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<document id="AF8472999D06E9EE100A2DDD82BF0498" ID-DOI="10.11646/zootaxa.3925.2.9" ID-GBIF-Dataset="265d544a-a242-43b2-b52b-4828c68d8744" ID-ISSN="1175-5326" ID-Zenodo-Dep="241484" ID-ZooBank="9EF51B4A-0199-43D1-8FD6-964986FEE573" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461002533302" checkinUser="plazi" docAuthor="Arbea, Javier I. &amp; Kahrarian, Morteza" docDate="2015" docId="039687CDA8249216FF7BFCD0FC37FBAC" docLanguage="en" docName="zt03925p290.pdf" docOrigin="Zootaxa 3925 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Folsomides halshinicus Arbea &amp; Kahrarian, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="285" masterDocId="FFAFFFB5A8269212FFECFFDFFFBAFFF1" masterDocTitle="The genus Folsomides Stach (Collembola, Isotomidae) in Kermanshah Province (W Iran) with the description of two new species" masterLastPageNumber="290" masterPageNumber="281" pageNumber="283" updateTime="1698564982372" updateUser="plazi">
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<mods:title id="7850C241E8CECDFF21E40F9A7FF2AED9">The genus Folsomides Stach (Collembola, Isotomidae) in Kermanshah Province (W Iran) with the description of two new species</mods:title>
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<mods:namePart id="73F1C10CA758779D6C1562A81583B5C2">Arbea, Javier I.</mods:namePart>
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<mods:namePart id="F84583CC0A374B9BD14A6F0446467E6D">Kahrarian, Morteza</mods:namePart>
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<mods:date id="FA590E2D2B91727B4536AC09AD6712EB">2015</mods:date>
<mods:detail id="42DDB686C33684D466F156B460700C24" type="volume">
<mods:number id="B64C4E66D4447B7F2C1AAC0FFB6B7DA5">3925</mods:number>
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<mods:identifier id="2F2D6AB23CF5924D1522B0C03EE724C5" type="DOI">10.11646/zootaxa.3925.2.9</mods:identifier>
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<treatment id="039687CDA8249216FF7BFCD0FC37FBAC" ID-DOI="http://doi.org/10.5281/zenodo.5686624" ID-GBIF-Taxon="119598534" ID-Zenodo-Dep="5686624" LSID="urn:lsid:plazi:treatment:039687CDA8249216FF7BFCD0FC37FBAC" httpUri="http://treatment.plazi.org/id/039687CDA8249216FF7BFCD0FC37FBAC" lastPageId="4" lastPageNumber="285" pageId="2" pageNumber="283">
<subSubSection id="C3256550A8249210FF7BFCD0FD93FCBA" pageId="2" pageNumber="283" type="nomenclature">
<paragraph id="8B8036DBA8249210FF7BFCD0FDA0FCDB" blockId="2.[151,553,783,843]" box="[151,538,783,810]" pageId="2" pageNumber="283">
<heading id="D0C881B7A8249210FF7BFCD0FDA0FCDB" bold="true" box="[151,538,783,810]" fontSize="11" level="1" pageId="2" pageNumber="283" reason="1">
<emphasis id="B94BEAC9A8249210FF7BFCD0FDA0FCDB" bold="true" box="[151,538,783,810]" pageId="2" pageNumber="283">
<taxonomicName id="4C3F4D58A8249210FF7BFCD0FE0FFCD8" box="[151,437,783,809]" class="Entognatha" family="Isotomidae" genus="Folsomides" kingdom="Animalia" order="Collembola" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="halshinicus" status="sp. nov.">
<emphasis id="B94BEAC9A8249210FF7BFCD0FE0FFCD8" bold="true" box="[151,437,783,809]" italics="true" pageId="2" pageNumber="283">Folsomides halshinicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857B2A8249210FE50FCCFFDA0FCDB" box="[444,538,784,810]" pageId="2" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8036DBA8249210FF7BFCEDFD93FCBA" blockId="2.[151,553,783,843]" box="[151,553,818,843]" pageId="2" pageNumber="283">
<figureCitation id="13042A5EA8249210FF7BFCEDFF40FCBA" box="[151,250,818,843]" captionStart="FIGURES 1 – 2" captionStartId="3.[151,264,1958,1981]" captionTargetBox="[268,1312,208,1919]" captionTargetId="figure@3.[254,1333,195,1935]" captionTargetPageId="3" captionText="FIGURES 1 – 2. Folsomides halshinicus sp. nov.: 1, habitus and chaetotaxy, lateral; 2, habitus and distribution of sensilla and macrochaetae. Scale: 0.2 mm." httpUri="https://zenodo.org/record/241485/files/figure.png" pageId="2" pageNumber="283">Figs 1–2</figureCitation>
,
<figureCitation id="13042A5EA8249210FEEAFCEDFEACFCBA" box="[262,278,818,843]" captionStart="FIGURES 5 – 8" captionStartId="6.[151,264,1888,1911]" captionTargetBox="[184,1399,207,1855]" captionTargetId="figure@6.[174,1413,193,1867]" captionTargetPageId="6" captionText="FIGURES 5 – 8. Folsomides spp.: 5, habitus and chaetotaxy in F. v i no s us, paratype from Canary Is.; 6 – 8, chaetotaxy of Abd IV – VI of F. halshinicus sp. nov. (6), F. subvinosus sp. nov. (7) and F. marchicus (8). Scales: 0.1 mm." httpUri="https://zenodo.org/record/241487/files/figure.png" pageId="2" pageNumber="283">6</figureCitation>
,
<figureCitation id="13042A5EA8249210FECEFCECFEC8FCBA" box="[290,370,819,843]" captionStart="FIGURES 9 – 14" captionStartId="7.[151,264,1144,1167]" captionTargetBox="[242,1341,206,1116]" captionTargetId="figure@7.[232,1355,193,1124]" captionTargetPageId="7" captionText="FIGURES 9 – 14. Folsomides spp.: 9 – 11, postantennal organ and omma in F. halshinicus sp. nov. (9), F. subvinosus sp. nov. (10) and F. marchicus (11); 12 – 14, mucrodens in F. subvinosus sp. nov. (12), F. halshinicus sp. nov. (13) and F. marchicus (14)." httpUri="https://zenodo.org/record/241488/files/figure.png" pageId="2" pageNumber="283">9 &amp; 13</figureCitation>
,
<tableCitation id="C6BD0360A8249210FE92FCEDFE66FCBA" box="[382,476,818,843]" captionStart="TABLE 1" captionStartId="1.[151,239,260,283]" captionText="TABLE 1. Diagnostic characters of Iranian Folsomides" pageId="2" pageNumber="283">Tables 1</tableCitation>
,
<tableCitation id="C6BD0360A8249210FE06FCECFE42FCBA" box="[490,504,819,843]" captionStart="TABLE 2" captionStartId="4.[151,239,152,175]" captionText="TABLE 2. Length of macrochaetae and ordinary chaetae in p-row in F. halshinicus sp. nov. (µm)." pageId="2" pageNumber="283">2</tableCitation>
&amp;
<tableCitation id="C6BD0360A8249210FDF7FCECFD93FCBA" box="[539,553,819,843]" captionStart="TABLE 4" captionStartId="8.[151,239,152,175]" captionTargetBox="[151,1402,306,883]" captionTargetPageId="8" captionText="TABLE 4. Comparison of the two new Iranian species of Folsomides with the species of F. angularis group, with reduced number of microsensilla on tergites." httpUri="http://table.plazi.org/id/DF406653A82E921AFF7BFF47FDC8FF3E" pageId="2" pageNumber="283" tableUuid="DF406653A82E921AFF7BFF47FDC8FF3E">4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3256550A8249210FF7BFCA5FE03FAEE" pageId="2" pageNumber="283" type="materials_examined">
<paragraph id="8B8036DBA8249210FF7BFCA5FE9DFC46" blockId="2.[151,1437,890,2031]" pageId="2" pageNumber="283">
<emphasis id="B94BEAC9A8249210FF7BFCA5FE82FC62" bold="true" box="[151,312,890,915]" pageId="2" pageNumber="283">
<typeStatus id="54848879A8249210FF7BFCA5FF68FC62" box="[151,210,890,915]" pageId="2" pageNumber="283">Type</typeStatus>
locality.
</emphasis>
Western
<collectingCountry id="F328764BA8249210FE49FCA4FE6CFC63" box="[421,470,891,914]" name="Iran" pageId="2" pageNumber="283">Iran</collectingCountry>
: Sarfirooz Abdb (Halshi), Paveh County, Kermanshah Province,
<geoCoordinate id="EE0B501CA8249210FB2DFCA5FA9DFC63" box="[1217,1319,890,915]" direction="north" orientation="latitude" pageId="2" pageNumber="283" precision="925" value="34.933334">34º56' N</geoCoordinate>
,
<geoCoordinate id="EE0B501CA8249210FAD9FCA5FA22FC63" box="[1333,1432,890,915]" direction="east" orientation="longitude" pageId="2" pageNumber="283" precision="925" value="46.45">46º27' E</geoCoordinate>
,
<quantity id="4CC79B3EA8249210FF7BFC41FF56FC47" box="[151,236,926,951]" metricMagnitude="3" metricUnit="m" metricValue="1.663" pageId="2" pageNumber="283" unit="m" value="1663.0">1663 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8B8036DBA8249210FF2BFC1DFD77FBD2" blockId="2.[151,1437,890,2031]" pageId="2" pageNumber="283">
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<typeStatus id="54848879A8249210FF2BFC1DFEB8FC2A" box="[199,258,962,987]" pageId="2" pageNumber="283">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54848879A8249210FE97FC1DFE59FC2A" box="[379,483,962,987]" pageId="2" pageNumber="283" type="holotype">Holotype</typeStatus>
female and 9
<typeStatus id="54848879A8249210FD6DFC1CFD56FC2A" box="[641,748,963,987]" pageId="2" pageNumber="283" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D39FD52A8249210FD10FC1DFCD1FC2A" box="[764,875,962,987]" pageId="2" pageNumber="283" type="female">5 females</specimenCount>
, 4 young specimens),
<collectingCountry id="F328764BA8249210FB8EFC1CFB28FC2B" box="[1122,1170,963,986]" name="Iran" pageId="2" pageNumber="283">Iran</collectingCountry>
, Kermanshah Province, Sarfirooz Abdb (Halshi), soil and leaf litter of oak jungle,
<date id="FF81101BA8249210FCA6FC39FC7AFC0E" box="[842,960,998,1023]" pageId="2" pageNumber="283" value="2013-02-06">06.II.2013</date>
, Morteza Kahrarian leg.
<typeStatus id="54848879A8249210FB04FC39FAEAFC0E" box="[1256,1360,998,1023]" pageId="2" pageNumber="283" type="holotype">Holotype</typeStatus>
and 7
<typeStatus id="54848879A8249210FF7BFBD4FEB8FBD2" box="[151,258,1035,1059]" pageId="2" pageNumber="283" type="paratype">paratypes</typeStatus>
deposited in
<collectionCode id="ED2EAE1EA8249210FE74FBD4FE5BFBD2" box="[408,481,1035,1059]" pageId="2" pageNumber="283">IAUK</collectionCode>
, 2
<typeStatus id="54848879A8249210FDEDFBD4FDD7FBD2" box="[513,621,1035,1059]" pageId="2" pageNumber="283" type="paratype">paratypes</typeStatus>
in
<collectionCode id="ED2EAE1EA8249210FD7CFBD4FD73FBD2" box="[656,713,1035,1059]" pageId="2" pageNumber="283">JIAP</collectionCode>
.
</paragraph>
<paragraph id="8B8036DBA8249210FF2BFBF1FE03FAEE" blockId="2.[151,1437,890,2031]" pageId="2" pageNumber="283">
<emphasis id="B94BEAC9A8249210FF2BFBF1FE3DFBB6" bold="true" box="[199,391,1070,1095]" pageId="2" pageNumber="283">Other material,</emphasis>
<collectingCountry id="F328764BA8249210FE78FBF0FE7EFBB7" box="[404,452,1071,1094]" name="Iran" pageId="2" pageNumber="283">Iran</collectingCountry>
, Kermanshah Province, Eslam Abad-e-Gharb County, Sia Khor Village,
<geoCoordinate id="EE0B501CA8249210FAC6FBF0FA22FBB7" box="[1322,1432,1071,1095]" direction="north" orientation="latitude" pageId="2" pageNumber="283" precision="925" value="34.116665">34º07’ N</geoCoordinate>
,
<geoCoordinate id="EE0B501CA8249210FF7BFB8DFEBBFB9B" box="[151,257,1106,1131]" direction="east" orientation="longitude" pageId="2" pageNumber="283" precision="925" value="46.6">46º36’ E</geoCoordinate>
,
<quantity id="4CC79B3EA8249210FEFDFB8CFED1FB9B" box="[273,363,1107,1131]" metricMagnitude="3" metricUnit="m" metricValue="1.4419999999999997" pageId="2" pageNumber="283" unit="m" value="1442.0">1442 m</quantity>
a.s.l., grassland soil,
<date id="FF81101BA8249210FD9CFB8CFD5FFB9A" box="[624,741,1107,1131]" pageId="2" pageNumber="283" value="2013-05-14">14.V.2013</date>
, Morteza Kahrarian leg.,
<specimenCount id="9D39FD52A8249210FBF4FB8CFBC4FB9A" box="[1048,1150,1106,1131]" pageId="2" pageNumber="283" type="female">1 female</specimenCount>
and 1 young specimen. Kermanshah County, Char zabar (Kor Kor Mountain),
<geoCoordinate id="EE0B501CA8249210FCEEFBA8FCD0FB7F" box="[770,874,1143,1167]" direction="north" orientation="latitude" pageId="2" pageNumber="283" precision="925" value="34.216667">34º13’ N</geoCoordinate>
,
<geoCoordinate id="EE0B501CA8249210FC9AFBA9FC60FB7F" box="[886,986,1142,1167]" direction="east" orientation="longitude" pageId="2" pageNumber="283" precision="925" value="46.666668">46º40’ E</geoCoordinate>
,
<quantity id="4CC79B3EA8249210FC0BFBA9FB87FB7F" box="[999,1085,1142,1167]" metricMagnitude="3" metricUnit="m" metricValue="1.5919999999999999" pageId="2" pageNumber="283" unit="m" value="1592.0">1592 m</quantity>
a.s.l., soil and leaf litter of oak jungle,
<date id="FF81101BA8249210FF07FB45FED8FB42" box="[235,354,1178,1203]" pageId="2" pageNumber="283" value="2013-02-15">15.II.2013</date>
, Morteza Kahrarian leg.,
<specimenCount id="9D39FD52A8249210FD6EFB44FD52FB42" box="[642,744,1178,1203]" pageId="2" pageNumber="283" type="female">1 female</specimenCount>
. Sar-e-pol-e-Zahab County, Patagh Area,
<geoCoordinate id="EE0B501CA8249210FB52FB45FA9DFB43" box="[1214,1319,1178,1203]" direction="north" orientation="latitude" pageId="2" pageNumber="283" precision="925" value="34.416668">34º25’ N</geoCoordinate>
,
<geoCoordinate id="EE0B501CA8249210FADFFB45FA2DFB43" box="[1331,1431,1178,1203]" direction="east" orientation="longitude" pageId="2" pageNumber="283" precision="925" value="46.016666">46º01’ E</geoCoordinate>
,
<quantity id="4CC79B3EA8249210FF7BFB60FF54FB27" box="[151,238,1215,1239]" metricMagnitude="3" metricUnit="m" metricValue="1.404" pageId="2" pageNumber="283" unit="m" value="1404.0">1404 m</quantity>
a.s.l., soil and leaf litter of oak jungle,
<date id="FF81101BA8249210FD5AFB60FC91FB26" box="[694,811,1215,1239]" pageId="2" pageNumber="283" value="2013-05-14">14.V.2013</date>
, Morteza Kahrarian leg.,
<specimenCount id="9D39FD52A8249210FBBDFB60FB79FB26" box="[1105,1219,1214,1239]" pageId="2" pageNumber="283" type="female">2 females</specimenCount>
. Kangavar County, Hassam Abad Village,
<geoCoordinate id="EE0B501CA8249210FE44FB3CFDAFFB0B" box="[424,533,1251,1275]" direction="north" orientation="latitude" pageId="2" pageNumber="283" precision="925" value="34.516666">34º31’ N</geoCoordinate>
,
<geoCoordinate id="EE0B501CA8249210FDC9FB3DFD34FB0B" box="[549,654,1250,1275]" direction="east" orientation="longitude" pageId="2" pageNumber="283" precision="925" value="47.916668">47º55’ E</geoCoordinate>
,
<quantity id="4CC79B3EA8249210FD73FB3CFD42FB0B" box="[671,760,1251,1275]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="283" unit="m" value="1480.0">1480 m</quantity>
a.s.l., Forest Plane (Button wood),
<date id="FF81101BA8249210FB4FFB3CFA94FB0A" box="[1187,1326,1251,1275]" pageId="2" pageNumber="283" value="2012-12-14">14.XII.2012</date>
, Morteza Kahrarian leg.,
<specimenCount id="9D39FD52A8249210FEA9FAD8FE0FFAEE" box="[325,437,1286,1311]" pageId="2" pageNumber="283" type="female">9 females</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256550A8249216FF2BFAF5FD63FCE9" lastPageId="4" lastPageNumber="285" pageId="2" pageNumber="283" type="description">
<paragraph id="8B8036DBA8249210FF2BFAF5FC66FA7A" blockId="2.[151,1437,890,2031]" pageId="2" pageNumber="283">
<emphasis id="B94BEAC9A8249210FF2BFAF5FEE3FAB2" bold="true" box="[199,345,1322,1347]" pageId="2" pageNumber="283">Description.</emphasis>
Size: 0.75–1.05 mm females (
<typeStatus id="54848879A8249210FD33FAF5FCF9FAB2" box="[735,835,1322,1347]" pageId="2" pageNumber="283" type="holotype">holotype</typeStatus>
0.88 mm). Colour: blue greyish; pigment more concentrated in eye fields. Body long and slender (
<figureCitation id="13042A5EA8249210FD3AFA91FC8BFA96" box="[726,817,1358,1383]" captionStart="FIGURES 1 – 2" captionStartId="3.[151,264,1958,1981]" captionTargetBox="[268,1312,208,1919]" captionTargetId="figure@3.[254,1333,195,1935]" captionTargetPageId="3" captionText="FIGURES 1 – 2. Folsomides halshinicus sp. nov.: 1, habitus and chaetotaxy, lateral; 2, habitus and distribution of sensilla and macrochaetae. Scale: 0.2 mm." httpUri="https://zenodo.org/record/241485/files/figure.png" pageId="2" pageNumber="283">Fig 1–2</figureCitation>
). Dorsal chaetae smooth and pointed. SA macrochaeta on Abd IV 0.30–0.35, on Abd V 0.45–0.50 times the tergite length (
<figureCitation id="13042A5EA8249210FC7CFAADFC74FA7A" box="[912,974,1394,1419]" captionStart="FIGURES 5 – 8" captionStartId="6.[151,264,1888,1911]" captionTargetBox="[184,1399,207,1855]" captionTargetId="figure@6.[174,1413,193,1867]" captionTargetPageId="6" captionText="FIGURES 5 – 8. Folsomides spp.: 5, habitus and chaetotaxy in F. v i no s us, paratype from Canary Is.; 6 – 8, chaetotaxy of Abd IV – VI of F. halshinicus sp. nov. (6), F. subvinosus sp. nov. (7) and F. marchicus (8). Scales: 0.1 mm." httpUri="https://zenodo.org/record/241487/files/figure.png" pageId="2" pageNumber="283">Fig 6</figureCitation>
).
</paragraph>
<paragraph id="8B8036DBA8249210FF2BFA48FCC1F992" blockId="2.[151,1437,890,2031]" pageId="2" pageNumber="283">Antennae shorter than cephalic diagonal (10:12–13). Ant IV with about 10 dorsal cylindrical sensilla, a dorsoexternal microsensillum and a subapical pit organ. Sensory organ of Ant III with a pair of small microsensilla and two guard sensilla shorter than surrounding chaetae, one on each side. Another microsensillum found in a ventrolateral location. Ant III with 23 common chaetae. Ant II with a small subcylindrical ventral sensillum and 18 common chaetae, and some of the basal ones short and fine. Ant I with two subcylindrical ventral sensilla and 12 common chaetae; one of the basal chaetae smaller than the others.</paragraph>
<paragraph id="8B8036DBA8249210FF2BF9B1FBC7F95A" blockId="2.[151,1437,890,2031]" pageId="2" pageNumber="283">
Five omma on each side of head. PAO long and narrow, 4.5–5.5 times as long as an omma, with two posterior chaetae (
<figureCitation id="13042A5EA8249210FF15F94DFE8DF95A" box="[249,311,1682,1707]" captionStart="FIGURES 9 – 14" captionStartId="7.[151,264,1144,1167]" captionTargetBox="[242,1341,206,1116]" captionTargetId="figure@7.[232,1355,193,1124]" captionTargetPageId="7" captionText="FIGURES 9 – 14. Folsomides spp.: 9 – 11, postantennal organ and omma in F. halshinicus sp. nov. (9), F. subvinosus sp. nov. (10) and F. marchicus (11); 12 – 14, mucrodens in F. subvinosus sp. nov. (12), F. halshinicus sp. nov. (13) and F. marchicus (14)." httpUri="https://zenodo.org/record/241488/files/figure.png" pageId="2" pageNumber="283">Fig 9</figureCitation>
). 2/5,5,4 labral chaetae. Maxillary palp bifurcate with three sublobal hairs.
</paragraph>
<paragraph id="8B8036DBA8249210FF2BF969FA26F8AE" blockId="2.[151,1437,890,2031]" pageId="2" pageNumber="283">
Dorsal chaetotaxy represented in
<figureCitation id="13042A5EA8249210FDB9F969FD03F93E" box="[597,697,1718,1743]" captionStart="FIGURES 1 – 2" captionStartId="3.[151,264,1958,1981]" captionTargetBox="[268,1312,208,1919]" captionTargetId="figure@3.[254,1333,195,1935]" captionTargetPageId="3" captionText="FIGURES 1 – 2. Folsomides halshinicus sp. nov.: 1, habitus and chaetotaxy, lateral; 2, habitus and distribution of sensilla and macrochaetae. Scale: 0.2 mm." httpUri="https://zenodo.org/record/241485/files/figure.png" pageId="2" pageNumber="283">Figure 1</figureCitation>
. Axial chaetotaxy of Th II-Abd V have 6,4/3,3,3,5,2 pairs of mesochaetae respectively; unpaired axial p-chaeta present on Abd IV. Macrochaetotaxy: 1,1/2,2,3,3,3, M on Abd I and SA on Abd III slightly longer than surrounding chaetae (
<tableCitation id="C6BD0360A8249210FC8FF921FC04F8E6" box="[867,958,1790,1815]" captionStart="TABLE 2" captionStartId="4.[151,239,152,175]" captionText="TABLE 2. Length of macrochaetae and ordinary chaetae in p-row in F. halshinicus sp. nov. (µm)." pageId="2" pageNumber="283">Table 2</tableCitation>
). Microsensillar chaetotaxy: 1,0/0,0,1 ms freely exposed on the body surface. Th II-Abd V with 3,3/2,2,2,2,4 sensilla slender and shorter than surrounding chaetae. The lower pair of sensilla on Abd V is shorter and thicker than the upper pair, set in front of p-row (
<figureCitation id="13042A5EA8249210FABCF899FA34F8AE" box="[1360,1422,1862,1887]" captionStart="FIGURES 5 – 8" captionStartId="6.[151,264,1888,1911]" captionTargetBox="[184,1399,207,1855]" captionTargetId="figure@6.[174,1413,193,1867]" captionTargetPageId="6" captionText="FIGURES 5 – 8. Folsomides spp.: 5, habitus and chaetotaxy in F. v i no s us, paratype from Canary Is.; 6 – 8, chaetotaxy of Abd IV – VI of F. halshinicus sp. nov. (6), F. subvinosus sp. nov. (7) and F. marchicus (8). Scales: 0.1 mm." httpUri="https://zenodo.org/record/241487/files/figure.png" pageId="2" pageNumber="283">Fig 6</figureCitation>
).
</paragraph>
<paragraph id="8B8036DBA8249210FF2BF8B5FCE6F81E" blockId="2.[151,1437,890,2031]" pageId="2" pageNumber="283">
Chaetotaxy of legs I to III: subcoxae I: 1,1,3; subcoxae II: 1,5,5; coxae: 4,8,9; trochanter: 8,8,6; femora: 13,14,14; tibiotarsi: 20,20,22 pointed chaetae respectively. Chaetae
<emphasis id="B94BEAC9A8249210FC5BF850FBB9F857" box="[951,1027,1935,1959]" italics="true" pageId="2" pageNumber="283">A1–A7</emphasis>
,
<emphasis id="B94BEAC9A8249210FBFBF850FBDEF857" box="[1047,1124,1935,1959]" italics="true" pageId="2" pageNumber="283">B1–B7</emphasis>
,
<emphasis id="B94BEAC9A8249210FB9BF850FB7CF857" box="[1143,1222,1935,1959]" italics="true" pageId="2" pageNumber="283">C1–C7</emphasis>
and
<emphasis id="B94BEAC9A8249210FAE5F850FAAFF857" box="[1289,1301,1935,1958]" italics="true" pageId="2" pageNumber="283">x</emphasis>
present on tibiotarsus of leg III. Tibiotarsi of legs I and II with single chaeta
<emphasis id="B94BEAC9A8249210FC82F86CFC11F83A" box="[878,939,1971,1995]" italics="true" pageId="2" pageNumber="283">B4/5.</emphasis>
Claw with no teeth on inner edge. Empodial appendage sharp; its length less than half internal edge of claw.
</paragraph>
<caption id="DF406653A8259211FF7BF879FE7DF82D" httpUri="https://zenodo.org/record/241485/files/figure.png" pageId="3" pageNumber="284" targetBox="[268,1312,208,1919]" targetPageId="3">
<paragraph id="8B8036DBA8259211FF7BF879FE7DF82D" blockId="3.[151,1435,1958,2012]" pageId="3" pageNumber="284">
<emphasis id="B94BEAC9A8259211FF7BF879FE81F84D" bold="true" box="[151,315,1958,1981]" pageId="3" pageNumber="284">FIGURES 1–2.</emphasis>
<taxonomicName id="4C3F4D58A8259211FEAEF878FD88F84C" box="[322,562,1959,1981]" class="Entognatha" family="Isotomidae" genus="Folsomides" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="284" phylum="Arthropoda" rank="species" species="halshinicus" status="sp. nov.">
<emphasis id="B94BEAC9A8259211FEAEF878FD88F84C" box="[322,562,1959,1981]" italics="true" pageId="3" pageNumber="284">Folsomides halshinicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857B2A8259211FDD4F878FD30F84D" box="[568,650,1959,1980]" pageId="3" pageNumber="284" rank="species">
<emphasis id="B94BEAC9A8259211FDD4F878FD30F84D" bold="true" box="[568,650,1959,1980]" pageId="3" pageNumber="284">sp. nov.</emphasis>
</taxonomicNameLabel>
: 1, habitus and chaetotaxy, lateral; 2, habitus and distribution of sensilla and macrochaetae. Scale: 0.2 mm.
</paragraph>
</caption>
<caption id="DF406653A8229216FF7BFF47FB17FF5E" box="[151,1197,151,175]" pageId="4" pageNumber="285">
<paragraph id="8B8036DBA8229216FF7BFF47FB17FF5E" blockId="4.[151,1197,151,175]" box="[151,1197,151,175]" pageId="4" pageNumber="285">
<emphasis id="B94BEAC9A8229216FF7BFF47FEB3FF5E" bold="true" box="[151,265,152,175]" pageId="4" pageNumber="285">TABLE 2.</emphasis>
Length of macrochaetae and ordinary chaetae in p-row in
<taxonomicName id="4C3F4D58A8229216FC97FF47FBB4FF5E" box="[891,1038,151,175]" class="Entognatha" family="Isotomidae" genus="Folsomides" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="halshinicus" status="sp. nov.">
<emphasis id="B94BEAC9A8229216FC97FF47FBB4FF5E" box="[891,1038,151,175]" italics="true" pageId="4" pageNumber="285">F. halshinicus</emphasis>
</taxonomicName>
<emphasis id="B94BEAC9A8229216FBF9FF47FBD0FF5E" bold="true" box="[1045,1130,152,175]" pageId="4" pageNumber="285">
<taxonomicNameLabel id="A27857B2A8229216FBF9FF47FBD0FF5E" box="[1045,1130,152,175]" pageId="4" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(µm).
</paragraph>
</caption>
<paragraph id="8B8036DBA8229216FE4CFF15FB4BFEFB" blockId="4.[159,1280,202,555]" pageId="4" pageNumber="285">Macrochaetae Ordinary L M SA chaetae</paragraph>
<paragraph id="8B8036DBA8229216FF73FEC2FB04FEC2" blockId="4.[159,1280,202,555]" box="[159,1214,285,307]" pageId="4" pageNumber="285">
Th
<date id="FF81101BA8229216FF2DFEC1FE00FEC2" box="[193,442,285,307]" pageId="4" pageNumber="285" value="1927-02">II 27</date>
̶ ̶ 18
</paragraph>
<paragraph id="8B8036DBA8229216FF73FE99FB04FEAD" blockId="4.[159,1280,202,555]" box="[159,1214,326,348]" pageId="4" pageNumber="285">Th III 28 ̶ ̶ 18</paragraph>
<paragraph id="8B8036DBA8229216FF73FEAFFB04FE77" blockId="4.[159,1280,202,555]" box="[159,1214,368,390]" pageId="4" pageNumber="285">
Abd
<date id="FF81101BA8229216FF3DFEAEFD01FE77" box="[209,699,368,390]" pageId="4" pageNumber="285" value="1924-01-31">I 31 24</date>
(15) 15
</paragraph>
<paragraph id="8B8036DBA8229216FF73FE46FB04FE5E" blockId="4.[159,1280,202,555]" box="[159,1214,409,431]" pageId="4" pageNumber="285">
Abd
<date id="FF81101BA8229216FF3DFE45FD01FE5E" box="[209,699,409,431]" pageId="4" pageNumber="285" value="1933-02-25">II 33 25</date>
(16) 16
</paragraph>
<paragraph id="8B8036DBA8229216FF73FE1DFB04FE29" blockId="4.[159,1280,202,555]" box="[159,1214,450,472]" pageId="4" pageNumber="285">
Abd
<date id="FF81101BA8229216FF3DFE1CFD01FE29" box="[209,699,450,472]" pageId="4" pageNumber="285" value="1937-03-30">III 37 30</date>
24 19
</paragraph>
<paragraph id="8B8036DBA8229216FF73FE33FB04FDF3" blockId="4.[159,1280,202,555]" box="[159,1214,492,514]" pageId="4" pageNumber="285">
Abd
<date id="FF81101BA8229216FF3DFE32FE00FDF3" box="[209,442,492,514]" pageId="4" pageNumber="285" value="1933-04">IV 33</date>
33 31 21
</paragraph>
<paragraph id="8B8036DBA8229216FF73FDCAFD63FCE9" blockId="4.[159,1280,202,555]" lastBlockId="4.[151,1437,623,1117]" pageId="4" pageNumber="285">
Abd
<date id="FF81101BA8229216FF3DFDC9FD01FDDA" box="[209,699,533,555]" pageId="4" pageNumber="285" value="1933-05-31">V 31 33</date>
36 19 Ventral chaetotaxy represented in
<figureCitation id="13042A5EA8229216FD89FDB0FD75FD79" box="[613,719,623,648]" captionStart="FIGURES 1 – 2" captionStartId="3.[151,264,1958,1981]" captionTargetBox="[268,1312,208,1919]" captionTargetId="figure@3.[254,1333,195,1935]" captionTargetPageId="3" captionText="FIGURES 1 – 2. Folsomides halshinicus sp. nov.: 1, habitus and chaetotaxy, lateral; 2, habitus and distribution of sensilla and macrochaetae. Scale: 0.2 mm." httpUri="https://zenodo.org/record/241485/files/figure.png" pageId="4" pageNumber="285">Figure 2</figureCitation>
. Head with 3+3 postlabial chaetae. Ventral tube with 3+3 laterodistal chaetae and 2 posterobasal ones. The retinacle with 3+3 teeth and one chaeta on corpus. Subcoxa of furca with 5–7 chaetae on anterior and 4(5) on posterior part. Furca well developed. Manubrium with 8+8 posterior and no anterior chaetae. Dens with two posterior and no anterior chaetae (
<figureCitation id="13042A5EA8229216FC3BFD03FB9EFD05" box="[983,1060,732,757]" captionStart="FIGURES 9 – 14" captionStartId="7.[151,264,1144,1167]" captionTargetBox="[242,1341,206,1116]" captionTargetId="figure@7.[232,1355,193,1124]" captionTargetPageId="7" captionText="FIGURES 9 – 14. Folsomides spp.: 9 – 11, postantennal organ and omma in F. halshinicus sp. nov. (9), F. subvinosus sp. nov. (10) and F. marchicus (11); 12 – 14, mucrodens in F. subvinosus sp. nov. (12), F. halshinicus sp. nov. (13) and F. marchicus (14)." httpUri="https://zenodo.org/record/241488/files/figure.png" pageId="4" pageNumber="285">Fig 13</figureCitation>
). The mucro with two teeth, with a small lateral lamella from base to subapical tooth.
</paragraph>
</subSubSection>
<subSubSection id="C3256550A8229216FF2BFCFBFBBCFBC9" pageId="4" pageNumber="285" type="discussion">
<paragraph id="8B8036DBA8229216FF2BFCFBFBBCFBC9" blockId="4.[151,1437,623,1117]" pageId="4" pageNumber="285">
<emphasis id="B94BEAC9A8229216FF2BFCFBFEF6FCCC" bold="true" box="[199,332,804,829]" pageId="4" pageNumber="285">Discussion.</emphasis>
Following
<bibRefCitation id="EFAE4B2AA8229216FE3BFCFBFD2EFCCD" author="Fjellberg" box="[471,660,804,829]" pageId="4" pageNumber="285" refString="Fjellberg, A. (1993) Revision of European and North African Folsomides Stach with special emphasis on the Canarian fauna (Collembola: Isotomidae). Entomologica Scandinavica, 23, 453 - 473." type="journal article" year="1993">Fjellberg (1993)</bibRefCitation>
,
<taxonomicName id="4C3F4D58A8229216FD49FCFBFC0BFCCD" box="[677,945,804,828]" class="Entognatha" family="Isotomidae" genus="Folsomides" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="halshinicus" status="sp. nov.">
<emphasis id="B94BEAC9A8229216FD49FCFBFC0BFCCD" box="[677,945,804,828]" italics="true" pageId="4" pageNumber="285">Folsomides halshinicus</emphasis>
</taxonomicName>
<emphasis id="B94BEAC9A8229216FC51FCFBFBA6FCCC" bold="true" box="[957,1052,804,829]" pageId="4" pageNumber="285">
<taxonomicNameLabel id="A27857B2A8229216FC51FCFBFBA6FCCC" box="[957,1052,804,829]" pageId="4" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the species group of
<taxonomicName id="4C3F4D58A8229216FF7BFC97FDF2FC91" authority="Axelson, 1905" authorityName="Axelson" authorityYear="1905" box="[151,584,839,864]" class="Entognatha" family="Isotomidae" genus="Folsomides" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B94BEAC9A8229216FF7BFC97FE30FC91" box="[151,394,840,864]" italics="true" pageId="4" pageNumber="285">Folsomides angularis</emphasis>
(
<bibRefCitation id="EFAE4B2AA8229216FE76FC98FDFAFC91" author="Axelson" box="[410,576,839,864]" pageId="4" pageNumber="285" refString="Axelson, W. M. (1905) Einige neue Collembolen aus Finland. Zoologischer Anzeiger, 28, 788 - 794." type="journal article" year="1905">Axelson, 1905</bibRefCitation>
)
</taxonomicName>
as it has 5+5 omma and dens with mucro. According to reduced number of ms on Th II-Abd III (10/001) this species is related to the subgroup formed by 13 species. Main differences between
<taxonomicName id="4C3F4D58A8229216FF17FC4EFE2DFC59" box="[251,407,912,936]" class="Entognatha" family="Isotomidae" genus="Folsomides" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="halshinicus" status="sp. nov.">
<emphasis id="B94BEAC9A8229216FF17FC4EFE2DFC59" box="[251,407,912,936]" italics="true" pageId="4" pageNumber="285">F. halshinicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857B2A8229216FE71FC50FE4DFC59" box="[413,503,911,936]" pageId="4" pageNumber="285" rank="species">
<emphasis id="B94BEAC9A8229216FE71FC50FE4DFC59" bold="true" box="[413,503,911,936]" pageId="4" pageNumber="285">sp. nov.</emphasis>
</taxonomicNameLabel>
and these species are summarized in
<tableCitation id="C6BD0360A8229216FC71FC50FC49FC59" box="[925,1011,911,936]" captionStart="TABLE 4" captionStartId="8.[151,239,152,175]" captionTargetBox="[151,1402,306,883]" captionTargetPageId="8" captionText="TABLE 4. Comparison of the two new Iranian species of Folsomides with the species of F. angularis group, with reduced number of microsensilla on tergites." httpUri="http://table.plazi.org/id/DF406653A82E921AFF7BFF47FDC8FF3E" pageId="4" pageNumber="285" tableUuid="DF406653A82E921AFF7BFF47FDC8FF3E">Table 4</tableCitation>
. The new species is most similar to
<taxonomicName id="4C3F4D58A8229216FA6BFC4EFE8CFC3D" class="Entognatha" family="Isotomidae" genus="Folsomides" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="chichinautzini">
<emphasis id="B94BEAC9A8229216FA6BFC4EFE8CFC3D" italics="true" pageId="4" pageNumber="285">F. chichinautzini</emphasis>
</taxonomicName>
. They differ in the body length and the dorsal chaetotaxy; 6 axial chaetae on Abd IV in
<taxonomicName id="4C3F4D58A8229216FA6BFC6AFE8CFC01" class="Entognatha" family="Isotomidae" genus="Folsomides" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="chichinautzini">
<emphasis id="B94BEAC9A8229216FA6BFC6AFE8CFC01" italics="true" pageId="4" pageNumber="285">F. chichinautzini</emphasis>
</taxonomicName>
vs
<quantity id="4CC79B3EA8229216FE8CFC08FE36FC01" box="[352,396,983,1008]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="285" unit="in" value="5.0">5 in</quantity>
the new species; 12233 macrochaetae on Abd I–V in
<taxonomicName id="4C3F4D58A8229216FC16FC06FB0CFC01" box="[1018,1206,984,1008]" class="Entognatha" family="Isotomidae" genus="Folsomides" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="chichinautzini">
<emphasis id="B94BEAC9A8229216FC16FC06FB0CFC01" box="[1018,1206,984,1008]" italics="true" pageId="4" pageNumber="285">F. chichinautzini</emphasis>
</taxonomicName>
(
<typeStatus id="54848879A8229216FB2AFC07FB42FC01" box="[1222,1272,984,1008]" pageId="4" pageNumber="285">type</typeStatus>
2 of
<bibRefCitation id="EFAE4B2AA8229216FADBFC08FF68FBE5" author="Fjellberg" pageId="4" pageNumber="285" refString="Fjellberg, A. (1993) Revision of European and North African Folsomides Stach with special emphasis on the Canarian fauna (Collembola: Isotomidae). Entomologica Scandinavica, 23, 453 - 473." type="journal article" year="1993">Fjellberg 1993</bibRefCitation>
, named
<typeStatus id="54848879A8229216FEDDFC22FEDBFBE4" box="[305,353,1021,1045]" pageId="4" pageNumber="285">type</typeStatus>
2b by us) vs
<quantity id="4CC79B3EA8229216FE10FC23FDDAFBE4" box="[508,608,1020,1045]" metricMagnitude="2" metricUnit="m" metricValue="5.672582" pageId="4" pageNumber="285" unit="in" value="22333.0">22333 in</quantity>
the new species (similar to
<typeStatus id="54848879A8229216FC73FC22FC75FBE4" box="[927,975,1021,1045]" pageId="4" pageNumber="285">type</typeStatus>
2 of
<bibRefCitation id="EFAE4B2AA8229216FBE1FC23FB0BFBE5" author="Fjellberg" box="[1037,1201,1020,1045]" pageId="4" pageNumber="285" refString="Fjellberg, A. (1993) Revision of European and North African Folsomides Stach with special emphasis on the Canarian fauna (Collembola: Isotomidae). Entomologica Scandinavica, 23, 453 - 473." type="journal article" year="1993">Fjellberg 1993</bibRefCitation>
but M on Abd I and SA on Abd III slightly longer than surrounding chaetae, named
<typeStatus id="54848879A8229216FC8DFBFFFC2BFBC9" box="[865,913,1056,1080]" pageId="4" pageNumber="285">type</typeStatus>
2a by us).
</paragraph>
</subSubSection>
<subSubSection id="C3256550A8229216FF2BFB9BFC37FBAC" box="[199,909,1092,1117]" pageId="4" pageNumber="285" type="etymology">
<paragraph id="8B8036DBA8229216FF2BFB9BFC37FBAC" blockId="4.[151,1437,623,1117]" box="[199,909,1092,1117]" pageId="4" pageNumber="285">
<emphasis id="B94BEAC9A8229216FF2BFB9BFEF4FBAC" bold="true" box="[199,334,1092,1117]" pageId="4" pageNumber="285">Etymology.</emphasis>
The species has been named after the
<typeStatus id="54848879A8229216FD10FB9AFC96FBAC" box="[764,812,1093,1117]" pageId="4" pageNumber="285">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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