229 lines
34 KiB
XML
229 lines
34 KiB
XML
<document id="7BBA4169A94287242B8DB4804F7DE5A1" ID="10.11646/zootaxa.4277.1.4SLASH11357" ID-CLB-Dataset="32837" ID-DOI="10.11646/zootaxa.4277.1.4" ID-GBIF-Dataset="034b7092-7aed-4e50-ace3-1318cb51c6ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="809053" ID-ZooBank="EED364DC-47FB-45A5-B0F5-CD71C41ECE2E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497512031586" checkinUser="plazi" docAuthor="Kimura, Seishi, Johnson, G. David, Peristiwady, Teguh & Matsuura, Keiichi" docDate="2017" docId="03EF4F051A693D39FF17FB1DE3F5FBD2" docLanguage="en" docName="zootaxa.4277.1.4SLASH11357.pdf" docOrigin="Zootaxa 4277 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Symphysanodontidae Katayama 1984" docType="treatment" docVersion="7" lastPageNumber="53" masterDocId="FFD6377D1A683D3BFF80FFEEE277FFAF" masterDocTitle="A new genus and species of the family Symphysanodontidae, Cymatognathus aureolateralis (Actinopterygii: Perciformes) from Indonesia" masterLastPageNumber="66" masterPageNumber="51" pageNumber="52" updateTime="1698440538984" updateUser="ExternalLinkService">
|
||
<mods:mods id="830BECC5F126C4A275B887E98F79B369" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="98AEBB83CAC81F6727ACFABF4882473B">
|
||
<mods:title id="6D2A3815FBBD25FE5932519ADC02019F">A new genus and species of the family Symphysanodontidae, Cymatognathus aureolateralis (Actinopterygii: Perciformes) from Indonesia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="A9E221595E0FA878AAE7155EAB60FFF3" type="personal">
|
||
<mods:role id="F38D5B838DD7C1416539876A5EF36758">
|
||
<mods:roleTerm id="FACD0C49FC04BB903C630B8FE512C991">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A6BA6441C09B9194527FE097A58CA5A4">Kimura, Seishi</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="31AC0ED5CCF825A560FD2B22D036C331" type="personal">
|
||
<mods:role id="24A46145ECC3AB2B0B0F2A221F714CB5">
|
||
<mods:roleTerm id="7A0D288F5AAAECF73AE0674CE1C41A2E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D4D065378C1A404E13A5D9329FF492F0">Johnson, G. David</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A4D85B473D77FCDD72DB37E73F33EA8A" type="personal">
|
||
<mods:role id="468C1C0DB6A0433F33443868114D4207">
|
||
<mods:roleTerm id="A6E6165DB5AFB4F76478FE1CDEFC63FE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BA1D49646C4E81E77214D52B9AAF672C">Peristiwady, Teguh</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="1F136D56820CD8A9C5E1338814868093" type="personal">
|
||
<mods:role id="9DF9C4851C835865C88211E1BA43B356">
|
||
<mods:roleTerm id="CCDB4E9D844E9BF1447BE8D42870D9D0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0904C7241C0E576A17A402231407B89C">Matsuura, Keiichi</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="20732F7504FDCA8ACF5458C0DE456983">text</mods:typeOfResource>
|
||
<mods:relatedItem id="7ACAA3CDFAE27B6743E2F5E5F69171A0" type="host">
|
||
<mods:titleInfo id="4D6DE229A74B4669884694A2088C4312">
|
||
<mods:title id="6AC8390CEA6FD1958AE55889D6B6C016">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="88E08B675E1D5A8C94ED8F5B8C46926D">
|
||
<mods:date id="6FB4B641A596A7CC6DC2C8F04317CF56">2017</mods:date>
|
||
<mods:detail id="5C0DEBD3F3369F1C48550BB974CC6D8A" type="volume">
|
||
<mods:number id="07C287328B4CA42FC8758F7E108E8804">4277</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="348F72CDF1896BE45E71335AF90A4579" type="issue">
|
||
<mods:number id="4AFC7B45DCDA61B1EF94D709C08181D3">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="9B0BCF6D03AA161D4257DAF79328040F" unit="page">
|
||
<mods:start id="108F7D7A50C56E63ED504DC662192053">51</mods:start>
|
||
<mods:end id="B96AF4AF176B768B983DB9258B94231E">66</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="5026720E6B53C8A436B113286F5FC7DA">journal article</mods:classification>
|
||
<mods:identifier id="198DC5D6D79274EF843D1491986E4A47" type="CLB-Dataset">32837</mods:identifier>
|
||
<mods:identifier id="C8440EDC4DC03818722017AD67395778" type="DOI">10.11646/zootaxa.4277.1.4</mods:identifier>
|
||
<mods:identifier id="C58CE0054CBF95D266DBAEAB7257405C" type="GBIF-Dataset">034b7092-7aed-4e50-ace3-1318cb51c6ce</mods:identifier>
|
||
<mods:identifier id="32A5C36CED0A1F7D282C53A7254D550C" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="327B3604B66BC79C084FEFC07FA93D5C" type="Zenodo-Dep">809053</mods:identifier>
|
||
<mods:identifier id="8FA005B4E16DA44D779579F338CC7861" type="ZooBank">EED364DC-47FB-45A5-B0F5-CD71C41ECE2E</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03EF4F051A693D39FF17FB1DE3F5FBD2" ID-DOI="http://doi.org/10.5281/zenodo.6029139" ID-GBIF-Taxon="131878644" ID-Zenodo-Dep="6029139" LSID="urn:lsid:plazi:treatment:03EF4F051A693D39FF17FB1DE3F5FBD2" httpUri="http://treatment.plazi.org/id/03EF4F051A693D39FF17FB1DE3F5FBD2" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
|
||
<subSubSection id="C35CAD981A693D3AFF17FB1DE370FA81" pageId="1" pageNumber="52" type="nomenclature">
|
||
<paragraph id="8BF9FE131A693D3AFF17FB1DE0B8FAA2" blockId="1.[151,719,1267,1326]" box="[151,719,1267,1293]" pageId="1" pageNumber="52">
|
||
<heading id="D0B1497F1A693D3AFF17FB1DE0B8FAA2" bold="true" box="[151,719,1267,1293]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
|
||
<emphasis id="B93222011A693D3AFF17FB1DE0B8FAA2" bold="true" box="[151,719,1267,1293]" pageId="1" pageNumber="52">
|
||
Family
|
||
<taxonomicName id="4C4685901A693D3AFF76FB1DE0B8FAA2" ID-CoL="626LW" ID-ENA="722446" authority="Katayama 1984" authorityName="Katayama" authorityYear="1984" box="[246,719,1267,1293]" class="Actinopterygii" family="Symphysanodontidae" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="52" phylum="Chordata" rank="family">
|
||
Symphysanodontidae
|
||
<bibRefCitation id="EFD783E21A693D3AFD8AFB1DE0B8FAA2" author="Katayama" box="[522,719,1267,1293]" pageId="1" pageNumber="52" refString="Katayama, M. (1984) Family Symphysanodontidae. In: Masuda, H., Amaoka, K., Araga, C., Uyeno, T. & Yoshino, T. (Eds.), The fishes of Japanese Archipelago (Text). Tokai University Press, Tokyo, pp. 138." type="book chapter" year="1984">Katayama 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF9FE131A693D3AFF17FAFBE370FA81" blockId="1.[151,719,1267,1326]" box="[151,263,1301,1326]" pageId="1" pageNumber="52">
|
||
(
|
||
<figureCitation id="137DE2961A693D3AFF1EFAFBE28BFA81" box="[158,252,1301,1326]" captionStart="FIGURE 1" captionStartId="3.[151,250,1938,1960]" captionTargetBox="[300,1286,195,1923]" captionTargetId="figure@3.[300,1286,195,1923]" captionTargetPageId="3" captionText="FIGURE 1. A. Cymatognathus aureolateralis sp. nov., holotype, MZB 19251, 181 mm SL, Bitung, North Sulawesi, Indonesia. B. Symphysanodon katayamai, FRLM 11738, 155 mm SL, Shima, Mie, Japan. 1. Fresh specimens (photographed by T. Peristiwady for C. aureolateralis and S. Kimura for S. katayamai). 2. X-ray photographs." httpUri="https://zenodo.org/record/809055/files/figure.png" pageId="1" pageNumber="52">Figure 1</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35CAD981A693D3AFF17FAB3E0F7FAD9" box="[151,640,1373,1398]" pageId="1" pageNumber="52" type="materials_examined">
|
||
<paragraph id="8BF9FE131A693D3AFF17FAB3E0F7FAD9" blockId="1.[151,1437,1373,2010]" box="[151,640,1373,1398]" pageId="1" pageNumber="52">
|
||
<emphasis id="B93222011A693D3AFF17FAB3E351FAD9" bold="true" box="[151,294,1373,1398]" pageId="1" pageNumber="52">Type genus:</emphasis>
|
||
<taxonomicName id="4C4685901A693D3AFEADFAB3E0F7FAD9" authority="Bleeker 1877" authorityName="Bleeker" authorityYear="1877" box="[301,640,1373,1398]" class="Actinopterygii" family="Symphysanodontidae" genus="Symphysanodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93222011A693D3AFEADFAB3E394FADA" box="[301,483,1373,1397]" italics="true" pageId="1" pageNumber="52">Symphysanodon</emphasis>
|
||
<bibRefCitation id="EFD783E21A693D3AFE6AFAB3E0F7FAD9" author="Bleeker" box="[490,640,1373,1398]" pageId="1" pageNumber="52" refString="Bleeker, P. (1877) Vischsoorten van Nieuw-Guinea, van Singapore, China, Japan en Mauritius. Processen-verbaal van de gewone vergaderingen der Koninklijke Akademie van Wetenschappen; Afdeeling Natuurkunde 1877 - 1878, 4, 2 - 3." type="journal article" year="1877">Bleeker 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35CAD981A693D3AFF47FA6FE707F83D" pageId="1" pageNumber="52" type="diagnosis">
|
||
<paragraph id="8BF9FE131A693D3AFF47FA6FE707F83D" blockId="1.[151,1437,1373,2010]" pageId="1" pageNumber="52">
|
||
<emphasis id="B93222011A693D3AFF47FA6FE336FA35" bold="true" box="[199,321,1409,1434]" pageId="1" pageNumber="52">Diagnosis.</emphasis>
|
||
Jaws subequal or upper jaw slightly projecting; both jaws with large and small conical teeth, no villiform teeth and no canines (
|
||
<figureCitation id="137DE2961A693D3AFD80FA4BE012FA11" box="[512,613,1445,1470]" captionStart="FIGURE 2" captionStartId="5.[151,250,1423,1445]" captionTargetBox="[151,1436,193,1401]" captionTargetId="figure@5.[151,1436,193,1401]" captionTargetPageId="5" captionText="FIGURE 2. Tooth patches on jaws. A. Cymatognathus aureolateralis sp. nov., paratype, FRLM 54552, 178 mm SL, Bitung, North Sulawesi, Indonesia. B. Symphysanodon katayamai, FRLM 11738, 155 mm SL, Shima, Mie, Japan. 1. Frontal view of both jaws. 2. Dorsal view of lower jaw. 3. Lateral view of both jaws. ETP = elevated tooth patch, LOT = outer tooth patch on lower jaw, UOT = outer tooth patch on upper jaw." httpUri="https://zenodo.org/record/809057/files/figure.png" pageId="1" pageNumber="52">Figure 2</figureCitation>
|
||
); anterior tips of both jaws with exposed outer patches of large conical teeth on sides, left and right tooth patches of upper jaw well separated but those of lower jaw close-set in adults (
|
||
<figureCitation id="137DE2961A693D3AFF1EFA03E376F9A9" box="[158,257,1517,1542]" captionStart="FIGURE 2" captionStartId="5.[151,250,1423,1445]" captionTargetBox="[151,1436,193,1401]" captionTargetId="figure@5.[151,1436,193,1401]" captionTargetPageId="5" captionText="FIGURE 2. Tooth patches on jaws. A. Cymatognathus aureolateralis sp. nov., paratype, FRLM 54552, 178 mm SL, Bitung, North Sulawesi, Indonesia. B. Symphysanodon katayamai, FRLM 11738, 155 mm SL, Shima, Mie, Japan. 1. Frontal view of both jaws. 2. Dorsal view of lower jaw. 3. Lateral view of both jaws. ETP = elevated tooth patch, LOT = outer tooth patch on lower jaw, UOT = outer tooth patch on upper jaw." httpUri="https://zenodo.org/record/809057/files/figure.png" pageId="1" pageNumber="52">Figure 2</figureCitation>
|
||
); outer tooth patches of the lower jaw fully or partly received into spaces between outer tooth patches of upper jaw; continuous tooth patch present along dorsal and medial surface of most of length of coronoid process of lower jaw (dentary) (
|
||
<figureCitation id="137DE2961A693D3AFE03F9DBE387F9E1" box="[387,496,1589,1614]" captionStart="FIGURE 2" captionStartId="5.[151,250,1423,1445]" captionTargetBox="[151,1436,193,1401]" captionTargetId="figure@5.[151,1436,193,1401]" captionTargetPageId="5" captionText="FIGURE 2. Tooth patches on jaws. A. Cymatognathus aureolateralis sp. nov., paratype, FRLM 54552, 178 mm SL, Bitung, North Sulawesi, Indonesia. B. Symphysanodon katayamai, FRLM 11738, 155 mm SL, Shima, Mie, Japan. 1. Frontal view of both jaws. 2. Dorsal view of lower jaw. 3. Lateral view of both jaws. ETP = elevated tooth patch, LOT = outer tooth patch on lower jaw, UOT = outer tooth patch on upper jaw." httpUri="https://zenodo.org/record/809057/files/figure.png" pageId="1" pageNumber="52">Figures 2</figureCitation>
|
||
,
|
||
<figureCitation id="137DE2961A693D3AFE7CF9D8E07CF9E1" box="[508,523,1590,1614]" captionStart="FIGURE 3" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[282,1323,685,1857]" captionTargetId="figure@6.[264,1323,625,1896]" captionTargetPageId="6" captionText="FIGURE 3. Medial view of lower jaw, showing “ elevated ” tooth patch (ETP) on dentary (D). A. Cymatognathus aureolateralis sp. nov., paratype, FRLM 54552, 178 mm SL, Bitung, North Sulawesi, Indonesia. B. Symphysanodon katayamai, FRLM 11738, 155 mm SL, Shima, Mie, Japan." httpUri="https://zenodo.org/record/809059/files/figure.png" pageId="1" pageNumber="52">3</figureCitation>
|
||
); palato-premaxillary ligament passing from anteriormost part of palatine, across head of maxilla, to ascending process of premaxilla (see
|
||
<bibRefCitation id="EFD783E21A693D3AFC90F9B7E1C3F9DD" author="Johnson" box="[784,948,1625,1650]" pageId="1" pageNumber="52" refString="Johnson, G. D. (1981) The limits and relationships of the Lutjanidae and associated families. Bulletin of the Scripps Institution of Oceanography, 24, 1 - 114." type="journal article" year="1981">Johnson, 1981</bibRefCitation>
|
||
;
|
||
<figureCitation id="137DE2961A693D3AFC40F9B7E657F9DD" box="[960,1056,1625,1650]" captionStart="FIGURE 4" captionStartId="7.[151,250,524,546]" captionTargetBox="[151,1436,193,502]" captionTargetId="figure@7.[151,1436,193,502]" captionTargetPageId="7" captionText="FIGURE 4. Palato-premaxillary ligament (arrows), lateral view of right side. A. Cymatognathus aureolateralis sp. nov., paratype, FRLM 54552, 178 mm SL, Bitung, North Sulawesi, Indonesia. B. Symphysanodon andersoni, USNM 435866, 200 mm SL, Salalah, Dhofar, Oman. C. S. katayamai, FRLM 11738, 155 mm SL, Shima, Mie, Japan." httpUri="https://zenodo.org/record/809061/files/figure.png" pageId="1" pageNumber="52">Figure 4</figureCitation>
|
||
); no supramaxilla; suborbital area narrow, first and second infraorbitals slightly covering upper margin of maxilla when mouth closed; fronto-parietal crest developed; preopercle with right-angled corner; posterior margin of preopercle smooth or very weakly serrated; opercle with two flat spines; a single, un-notched dorsal fin with nine or ten spines and nine to eleven soft rays [almost always IX, 10]; anal fin with three spines and seven or eight soft rays; pelvic axillary scale lanceolate; 10 + 15 vertebrae; supraneural and dorsal-fin pterygiophore insertion pattern 0/0/0+2+1/1/ (
|
||
<taxonomicName id="4C4685901A693D3AFB76F8E0E339F8E5" authority="Anderson, 1970" authorityName="Anderson" authorityYear="1970" class="Actinopterygii" family="Symphysanodontidae" genus="Symphysanodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="52" phylum="Chordata" rank="species" species="octoactinus">
|
||
<emphasis id="B93222011A693D3AFB76F8E0E7EBF88A" box="[1270,1436,1806,1829]" italics="true" pageId="1" pageNumber="52">S. octoactinus</emphasis>
|
||
<bibRefCitation id="EFD783E21A693D3AFF17F8DFE339F8E5" author="Anderson" box="[151,334,1841,1866]" pageId="1" pageNumber="52" refString="Anderson, W. D. Jr. (1970) Revision of the genus Symphysanodon (Pisces: Lutjanidae) with descriptions of four new species. Fishery Bulletin, 68, 325 - 346." type="journal article" year="1970">Anderson, 1970</bibRefCitation>
|
||
</taxonomicName>
|
||
exceptional in having 0/0/0+2/1+1/1/); first supraneural distinctly T-shaped; dorsal procurrent caudal-fin rays 12–14; ventral procurrent caudal-fin rays 10–12; base of penultimate ventral procurrent caudal-fin ray foreshortened; no procurrent spur (
|
||
<emphasis id="B93222011A693D3AFDC9F894E0F0F83E" box="[585,647,1914,1937]" italics="true" pageId="1" pageNumber="52">sensu</emphasis>
|
||
<bibRefCitation id="EFD783E21A693D3AFD0FF897E15DF83D" author="Johnson" box="[655,810,1913,1938]" pageId="1" pageNumber="52" refString="Johnson, G. D. (1975) The procurrent spur: an undescribed perciform caudal character and its phylogenetic implications. Occasional papers of the California Academy of Sciences, 121, 1 - 23." type="journal article" year="1975">Johnson 1975</bibRefCitation>
|
||
) on posteriormost ventral procurrent caudal-fin ray.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35CAD981A693D39FF47F873E3F5FBD2" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52" type="discussion">
|
||
<paragraph id="8BF9FE131A693D39FF47F873E351FF7B" blockId="1.[151,1437,1373,2010]" lastBlockId="2.[151,1437,151,1149]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
|
||
<emphasis id="B93222011A693D3AFF47F873E34CF819" bold="true" box="[199,315,1949,1974]" pageId="1" pageNumber="52">Remarks.</emphasis>
|
||
Prior to Katayama’s (1984) establishment of the
|
||
<taxonomicName id="4C4685901A693D3AFCE3F873E74FF819" authority="Fourmanoir (1981)" authorityName="Fourmanoir" authorityYear="1981" box="[867,1336,1949,1974]" class="Actinopterygii" family="Symphysanodontidae" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="52" phylum="Chordata" rank="family">
|
||
Symphysanodontidae,
|
||
<bibRefCitation id="EFD783E21A693D3AFBE2F873E74FF819" author="Fourmanoir" box="[1122,1336,1949,1974]" pageId="1" pageNumber="52" refString="Fourmanoir, P. (1981) Poissons (premiere liste). Resultats des Campagnes Musorstom. I. Philippines (18 - 28 mars 1976). Memoires de l'ORSTOM, 91 (3), 85 - 102." type="journal article" year="1981">Fourmanoir (1981)</bibRefCitation>
|
||
</taxonomicName>
|
||
used the family name
|
||
<taxonomicName id="4C4685901A693D3AFEBAF82FE05CF875" authorityName="Fourmanoir" authorityYear="1981" box="[314,555,1985,2010]" class="Actinopterygii" family="Symphysanodontidae" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="52" phylum="Chordata" rank="family">Symphysanodontidae</taxonomicName>
|
||
for
|
||
<taxonomicName id="4C4685901A693D3AFDE9F82FE168F876" authorityName="Bleeker" authorityYear="1877" box="[617,799,1985,2009]" class="Actinopterygii" family="Symphysanodontidae" genus="Symphysanodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93222011A693D3AFDE9F82FE168F876" box="[617,799,1985,2009]" italics="true" pageId="1" pageNumber="52">Symphysanodon</emphasis>
|
||
</taxonomicName>
|
||
. However, Fourmanoir’s
|
||
<taxonomicName id="4C4685901A693D3AFBD7F82FE73FF875" authorityName="Fourmanoir" authorityYear="1981" box="[1111,1352,1985,2010]" class="Actinopterygii" family="Symphysanodontidae" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="52" phylum="Chordata" rank="family">Symphysanodontidae</taxonomicName>
|
||
is not available because he did not describe any characters of his
|
||
<taxonomicName id="4C4685901A6A3D39FCABFF79E66BFF1F" authorityName="Fourmanoir" authorityYear="1981" box="[811,1052,151,176]" class="Actinopterygii" family="Symphysanodontidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Symphysanodontidae</taxonomicName>
|
||
(Art. 13.1.1,
|
||
<bibRefCitation id="EFD783E21A6A3D39FB33FF76E74AFF1F" author="ICZN" box="[1203,1341,152,176]" pageId="2" pageNumber="53" refString="ICZN (1999) International code of zoological nomenclature, 4 th edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN, 1999</bibRefCitation>
|
||
; van der Laan, 2014).
|
||
</paragraph>
|
||
<paragraph id="8BF9FE131A6A3D39FF47FF0EE62FFCF3" blockId="2.[151,1437,151,1149]" pageId="2" pageNumber="53">
|
||
As stated above, members of the
|
||
<taxonomicName id="4C4685901A6A3D39FDE7FF31E12FFF57" authorityName="Fourmanoir" authorityYear="1981" box="[615,856,223,248]" class="Actinopterygii" family="Symphysanodontidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Symphysanodontidae</taxonomicName>
|
||
are generally similar to the
|
||
<taxonomicName id="4C4685901A6A3D39FB3DFF31E731FF57" box="[1213,1350,223,248]" class="Actinopterygii" family="Serranidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="subFamily" subFamily="Anthiadinae">Anthiadinae</taxonomicName>
|
||
of the
|
||
<taxonomicName id="4C4685901A6A3D39FF17FEEAE365FEB2" box="[151,274,260,285]" class="Actinopterygii" family="Serranidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Serranidae</taxonomicName>
|
||
, and
|
||
<bibRefCitation id="EFD783E21A6A3D39FED1FEEAE383FEB3" author="Fowler" box="[337,500,260,285]" pageId="2" pageNumber="53" refString="Fowler, H. W. (1928) The fishes of Oceania. Memoirs of the Bernice P. Bishop Museum, 10, 1 - 540, pls. 1 - 49." type="journal article" year="1928">Fowler (1928)</bibRefCitation>
|
||
placed
|
||
<taxonomicName id="4C4685901A6A3D39FDD0FEEAE171FEB3" authorityName="Bleeker" authorityYear="1877" box="[592,774,260,284]" class="Actinopterygii" family="Symphysanodontidae" genus="Symphysanodon" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93222011A6A3D39FDD0FEEAE171FEB3" box="[592,774,260,284]" italics="true" pageId="2" pageNumber="53">Symphysanodon</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<taxonomicName id="4C4685901A6A3D39FCDBFEEAE19FFEB2" box="[859,1000,260,285]" class="Actinopterygii" family="Serranidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="subFamily" subFamily="Anthiadinae">Anthiadinae</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFD783E21A6A3D39FC74FEEBE6E1FEB2" author="Munro" box="[1012,1174,260,285]" pageId="2" pageNumber="53" refString="Munro, I. S. R. (1967) The Fishes of New Guinea. Department of Agriculture, Stock and Fisheries, Port Moresby, xxxvii + 651 pp." type="book" year="1967">Munro (1967)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD783E21A6A3D39FB25FEEBE71DFEB2" author="Katayama" box="[1189,1386,260,285]" pageId="2" pageNumber="53" refString="Katayama, M. (1968) Notes on the osteology and systematic position of Symphysanodon typus Bleeker. Bulletin of the Faculty of Education, Yamaguchi University, Natural Science, 17 (2), 105 - 111." type="journal article" year="1968">Katayama (1968)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFD783E21A6A3D39FF17FEC9E320FEEF" author="Anderson" box="[151,343,295,320]" pageId="2" pageNumber="53" refString="Anderson, W. D. Jr. (1970) Revision of the genus Symphysanodon (Pisces: Lutjanidae) with descriptions of four new species. Fishery Bulletin, 68, 325 - 346." type="journal article" year="1970">Anderson (1970)</bibRefCitation>
|
||
included
|
||
<taxonomicName id="4C4685901A6A3D39FE4AFEC6E0F7FEEF" authorityName="Bleeker" authorityYear="1877" box="[458,640,296,320]" class="Actinopterygii" family="Symphysanodontidae" genus="Symphysanodon" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93222011A6A3D39FE4AFEC6E0F7FEEF" box="[458,640,296,320]" italics="true" pageId="2" pageNumber="53">Symphysanodon</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<taxonomicName id="4C4685901A6A3D39FD53FEC9E138FEEF" box="[723,847,295,320]" class="Actinopterygii" family="Lutjanidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Lutjanidae</taxonomicName>
|
||
. However,
|
||
<bibRefCitation id="EFD783E21A6A3D39FC51FEC9E6F7FEEF" author="Johnson" box="[977,1152,295,320]" pageId="2" pageNumber="53" refString="Johnson, G. D. (1975) The procurrent spur: an undescribed perciform caudal character and its phylogenetic implications. Occasional papers of the California Academy of Sciences, 121, 1 - 23." type="journal article" year="1975">Johnson (1975)</bibRefCitation>
|
||
found no evidence from osteology and cheek myology for a close relationship between
|
||
<taxonomicName id="4C4685901A6A3D39FCE4FEA2E66DFECB" authorityName="Bleeker" authorityYear="1877" box="[868,1050,332,356]" class="Actinopterygii" family="Symphysanodontidae" genus="Symphysanodon" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93222011A6A3D39FCE4FEA2E66DFECB" box="[868,1050,332,356]" italics="true" pageId="2" pageNumber="53">Symphysanodon</emphasis>
|
||
</taxonomicName>
|
||
and the
|
||
<taxonomicName id="4C4685901A6A3D39FBFEFEA2E68DFECA" box="[1150,1274,332,357]" class="Actinopterygii" family="Lutjanidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Lutjanidae</taxonomicName>
|
||
. Moreover, he noted that the base of the penultimate ventral procurrent caudal-fin ray is foreshortened in
|
||
<taxonomicName id="4C4685901A6A3D39FB5FFE9EE7E2FE27" authorityName="Bleeker" authorityYear="1877" box="[1247,1429,368,392]" class="Actinopterygii" family="Symphysanodontidae" genus="Symphysanodon" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93222011A6A3D39FB5FFE9EE7E2FE27" box="[1247,1429,368,392]" italics="true" pageId="2" pageNumber="53">Symphysanodon</emphasis>
|
||
</taxonomicName>
|
||
, although there is no spur on the last ventral procurrent caudal-fin ray. He suggested that
|
||
<taxonomicName id="4C4685901A6A3D39FB0AFE7AE737FE03" authorityName="Bleeker" authorityYear="1877" box="[1162,1344,404,428]" class="Actinopterygii" family="Symphysanodontidae" genus="Symphysanodon" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93222011A6A3D39FB0AFE7AE737FE03" box="[1162,1344,404,428]" italics="true" pageId="2" pageNumber="53">Symphysanodon</emphasis>
|
||
</taxonomicName>
|
||
is not a lutjanid and may occupy a very primitive position within the Percoidei. Subsequently
|
||
<bibRefCitation id="EFD783E21A6A3D39FBECFE59E76CFE7F" author="Johnson" box="[1132,1307,439,464]" pageId="2" pageNumber="53" refString="Johnson, G. D. (1981) The limits and relationships of the Lutjanidae and associated families. Bulletin of the Scripps Institution of Oceanography, 24, 1 - 114." type="journal article" year="1981">Johnson (1981)</bibRefCitation>
|
||
provided a detailed description of
|
||
<taxonomicName id="4C4685901A6A3D39FE33FE32E01EFE5B" authorityName="Bleeker" authorityYear="1877" box="[435,617,476,500]" class="Actinopterygii" family="Symphysanodontidae" genus="Symphysanodon" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93222011A6A3D39FE33FE32E01EFE5B" box="[435,617,476,500]" italics="true" pageId="2" pageNumber="53">Symphysanodon</emphasis>
|
||
</taxonomicName>
|
||
and treated the genus as
|
||
<emphasis id="B93222011A6A3D39FC37FE33E62CFE5B" box="[951,1115,476,500]" italics="true" pageId="2" pageNumber="53">incertae sedis</emphasis>
|
||
. Our examinations of the
|
||
<taxonomicName id="4C4685901A6A3D39FF17FE11E3FCFDB7" authorityName="Fourmanoir" authorityYear="1981" box="[151,395,511,536]" class="Actinopterygii" family="Symphysanodontidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Symphysanodontidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4685901A6A3D39FE18FE11E067FDB7" box="[408,528,511,536]" class="Actinopterygii" family="Lutjanidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Lutjanidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4685901A6A3D39FDCAFE11E0B5FDB7" box="[586,706,511,536]" class="Actinopterygii" family="Serranidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Serranidae</taxonomicName>
|
||
have further revealed that the
|
||
<taxonomicName id="4C4685901A6A3D39FBA5FE11E761FDB7" authorityName="Fourmanoir" authorityYear="1981" box="[1061,1302,511,536]" class="Actinopterygii" family="Symphysanodontidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Symphysanodontidae</taxonomicName>
|
||
differ from the latter two families in having exposed outer tooth patches on the upper and lower jaws in adults (vs. no outer tooth patches on either jaw in the latter two families), continuous tooth patch present along the dorsal and medial surface of most of the length of the coronoid process of the lower jaw, a lanceolate pelvic axillary scale (vs. triangular or rectangular scale or an undeveloped scale), a unique (
|
||
<taxonomicName id="4C4685901A6A3D39FC20FD7EE63FFD08" box="[928,1096,656,679]" class="Actinopterygii" family="Symphysanodontidae" genus="Symphysanodon" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="octoactinius">
|
||
<emphasis id="B93222011A6A3D39FC20FD7EE63FFD08" box="[928,1096,656,679]" italics="true" pageId="2" pageNumber="53">S. octoactinius</emphasis>
|
||
</taxonomicName>
|
||
exceptional) supraneural and dorsal-fin pterygiophore insertion pattern 0/0/0+2+1/1/, and 25 total vertebrae [vs. 24 or 26–28 (
|
||
<bibRefCitation id="EFD783E21A6A3D39FA8CFD5AE339FD5F" author="Anderson" pageId="2" pageNumber="53" refString="Anderson, W. D. Jr. & Heemstra, P. C. (1989) Ellerkeldia, a junior synonym of Hypoplectrodes, with redescriptions of the type species of the genera (Pisces: Serranidae: Anthiinae). Proceedings of the Biological Society of Washington, 102 (4), 1001 - 1017." type="journal article" year="1989">Anderson & Heemstra, 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD783E21A6A3D39FEDBFD39E021FD5F" author="Anderson" box="[347,598,727,752]" pageId="2" pageNumber="53" refString="Anderson, W. D. Jr., Parin, N. V. & Randall, J. E. (1990) A new genus and species of anthiine fish (Pisces: Serranidae) from the eastern South Pacific with comments on anthiine relationships. Proceedings of the Biological Society of Washington, 103 (4), 922 - 930." type="journal article" year="1990">
|
||
Anderson
|
||
<emphasis id="B93222011A6A3D39FE51FD37E07CFD5F" box="[465,523,728,752]" italics="true" pageId="2" pageNumber="53">et al.</emphasis>
|
||
, 1990
|
||
</bibRefCitation>
|
||
; present observation)]. The
|
||
<taxonomicName id="4C4685901A6A3D39FC17FD39E6FFFD5F" authorityName="Fourmanoir" authorityYear="1981" box="[919,1160,727,752]" class="Actinopterygii" family="Symphysanodontidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Symphysanodontidae</taxonomicName>
|
||
are also distinguishable from the
|
||
<taxonomicName id="4C4685901A6A3D39FF7CFD12E303FCBA" box="[252,372,764,789]" class="Actinopterygii" family="Serranidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Serranidae</taxonomicName>
|
||
by having a well-developed fronto-parietal crest (vs. undeveloped) and from the
|
||
<taxonomicName id="4C4685901A6A3D39FB7DFD12E702FCBA" box="[1277,1397,764,789]" class="Actinopterygii" family="Lutjanidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Lutjanidae</taxonomicName>
|
||
by, among other things, having a right-angled preopercular corner (vs. round), as already stated by
|
||
<bibRefCitation id="EFD783E21A6A3D39FB51FCCEE7E0FC97" author="Katayama" box="[1233,1431,800,824]" pageId="2" pageNumber="53" refString="Katayama, M. (1984) Family Symphysanodontidae. In: Masuda, H., Amaoka, K., Araga, C., Uyeno, T. & Yoshino, T. (Eds.), The fishes of Japanese Archipelago (Text). Tokai University Press, Tokyo, pp. 138." type="book chapter" year="1984">Katayama (1984)</bibRefCitation>
|
||
, and no attachment of the adductor mandibulae to the subocular shelf (
|
||
<bibRefCitation id="EFD783E21A6A3D39FC25FCAAE63BFCF3" author="Johnson" box="[933,1100,836,861]" pageId="2" pageNumber="53" refString="Johnson, G. D. (1981) The limits and relationships of the Lutjanidae and associated families. Bulletin of the Scripps Institution of Oceanography, 24, 1 - 114." type="journal article" year="1981">Johnson, 1981</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF9FE131A6A3D39FF47FC89E3F5FBD2" blockId="2.[151,1437,151,1149]" pageId="2" pageNumber="53">
|
||
The supraneural and dorsal-fin pterygiophore insertion pattern (0/0/0+2+1/1/) that characterizes all but one species of
|
||
<taxonomicName id="4C4685901A6A3D39FE94FC62E072FC0A" authorityName="Fourmanoir" authorityYear="1981" box="[276,517,908,933]" class="Actinopterygii" family="Symphysanodontidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Symphysanodontidae</taxonomicName>
|
||
is also found in the African percoid family
|
||
<taxonomicName id="4C4685901A6A3D39FB8EFC62E6D2FC0A" authorityName="Heemstra & Hecht" authorityYear="1986" box="[1038,1189,908,933]" class="Actinopterygii" family="Dinopercidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="53" phylum="Chordata" rank="family">Dinopercidae</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFD783E21A6A3D39FB67FC62E7E7FC0B" author="Johnson" box="[1255,1424,908,933]" pageId="2" pageNumber="53" refString="Johnson, G. D. (1984) Percoidei: development and relationships. In: Moser, H. G., Richards, W. J., Cohen, D. M., Fahay, M. P., Kendall, A. W. Jr. & Richardson, S. L. (Eds.), Ontogeny and systematics of fishes. Special Publication 1. American Society of Ichthyologists and Herpetologists, Lawrence, pp. 464 - 498." type="journal article" year="1984">Johnson, 1984</bibRefCitation>
|
||
). However, a relationship between those two families is probably rather distant, because the latter differs from the former as follows: lower jaw projecting; no outer patches of teeth at anterior tips of jaws; preopercle with round corner; posterior margin of preopercle serrated; opercle with one flat spine; a single dorsal fin notched with 11 spines and 17–19 soft rays; anal fin with three spines and 11–13 soft rays; no axillary scale; 10 + 16 vertebrae; first supraneural reverse L-shaped; procurrent spur on posteriormost ventral procurrent caudal-fin ray (
|
||
<bibRefCitation id="EFD783E21A6A3D39FB6FFBD1E300FBD2" author="Johnson" pageId="2" pageNumber="53" refString="Johnson, G. D. (1984) Percoidei: development and relationships. In: Moser, H. G., Richards, W. J., Cohen, D. M., Fahay, M. P., Kendall, A. W. Jr. & Richardson, S. L. (Eds.), Ontogeny and systematics of fishes. Special Publication 1. American Society of Ichthyologists and Herpetologists, Lawrence, pp. 464 - 498." type="journal article" year="1984">Johnson, 1984; present observation</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |