152 lines
16 KiB
XML
152 lines
16 KiB
XML
<document id="A0EC273E9B8D29510EDA262E03FD92FB" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. & Marek, Paul E." docDate="2021" docId="03B18797FFC99D294BA506E9DB3725C0" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Benedictesmus Shear & Marek 2021, new genus" docType="treatment" docVersion="6" lastPageNumber="113" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="113" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="39875CD115AE35C322A0312C2DBB0406" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3DD003969A0FB4B902DF0C7F749BAFAB">
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<mods:title id="A509D00BFFCB8CD9F4298AA456695B64">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
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<mods:name id="7CBB86CE5161A1694590ECCA13DE0447" type="personal">
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<mods:role id="EC2DDE1ECC5AA15FEEAEC478CBC26E05">
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<mods:roleTerm id="32F8BA5F14AC1A6B09E6179C40BC5373">Author</mods:roleTerm>
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<mods:namePart id="39C810DAF2FD4224E8964445B87E79CC">Shear, William A.</mods:namePart>
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<mods:affiliation id="638B2247FE1D57A1CA0BABBFC0521C21">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
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<mods:name id="0CA2ACD28153B810B177BEEB683DA0AE" type="personal">
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<mods:roleTerm id="A00A385ECAF36447CD982FE18FF6C20C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D149A87DA414DE4C6676083EAC1A41ED">Marek, Paul E.</mods:namePart>
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<mods:nameIdentifier id="E21B1049402D5E8766779472D67F9489" type="ORCID">0000-0002-7048-2514</mods:nameIdentifier>
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<mods:affiliation id="3095E96F9D7289C2E6BD373475ABFF84">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
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<mods:nameIdentifier id="CCBBBB1CC8AEACCB95332BDCCA292966" type="email">pmarek@vt.edu</mods:nameIdentifier>
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<mods:typeOfResource id="45B746DDD31D22F698F49D2A53261887">text</mods:typeOfResource>
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<mods:titleInfo id="3F7529FE772FB6A03B5A856BCA2AA371">
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<mods:title id="6F0F66ACE96F8B436344A4DBDC97233E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="07FDE72B402E56EEC5C612F941D31DF7">
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<mods:date id="403A33EE0EE5D929105FA8357AD3C762">2021</mods:date>
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<mods:number id="21CBDA390213A69F4970D46BFBDDA592">2021-05-24</mods:number>
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<mods:detail id="772F764B31F388DDC672D09871230C21" type="volume">
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<mods:number id="BBFF88D242912DC81CC9E5718F1D69BA">4975</mods:number>
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<mods:detail id="7892B90B461E4475A58C2B2F69299A57" type="issue">
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<mods:number id="099168038F666F7A400AF89F57B04E6B">1</mods:number>
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<mods:classification id="602ADFE2BB854373DC0104B5DBDA2A4E">journal article</mods:classification>
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<mods:identifier id="5CAB0C993AC3C0A02052D1D5FCFA323A" type="DOI">10.11646/zootaxa.4975.1.3</mods:identifier>
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</mods:mods>
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<treatment id="03B18797FFC99D294BA506E9DB3725C0" ID-DOI="http://doi.org/10.5281/zenodo.4805051" ID-GBIF-Taxon="182644915" ID-Zenodo-Dep="4805051" LSID="urn:lsid:plazi:treatment:03B18797FFC99D294BA506E9DB3725C0" httpUri="http://treatment.plazi.org/id/03B18797FFC99D294BA506E9DB3725C0" lastPageNumber="113" pageId="32" pageNumber="113">
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<subSubSection id="C302650AFFC99D294BA506E9DBEB23A0" box="[151,680,979,1006]" pageId="32" pageNumber="113" type="nomenclature">
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<paragraph id="8BA73681FFC99D294BA506E9DBEB23A0" blockId="32.[151,680,979,1006]" box="[151,680,979,1006]" pageId="32" pageNumber="113">
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<heading id="D0EF81EDFFC99D294BA506E9DBEB23A0" bold="true" box="[151,680,979,1006]" fontSize="11" level="1" pageId="32" pageNumber="113" reason="1">
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<emphasis id="B96CEA93FFC99D294BA506E9DBEB23A0" bold="true" box="[151,680,979,1006]" pageId="32" pageNumber="113">
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<taxonomicName id="4C184D02FFC99D294BA506E9DB5A23A0" authority="Shear & Marek" authorityName="Shear & Marek" authorityYear="2021" box="[151,537,979,1006]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B96CEA93FFC99D294BA506E9D80F23A3" bold="true" box="[151,332,979,1006]" italics="true" pageId="32" pageNumber="113">Benedictesmus</emphasis>
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Shear & Marek
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25F57E8FFC99D29491406E9DBEB23A0" box="[550,680,979,1005]" pageId="32" pageNumber="113" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C302650AFFC99D294BA50122DD8A2420" pageId="32" pageNumber="113" type="materials_examined">
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<paragraph id="8BA73681FFC99D294BA50122DBAE2462" blockId="32.[151,749,1048,1072]" box="[151,749,1048,1072]" pageId="32" pageNumber="113">
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<emphasis id="B96CEA93FFC99D294BA50122D85A2462" box="[151,281,1048,1071]" italics="true" pageId="32" pageNumber="113">
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<typeStatus id="54A38823FFC99D294BA50122D9852462" box="[151,198,1048,1071]" pageId="32" pageNumber="113">Type</typeStatus>
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Species
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</emphasis>
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:
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<taxonomicName id="4C184D02FFC99D294A140122DBEE247D" authority="Shear & Marek" authorityName="Shear & Marek" authorityYear="2021" box="[294,685,1048,1072]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="species" species="aureus" status="sp. nov.">
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<emphasis id="B96CEA93FFC99D294A140122DB442462" box="[294,519,1048,1072]" italics="true" pageId="32" pageNumber="113">Benedictesmus aureus</emphasis>
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Shear & Marek
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25F57E8FFC99D2949840122DBAE2462" box="[694,749,1048,1071]" pageId="32" pageNumber="113" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BA73681FFC99D294BA5016FDD8A2420" blockId="32.[151,1225,1108,1133]" box="[151,1225,1108,1133]" pageId="32" pageNumber="113">
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<emphasis id="B96CEA93FFC99D294BA5016FD8032420" box="[151,320,1109,1133]" italics="true" pageId="32" pageNumber="113">Species included</emphasis>
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:
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<emphasis id="B96CEA93FFC99D294A7C016EDAB12420" bold="true" box="[334,1010,1108,1133]" pageId="32" pageNumber="113">
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<emphasis id="B96CEA93FFC99D294A7C016EDB702421" bold="true" box="[334,563,1108,1132]" italics="true" pageId="32" pageNumber="113">
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<taxonomicName id="4C184D02FFC99D294A7C016EDB6D2421" authority="Shear & Marek, 2021" authorityName="Shear & Marek" authorityYear="2021" box="[334,558,1108,1132]" class="Diplopoda" family="Polydesmidae" genus="Bendictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="species" species="aureus" status="sp. nov.">Bendictesmus aureus</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A25F57E8FFC99D294908016FDB372420" box="[570,628,1109,1133]" pageId="32" pageNumber="113" rank="species">n. sp.</taxonomicNameLabel>
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,
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<emphasis id="B96CEA93FFC99D2949B2016FDBAD2421" bold="true" box="[640,750,1108,1132]" italics="true" pageId="32" pageNumber="113">
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<taxonomicName id="4C184D02FFC99D2949B2016FDBAA2421" authority="Shear & Marek, 2021" authorityName="Shear & Marek" authorityYear="2021" box="[640,745,1108,1132]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="species" species="ellenae" status="sp. nov.">B. ellenae</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A25F57E8FFC99D2949C6016FDA6D2420" box="[756,814,1109,1133]" pageId="32" pageNumber="113" rank="species">n. sp.</taxonomicNameLabel>
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,
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<taxonomicName id="4C184D02FFC99D294808016FDAEF2420" authority="Shear & Marek, 2021" authorityName="Shear & Marek" authorityYear="2021" box="[826,940,1109,1133]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="species" species="yaquina" status="sp. nov.">
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<emphasis id="B96CEA93FFC99D294808016FDAEF2420" bold="true" box="[826,940,1109,1133]" italics="true" pageId="32" pageNumber="113">B. yaquina</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25F57E8FFC99D29488B016FDAB12420" box="[953,1010,1109,1133]" pageId="32" pageNumber="113" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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and
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<emphasis id="B96CEA93FFC99D294F11016FDD8A2420" bold="true" box="[1059,1225,1108,1133]" pageId="32" pageNumber="113">
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<emphasis id="B96CEA93FFC99D294F11016FDDCA2421" bold="true" box="[1059,1161,1108,1132]" italics="true" pageId="32" pageNumber="113">
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<taxonomicName id="4C184D02FFC99D294F11016FDDC72421" authority="Shear & Marek, 2021" authorityName="Shear & Marek" authorityYear="2021" box="[1059,1156,1108,1132]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="species" species="timber" status="sp. nov.">B. timber</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A25F57E8FFC99D294FBD016FDD8A2420" box="[1167,1225,1109,1133]" pageId="32" pageNumber="113" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C302650AFFC99D294BA501A1D89A24B3" pageId="32" pageNumber="113" type="diagnosis">
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<paragraph id="8BA73681FFC99D294BA501A1D89A24B3" blockId="32.[151,1437,1179,1422]" pageId="32" pageNumber="113">
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<emphasis id="B96CEA93FFC99D294BA501A1D85124F8" bold="true" box="[151,274,1179,1205]" pageId="32" pageNumber="113">Diagnosis.</emphasis>
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Males of some
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<taxonomicName id="4C184D02FFC99D294AE201A1DB9124F8" box="[464,722,1179,1205]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B96CEA93FFC99D294AE201A1DB3524F8" box="[464,630,1179,1205]" italics="true" pageId="32" pageNumber="113">Benedictesmus</emphasis>
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species
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</taxonomicName>
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have only 18 postcephalic rings, while females have 19. The short, simple gonopods have a bifurcate terminal zone (
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<emphasis id="B96CEA93FFC99D2948330185DA5D2494" bold="true" box="[769,798,1215,1241]" pageId="32" pageNumber="113">tz,</emphasis>
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<figureCitation id="13232A04FFC99D2948160185DA832497" box="[804,960,1215,1242]" captionStart="FIGURES 101–106" captionStartId="36.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1828]" captionTargetId="figure-24@36.[151,1436,181,1828]" captionTargetPageId="36" captionText="FIGURES 101–106. Species of Benedictesmus, n. gen. 101, Gonopods of Benedictesmus aureus, n. sp., ventral view. SEM stub WS33-6. 102–106. Benedictesmus ellenae, n. sp. 102. Head and anterior rings, dorsal view. 103. Midbody ring, dorsal view. 104. Pygidium, lateral view. 105. Gonopods in situ, lateroventral view. 106. Pulvillus of gonopod. SEM stub WS33- 13.Abbreviations: p, pulvillus; pp, pulvillar process; ppf, prefemorite pore field; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805047" httpUri="https://zenodo.org/record/4805047/files/figure.png" pageId="32" pageNumber="113">Figs 101, 105</figureCitation>
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); the pulvillus may be extended into a tube (
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<emphasis id="B96CEA93FFC99D294B9301DED85424B0" box="[161,279,1251,1277]" italics="true" pageId="32" pageNumber="113">
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<taxonomicName id="4C184D02FFC99D294B9301DED85724B0" authority="Shear & Marek, 2021" authorityName="Shear & Marek" authorityYear="2021" box="[161,276,1251,1277]" class="Diplopoda" family="Polydesmidae" genus="Benedictesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="113" phylum="Arthropoda" rank="species" species="ellenae" status="sp. nov.">B. ellenae</taxonomicName>
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,
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</emphasis>
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<emphasis id="B96CEA93FFC99D294A2C01D9D81E24B0" bold="true" box="[286,349,1251,1277]" pageId="32" pageNumber="113">
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<taxonomicNameLabel id="A25F57E8FFC99D294A2C01D9D81E24B0" box="[286,349,1251,1277]" pageId="32" pageNumber="113" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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,
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<figureCitation id="13232A04FFC99D294A5901D9D88E24B3" box="[363,461,1251,1278]" captionStart="FIGURES 101–106" captionStartId="36.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1828]" captionTargetId="figure-24@36.[151,1436,181,1828]" captionTargetPageId="36" captionText="FIGURES 101–106. Species of Benedictesmus, n. gen. 101, Gonopods of Benedictesmus aureus, n. sp., ventral view. SEM stub WS33-6. 102–106. Benedictesmus ellenae, n. sp. 102. Head and anterior rings, dorsal view. 103. Midbody ring, dorsal view. 104. Pygidium, lateral view. 105. Gonopods in situ, lateroventral view. 106. Pulvillus of gonopod. SEM stub WS33- 13.Abbreviations: p, pulvillus; pp, pulvillar process; ppf, prefemorite pore field; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805047" httpUri="https://zenodo.org/record/4805047/files/figure.png" pageId="32" pageNumber="113">Fig. 106</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C302650AFFC99D294BF5003DD8DB2524" pageId="32" pageNumber="113" type="etymology">
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<paragraph id="8BA73681FFC99D294BF5003DD8DB2524" blockId="32.[151,1437,1179,1422]" pageId="32" pageNumber="113">
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<emphasis id="B96CEA93FFC99D294BF5003DD80D256C" bold="true" box="[199,334,1287,1313]" pageId="32" pageNumber="113">Etymology.</emphasis>
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The genus is named for the late Ellen M. Benedict (1931–2005), a pseudoscorpion specialist who provided much of the material for this study. Dr. Benedict taught at Portland State University, Malhuer Field Station and Pacific University.
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</paragraph>
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</subSubSection>
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<subSubSection id="C302650AFFC99D294BF50049DB3725C0" box="[199,628,1395,1422]" pageId="32" pageNumber="113" type="distribution">
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<paragraph id="8BA73681FFC99D294BF50049DB3725C0" blockId="32.[151,1437,1179,1422]" box="[199,628,1395,1422]" pageId="32" pageNumber="113">
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<emphasis id="B96CEA93FFC99D294BF50049D82225C0" bold="true" box="[199,353,1395,1421]" pageId="32" pageNumber="113">Distribution.</emphasis>
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Coastal eastern
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<collectingRegion id="49DCF863FFC99D29492B0049DB3225C0" box="[537,625,1395,1421]" country="United States of America" name="Oregon" pageId="32" pageNumber="113">Oregon</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |