treatments-xml/data/03/B1/87/03B18797FFCA9D2C4BA500D4DDB821B8.xml

<document id="564B466CD20A561E294936479777EE81" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2021" docId="03B18797FFCA9D2C4BA500D4DDB821B8" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Benedictesmus yaquina Shear &amp; Marek 2021, n. sp." docType="treatment" docVersion="7" lastPageNumber="118" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="116" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BADA653D25E105B8FCA44108E018ED97">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
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<mods:namePart id="1512C715E1FA19D9E8DFC09D9F3B3BEE">Shear, William A.</mods:namePart>
<mods:affiliation id="C8195ADBD4D9E2230FF2186E8CBD5059">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
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<mods:namePart id="8F87C8E3A7730EA621E344FA5D2E487E">Marek, Paul E.</mods:namePart>
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<mods:affiliation id="866C371E63AE6CE0B961CC777CA215D0">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
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<treatment id="03B18797FFCA9D2C4BA500D4DDB821B8" ID-DOI="http://doi.org/10.5281/zenodo.4923607" ID-GBIF-Taxon="182644917" ID-Zenodo-Dep="4923607" LSID="urn:lsid:plazi:treatment:03B18797FFCA9D2C4BA500D4DDB821B8" httpUri="http://treatment.plazi.org/id/03B18797FFCA9D2C4BA500D4DDB821B8" lastPageId="37" lastPageNumber="118" pageId="35" pageNumber="116">
<subSubSection id="C302650AFFCA9D2A4BA500D4DB912644" box="[151,722,1518,1545]" pageId="35" pageNumber="116" type="nomenclature">
<paragraph id="8BA73681FFCA9D2A4BA500D4DB912644" blockId="35.[151,722,1518,1581]" box="[151,722,1518,1545]" pageId="35" pageNumber="116">
<heading id="D0EF81EDFFCA9D2A4BA500D4DB912644" bold="true" box="[151,722,1518,1545]" fontSize="11" level="1" pageId="35" pageNumber="116" reason="1">
<emphasis id="B96CEA93FFCA9D2A4BA500D4DB912644" bold="true" box="[151,722,1518,1545]" pageId="35" pageNumber="116">
<taxonomicName id="4C184D02FFCA9D2A4BA500D4DBC12644" authority="Shear &amp; Marek" authorityName="Shear &amp; Marek" authorityYear="2021" box="[151,642,1518,1545]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="35" pageNumber="116" phylum="Arthropoda" rank="species" species="yaquina" status="sp. nov.">
<emphasis id="B96CEA93FFCA9D2A4BA500D4D8F62644" bold="true" box="[151,437,1518,1545]" italics="true" pageId="35" pageNumber="116">Benedictesmus yaquina</emphasis>
Shear &amp; Marek
</taxonomicName>
, 
<taxonomicNameLabel id="A25F57E8FFCA9D2A49BD00D5DB912644" box="[655,722,1519,1545]" pageId="35" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C302650AFFCA9D2A4BA50329DC2926D4" pageId="35" pageNumber="116" type="description">
<paragraph id="8BA73681FFCA9D2A4BA50329D8262660" blockId="35.[151,722,1518,1581]" box="[151,357,1554,1581]" pageId="35" pageNumber="116">
<figureCitation id="13232A04FFCA9D2A4BA50329D8732660" box="[151,304,1555,1581]" captionStart-0="FIGURES 111–116" captionStart-1="FIGURES 117–120" captionStartId-0="38.[151,264,1943,1967]" captionStartId-1="39.[151,264,1199,1223]" captionTargetBox-0="[170,1418,182,1917]" captionTargetBox-1="[151,1436,181,1176]" captionTargetId-0="figure-24@38.[170,1418,181,1919]" captionTargetId-1="figure-17@39.[151,1436,181,1176]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionText-0="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." captionText-1="FIGURES 117–120. Species of Benedictesmus, n. gen. 117. Gonopod tips of Benedictesmus yaquina, n. sp., posterior view. SEM stub WS33-14. Figs 118–120. Benedictesmus timber, n. sp. 118. Head and anterior rings, dorsal view. 119. Midbody ring, dorsal view. 120. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi-0="http://doi.org/10.5281/zenodo.4805063" figureDoi-1="http://doi.org/10.5281/zenodo.4805069" httpUri-0="https://zenodo.org/record/4805063/files/figure.png" httpUri-1="https://zenodo.org/record/4805069/files/figure.png" pageId="35" pageNumber="116">Figs 111–117</figureCitation>
, 
<figureCitation id="13232A04FFCA9D2A4A090328D8262660" box="[315,357,1554,1581]" captionStart="FIGURES 133–137" captionStartId="42.[151,264,1252,1276]" captionTargetBox="[160,1429,181,1218]" captionTargetId="figure-24@42.[151,1436,181,1228]" captionTargetPageId="42" captionText="FIGURES 133–137. Drawings of gonopods of polydesmid species. 133. Right gonopod of Benedictesmus aureus, n. sp., mesal view. 134. Right gonopod of Benedictesmus ellenae,n. sp., lateral view. 135, Right gonopod of Benedictesmus yaquina, n. sp., lateral view. Fig. 136, 137, Benedictesmus timber.n. sp. 136, Right gonopod, lateral view. 137, Tip of right gonopod, mesal view. Abbreviation: p, pulvillus." figureDoi="http://doi.org/10.5281/zenodo.4805079" httpUri="https://zenodo.org/record/4805079/files/figure.png" pageId="35" pageNumber="116">135</figureCitation>
</paragraph>
<paragraph id="8BA73681FFCA9D2A4BA50361DC2926D4" blockId="35.[151,1437,1627,2013]" pageId="35" pageNumber="116">
<emphasis id="B96CEA93FFCA9D2A4BA50361D9A72638" bold="true" box="[151,228,1627,1653]" pageId="35" pageNumber="116">Types.</emphasis>
<materialsCitation id="3B703CDCFFCA9D2A4BDC0361DC2526D4" ID-GBIF-Occurrence="3128519339" collectingDate="1971-12-20" collectionCode="CAS" collectorName="E. M. Benedict. All" county="Lincoln Co." elevation="6" location="Elk City" municipality="Male" pageId="35" pageNumber="116" specimenCount="2" stateProvince="Oregon" typeStatus="holotype">
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and male and female 
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from 
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<elevation id="0035D1B2FFCA9D2A487C0361DAD22638" box="[846,913,1627,1653]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="35" pageNumber="116" unit="m" value="0.6">0.6 m</elevation>
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west of 
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, 
<location id="8EC7605AFFCA9D2A4F5A0361DC4D2638" LSID="urn:lsid:plazi:treatment:03B18797FFCA9D2C4BA500D4DDB821B8:8EC7605AFFCA9D2A4F5A0361DC4D2638" box="[1128,1294,1627,1653]" county="Lincoln Co." municipality="Male" name="Yaquina River" pageId="35" pageNumber="116" stateProvince="Oregon">Yaquina River</location>
, 44.6233°, - 123.8918°, 
<collectingCounty id="62C64E0DFFCA9D2A4A2B0345D8E126D4" box="[281,418,1663,1689]" pageId="35" pageNumber="116">Lincoln Co.</collectingCounty>
, 
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, collected 
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<collectingDate id="EFE2E9A9FFCA9D2A49490345DA1326D4" box="[635,848,1663,1689]" pageId="35" pageNumber="116" value="1971-12-20">20 December 1971</collectingDate>
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by 
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types deposited in 
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.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFCA9D2A4BF50398D9922748" pageId="35" pageNumber="116" type="diagnosis">
<paragraph id="8BA73681FFCA9D2A4BF50398D9922748" blockId="35.[151,1437,1627,2013]" pageId="35" pageNumber="116">
<emphasis id="B96CEA93FFCA9D2A4BF50398D80126F1" bold="true" box="[199,322,1698,1724]" pageId="35" pageNumber="116">Diagnosis.</emphasis>
Distinct from other species of the genus in having an unenlarged, straight epiproct (
<figureCitation id="13232A04FFCA9D2A4E310399DC2026F0" box="[1283,1379,1699,1725]" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="35" pageNumber="116">Fig. 113</figureCitation>
) and four rows of setae on the anterior metazonites (
<figureCitation id="13232A04FFCA9D2A499C03FDDA4C26AC" box="[686,783,1735,1761]" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="35" pageNumber="116">Fig. 112</figureCitation>
), increasing to five rows on the most posterior rings (
<figureCitation id="13232A04FFCA9D2A4E5D03FDD9872748" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="35" pageNumber="116">Fig. 113</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFCA9D2A4BF50235DC1C2764" box="[199,1375,1807,1833]" pageId="35" pageNumber="116" type="etymology">
<paragraph id="8BA73681FFCA9D2A4BF50235DC1C2764" blockId="35.[151,1437,1627,2013]" box="[199,1375,1807,1833]" pageId="35" pageNumber="116">
<emphasis id="B96CEA93FFCA9D2A4BF50235D80D2764" bold="true" box="[199,334,1807,1833]" pageId="35" pageNumber="116">Etymology.</emphasis>
The species epithet, a noun in apposition, refers to the 
<typeStatus id="54A38823FFCA9D2A488A0235DAAB2764" box="[952,1000,1807,1833]" pageId="35" pageNumber="116">type</typeStatus>
locality along the Yaquina River.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFCA9D2C4BF50208DDD12150" lastPageId="37" lastPageNumber="118" pageId="35" pageNumber="116" type="description">
<paragraph id="8BA73681FFCA9D2C4BF50208DDD12150" blockId="35.[151,1437,1627,2013]" lastBlockId="37.[151,1437,150,501]" lastPageId="37" lastPageNumber="118" pageId="35" pageNumber="116">
<emphasis id="B96CEA93FFCA9D2A4BF50208D81A2701" bold="true" box="[199,345,1842,1868]" pageId="35" pageNumber="116">Description.</emphasis>
<typeStatus id="54A38823FFCA9D2A4A520209D88B2700" box="[352,456,1843,1869]" pageId="35" pageNumber="116" type="holotype">Holotype</typeStatus>
male. Nineteen rings (
<figureCitation id="13232A04FFCA9D2A49FA0209DA6B2700" box="[712,808,1843,1869]" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="35" pageNumber="116">Fig. 111</figureCitation>
; the telson or 19th ring is concealed in the figure by the crassate anterior legs). Length about 
<quantity id="4CE09B64FFCA9D2A490A026CDBD3273C" box="[568,656,1878,1905]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="35" pageNumber="116" unit="mm" value="5.5">5.5 mm</quantity>
, greatest width 
<quantity id="4CE09B64FFCA9D2A4870026DDAE9273C" box="[834,938,1879,1905]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="35" pageNumber="116" unit="mm" value="0.62">0.62 mm</quantity>
. Head densely setose, cuticle alveolate (
<figureCitation id="13232A04FFCA9D2A4E5D026DD98727D8" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="35" pageNumber="116">Fig. 112</figureCitation>
). Collum with anterior marginal row of 20 setae (
<figureCitation id="13232A04FFCA9D2A49C20241DA1127D8" box="[752,850,1915,1941]" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="35" pageNumber="116">Fig. 112</figureCitation>
). Anterior metazonites with four rows of setae, transitioning to four rows (
<figureCitation id="13232A04FFCA9D2A4AAB02A5D8B427F4" box="[409,503,1951,1977]" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="35" pageNumber="116">Fig. 113</figureCitation>
) about ring 15, setal tubercles prominent throughout. Alveolate cuticle absent from metazonites posterior to collum. Epiproct (
<figureCitation id="13232A04FFCA9D2A494B02F9DB9B2790" box="[633,728,1987,2013]" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="35" pageNumber="116">Fig. 114</figureCitation>
) not swollen, relatively long, straight, with extensive alveolate cuticle. Anterior legs crassate (
<figureCitation id="13232A04FFCC9D2C4AC805ADDB1F20FC" box="[506,604,151,177]" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="37" pageNumber="118">Fig. 111</figureCitation>
), tarsi with sphaerotrichomes. Gonopod (
<figureCitation id="13232A04FFCC9D2C4F0505ADDD9620FC" box="[1079,1237,150,177]" captionStart-0="FIGURES 111–116" captionStart-1="FIGURES 117–120" captionStartId-0="38.[151,264,1943,1967]" captionStartId-1="39.[151,264,1199,1223]" captionTargetBox-0="[170,1418,182,1917]" captionTargetBox-1="[151,1436,181,1176]" captionTargetId-0="figure-24@38.[170,1418,181,1919]" captionTargetId-1="figure-17@39.[151,1436,181,1176]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionText-0="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." captionText-1="FIGURES 117–120. Species of Benedictesmus, n. gen. 117. Gonopod tips of Benedictesmus yaquina, n. sp., posterior view. SEM stub WS33-14. Figs 118–120. Benedictesmus timber, n. sp. 118. Head and anterior rings, dorsal view. 119. Midbody ring, dorsal view. 120. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi-0="http://doi.org/10.5281/zenodo.4805063" figureDoi-1="http://doi.org/10.5281/zenodo.4805069" httpUri-0="https://zenodo.org/record/4805063/files/figure.png" httpUri-1="https://zenodo.org/record/4805069/files/figure.png" pageId="37" pageNumber="118">Figs 115–117</figureCitation>
, 
<figureCitation id="13232A04FFCC9D2C4FD105ACDC4D20FC" box="[1251,1294,150,177]" captionStart="FIGURES 133–137" captionStartId="42.[151,264,1252,1276]" captionTargetBox="[160,1429,181,1218]" captionTargetId="figure-24@42.[151,1436,181,1228]" captionTargetPageId="42" captionText="FIGURES 133–137. Drawings of gonopods of polydesmid species. 133. Right gonopod of Benedictesmus aureus, n. sp., mesal view. 134. Right gonopod of Benedictesmus ellenae,n. sp., lateral view. 135, Right gonopod of Benedictesmus yaquina, n. sp., lateral view. Fig. 136, 137, Benedictesmus timber.n. sp. 136, Right gonopod, lateral view. 137, Tip of right gonopod, mesal view. Abbreviation: p, pulvillus." figureDoi="http://doi.org/10.5281/zenodo.4805079" httpUri="https://zenodo.org/record/4805079/files/figure.png" pageId="37" pageNumber="118">135</figureCitation>
) with rather flattened prefemorite. Acropodite short, robust. Pulvillus (
<emphasis id="B96CEA93FFCC9D2C48180581DA022098" bold="true" box="[810,833,187,213]" pageId="37" pageNumber="118">p,</emphasis>
<figureCitation id="13232A04FFCC9D2C487B0581DAEA2098" box="[841,937,187,213]" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="37" pageNumber="118">Fig. 116</figureCitation>
, 
<figureCitation id="13232A04FFCC9D2C48850581DAA22098" box="[951,993,187,213]" captionStart="FIGURES 117–120" captionStartId="39.[151,264,1199,1223]" captionTargetBox="[151,1436,181,1176]" captionTargetId="figure-17@39.[151,1436,181,1176]" captionTargetPageId="39" captionText="FIGURES 117–120. Species of Benedictesmus, n. gen. 117. Gonopod tips of Benedictesmus yaquina, n. sp., posterior view. SEM stub WS33-14. Figs 118–120. Benedictesmus timber, n. sp. 118. Head and anterior rings, dorsal view. 119. Midbody ring, dorsal view. 120. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805069" httpUri="https://zenodo.org/record/4805069/files/figure.png" pageId="37" pageNumber="118">117</figureCitation>
) just distal to midlength of acropodite, somewhat elongate. Pulvillar process (
<emphasis id="B96CEA93FFCC9D2C497905E5DB3120B4" bold="true" box="[587,626,223,249]" pageId="37" pageNumber="118">pp,</emphasis>
<figureCitation id="13232A04FFCC9D2C494B05E5DB9B20B4" box="[633,728,223,249]" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="37" pageNumber="118">Fig. 116</figureCitation>
, 
<figureCitation id="13232A04FFCC9D2C49D705E5DA4C20B4" box="[741,783,223,249]" captionStart="FIGURES 117–120" captionStartId="39.[151,264,1199,1223]" captionTargetBox="[151,1436,181,1176]" captionTargetId="figure-17@39.[151,1436,181,1176]" captionTargetPageId="39" captionText="FIGURES 117–120. Species of Benedictesmus, n. gen. 117. Gonopod tips of Benedictesmus yaquina, n. sp., posterior view. SEM stub WS33-14. Figs 118–120. Benedictesmus timber, n. sp. 118. Head and anterior rings, dorsal view. 119. Midbody ring, dorsal view. 120. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805069" httpUri="https://zenodo.org/record/4805069/files/figure.png" pageId="37" pageNumber="118">117</figureCitation>
) with short tooth on lateral side. Terminal zone entire (
<emphasis id="B96CEA93FFCC9D2C4E4D05E5DCDF20B4" bold="true" box="[1407,1436,223,249]" pageId="37" pageNumber="118">tz,</emphasis>
<figureCitation id="13232A04FFCC9D2C4BA50439D9B52150" box="[151,246,259,285]" captionStart="FIGURES 111–116" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[170,1418,182,1917]" captionTargetId="figure-24@38.[170,1418,181,1919]" captionTargetPageId="38" captionText="FIGURES 111–116. Benedictesmus yaquina, n. gen., n. sp. 111. Male, lateral view. 112. Head and anterior rings, dorsal view. 113. Midbody ring, dorsal view. 114. Pygidium, lateral view. 115. Gonopods in situ, ventral view. 116. Same, lateral view. SEM stub WS33-14. Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805063" httpUri="https://zenodo.org/record/4805063/files/figure.png" pageId="37" pageNumber="118">Fig. 116</figureCitation>
, 
<figureCitation id="13232A04FFCC9D2C4A330439D86E2150" box="[257,301,259,285]" captionStart="FIGURES 117–120" captionStartId="39.[151,264,1199,1223]" captionTargetBox="[151,1436,181,1176]" captionTargetId="figure-17@39.[151,1436,181,1176]" captionTargetPageId="39" captionText="FIGURES 117–120. Species of Benedictesmus, n. gen. 117. Gonopod tips of Benedictesmus yaquina, n. sp., posterior view. SEM stub WS33-14. Figs 118–120. Benedictesmus timber, n. sp. 118. Head and anterior rings, dorsal view. 119. Midbody ring, dorsal view. 120. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805069" httpUri="https://zenodo.org/record/4805069/files/figure.png" pageId="37" pageNumber="118">117</figureCitation>
), with distinct apical tooth. Females similar to males in nonsexual characters.
</paragraph>
</subSubSection>
<caption id="DF676609FFCD9D2D4BA50206DDC1278D" ID-DOI="http://doi.org/10.5281/zenodo.4805047" ID-Zenodo-Dep="4805047" httpUri="https://zenodo.org/record/4805047/files/figure.png" pageId="36" pageNumber="117" startId="36.[151,264,1852,1876]" targetBox="[151,1436,181,1828]" targetPageId="36">
<paragraph id="8BA73681FFCD9D2D4BA50206DDC1278D" blockId="36.[151,1436,1852,1984]" pageId="36" pageNumber="117">
<emphasis id="B96CEA93FFCD9D2D4BA50206D82B2719" bold="true" box="[151,360,1852,1876]" pageId="36" pageNumber="117">FIGURES 101–106</emphasis>
. Species of 
<taxonomicName id="4C184D02FFCD9D2D4AD40206DBC02719" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[486,643,1852,1876]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="36" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">Benedictesmus</taxonomicName>
, 
<taxonomicNameLabel id="A25F57E8FFCD9D2D49A20206DB942719" box="[656,727,1852,1876]" pageId="36" pageNumber="117" rank="genus">
<emphasis id="B96CEA93FFCD9D2D49A20206DB922719" bold="true" box="[656,721,1852,1876]" pageId="36" pageNumber="117">n. gen</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B96CEA93FFCD9D2D49ED0206DA462719" bold="true" box="[735,773,1852,1876]" pageId="36" pageNumber="117">101</emphasis>
, Gonopods of 
<emphasis id="B96CEA93FFCD9D2D48AC0206DD932719" bold="true" box="[926,1232,1852,1876]" pageId="36" pageNumber="117">
<taxonomicName id="4C184D02FFCD9D2D48AC0206DDCA2719" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[926,1161,1852,1876]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="36" pageNumber="117" phylum="Arthropoda" rank="species" species="aureus" status="sp. nov.">
<emphasis id="B96CEA93FFCD9D2D48AC0206DDCA2719" bold="true" box="[926,1161,1852,1876]" italics="true" pageId="36" pageNumber="117">Benedictesmus aureus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25F57E8FFCD9D2D4FA40206DD932719" box="[1174,1232,1852,1876]" pageId="36" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, ventral view. SEM stub WS33-6. 
<emphasis id="B96CEA93FFCD9D2D4A1E025AD8C62735" bold="true" box="[300,389,1888,1912]" pageId="36" pageNumber="117">102–106</emphasis>
. 
<emphasis id="B96CEA93FFCD9D2D4AA1025ADBBA2735" bold="true" box="[403,761,1888,1912]" pageId="36" pageNumber="117">
<taxonomicName id="4C184D02FFCD9D2D4AA1025ADBC12735" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[403,642,1888,1912]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="36" pageNumber="117" phylum="Arthropoda" rank="species" species="ellenae" status="sp. nov.">
<emphasis id="B96CEA93FFCD9D2D4AA1025ADBC12735" bold="true" box="[403,642,1888,1912]" italics="true" pageId="36" pageNumber="117">Benedictesmus ellenae</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25F57E8FFCD9D2D49A2025ADB882735" box="[656,715,1888,1912]" pageId="36" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
102
</emphasis>
. Head and anterior rings, dorsal view. 
<emphasis id="B96CEA93FFCD9D2D4FBC025ADDF72735" bold="true" box="[1166,1204,1888,1912]" pageId="36" pageNumber="117">103</emphasis>
. Midbody ring, dorsal view. 
<emphasis id="B96CEA93FFCD9D2D4BE402BED9BF27D1" bold="true" box="[214,252,1924,1948]" pageId="36" pageNumber="117">104</emphasis>
. Pygidium, lateral view. 
<emphasis id="B96CEA93FFCD9D2D493702BEDB6827D1" bold="true" box="[517,555,1924,1948]" pageId="36" pageNumber="117">105</emphasis>
. Gonopods 
<emphasis id="B96CEA93FFCD9D2D499E02BFDBAD27D1" box="[684,750,1925,1948]" italics="true" pageId="36" pageNumber="117">in situ</emphasis>
, lateroventral view. 
<emphasis id="B96CEA93FFCD9D2D48F702BEDAA827D1" bold="true" box="[965,1003,1924,1948]" pageId="36" pageNumber="117">106</emphasis>
. Pulvillus of gonopod. SEM stub WS33- 13.Abbreviations: 
<emphasis id="B96CEA93FFCD9D2D4A660292D821278D" bold="true" box="[340,354,1960,1984]" pageId="36" pageNumber="117">p</emphasis>
, pulvillus; 
<emphasis id="B96CEA93FFCD9D2D4AE70292D8B2278D" bold="true" box="[469,497,1960,1984]" pageId="36" pageNumber="117">pp</emphasis>
, pulvillar process; 
<emphasis id="B96CEA93FFCD9D2D49810292DB9B278D" bold="true" box="[691,728,1960,1984]" pageId="36" pageNumber="117">ppf</emphasis>
, prefemorite pore field; 
<emphasis id="B96CEA93FFCD9D2D48E20292DAA7278D" bold="true" box="[976,996,1960,1984]" pageId="36" pageNumber="117">tz</emphasis>
, terminal zone.
</paragraph>
</caption>
<subSubSection id="C302650AFFCC9D2C4BF5041CDDB821B8" pageId="37" pageNumber="118" type="distribution">
<paragraph id="8BA73681FFCC9D2C4BF5041CDA17210C" blockId="37.[151,1437,150,501]" box="[199,852,294,321]" pageId="37" pageNumber="118">
<emphasis id="B96CEA93FFCC9D2C4BF5041CD822210D" bold="true" box="[199,353,294,320]" pageId="37" pageNumber="118">Distribution.</emphasis>
Benton, Douglas and Lincoln Cos., 
<collectingRegion id="49DCF863FFCC9D2C49CB041DDA12210C" box="[761,849,295,321]" country="United States of America" name="Oregon" pageId="37" pageNumber="118">Oregon</collectingRegion>
.
</paragraph>
<paragraph id="8BA73681FFCC9D2C4BF50471DDB821B8" blockId="37.[151,1437,150,501]" pageId="37" pageNumber="118">
<emphasis id="B96CEA93FFCC9D2C4BF50471D8612128" box="[199,290,331,357]" italics="true" pageId="37" pageNumber="118">Records</emphasis>
: 
<collectingRegion id="49DCF863FFCC9D2C4A020471D8EA2128" box="[304,425,331,357]" country="United States of America" name="Oregon" pageId="37" pageNumber="118">OREGON</collectingRegion>
: 
<emphasis id="B96CEA93FFCC9D2C4A810471DB722128" box="[435,561,331,357]" italics="true" pageId="37" pageNumber="118">Benton Co.</emphasis>
: 
<quantity id="4CE09B64FFCC9D2C490D0471DBC42128" box="[575,647,331,357]" metricMagnitude="3" metricUnit="m" metricValue="3.7014912" pageId="37" pageNumber="118" unit="mi" value="2.3">2.3 mi</quantity>
northwest of Glenbrook on South Fork of the Alsea River access road, 44.3313°, -123.4314°, 
<quantity id="4CE09B64FFCC9D2C4AA70455D8A721C4" box="[405,484,367,393]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="37" pageNumber="118" unit="ft" value="1200.0">1200 ft</quantity>
asl, 
<date id="FFA61041FFCC9D2C49250455DB9D21C4" box="[535,734,367,393]" pageId="37" pageNumber="118" value="1971-12-04">4 December 1971</date>
, E. M. Benedict, mm; Clemens Park, Seely Creek Road, 
<quantity id="4CE09B64FFCC9D2C4E640455DCDE21C4" box="[1366,1437,367,393]" metricMagnitude="2" metricUnit="m" metricValue="4.828032" pageId="37" pageNumber="118" unit="mi" value="0.3">0.3 mi</quantity>
from SR34, 44.4092°, -123.4644°, 
<quantity id="4CE09B64FFCC9D2C491004A9DB2721E0" box="[546,612,403,429]" metricMagnitude="2" metricUnit="m" metricValue="1.2192" pageId="37" pageNumber="118" unit="ft" value="400.0">400 ft</quantity>
asl, 
<date id="FFA61041FFCC9D2C49AB04A9DA2721E0" box="[665,868,402,429]" pageId="37" pageNumber="118" value="2005-12-04">4 December 2005</date>
, W. Leonard, C. Richart, mm, ff. 
<emphasis id="B96CEA93FFCC9D2C4FEE04A9DC2921E0" box="[1244,1386,403,429]" italics="true" pageId="37" pageNumber="118">Douglas Co.</emphasis>
: 
<quantity id="4CE09B64FFCC9D2C4E4B04A9D9F6219C" metricMagnitude="3" metricUnit="m" metricValue="5.1499008" pageId="37" pageNumber="118" unit="mi" value="3.2">3.2 mi</quantity>
northeast of Scottsburg, 43.6617°, -123.7877°, 
<quantity id="4CE09B64FFCC9D2C49F7048DDA45219C" box="[709,774,439,465]" metricMagnitude="2" metricUnit="m" metricValue="1.2192" pageId="37" pageNumber="118" unit="ft" value="400.0">400 ft</quantity>
asl, 
<date id="FFA61041FFCC9D2C4805048DDD48219C" box="[823,1035,439,465]" pageId="37" pageNumber="118" value="1971-12-11">11 December 1971</date>
, E. M.Benedict, mm, f. 
<emphasis id="B96CEA93FFCC9D2C4E20048DDCD6219C" box="[1298,1429,439,465]" italics="true" pageId="37" pageNumber="118">Lincoln Co.</emphasis>
: 
<quantity id="4CE09B64FFCC9D2C4BA504E1D99C21B8" box="[151,223,475,501]" metricMagnitude="3" metricUnit="m" metricValue="2.2530816" pageId="37" pageNumber="118" unit="mi" value="1.4">1.4 mi</quantity>
west of Nashville, 44.6716°, -123.6100°, 
<quantity id="4CE09B64FFCC9D2C498504E1DBBA21B8" box="[695,761,475,501]" metricMagnitude="2" metricUnit="m" metricValue="1.8288" pageId="37" pageNumber="118" unit="ft" value="600.0">600 ft</quantity>
asl, 
<date id="FFA61041FFCC9D2C481C04E1DD4621B8" box="[814,1029,475,501]" pageId="37" pageNumber="118" value="1971-12-20">20 December 1971</date>
, E. M. Benedict, mm.
</paragraph>
</subSubSection>
</treatment>
</document>