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<document id="199AE7E1651C3F6833A53EFAE6CCE059" ID-CLB-Dataset="7631" ID-DOI="10.1206/0003-0090-423.1.1" ID-GBIF-Dataset="2e886aea-b59a-45a6-aeaa-2427d584a894" ID-ISSN="0003-0090" ID-Zenodo-Dep="4613008" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615995240537" checkinUser="felipe" docAuthor="Grimaldi, David A." docDate="2018" docId="03B187A8FFC5FFBAFF633CEA7C79FC99" docLanguage="en" docName="BulAmeMusNatHis.2018.423.1-97.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 2018 (423)" docSource="http://www.bioone.org/doi/10.1206/0003-0090-423.1.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Proironia gibbera Grimaldi 2018, new species" docType="treatment" docVersion="8" lastPageNumber="97" masterDocId="FF88FFD0FF86FFFEFFAA39537F49FFF7" masterDocTitle="Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi" masterLastPageNumber="97" masterPageNumber="1" pageNumber="1" updateTime="1698933720305" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8B6C3CCC45486A19A5C1EAAB747547D1">Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi</mods:title>
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<mods:namePart id="DBDCAFCE935F90ADFB49445C52F4705A">Grimaldi, David A.</mods:namePart>
<mods:affiliation id="F569778B8E0F4FFF3E49BD32EBC246FD">Division of Invertebrate Zoology American Museum of Natural History, New York</mods:affiliation>
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<mods:title id="C802CF0FF114228D353903B0962F48CA">Bulletin of the American Museum of Natural History</mods:title>
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<mods:date id="CD52BC267EB5C45162012654393DF218">2018</mods:date>
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<mods:number id="2532B300136A9D0673456A32C7C8DE86">2018-10-24</mods:number>
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<mods:number id="84381615298BFFAE006B247960E15543">2018</mods:number>
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<mods:number id="4909314622347804541555B25ACB57F2">423</mods:number>
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<mods:classification id="3D21B0A479BDFC06F08E68D8117D115E">journal article</mods:classification>
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<mods:identifier id="375E2B37C8598FDE73EE5A2025BBA1A0" type="DOI">10.1206/0003-0090-423.1.1</mods:identifier>
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<treatment id="03B187A8FFC5FFBAFF633CEA7C79FC99" ID-DOI="http://doi.org/10.5281/zenodo.4612823" ID-GBIF-Taxon="180670781" ID-Zenodo-Dep="4612823" LSID="urn:lsid:plazi:treatment:03B187A8FFC5FFBAFF633CEA7C79FC99" httpUri="http://treatment.plazi.org/id/03B187A8FFC5FFBAFF633CEA7C79FC99" lastPageId="68" pageId="67">
<subSubSection id="C3026535FFC5FFBDFF633CEA7D56FA26" box="[201,543,1465,1489]" pageId="67" type="nomenclature">
<paragraph id="8BA736BEFFC5FFBDFF633CEA7D56FA26" blockId="67.[201,543,1465,1489]" box="[201,543,1465,1489]" pageId="67">
<heading id="D0EF81D2FFC5FFBDFF633CEA7D56FA26" box="[201,543,1465,1489]" centered="true" fontSize="9" level="2" pageId="67" reason="2">
<taxonomicName id="4C184D3DFFC5FFBDFF633CEA7ED9FA26" ID-CoL="8DGYD" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[201,400,1465,1489]" class="Insecta" family="Ironomyiidae" genus="Proironia" kingdom="Animalia" order="Diptera" pageId="67" phylum="Arthropoda" rank="species" species="gibbera" status="sp. nov.">
<emphasis id="B96CEAACFFC5FFBDFF633CEA7ED9FA26" bold="true" box="[201,400,1465,1489]" italics="true" pageId="67">Proironia gibbera</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57D7FFC5FFBDFE343CE97D56FA26" box="[414,543,1465,1489]" pageId="67" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFC5FFBDFE8D3CB87E88F9F4" box="[295,449,1515,1539]" pageId="67" type="description">
<paragraph id="8BA736BEFFC5FFBDFE8D3CB87E88F9F4" blockId="67.[295,449,1515,1539]" box="[295,449,1515,1539]" pageId="67">
<heading id="D0EF81D2FFC5FFBDFE8D3CB87E88F9F4" box="[295,449,1515,1539]" centered="true" fontSize="9" level="2" pageId="67" reason="2">
<figureCitation id="13232A3BFFC5FFBDFE8D3CB87ED2F9F4" box="[295,411,1515,1539]" captionStart="FIG" captionStartId="20.[111,153,1606,1627]" captionTargetBox="[110,1214,228,1574]" captionTargetId="figure-10@20.[110,1214,228,1575]" captionTargetPageId="20" captionText="FIG. 13. Proironia gibbera, n. gen., n. sp. (Ironomyiidae), holotype female in Burmese amber, AMNH Bu-SE7. AC. Habitus, in ventral (A), right lateral (B), and left lateral (C) views. D, E. Head, in frontal and ventral views, respectively. F. Wing. G. Terminalia, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.4613039" httpUri="https://zenodo.org/record/4613039/files/figure.png" pageId="67">Figures 13</figureCitation>
,
<figureCitation id="13232A3BFFC5FFBDFE0D3CB87E88F9F4" box="[423,449,1515,1539]" captionStart="FIG" captionStartId="67.[112,152,1089,1110]" captionTargetBox="[110,1209,232,1044]" captionTargetId="figure-243@67.[108,1212,224,1056]" captionTargetPageId="67" captionText="FIG. 41. Proironia gibbera, n. gen., n. sp. (Ironomyiidae), holotype female in Burmese amber, AMNH Bu-SE7. A. Frontal view of head. B. Left lateral view of thorax. C. Abdominal apex. D. Proleg: femur and tibia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4613111" httpUri="https://zenodo.org/record/4613111/files/figure.png" pageId="67">41</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFC5FFBDFF263F4E7C8EFA03" pageId="67" type="diagnosis">
<paragraph id="8BA736BEFFC5FFBDFF263F4E7C8EFA03" blockId="67.[108,636,1564,1721]" lastBlockId="67.[684,1213,1170,1723]" pageId="67">
DIAGNOSIS: Differs from the other species of the genus, also in Burmese amber, by the following: Head broader, concavity of occiput not nearly as deep; pedicel lobe long, more fingerlike, extended&gt;0.5× the length of basal flagellomere (vs. less than this, lobe more triangular); notopleural, postalar setae shorter; thorax deeper, scutum more dome shaped (vs. slightly flattened on top); pro- and mesocoxae not setose (vs. with dense, fine setae); mesofemur with longitudinal groove, nearly length of femur (vs. without); wing with costal edge convex (vs. flat), apical fork of Sc-R
<subScript id="179C34FBFFC5FFBDFCEF3CD67C07FA61" attach="left" box="[837,846,1413,1430]" fontSize="7" pageId="67">1</subScript>
larger; apex of wing broadly rounded (vs. acute); fork of M
<subScript id="179C34FBFFC5FFBDFBBA3CF57B50FA40" attach="left" box="[1040,1049,1446,1463]" fontSize="7" pageId="67">1</subScript>
-M
<subScript id="179C34FBFFC5FFBDFB913CF57B0DFA40" attach="left" box="[1083,1092,1446,1463]" fontSize="7" pageId="67">2</subScript>
with very short stem; crossvein dm-cu oblique to M
<subScript id="179C34FBFFC5FFBDFB283C947BC2FA2F" attach="left" box="[1154,1163,1479,1496]" fontSize="7" pageId="67">2</subScript>
and CuA
<subScript id="179C34FBFFC5FFBDFD753CBB7DA1FA0E" attach="left" box="[735,744,1512,1529]" fontSize="7" pageId="67">1</subScript>
(vs. perpendicular).
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFC5FFBAFD663CAD7C55FCDB" lastPageId="68" pageId="67" type="description">
<paragraph id="8BA736BEFFC5FFBAFD663CAD7E74FA15" blockId="67.[684,1213,1170,1723]" lastBlockId="68.[108,639,226,1738]" lastPageId="68" pageId="67">
DESCRIPTION:
<emphasis id="B96CEAACFFC5FFBDFCDC3CAD7CF2F9E1" bold="true" box="[886,955,1534,1558]" pageId="67">
<collectorName id="26ED5368FFC5FFBDFCDC3CAD7CFFF9E1" box="[886,950,1534,1558]" pageId="67">Head</collectorName>
:
</emphasis>
<collectorName id="26ED5368FFC5FFBDFC6D3CAD7B5BF9E1" box="[967,1042,1534,1558]" pageId="67">Rather</collectorName>
broad, depth/ width 0.66.
<collectorName id="26ED5368FFC5FFBDFC923F4C7CD9F9C0" box="[824,912,1567,1591]" pageId="67">Occiput</collectorName>
near posterior margin of eyes rimmed with row of stiff, whitish setae.
<emphasis id="B96CEAACFFC5FFBDFD063F327DAFF98E" bold="true" box="[684,742,1633,1657]" pageId="67">
<location id="8EC76065FFC5FFBDFD063F327DA8F98E" LSID="urn:lsid:plazi:treatment:03B187A8FFC5FFBAFF633CEA7C79FC99:8EC76065FFC5FFBDFD063F327DA8F98E" box="[684,737,1633,1657]" municipality="Bare" name="Eyes" pageId="67">Eyes</location>
:
</emphasis>
<collectingMunicipality id="6BC3ACC4FFC5FFBDFD473F327C69F98F" box="[749,800,1633,1656]" pageId="67">Bare</collectingMunicipality>
, fully separated in female by distance slightly greater than outside width of lateral ocelli; no differentiation of facets. Frons: Dark brown, largely bare, with a few fine, scattered, minute setulae; lateral margins parallel, modest emarginations around bases of antennae; median furrow present, faint;
<materialsCitation id="3B703CE3FFC2FFBAFECE38167FA0FE68" ID-GBIF-Occurrence="3059949309" location="Face" pageId="68" specimenCount="1">
no ocellar, postocellar or frontal setae present.
<location id="8EC76065FFC2FFBAFEF138357EC5FE8A" LSID="urn:lsid:plazi:treatment:03B187A8FFC5FFBAFF633CEA7C79FC99:8EC76065FFC2FFBAFEF138357EC5FE8A" box="[347,396,358,381]" name="Face" pageId="68">Face</location>
below antennae relatively short
</materialsCitation>
; oral cavity exposed anteriorly; clypeus broad but narrow;
<materialsCitation id="3B703CE3FFC2FFBAFED838FB7E13FDF5" ID-GBIF-Occurrence="3059949310" collectorName="Vibrissa" location="Antenna" municipality="Bases" pageId="68" specimenCount="1">
oral margin rimmed by protruding membrane.
<collectorName id="26ED5368FFC2FFBAFECD389A7EF7FE16" box="[359,446,457,481]" pageId="68">Vibrissa</collectorName>
absent.
<emphasis id="B96CEAACFFC2FFBAFDB9389A7D35FE16" bold="true" box="[531,636,457,481]" pageId="68">
<location id="8EC76065FFC2FFBAFDB9389A7D3FFE16" LSID="urn:lsid:plazi:treatment:03B187A8FFC5FFBAFF633CEA7C79FC99:8EC76065FFC2FFBAFDB9389A7D3FFE16" box="[531,630,457,481]" municipality="Bases" name="Antenna" pageId="68">Antenna</location>
:
</emphasis>
<collectingMunicipality id="6BC3ACC4FFC2FFBAFFC638B87FE1FDF5" box="[108,168,491,514]" pageId="68">Bases</collectingMunicipality>
very close, oval
</materialsCitation>
; scape very small, conical; pedicel with fingerlike lobe inserted into lateral and mesal surfaces of basal flagellomere; basal flagellomere subcircular, laterally flattened; arista terminal, with 1 or 2 small basal articles (not clearly observed);
<materialsCitation id="3B703CE3FFC2FFBAFE853BDC7D31FD1D" ID-GBIF-Occurrence="3059949315" collectorName="Mouthparts &amp; Compact" location="Palp" pageId="68" specimenCount="1">
arista bare.
<emphasis id="B96CEAACFFC2FFBAFE003BC37D7EFD5F" bold="true" box="[426,567,656,680]" pageId="68">
<collectorName id="26ED5368FFC2FFBAFE003BC37D7BFD5F" box="[426,562,656,680]" pageId="68">Mouthparts</collectorName>
:
</emphasis>
<collectorName id="26ED5368FFC2FFBAFD963BC37FD5FD3F" pageId="68">Compact</collectorName>
, not protruding beyond level of basal flagellomeres.
<location id="8EC76065FFC2FFBAFF723B817E40FD1D" LSID="urn:lsid:plazi:treatment:03B187A8FFC5FFBAFF633CEA7C79FC99:8EC76065FFC2FFBAFF723B817E40FD1D" box="[216,265,722,746]" name="Palp" pageId="68">Palp</location>
small, globular, light in color
</materialsCitation>
;
<materialsCitation id="3B703CE3FFC2FFBAFFC63BA07E35FCBA" ID-GBIF-Occurrence="3059949312" location="Very" pageId="68" specimenCount="1">
labellum small, laterally flattened, extended slightly beyond level of palps.
<emphasis id="B96CEAACFFC2FFBAFE073A477D4CFCDB" bold="true" box="[429,517,788,812]" pageId="68">Thorax:</emphasis>
<location id="8EC76065FFC2FFBAFDA03A477D74FCDB" LSID="urn:lsid:plazi:treatment:03B187A8FFC5FFBAFF633CEA7C79FC99:8EC76065FFC2FFBAFDA03A477D74FCDB" box="[522,573,788,812]" name="Very" pageId="68">Very</location>
deep, depth ca. 1.1× the length
</materialsCitation>
; scutum arched, almost hemispherical in profile. Scutum with scattered, very fine, decumbent setulae (barely visible); one pair of dorsocentral setae, short, stout, very close to anterior margin of scutellum; scutellum with 4 pairs of short, stout setae; 4 notopleural setae, located just above pleural cleft and anterior to transverse suture; 2 postalar setae; prescutellum well developed, lenticular.
<emphasis id="B96CEAACFFC2FFBAFE1C3D6E7EB4FBA2" bold="true" box="[438,509,1085,1109]" pageId="68">Wing:</emphasis>
Relatively broad, width/length 0.51, apex broadly rounded, anterior (costal) edge convex (not flat);
<materialsCitation id="3B703CE3FFC2FFBAFD903DD37F8BFACA" ID-GBIF-Occurrence="3059949313" collectionCode="R" collectorName="Vein C &amp; Sc" pageId="68" specimenCode="R4, R1, R2" specimenCount="1">
membrane entirely glassy, no microtrichia (even near wing margin).
<collectorName id="26ED5368FFC2FFBAFEBD3D917E2CFB2D" box="[279,357,1218,1242]" pageId="68">Vein C</collectorName>
ends at apex of
<specimenCode id="DBBE9EC5FFC2FFBAFD8D3D917D09FB29" box="[551,576,1218,1246]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="68">
R
<subScript id="179C34FBFFC2FFBAFD9D3D9E7D09FB29" attach="left" box="[567,576,1229,1246]" fontSize="7" pageId="68">4</subScript>
</specimenCode>
+5.
<collectorName id="26ED5368FFC2FFBAFDC93D917D35FB2D" box="[611,636,1218,1242]" pageId="68">Sc</collectorName>
long, fused to
<specimenCode id="DBBE9EC5FFC2FFBAFEA03DB07E6DFB08" box="[266,292,1251,1279]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="68">
R
<subScript id="179C34FBFFC2FFBAFEB13DBD7E6DFB08" attach="left" box="[283,292,1262,1279]" fontSize="7" pageId="68">1</subScript>
</specimenCode>
for 0.62× its length, both veins free at base and apex.
<specimenCode id="DBBE9EC5FFC2FFBAFEC93C577E32FAD7" box="[355,379,1284,1312]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="68">
R
<subScript id="179C34FBFFC2FFBAFED83C5C7E32FAD7" attach="left" box="[370,379,1295,1312]" fontSize="7" pageId="68">2</subScript>
</specimenCode>
+3 and
<specimenCode id="DBBE9EC5FFC2FFBAFE6C3C577EA9FAD7" box="[454,480,1284,1312]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="68">
R
<subScript id="179C34FBFFC2FFBAFE7D3C5C7EA9FAD7" attach="left" box="[471,480,1295,1312]" fontSize="7" pageId="68">4</subScript>
</specimenCode>
+5 long, nearly parallel
</materialsCitation>
; crossvein br-m very short; cell d long and narrow (L/
<geoCoordinate id="EE2C5079FFC2FFBAFEB13C157E3AFAA9" box="[283,371,1350,1374]" degrees="5.55" direction="west" orientation="longitude" pageId="68" precision="555" value="-5.55">W 5.55</geoCoordinate>
), apex acute (crossvein dm-cu oblique). M
<subScript id="179C34FBFFC2FFBAFEEF3C217E07FA74" attach="left" box="[325,334,1394,1411]" fontSize="7" pageId="68">1</subScript>
and M
<subScript id="179C34FBFFC2FFBAFE0F3C217EE7FA74" attach="left" box="[421,430,1394,1411]" fontSize="7" pageId="68">2</subScript>
joined at base by very short stem connecting to cell d; cell cup narrow, apex acute. A
<subScript id="179C34FBFFC2FFBAFEC83CE67E22FA31" attach="left" box="[354,363,1461,1478]" fontSize="7" pageId="68">1</subScript>
, A
<subScript id="179C34FBFFC2FFBAFE273CE67EDFFA31" attach="left" box="[397,406,1461,1478]" fontSize="7" pageId="68">2</subScript>
, anal lobe and alula not entirely visible.
</paragraph>
<paragraph id="8BA736BEFFC2FFBAFF273CB87BF2FE37" blockId="68.[108,639,226,1738]" lastBlockId="68.[684,1213,226,878]" pageId="68">
<emphasis id="B96CEAACFFC2FFBAFF273CB87F81F9F4" bold="true" box="[141,200,1515,1539]" pageId="68">Legs:</emphasis>
Slender. All coxae with a few, fine, light setae; femora without macrosetae. Protibia with thick, short, preapical seta on mesal surface, ca. 10 short transverse, ctenidial-like rows of fine setae, apical row long and comblike; tarsomeres 14 with short ctenidial combs. Mesofemur with narrow but well-defined longitudinal groove on ventral surface (nearly length of femur), to which tibia must fit against. Mesotibia with minute, spurlike lobe on ventral surface, spurlike ventroapical sea; tarsomeres 14 with short rows transverse ctenidia. Metatarsus same as mesotarsus. Pretarsus with claws well developed, pulvilli slightly shorter than claws; empodium setiform.
</paragraph>
<paragraph id="8BA736BEFFC2FFBAFD67389A7B5CFD3F" blockId="68.[684,1213,226,878]" pageId="68">
<emphasis id="B96CEAACFFC2FFBAFD67389A7C0FFE16" bold="true" box="[717,838,457,481]" pageId="68">Abdomen:</emphasis>
Relatively short, tapered apicad; with fine, short, scattered setulae on tergites and sternites, no macrosetae. Eight tergites and 7 sternites visible, well developed, narrow pleural membrane exposed. Dorsal margin of tergite 7 very short, hidden beneath posterior margin of tergite 6. Cercus 1-segmented.
<emphasis id="B96CEAACFFC2FFBAFC513BC37D86FD3F" bold="true" pageId="68">Female Terminalia:</emphasis>
Largely retracted, not visible.
</paragraph>
<paragraph id="8BA736BEFFC2FFBAFD663B817C55FCDB" blockId="68.[684,1213,226,878]" pageId="68">
<materialsCitation id="3B703CE3FFC2FFBAFD663B817C55FCDB" ID-GBIF-Occurrence="3059949316" collectionCode="AMNH" pageId="68" specimenCount="1" typeStatus="holotype">
SPECIMENS EXAMINED:
<typeStatus id="54A3881CFFC2FFBAFC413B817B1FFD1D" box="[1003,1110,722,746]" pageId="68" type="holotype">Holotype</typeStatus>
, female,
<collectionCode id="ED09AE7BFFC2FFBAFD063BA07C49FCFC" box="[684,768,755,779]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="68">AMNH</collectionCode>
Bu-SE07, in Burmese amber. In excellent condition.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFC2FFBAFD663A667C79FC99" pageId="68" type="etymology">
<paragraph id="8BA736BEFFC2FFBAFD663A667C79FC99" blockId="68.[684,1213,226,878]" pageId="68">
ETYMOLOGY: From Latin
<emphasis id="B96CEAACFFC2FFBAFB983A667B30FCBA" box="[1074,1145,821,845]" italics="true" pageId="68">gibber</emphasis>
(L., “hunched”).
</paragraph>
</subSubSection>
</treatment>
</document>