130 lines
16 KiB
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130 lines
16 KiB
XML
<document id="84B9129A3A5822A32242FF22CFFA2F15" ID-CLB-Dataset="7631" ID-DOI="10.1206/0003-0090-423.1.1" ID-GBIF-Dataset="2e886aea-b59a-45a6-aeaa-2427d584a894" ID-ISSN="0003-0090" ID-Zenodo-Dep="4613008" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615995240537" checkinUser="felipe" docAuthor="Grimaldi, David A." docDate="2018" docId="03B187A8FFCAFFB3FD103DDB7EA5FB81" docLanguage="en" docName="BulAmeMusNatHis.2018.423.1-97.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 2018 (423)" docSource="http://www.bioone.org/doi/10.1206/0003-0090-423.1.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Hennigophora pria Grimaldi 2018" docType="treatment" docVersion="6" lastPageNumber="97" masterDocId="FF88FFD0FF86FFFEFFAA39537F49FFF7" masterDocTitle="Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi" masterLastPageNumber="97" masterPageNumber="1" pageNumber="1" updateTime="1698933720305" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="FBB194BB3D98A46D862A0CFC8565971A">
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<mods:title id="4027DC772DE357F502A6291672E5CC7B">Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi</mods:title>
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<mods:name id="44C30CAEAB6AED9252DF86556CA372B4" type="personal">
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<mods:roleTerm id="C801F11D2D2DFBEF885ED4C356C1DC53">Author</mods:roleTerm>
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<mods:namePart id="0B1C3E7E975CD0465B0187ED1A7AD647">Grimaldi, David A.</mods:namePart>
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<mods:affiliation id="AA2009B605308AE2B2F22C37EBC24DA2">Division of Invertebrate Zoology American Museum of Natural History, New York</mods:affiliation>
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<mods:typeOfResource id="9311E09352523FF852F540AD59064969">text</mods:typeOfResource>
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<mods:title id="061F669108C03679550A9634D54D69C4">Bulletin of the American Museum of Natural History</mods:title>
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<mods:part id="A88DCE45819FA4C4336D8767A550FC2B">
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<mods:date id="741FC60753D14CC0B07E5ABD7500E347">2018</mods:date>
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<mods:number id="E6DE838317A41121FFADD29F8F9F0ECF">2018-10-24</mods:number>
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<mods:detail id="DC161B9F2347EC3F42AF9DDDF8B99408" type="volume">
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<mods:number id="D09EEEDB55E6629A488E90E08F797A2D">2018</mods:number>
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<mods:number id="41190B3A3BAC545FE9BCCDD7CAF5813F">423</mods:number>
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<mods:start id="C64D6297FD1885E8212E01F303BB9032">1</mods:start>
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<mods:end id="8162916F6EC1F2FC6B52A3FA596AED51">97</mods:end>
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<mods:url id="66EE06960463098EF84359CE353B1833">http://www.bioone.org/doi/10.1206/0003-0090-423.1.1</mods:url>
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<mods:classification id="AB9A755BAD52129F9B8852B4C848062F">journal article</mods:classification>
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<mods:identifier id="AB05C723D458F834BC6431678E6D96E1" type="CLB-Dataset">7631</mods:identifier>
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<mods:identifier id="6BAF42C196086435DF03D626DED730C7" type="DOI">10.1206/0003-0090-423.1.1</mods:identifier>
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<mods:identifier id="D0E5DBA97864629FE243B79B8BD95768" type="GBIF-Dataset">2e886aea-b59a-45a6-aeaa-2427d584a894</mods:identifier>
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<mods:identifier id="52E9D290AE49B957FB09DC21F010A781" type="ISSN">0003-0090</mods:identifier>
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<treatment id="03B187A8FFCAFFB3FD103DDB7EA5FB81" ID-DOI="http://doi.org/10.5281/zenodo.4631004" ID-GBIF-Taxon="180670774" ID-Zenodo-Dep="4631004" LSID="urn:lsid:plazi:treatment:03B187A8FFCAFFB3FD103DDB7EA5FB81" httpUri="http://treatment.plazi.org/id/03B187A8FFCAFFB3FD103DDB7EA5FB81" lastPageId="77" pageId="76">
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<subSubSection id="C3026535FFCAFFB2FD103DDB7BE6FB57" box="[698,1199,1160,1184]" pageId="76" type="nomenclature">
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<paragraph id="8BA736BEFFCAFFB2FD103DDB7BE6FB57" blockId="76.[698,1199,1160,1218]" box="[698,1199,1160,1184]" pageId="76">
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<heading id="D0EF81D2FFCAFFB2FD103DDB7BE6FB57" box="[698,1199,1160,1184]" centered="true" fontSize="9" level="2" pageId="76" reason="2">
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<taxonomicName id="4C184D3DFFCAFFB2FD103DDB7BE2FB57" authority="(Grimaldi and Cumming)" authorityName="Grimaldi" authorityYear="2018" baseAuthorityName="Grimaldi and Cumming" baseAuthorityYear="2018" box="[698,1195,1160,1184]" class="Insecta" family="Sciadoceridae" genus="Hennigophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="76" phylum="Arthropoda" rank="species" species="pria" status="comb. nov.">
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<emphasis id="B96CEAACFFCAFFB2FD103DDB7CCCFB57" box="[698,901,1160,1184]" italics="true" pageId="76">Hennigophora pria</emphasis>
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(Grimaldi and Cumming)
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</taxonomicName>
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,
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3026535FFCAFFB2FCFC3DF97B60FB04" pageId="76" type="description">
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<paragraph id="8BA736BEFFCAFFB2FCFC3DF97B5DFB35" blockId="76.[698,1199,1160,1218]" box="[854,1044,1194,1218]" pageId="76">new combination</paragraph>
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<subSection id="E2972F55FFCAFFB2FCEA3D887B60FB04" box="[832,1065,1243,1267]" pageId="76" type="multiple">
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<paragraph id="8BA736BEFFCAFFB2FCEA3D887B60FB04" blockId="76.[832,1065,1243,1267]" box="[832,1065,1243,1267]" pageId="76">
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<figureCitation id="13232A3BFFCAFFB2FCEA3D887CFDFB04" box="[832,948,1243,1267]" captionStart="FIG" captionStartId="25.[108,150,1548,1569]" captionTargetBox="[108,1212,225,1520]" captionTargetId="figure-7@25.[108,1212,225,1520]" captionTargetPageId="25" captionText="FIG. 18. Hennigophora pria (Grimaldi and Cumming) (Phoridae), new female specimen in Late Cretaceous amber from New Jersey, AMNH NJ-Ch1. A. Right lateral habitus. B. Head and thorax, lateral. C. Wing, abdomen, legs, left lateral. Legs and thorax appear to be natural coloration; light abdomen is due to milky coating." figureDoi="http://doi.org/10.5281/zenodo.4613057" httpUri="https://zenodo.org/record/4613057/files/figure.png" pageId="76">Figures 18</figureCitation>
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,
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<figureCitation id="13232A3BFFCAFFB2FC6B3D887CA6FB04" box="[961,1007,1243,1267]" captionStart="FIG" captionStartId="78.[108,150,1209,1230]" captionTargetBox="[160,1112,234,1176]" captionTargetId="figure-182@78.[108,1212,224,1180]" captionTargetPageId="78" captionText="FIG. 46. Sciadocerinae in Eocene Baltic amber (A, B) and Late Cretaceous amber from New Jersey (C). A. Eosciadocera pauciseta, AMNH Ba-Je565, frontal view of head. B. Hennigophora robusta, AMNH Ba-JH58, oblique frontal view of head. C. Hennigophora pria, AMNH NJ-Ch1, lateral view of head and thorax." figureDoi="http://doi.org/10.5281/zenodo.4613127" httpUri="https://zenodo.org/record/4613127/files/figure.png" pageId="76">46C</figureCitation>
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,
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<figureCitation id="13232A3BFFCAFFB2FC563D887B60FB04" box="[1020,1065,1243,1267]" captionStart="FIG" captionStartId="79.[108,150,1641,1662]" captionTargetBox="[243,1096,233,1603]" captionTargetId="figure-7@79.[108,1212,224,1612]" captionTargetPageId="79" captionText="FIG. 47. Wings of Sciadocerinae preserved in Eocene Baltic amber (B, D), and Cretaceous ambers (A, C), to different scales.A. Hennigophora pria in New Jersey amber, AMNH NJ-Ch1. B. Hennigophora robusta in Baltic amber, AMNH Ba-JH51. C. Prophora dimorion in Burmese amber, JZC Bu242. D. Eosciadocera pauciseta in Baltic amber, AMNH Ba-JVe565." figureDoi="http://doi.org/10.5281/zenodo.4613129" httpUri="https://zenodo.org/record/4613129/files/figure.png" pageId="76">47A</figureCitation>
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</paragraph>
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</subSection>
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</subSubSection>
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<subSubSection id="C3026535FFCAFFB2FD063C427BFFFADE" box="[684,1206,1297,1321]" pageId="76" type="reference_group">
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<paragraph id="8BA736BEFFCAFFB2FD063C427BFFFADE" blockId="76.[684,1206,1297,1321]" box="[684,1206,1297,1321]" pageId="76">
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<heading id="D0EF81D2FFCAFFB2FD063C427BFFFADE" box="[684,1206,1297,1321]" centered="true" fontSize="9" level="2" pageId="76" reason="6">
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<treatmentCitationGroup id="AB081190FFCAFFB2FD063C427BFFFADE" box="[684,1206,1297,1321]" pageId="76">
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<treatmentCitation id="0AB910AFFFCAFFB2FD063C427BF8FADE" author="Grimaldi, D. A. & J. M. Cumming" box="[684,1201,1297,1321]" page="97" pageId="76" year="1999">
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<taxonomicName id="4C184D3DFFCAFFB2FD063C427BF8FADE" ID-CoL="8B3F8" authority="Grimaldi and Cumming, 1999: 97" authorityName="Grimaldi and Cumming" authorityPageNumber="97" authorityYear="1999" box="[684,1201,1297,1321]" class="Insecta" family="Phoridae" genus="Archiphora" kingdom="Animalia" order="Diptera" pageId="76" phylum="Arthropoda" rank="species" species="pria">
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<emphasis id="B96CEAACFFCAFFB2FD063C427C02FADE" box="[684,843,1297,1321]" italics="true" pageId="76">Archiphora pria</emphasis>
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<bibRefCitation id="EF894B4FFFCAFFB2FCF83C427BF8FADE" author="Grimaldi, D. A. & J. M. Cumming" box="[850,1201,1297,1321]" pageId="76" pagination="1 - 124" refId="ref39017" refString="Grimaldi, D. A., and J. M. Cumming. 1999. Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History 239: 1 - 124." type="journal article" year="1999">Grimaldi and Cumming, 1999: 97</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3026535FFCAFFB3FD663C167EA5FB81" lastPageId="77" pageId="76" type="discussion">
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<paragraph id="8BA736BEFFCAFFB3FD663C167EEEFCFC" blockId="76.[684,1214,1349,1737]" lastBlockId="77.[108,638,226,1142]" lastPageId="77" pageId="76">
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SUMMARY: The original (
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<typeStatus id="54A3881CFFCAFFB2FC653C167B66FAAA" box="[975,1071,1349,1373]" pageId="76" type="holotype">holotype</typeStatus>
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) specimen of
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<taxonomicName id="4C184D3DFFCAFFB2FD063C347C49FA89" box="[684,768,1383,1406]" class="Insecta" family="Phoridae" genus="Archiphora" kingdom="Animalia" order="Diptera" pageId="76" phylum="Arthropoda" rank="species" species="pria">
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<emphasis id="B96CEAACFFCAFFB2FD063C347C49FA89" box="[684,768,1383,1406]" italics="true" pageId="76">H. pria</emphasis>
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</taxonomicName>
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is covered with a milky coating that obscures many surface details, critically the acrostichals, as well as the presence/absence of a median furrow on the frons, and terminalia, though portions of larger macrosetae protrude through the coating. A new specimen in
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<collectingRegion id="49DCF85CFFCAFFB2FBF43F5F7D87F9B2" country="United States of America" name="New Jersey" pageId="76">New Jersey</collectingRegion>
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amber, AMNH NJ-Ch1, has some critical features that are better preserved, but this specimen lost the left side of the head and thorax before or during fossilization, and the abdomen has a thin milky coating. A few acrostichals can be seen in left profile, which appear to be in one (or possibly several) rows and are significantly smaller than those in
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<taxonomicName id="4C184D3DFFCBFFB3FF6138777E73FECB" baseAuthorityName="Meunier" baseAuthorityYear="1905" box="[203,314,292,316]" class="Insecta" family="Sciadoceridae" genus="Hennigophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="77" phylum="Arthropoda" rank="species" species="robusta">
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<emphasis id="B96CEAACFFCBFFB3FF6138777E73FECB" box="[203,314,292,316]" italics="true" pageId="77">H. robusta</emphasis>
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</taxonomicName>
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(whose acrostichals are about as large as the flanking dorsocentrals; fig. 19). Male genitalic structure cannot be compared to
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<taxonomicName id="4C184D3DFFCBFFB3FDCB38347FF0FE68" baseAuthorityName="Meunier" baseAuthorityYear="1905" class="Insecta" family="Sciadoceridae" genus="Hennigophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="77" phylum="Arthropoda" rank="species" species="robusta">
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<emphasis id="B96CEAACFFCBFFB3FDCB38347FF0FE68" italics="true" pageId="77">H. robusta</emphasis>
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</taxonomicName>
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since this specimen is female. Most importantly, the stridulatum is present in AMNH NJ-Ch1, with a scraper very visible on the mesal surface of the right profemur, and a file on the lateral surface of the left procoxa. The location, arc, and length of the stridulatum are just as in living sciadocerines, but the number and structure of ridges and teeth are not visible. A stridulatum was also observed in
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<taxonomicName id="4C184D3DFFCBFFB3FEE73BC37EF5FD50" baseAuthorityName="Meunier" baseAuthorityYear="1905" box="[333,444,655,679]" class="Insecta" family="Sciadoceridae" genus="Hennigophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="77" phylum="Arthropoda" rank="species" species="robusta">
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<emphasis id="B96CEAACFFCBFFB3FEE73BC37EF5FD50" box="[333,444,655,679]" italics="true" pageId="77">H. robusta</emphasis>
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</taxonomicName>
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based on a file on the right procoxa of a male specimen, AMNH Ba-JH58 (fig. 19) (this specimen also lacks a median furrow on the frons).
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</paragraph>
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<paragraph id="8BA736BEFFCBFFB3FF263A477EA5FB81" blockId="77.[108,638,226,1142]" pageId="77">
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<taxonomicName id="4C184D3DFFCBFFB3FF263A477E11FCDC" authorityName="Grimaldi" authorityYear="2018" baseAuthorityName="Grimaldi and Cumming" baseAuthorityYear="2018" box="[140,344,788,812]" class="Insecta" family="Sciadoceridae" genus="Hennigophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="77" phylum="Arthropoda" rank="species" species="pria">
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<emphasis id="B96CEAACFFCBFFB3FF263A477E11FCDC" box="[140,344,788,812]" italics="true" pageId="77">Hennigophora pria</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName id="4C184D3DFFCBFFB3FE473A477D29FCDB" baseAuthorityName="Meunier" baseAuthorityYear="1905" box="[493,608,788,812]" class="Insecta" family="Sciadoceridae" genus="Hennigophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="77" phylum="Arthropoda" rank="species" species="robusta">
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<emphasis id="B96CEAACFFCBFFB3FE473A477D29FCDB" box="[493,608,788,812]" italics="true" pageId="77">H. robusta</emphasis>
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</taxonomicName>
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in venation (proportions and lengths of main veins, a tubular stem of M; fig. 47A, B), setation of legs; and a broad frons (judging from the preserved right half in AMNH NJ-Ch1). It differs from
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<taxonomicName id="4C184D3DFFCBFFB3FDC93ACA7FF2FC26" class="Insecta" family="Sciadoceridae" genus="Hennigophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="77" phylum="Arthropoda" rank="species" species="robutsa">
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<emphasis id="B96CEAACFFCBFFB3FDC93ACA7FF2FC26" italics="true" pageId="77">H. robutsa</emphasis>
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</taxonomicName>
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by having a pair of reclinate interfrontals, near the frontal margin of the frons (vs. absent); also, setae are shorter, especially the acrostichals, supraalars, postalars, and scutellars. In agreement with
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<bibRefCitation id="EF894B4FFFCBFFB3FF493D6D7EC1FBA2" author="Brown, B. V." box="[227,392,1085,1109]" pageId="77" pagination="1 - 14" refId="ref38183" refString="Brown, B. V. 2007 a. A further new genus of primitive phorid fly (Diptera: Phoridae) from Baltic amber and its phylogenetic implications. Contributions in Science, Natural History Museum of Los Angeles County 513: 1 - 14." type="journal article" year="2007">Brown (2007a)</bibRefCitation>
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,
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<taxonomicName id="4C184D3DFFCBFFB3FE3E3D6E7D45FBA2" box="[404,524,1085,1109]" class="Insecta" family="Phoridae" genus="Archiphora" kingdom="Animalia" order="Diptera" pageId="77" phylum="Arthropoda" rank="genus">
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<emphasis id="B96CEAACFFCBFFB3FE3E3D6E7D45FBA2" box="[404,524,1085,1109]" italics="true" pageId="77">Archiphora</emphasis>
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</taxonomicName>
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should be restricted to the sole living species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |