" pageId="19" pageNumber="320" value="53.0">53</number>
|
||
</paragraph>
|
||
<paragraph blockId="19.[159,1367,243,1524]" pageId="19" pageNumber="320">
|
||
Standard length (mm)
|
||
<number box="[478,522,397,419]" pageId="19" pageNumber="320" value="95.1">95.1</number>
|
||
<number box="[644,758,397,419]" pageId="19" pageNumber="320" value="91.1" valueMax="107.3" valueMin="74.9">74.9‒107.3</number>
|
||
<number box="[956,1000,397,419]" pageId="19" pageNumber="320" value="89.7">89.7</number>
|
||
<number box="[1198,1312,397,419]" pageId="19" pageNumber="320" value="98.6" valueMax="125.9" valueMin="71.3">71.3‒125.9</number>
|
||
<collectingCountry box="[159,188,480,502]" name="American Samoa" pageId="19" pageNumber="320">As</collectingCountry>
|
||
% of standard length
|
||
</paragraph>
|
||
<paragraph blockId="19.[159,1367,243,1524]" pageId="19" pageNumber="320">
|
||
Head length
|
||
<number box="[478,522,521,543]" pageId="19" pageNumber="320" value="30.2">30.2</number>
|
||
<number box="[644,688,521,543]" pageId="19" pageNumber="320" value="29.1">29.1</number>
|
||
‒32.0 (
|
||
<number box="[761,805,521,543]" pageId="19" pageNumber="320" value="30.6">30.6</number>
|
||
)
|
||
<number box="[956,1003,521,543]" pageId="19" pageNumber="320" value="28.4">28.4</number>
|
||
, 30.0
|
||
<number box="[1198,1242,521,543]" pageId="19" pageNumber="320" value="27.9">27.9</number>
|
||
‒33.0 (
|
||
<number box="[1313,1360,521,543]" pageId="19" pageNumber="320" value="29.7">29.7</number>
|
||
) Head depth
|
||
<number box="[478,522,563,585]" pageId="19" pageNumber="320" value="33.3">33.3</number>
|
||
<number box="[644,745,563,585]" pageId="19" pageNumber="320" value="34.2" valueMax="37.1" valueMin="31.3">31.3‒37.1</number>
|
||
(
|
||
<number box="[761,805,563,585]" pageId="19" pageNumber="320" value="34.3">34.3</number>
|
||
)
|
||
<number box="[956,1003,563,585]" pageId="19" pageNumber="320" value="30.7">30.7</number>
|
||
,
|
||
<number box="[1012,1056,563,585]" pageId="19" pageNumber="320" value="31.1">31.1</number>
|
||
<number box="[1198,1300,563,585]" pageId="19" pageNumber="320" value="33.5" valueMax="36.4" valueMin="30.6">30.6‒36.4</number>
|
||
(
|
||
<number box="[1313,1360,563,585]" pageId="19" pageNumber="320" value="33.4">33.4</number>
|
||
) Snout length
|
||
<number box="[478,510,604,626]" pageId="19" pageNumber="320" value="7.3">7.3</number>
|
||
6.0‒8.0 (
|
||
<number box="[734,768,604,626]" pageId="19" pageNumber="320" value="6.9">6.9</number>
|
||
)
|
||
<number box="[956,988,604,626]" pageId="19" pageNumber="320" value="6.7">6.7</number>
|
||
<number box="[1198,1274,604,626]" pageId="19" pageNumber="320" value="7.0" valueMax="8.4" valueMin="5.6">5.6‒8.4</number>
|
||
(
|
||
<number box="[1288,1322,604,626]" pageId="19" pageNumber="320" value="6.8">6.8</number>
|
||
) Eye diameter
|
||
<number box="[478,522,645,667]" pageId="19" pageNumber="320" value="12.5">12.5</number>
|
||
<number box="[644,688,645,667]" pageId="19" pageNumber="320" value="12.2">12.2</number>
|
||
‒15.0 (13.0)
|
||
<number box="[956,1003,645,667]" pageId="19" pageNumber="320" value="12.2">12.2</number>
|
||
,
|
||
<number box="[1012,1056,645,667]" pageId="19" pageNumber="320" value="12.5">12.5</number>
|
||
<number box="[1198,1300,645,667]" pageId="19" pageNumber="320" value="12.399999999999999" valueMax="14.2" valueMin="10.6">10.6‒14.2</number>
|
||
(12.0) Interorbital width
|
||
<number box="[478,510,686,708]" pageId="19" pageNumber="320" value="8.3">8.3</number>
|
||
<number box="[644,674,686,708]" pageId="19" pageNumber="320" value="8.1">8.1</number>
|
||
‒10.0 (
|
||
<number box="[747,780,686,708]" pageId="19" pageNumber="320" value="9.1">9.1</number>
|
||
) 8.0,
|
||
<number box="[1000,1032,686,708]" pageId="19" pageNumber="320" value="8.9">8.9</number>
|
||
<number box="[1198,1274,686,708]" pageId="19" pageNumber="320" value="8.5" valueMax="9.9" valueMin="7.1">7.1‒9.9</number>
|
||
(
|
||
<number box="[1288,1321,686,708]" pageId="19" pageNumber="320" value="8.1">8.1</number>
|
||
) Upper jaw length
|
||
<number box="[478,522,728,750]" pageId="19" pageNumber="320" value="16.7">16.7</number>
|
||
<number box="[644,745,728,750]" pageId="19" pageNumber="320" value="16.6" valueMax="17.3" valueMin="15.9">15.9‒17.3</number>
|
||
(
|
||
<number box="[759,805,728,750]" pageId="19" pageNumber="320" value="16.4">16.4</number>
|
||
)
|
||
<number box="[956,1003,728,750]" pageId="19" pageNumber="320" value="15.9">15.9</number>
|
||
,
|
||
<number box="[1012,1056,728,750]" pageId="19" pageNumber="320" value="15.6">15.6</number>
|
||
<number box="[1198,1241,728,750]" pageId="19" pageNumber="320" value="14.3">14.3</number>
|
||
‒17.0 (
|
||
<number box="[1313,1359,728,750]" pageId="19" pageNumber="320" value="15.6">15.6</number>
|
||
) Predorsal length
|
||
<number box="[478,522,769,791]" pageId="19" pageNumber="320" value="38.6">38.6</number>
|
||
<number box="[644,745,769,791]" pageId="19" pageNumber="320" value="39.25" valueMax="40.1" valueMin="38.4">38.4‒40.1</number>
|
||
(
|
||
<number box="[759,806,769,791]" pageId="19" pageNumber="320" value="39.6">39.6</number>
|
||
)
|
||
<number box="[956,1003,769,791]" pageId="19" pageNumber="320" value="39.8">39.8</number>
|
||
. 40.0 37.0‒
|
||
<number box="[1256,1299,769,791]" pageId="19" pageNumber="320" value="41.1">41.1</number>
|
||
(39.0) Prepelvic length
|
||
<number box="[478,522,811,833]" pageId="19" pageNumber="320" value="40.6">40.6</number>
|
||
39.0‒
|
||
<number box="[701,746,811,833]" pageId="19" pageNumber="320" value="43.2">43.2</number>
|
||
(
|
||
<number box="[759,806,811,833]" pageId="19" pageNumber="320" value="40.9">40.9</number>
|
||
)
|
||
<number box="[956,1003,811,833]" pageId="19" pageNumber="320" value="39.8">39.8</number>
|
||
, 40.0 38.0‒43.0 (
|
||
<number box="[1313,1360,811,833]" pageId="19" pageNumber="320" value="40.4">40.4</number>
|
||
) Preanal length
|
||
<number box="[478,522,852,874]" pageId="19" pageNumber="320" value="54.2">54.2</number>
|
||
53.0‒
|
||
<number box="[702,746,852,874]" pageId="19" pageNumber="320" value="59.8">59.8</number>
|
||
(
|
||
<number box="[759,806,852,874]" pageId="19" pageNumber="320" value="55.5">55.5</number>
|
||
)
|
||
<number box="[956,1003,852,874]" pageId="19" pageNumber="320" value="52.3">52.3</number>
|
||
,
|
||
<number box="[1012,1056,852,874]" pageId="19" pageNumber="320" value="51.1">51.1</number>
|
||
<number box="[1198,1299,852,874]" pageId="19" pageNumber="320" value="54.55" valueMax="59.6" valueMin="49.5">49.5‒59.6</number>
|
||
(
|
||
<number box="[1313,1360,852,874]" pageId="19" pageNumber="320" value="54.4">54.4</number>
|
||
) Body depth
|
||
<number box="[478,522,893,915]" pageId="19" pageNumber="320" value="42.7">42.7</number>
|
||
40.0‒
|
||
<number box="[702,746,893,915]" pageId="19" pageNumber="320" value="47.7">47.7</number>
|
||
(
|
||
<number box="[759,806,893,915]" pageId="19" pageNumber="320" value="43.5">43.5</number>
|
||
)
|
||
<number box="[956,1003,893,915]" pageId="19" pageNumber="320" value="41.1">41.1</number>
|
||
, 42.0
|
||
<number box="[1198,1299,893,915]" pageId="19" pageNumber="320" value="43.7" valueMax="47.2" valueMin="40.2">40.2‒47.2</number>
|
||
(
|
||
<number box="[1313,1360,893,915]" pageId="19" pageNumber="320" value="43.6">43.6</number>
|
||
) Dorsal-fin length
|
||
<number box="[478,522,934,956]" pageId="19" pageNumber="320" value="22.9">22.9</number>
|
||
<number box="[644,746,934,956]" pageId="19" pageNumber="320" value="23.450000000000003" valueMax="25.6" valueMin="21.3">21.3‒25.6</number>
|
||
(23.0) damaged 20.0‒
|
||
<number box="[1256,1300,934,956]" pageId="19" pageNumber="320" value="26.8">26.8</number>
|
||
(
|
||
<number box="[1313,1359,934,956]" pageId="19" pageNumber="320" value="22.1">22.1</number>
|
||
) Anal-fin length
|
||
<number box="[478,522,976,998]" pageId="19" pageNumber="320" value="13.5">13.5</number>
|
||
<number box="[644,676,976,998]" pageId="19" pageNumber="320" value="9.4">9.4</number>
|
||
‒14.0 (
|
||
<number box="[747,793,976,998]" pageId="19" pageNumber="320" value="12.3">12.3</number>
|
||
) damaged
|
||
<number box="[1198,1287,976,998]" pageId="19" pageNumber="320" value="12.45" valueMax="15.5" valueMin="9.4">9.4‒15.5</number>
|
||
(
|
||
<number box="[1300,1346,976,998]" pageId="19" pageNumber="320" value="13.2">13.2</number>
|
||
) Pectoral-fin length
|
||
<number box="[478,522,1017,1039]" pageId="19" pageNumber="320" value="27.1">27.1</number>
|
||
<number box="[644,746,1017,1039]" pageId="19" pageNumber="320" value="26.95" valueMax="28.4" valueMin="25.5">25.5‒28.4</number>
|
||
(
|
||
<number box="[759,805,1017,1039]" pageId="19" pageNumber="320" value="27.1">27.1</number>
|
||
) Damaged
|
||
<number box="[1198,1299,1017,1039]" pageId="19" pageNumber="320" value="26.5" valueMax="29.6" valueMin="23.4">23.4‒29.6</number>
|
||
(
|
||
<number box="[1313,1360,1017,1039]" pageId="19" pageNumber="320" value="25.4">25.4</number>
|
||
) Pelvic-fin length
|
||
<number box="[478,522,1059,1081]" pageId="19" pageNumber="320" value="13.5">13.5</number>
|
||
<number box="[644,746,1059,1081]" pageId="19" pageNumber="320" value="12.25" valueMax="14.1" valueMin="10.4">10.4‒14.1</number>
|
||
(13.0) Damaged
|
||
<number box="[1198,1287,1059,1081]" pageId="19" pageNumber="320" value="12.0" valueMax="14.1" valueMin="9.9">9.9‒14.1</number>
|
||
(
|
||
<number box="[1300,1346,1059,1081]" pageId="19" pageNumber="320" value="12.8">12.8</number>
|
||
) Dorsal-fin base
|
||
<number box="[478,522,1100,1122]" pageId="19" pageNumber="320" value="15.6">15.6</number>
|
||
<number box="[644,746,1100,1122]" pageId="19" pageNumber="320" value="16.9" valueMax="18.2" valueMin="15.6">15.6‒18.2</number>
|
||
(
|
||
<number box="[759,805,1100,1122]" pageId="19" pageNumber="320" value="16.4">16.4</number>
|
||
)
|
||
<number box="[956,1002,1100,1122]" pageId="19" pageNumber="320" value="16.7">16.7</number>
|
||
, 17.0 14.0‒
|
||
<number box="[1256,1300,1100,1122]" pageId="19" pageNumber="320" value="17.9">17.9</number>
|
||
(16.0) Anal-fin base 49.0
|
||
<number box="[644,746,1141,1163]" pageId="19" pageNumber="320" value="50.3" valueMax="54.7" valueMin="45.9">45.9‒54.7</number>
|
||
(
|
||
<number box="[759,806,1141,1163]" pageId="19" pageNumber="320" value="49.5">49.5</number>
|
||
)
|
||
<number box="[956,1003,1141,1163]" pageId="19" pageNumber="320" value="48.9">48.9</number>
|
||
, 50.0
|
||
<number box="[1198,1300,1141,1163]" pageId="19" pageNumber="320" value="49.849999999999994" valueMax="53.8" valueMin="45.9">45.9‒53.8</number>
|
||
(
|
||
<number box="[1313,1360,1141,1163]" pageId="19" pageNumber="320" value="50.4">50.4</number>
|
||
) Caudal-peduncle length
|
||
<number box="[478,522,1182,1204]" pageId="19" pageNumber="320" value="10.4">10.4</number>
|
||
<number box="[644,734,1182,1204]" pageId="19" pageNumber="320" value="9.6" valueMax="10.6" valueMin="8.6">8.6‒10.6</number>
|
||
(
|
||
<number box="[747,781,1182,1204]" pageId="19" pageNumber="320" value="9.6">9.6</number>
|
||
)
|
||
<number box="[956,990,1182,1204]" pageId="19" pageNumber="320" value="8.9">8.9</number>
|
||
,
|
||
<number box="[1198,1287,1182,1204]" pageId="19" pageNumber="320" value="8.899999999999999" valueMax="10.2" valueMin="7.6">7.6‒10.2</number>
|
||
(9.0)
|
||
</paragraph>
|
||
<paragraph blockId="19.[159,1367,243,1524]" box="[956,987,1213,1235]" pageId="19" pageNumber="320">
|
||
<number box="[956,987,1213,1235]" pageId="19" pageNumber="320" value="9.1">9.1</number>
|
||
</paragraph>
|
||
<paragraph blockId="19.[159,1367,243,1524]" pageId="19" pageNumber="320">
|
||
Caudal-peduncle depth
|
||
<number box="[478,510,1254,1276]" pageId="19" pageNumber="320" value="8.3">8.3</number>
|
||
<number box="[644,733,1254,1276]" pageId="19" pageNumber="320" value="9.15" valueMax="10.5" valueMin="7.8">7.8‒10.5</number>
|
||
(
|
||
<number box="[747,780,1254,1276]" pageId="19" pageNumber="320" value="8.7">8.7</number>
|
||
)
|
||
<number box="[956,990,1254,1276]" pageId="19" pageNumber="320" value="7.8">7.8</number>
|
||
,
|
||
<number box="[1000,1031,1254,1276]" pageId="19" pageNumber="320" value="9.1">9.1</number>
|
||
<number box="[1198,1287,1254,1276]" pageId="19" pageNumber="320" value="8.75" valueMax="10.1" valueMin="7.4">7.4‒10.1</number>
|
||
(
|
||
<number box="[1300,1334,1254,1276]" pageId="19" pageNumber="320" value="8.6">8.6</number>
|
||
)
|
||
<collectingCountry box="[159,188,1337,1359]" name="American Samoa" pageId="19" pageNumber="320">As</collectingCountry>
|
||
% of head length
|
||
</paragraph>
|
||
<paragraph blockId="19.[159,1367,243,1524]" lastBlockId="19.[151,1436,1596,2011]" pageId="19" pageNumber="320">
|
||
Snout length
|
||
<number box="[478,522,1379,1401]" pageId="19" pageNumber="320" value="24.1">24.1</number>
|
||
<number box="[644,745,1379,1401]" pageId="19" pageNumber="320" value="23.4" valueMax="26.1" valueMin="20.7">20.7‒26.1</number>
|
||
(
|
||
<number box="[761,805,1379,1401]" pageId="19" pageNumber="320" value="22.9">22.9</number>
|
||
)
|
||
<number box="[956,1003,1379,1401]" pageId="19" pageNumber="320" value="22.2">22.2</number>
|
||
, 24.0
|
||
<number box="[1198,1300,1379,1401]" pageId="19" pageNumber="320" value="22.75" valueMax="27.3" valueMin="18.2">18.2‒27.3</number>
|
||
(23.0) Eye diameter
|
||
<number box="[478,522,1420,1442]" pageId="19" pageNumber="320" value="41.4">41.4</number>
|
||
40.0‒
|
||
<number box="[702,746,1420,1442]" pageId="19" pageNumber="320" value="46.9">46.9</number>
|
||
(
|
||
<number box="[761,805,1420,1442]" pageId="19" pageNumber="320" value="42.8">42.8</number>
|
||
)
|
||
<number box="[956,1003,1420,1442]" pageId="19" pageNumber="320" value="40.7">40.7</number>
|
||
, 44.0
|
||
<number box="[1198,1300,1420,1442]" pageId="19" pageNumber="320" value="40.5" valueMax="45.5" valueMin="35.5">35.5‒45.5</number>
|
||
(
|
||
<number box="[1313,1360,1420,1442]" pageId="19" pageNumber="320" value="40.4">40.4</number>
|
||
) Interorbital width
|
||
<number box="[478,522,1461,1483]" pageId="19" pageNumber="320" value="27.6">27.6</number>
|
||
<number box="[644,746,1461,1483]" pageId="19" pageNumber="320" value="30.45" valueMax="33.3" valueMin="27.6">27.6‒33.3</number>
|
||
(
|
||
<number box="[759,806,1461,1483]" pageId="19" pageNumber="320" value="29.9">29.9</number>
|
||
) 28.0,
|
||
<number box="[1012,1056,1461,1483]" pageId="19" pageNumber="320" value="29.6">29.6</number>
|
||
<number box="[1198,1300,1461,1483]" pageId="19" pageNumber="320" value="27.799999999999997" valueMax="32.3" valueMin="23.3">23.3‒32.3</number>
|
||
(
|
||
<number box="[1313,1360,1461,1483]" pageId="19" pageNumber="320" value="27.3">27.3</number>
|
||
) Upper jaw length
|
||
<number box="[478,522,1502,1524]" pageId="19" pageNumber="320" value="55.2">55.2</number>
|
||
50.0‒
|
||
<number box="[702,746,1502,1524]" pageId="19" pageNumber="320" value="56.7">56.7</number>
|
||
(
|
||
<number box="[759,806,1502,1524]" pageId="19" pageNumber="320" value="53.8">53.8</number>
|
||
)
|
||
<number box="[956,1003,1502,1524]" pageId="19" pageNumber="320" value="51.9">51.9</number>
|
||
, 56.0
|
||
<number box="[1198,1281,1502,1524]" pageId="19" pageNumber="320" value="53.35" valueMax="56.7" valueMin="50.0">50‒56.7</number>
|
||
(
|
||
<number box="[1294,1340,1502,1524]" pageId="19" pageNumber="320" value="52.8">52.8</number>
|
||
)
|
||
<emphasis bold="true" box="[199,365,1596,1621]" pageId="19" pageNumber="320">Comparisons.</emphasis>
|
||
Compare with the other
|
||
<taxonomicName box="[662,782,1596,1620]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="genus">
|
||
<emphasis box="[662,782,1596,1620]" italics="true" pageId="19" pageNumber="320">Pempheris</emphasis>
|
||
</taxonomicName>
|
||
species,
|
||
<taxonomicName box="[891,1036,1597,1620]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[891,1036,1597,1620]" italics="true" pageId="19" pageNumber="320">P. tominagai</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<emphasis box="[1196,1334,1596,1620]" italics="true" pageId="19" pageNumber="320">P. s c h w e n ki i</emphasis>
|
||
in most counts and measurement, and as a result has been frequently misidentified as
|
||
<taxonomicName box="[1030,1168,1632,1656]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="schwenkii">
|
||
<emphasis box="[1030,1168,1632,1656]" italics="true" pageId="19" pageNumber="320">P. schwenkii</emphasis>
|
||
</taxonomicName>
|
||
. These two species can distinguish from the other species in genus
|
||
<taxonomicName box="[639,759,1668,1692]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="genus">
|
||
<emphasis box="[639,759,1668,1692]" italics="true" pageId="19" pageNumber="320">Pempheris</emphasis>
|
||
</taxonomicName>
|
||
in having lower scale counts (less than
|
||
<number box="[1208,1236,1668,1693]" pageId="19" pageNumber="320" value="53.0">53</number>
|
||
pored lateral-line scales; less than
|
||
<number box="[336,363,1704,1729]" pageId="19" pageNumber="320" value="41.0">41</number>
|
||
/
|
||
<number box="[371,384,1704,1729]" pageId="19" pageNumber="320" value="2.0">2</number>
|
||
scale rows above lateral line; circumpeduncular scales
|
||
<number box="[999,1072,1704,1728]" pageId="19" pageNumber="320" value="11.0" valueMax="12.0" valueMin="10.0">10–12</number>
|
||
), lower body depth 40.0–
|
||
<number box="[1360,1411,1704,1728]" pageId="19" pageNumber="320" value="47.7">47.7</number>
|
||
%, and anal fin with base zonal black. Between these two species,
|
||
<taxonomicName box="[855,996,1741,1764]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[855,996,1741,1764]" italics="true" pageId="19" pageNumber="320">P. tominagai</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<emphasis box="[1298,1432,1740,1764]" italics="true" pageId="19" pageNumber="320">P. s c h w e n ki i</emphasis>
|
||
by the following characters: scale rows above lateral line
|
||
<number box="[794,808,1776,1800]" pageId="19" pageNumber="320" value="4.0">4</number>
|
||
<number box="[815,825,1776,1800]" pageId="19" pageNumber="320" value="1.0">1</number>
|
||
/
|
||
<number box="[835,851,1776,1800]" pageId="19" pageNumber="320" value="2.0">2</number>
|
||
(vs.
|
||
<number box="[906,920,1776,1800]" pageId="19" pageNumber="320" value="3.0">3</number>
|
||
<number box="[927,937,1776,1800]" pageId="19" pageNumber="320" value="1.0">1</number>
|
||
/
|
||
<quantity box="[947,991,1776,1801]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="19" pageNumber="320" unit="in" value="2.0">2 in</quantity>
|
||
<emphasis box="[997,1131,1776,1800]" italics="true" pageId="19" pageNumber="320">P. s c h w e n ki i</emphasis>
|
||
); usually yellowish caudal fin in fresh specimens (vs. pinkish); posterior nostril usually open (
|
||
<figureCitation box="[926,995,1812,1837]" captionStart="FIGURE 8" captionStartId="17.[151,250,1637,1659]" captionTargetBox="[390,1196,1283,1621]" captionTargetId="figure@17.[390,1196,1283,1622]" captionTargetPageId="17" captionText="FIGURE 8. Illustrations of opened posterior nostril (a), and compressed posterior nostril (b)." httpUri="https://zenodo.org/record/229637/files/figure.png" pageId="19" pageNumber="320">
|
||
Fig.
|
||
<number box="[981,995,1812,1836]" pageId="19" pageNumber="320" value="8.0">8</number>
|
||
</figureCitation>
|
||
a; vs. usually compressed:
|
||
<figureCitation box="[1305,1373,1812,1837]" captionStart="FIGURE 8" captionStartId="17.[151,250,1637,1659]" captionTargetBox="[390,1196,1283,1621]" captionTargetId="figure@17.[390,1196,1283,1622]" captionTargetPageId="17" captionText="FIGURE 8. Illustrations of opened posterior nostril (a), and compressed posterior nostril (b)." httpUri="https://zenodo.org/record/229637/files/figure.png" pageId="19" pageNumber="320">
|
||
Fig.
|
||
<number box="[1359,1373,1812,1837]" pageId="19" pageNumber="320" value="8.0">8</number>
|
||
</figureCitation>
|
||
b). In addition to these morphological differences, the adequate genetic divergence compared with other species, strongly indicated that
|
||
<taxonomicName box="[309,451,1885,1908]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[309,451,1885,1908]" italics="true" pageId="19" pageNumber="320">P. tominagai</emphasis>
|
||
</taxonomicName>
|
||
is a validated species distribute in the western Indian Ocean and the Red Sea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="20" lastPageNumber="321" pageId="19" pageNumber="320" type="discussion">
|
||
<paragraph blockId="19.[151,1436,1596,2011]" pageId="19" pageNumber="320">
|
||
<emphasis bold="true" box="[199,315,1920,1945]" pageId="19" pageNumber="320">Remarks.</emphasis>
|
||
<bibRefCitation author="Heemstra" box="[330,688,1920,1944]" pageId="19" pageNumber="320" refString="Heemstra, P. & Heemstra, E. (2004) Coastal fishes of Southern Africa. NISC and SAIAB, Grahamstown (South Africa), 488 pp." type="book" year="2004">
|
||
Heemstra & Heemstra (
|
||
<number box="[620,679,1920,1944]" pageId="19" pageNumber="320" value="2004.0">2004</number>
|
||
)
|
||
</bibRefCitation>
|
||
reported
|
||
<taxonomicName box="[812,958,1920,1944]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="schwenkii">
|
||
<emphasis box="[812,958,1920,1944]" italics="true" pageId="19" pageNumber="320">P. schwenkii</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry box="[1044,1198,1920,1945]" name="South Africa" pageId="19" pageNumber="320">South Africa</collectingCountry>
|
||
, but it might be a misidentification of
|
||
<taxonomicName box="[385,529,1954,1977]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[385,529,1954,1977]" italics="true" pageId="19" pageNumber="320">P. tominagai</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Mouneimne" box="[547,767,1953,1978]" pageId="19" pageNumber="320" refString="Mouneimne, N. (1979) Poissons nouveaux pour les cotes Libanaises. Cybium, 6, 105 - 110." type="journal article" year="1979">
|
||
Mouneimne (
|
||
<number box="[701,760,1954,1978]" pageId="19" pageNumber="320" value="1979.0">1979</number>
|
||
)
|
||
</bibRefCitation>
|
||
reported
|
||
<taxonomicName box="[882,1007,1953,1977]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="species" species="molucca">
|
||
<emphasis box="[882,1007,1953,1977]" italics="true" pageId="19" pageNumber="320">P. molucca</emphasis>
|
||
</taxonomicName>
|
||
from the Mediterranean Sea, off the coast of
|
||
<collectingCountry box="[256,354,1986,2011]" name="Lebanon" pageId="19" pageNumber="320">Lebanon</collectingCountry>
|
||
(the first record of
|
||
<taxonomicName box="[597,717,1987,2011]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="320" phylum="Chordata" rank="genus">
|
||
<emphasis box="[597,717,1987,2011]" italics="true" pageId="19" pageNumber="320">Pempheris</emphasis>
|
||
</taxonomicName>
|
||
in this region), but, the illustration seems consistent with
|
||
<emphasis box="[1415,1434,1987,2010]" italics="true" pageId="19" pageNumber="320">P.</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="20.[151,1436,151,546]" pageId="20" pageNumber="321">
|
||
<taxonomicName box="[151,265,152,175]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[151,265,152,175]" italics="true" pageId="20" pageNumber="321">tominagai</emphasis>
|
||
</taxonomicName>
|
||
. However, no specimens of
|
||
<taxonomicName box="[602,748,152,176]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[602,748,152,176]" italics="true" pageId="20" pageNumber="321">P. tominagai</emphasis>
|
||
</taxonomicName>
|
||
collected from the Mediterranean Sea were verified in the present study. More
|
||
<taxonomicName box="[378,498,185,209]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="321" phylum="Chordata" rank="genus">
|
||
<emphasis box="[378,498,185,209]" italics="true" pageId="20" pageNumber="321">Pempheris</emphasis>
|
||
</taxonomicName>
|
||
materials from the Mediterranean Sea are required to substantiate the presence of
|
||
<taxonomicName class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis italics="true" pageId="20" pageNumber="321">P. tominagai</emphasis>
|
||
</taxonomicName>
|
||
in this region.
|
||
<bibRefCitation author="Randall" box="[445,617,218,243]" pageId="20" pageNumber="321" refString="Randall, J. E. (1995) Coastal fishes of Oman. University of Hawaii Press, Honolulu, 439 pp." type="book" year="1995">
|
||
Randall (
|
||
<number box="[551,609,218,243]" pageId="20" pageNumber="321" value="1995.0">1995</number>
|
||
)
|
||
</bibRefCitation>
|
||
reported
|
||
<taxonomicName box="[730,900,219,243]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="vanicolensis">
|
||
<emphasis box="[730,900,219,243]" italics="true" pageId="20" pageNumber="321">P. vanicolensis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry box="[973,1044,219,243]" name="Oman" pageId="20" pageNumber="321">Oman</collectingCountry>
|
||
, providing two color photographs (Fig.
|
||
<number box="[217,259,252,277]" pageId="20" pageNumber="321" value="637.0">637</number>
|
||
and
|
||
<quantity box="[325,401,252,277]" metricMagnitude="1" metricUnit="m" metricValue="1.6205199999999997" pageId="20" pageNumber="321" unit="in" value="638.0">638 in</quantity>
|
||
the literature). One of the photograph (Fig.
|
||
<number box="[939,982,252,277]" pageId="20" pageNumber="321" value="638.0">638</number>
|
||
) seems to depict an assemblage of
|
||
<taxonomicName class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis italics="true" pageId="20" pageNumber="321">P. tominagai</emphasis>
|
||
</taxonomicName>
|
||
, based on the blackish band on the anal fin base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="20" pageNumber="321" type="distribution">
|
||
<paragraph blockId="20.[151,1436,151,546]" pageId="20" pageNumber="321">
|
||
<emphasis bold="true" box="[199,352,318,343]" pageId="20" pageNumber="321">Distribution.</emphasis>
|
||
<taxonomicName box="[359,600,318,342]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[359,600,318,342]" italics="true" pageId="20" pageNumber="321">Pempheris tominagai</emphasis>
|
||
</taxonomicName>
|
||
is widely distributed in the Red Sea and the Indian Ocean. This species is also distributed in the north western coast of
|
||
<collectingCountry box="[658,763,351,376]" name="Australia" pageId="20" pageNumber="321">Australia</collectingCountry>
|
||
, but not along the coasts and islands of
|
||
<collectingCountry box="[1213,1322,351,376]" name="Indonesia" pageId="20" pageNumber="321">Indonesia</collectingCountry>
|
||
and north eastern
|
||
<collectingCountry box="[238,344,385,410]" name="Australia" pageId="20" pageNumber="321">Australia</collectingCountry>
|
||
. Specimens were collected from
|
||
<quantity box="[722,808,385,409]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="20" pageNumber="321" unit="m" value="5.0" valueMax="10.0" valueMin="0.0">0–10 m</quantity>
|
||
depth.
|
||
<taxonomicName box="[894,1132,385,409]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="schwenkii">
|
||
<emphasis box="[894,1132,385,409]" italics="true" pageId="20" pageNumber="321">Pempheris schwenkii</emphasis>
|
||
</taxonomicName>
|
||
which the sister species is widely distributed in the Pacific Ocean, but is not overlapped with that of
|
||
<taxonomicName box="[1009,1153,419,442]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[1009,1153,419,442]" italics="true" pageId="20" pageNumber="321">P. tominagai</emphasis>
|
||
</taxonomicName>
|
||
, and neither species had never collected from the Andaman Sea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="20" pageNumber="321" type="etymology">
|
||
<paragraph blockId="20.[151,1436,151,546]" pageId="20" pageNumber="321">
|
||
<emphasis bold="true" box="[199,326,485,510]" pageId="20" pageNumber="321">Etymology</emphasis>
|
||
. The specific name,
|
||
<taxonomicName box="[597,710,486,509]" class="Actinopterygii" family="Pempheridae" genus="Pempheris" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="tominagai">
|
||
<emphasis box="[597,710,486,509]" italics="true" pageId="20" pageNumber="321">tominagai</emphasis>
|
||
</taxonomicName>
|
||
, honors the late Yoshiaki Tominaga, pre-eminent Japanese ichthyologist who contributed to taxonomy and morphology of
|
||
<taxonomicName box="[863,1013,521,546]" class="Actinopterygii" family="Pempheridae" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="321" phylum="Chordata" rank="family">Pempheridae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |