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<document id="E8D8963503C0DE0A92C3B07A7AC977F9" ID-DOI="10.5281/zenodo.194895" ID-GBIF-Dataset="342f6766-55cc-413b-a7e1-e64b5002e1d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="194895" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460219263375" checkinUser="plazi" docAuthor="Gonçalves, Inês C., Da-Silva, Elidiomar R. &amp; Nessimian, Jorge L." docDate="2010" docId="039687B7FA5AFFD6FF03E2315EFB1C5B" docLanguage="en" docName="zt02438p068.pdf" docOrigin="Zootaxa 2438" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Thraulodes jones Gonçalves, Da-Silva &amp; Nessimian, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="67" masterDocId="FFAFFFCFFA5BFFD0FF94E01E5A5E1A2B" masterDocTitle="A new species of Thraulodes Ulmer (Ephemeroptera: Leptophlebiidae) from Southeastern Brazil" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698237393201" updateUser="plazi">
<mods:mods id="B9F9E33FE32768A88CC3A21DA9915EFD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="223830C64CEA4F7D67FF886B47D43B5A">
<mods:title id="E474BE5BD7464C1E1F22A533AB548EE2">A new species of Thraulodes Ulmer (Ephemeroptera: Leptophlebiidae) from Southeastern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="0B0A10296A3BE551C0FC06203FA929B8" type="personal">
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<mods:namePart id="C1B0330E9351100A0C9B6FDC360AA241">Gonçalves, Inês C.</mods:namePart>
</mods:name>
<mods:name id="CB80D1F9E9B21200AF02EDE66B9841AD" type="personal">
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<mods:namePart id="08CA1FAC97C28692D38B0BB9A8D3C7C3">Da-Silva, Elidiomar R.</mods:namePart>
</mods:name>
<mods:name id="D73D6AFDB60DBDFE6EB0FF9BF1AC57AF" type="personal">
<mods:role id="A545888F8F6B86A41E4CA6B491871837">
<mods:roleTerm id="FBBD88E870DCA57AD83D119D8C8B4C1A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="79CE084EB4C4107E8A6CC323FC1712BA">Nessimian, Jorge L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F317E0FF55F9D4ADD22BC6666F904658">text</mods:typeOfResource>
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<mods:title id="912F9B5491FEBB7EDA0B48460284A91A">Zootaxa</mods:title>
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<mods:part id="B5BACB1D42EFAF77ABC7B08CCB32DD8A">
<mods:date id="C12A0CDBD16C5A9696E93FB8F8E90743">2010</mods:date>
<mods:detail id="B27AB6E61F81ABDE7EC4F4032BB92306" type="volume">
<mods:number id="6163B3F7F2B7040896C70EC785D9817F">2438</mods:number>
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<mods:identifier id="495410BDE1481EEE7761AC51F1EDC22B" type="DOI">10.5281/zenodo.194895</mods:identifier>
<mods:identifier id="ED0C7CBD469AA7F786D146E587DB6B76" type="GBIF-Dataset">342f6766-55cc-413b-a7e1-e64b5002e1d1</mods:identifier>
<mods:identifier id="8D6B919185573E14D0E6A2ECFDAB5121" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D0C229B079ECC3C99D33D72A654FF912" type="Zenodo-Dep">194895</mods:identifier>
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<treatment id="039687B7FA5AFFD6FF03E2315EFB1C5B" ID-DOI="http://doi.org/10.5281/zenodo.6200175" ID-GBIF-Taxon="119384994" ID-Zenodo-Dep="6200175" LSID="urn:lsid:plazi:treatment:039687B7FA5AFFD6FF03E2315EFB1C5B" httpUri="http://treatment.plazi.org/id/039687B7FA5AFFD6FF03E2315EFB1C5B" lastPageId="6" lastPageNumber="67" pageId="1" pageNumber="62">
<subSubSection id="C325652AFA5AFFD1FF03E23159291862" box="[151,887,559,585]" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="8B8036A1FA5AFFD1FF03E23159291862" blockId="1.[151,887,559,585]" box="[151,887,559,585]" pageId="1" pageNumber="62">
<heading id="D0C881CDFA5AFFD1FF03E23159291862" bold="true" box="[151,887,559,585]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
<emphasis id="B94BEAB3FA5AFFD1FF03E23159291862" bold="true" box="[151,887,559,585]" pageId="1" pageNumber="62">
<taxonomicName id="4C3F4D22FA5AFFD1FF03E2315B361862" ID-CoL="56LKH" box="[151,360,559,585]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="jones" status="sp. nov.">
<emphasis id="B94BEAB3FA5AFFD1FF03E2315B361862" bold="true" box="[151,360,559,585]" italics="true" pageId="1" pageNumber="62">Thraulodes jones</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857C8FA5AFFD1FEFBE2315B931862" box="[367,461,559,585]" pageId="1" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
Gonçalves, Da-Silva &amp; Nessimian
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C325652AFA5AFFD6FF03E26959221867" lastPageId="6" lastPageNumber="67" pageId="1" pageNumber="62" type="description">
<paragraph id="8B8036A1FA5AFFD1FF03E2695EE2189C" blockId="1.[151,1437,631,2002]" pageId="1" pageNumber="62">
<emphasis id="B94BEAB3FA5AFFD1FF03E2695BE018BA" box="[151,446,631,657]" italics="true" pageId="1" pageNumber="62">
<emphasis id="B94BEAB3FA5AFFD1FF03E2695B7218BA" bold="true" box="[151,300,631,657]" italics="true" pageId="1" pageNumber="62">Male Imago</emphasis>
(
<typeStatus id="54848803FA5AFFD1FEA9E2665BF018BA" box="[317,430,632,657]" pageId="1" pageNumber="62" type="holotype">Holotype</typeStatus>
):
</emphasis>
Length: body 12.0 mm; forewing
<quantity id="4CC79B44FA5AFFD1FCCEE266599918B9" box="[858,967,632,658]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="1" pageNumber="62" unit="mm" value="12.4">12.4 mm</quantity>
; hind wing 2.0 mm. General coloration light brown, with dark brown markings. First abdominal segments whitish. Wings hyaline.
</paragraph>
<paragraph id="8B8036A1FA5AFFD1FF50E2DC5ED7190C" blockId="1.[151,1437,631,2002]" pageId="1" pageNumber="62">
<emphasis id="B94BEAB3FA5AFFD1FF50E2DC5B5318F0" box="[196,269,706,731]" italics="true" pageId="1" pageNumber="62">Head:</emphasis>
Area between eyes and ocelli dark brown. Anterior margin of head yellowish, translucent. Ocelli white surrounded by black ring. Upper portion of turbinate eyes orange brown. Base of antenna surrounded by dark brown; scape with dark brown ring and pedicel dark brown; flagellum yellowish.
</paragraph>
<paragraph id="8B8036A1FA5AFFD1FF50E32C5F1A1997" blockId="1.[151,1437,631,2002]" pageId="1" pageNumber="62">
<emphasis id="B94BEAB3FA5AFFD1FF50E32C5B7D1960" box="[196,291,818,843]" italics="true" pageId="1" pageNumber="62">Thorax:</emphasis>
Pronotum yellowish bearing two median dark marks extending to posterior margin. General color of mesonotum and metanotum light brown. Posterior area of mesonotum yellowish, apex dark brown. Prosternum and mesosternum dark brown. Mesosternum with two median longitudinal light brown lines. Metasternum whitish with two dark brown markings. Thoracic pleura whitish with dark brown marks.
</paragraph>
<paragraph id="8B8036A1FA5AFFD1FF50E3D658FF1EB7" blockId="1.[151,1437,631,2002]" pageId="1" pageNumber="62">
<emphasis id="B94BEAB3FA5AFFD1FF50E3D65B5919CA" box="[196,263,968,993]" italics="true" pageId="1" pageNumber="62">Legs:</emphasis>
Femora of foreleg whitish with a blackish subtriangular basal mark; blackish median band, well marked centrally and inconspicuous toward margins; inner margin of femora with apical half blackish; apical margin of femora blackish; area between median band and apex orange brown. Tibia light orange brown, distal fourth blackish. Tarsus brown with base and apex whitish; last tarsal segment and tarsal claws blackish (
<figureCitation id="13042A24FA5AFFD1FF34E4435AB81E5C" box="[160,230,1117,1143]" captionStart="FIGURES 1 5" captionStartId="2.[151,269,1744,1768]" captionTargetBox="[159,1434,211,1722]" captionTargetId="figure@2.[151,1436,194,1723]" captionTargetPageId="2" captionText="FIGURES 1 5. Thraulodes jones sp. nov. Male imago. 1. Fore leg. 2. Forewing. 3. Hind wing. 4 — Abdominal terga II— X. 5. Genitalia (scale: 1.0 mm)." httpUri="https://zenodo.org/record/194896/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
). Median and hind legs similar to foreleg in color, except for lack of basal blackish mark on femora; tarsi yellowish, with last tarsomere blackish.
</paragraph>
<paragraph id="8B8036A1FA5AFFD1FF50E4B65B881F27" blockId="1.[151,1437,631,2002]" pageId="1" pageNumber="62">
<emphasis id="B94BEAB3FA5AFFD1FF50E4B65B481EEA" box="[196,278,1192,1217]" italics="true" pageId="1" pageNumber="62">Wings:</emphasis>
Membrane of wings hyaline with dark brown stain at base (
<figureCitation id="13042A24FA5AFFD1FC74E4B65E0D1EE9" box="[992,1107,1192,1218]" captionStart="FIGURES 1 5" captionStartId="2.[151,269,1744,1768]" captionTargetBox="[159,1434,211,1722]" captionTargetId="figure@2.[151,1436,194,1723]" captionTargetPageId="2" captionText="FIGURES 1 5. Thraulodes jones sp. nov. Male imago. 1. Fore leg. 2. Forewing. 3. Hind wing. 4 — Abdominal terga II— X. 5. Genitalia (scale: 1.0 mm)." httpUri="https://zenodo.org/record/194896/files/figure.png" pageId="1" pageNumber="62">Figs. 23</figureCitation>
). Forewing veins brownish, hind wing veins whitish. Area between C, Sc and R1 of forewing whitish toward apex. Five basal and 1718 cross-veins distal to bullae.
</paragraph>
<paragraph id="8B8036A1FA5AFFD1FF50E5065B191FC7" blockId="1.[151,1437,631,2002]" pageId="1" pageNumber="62">
<emphasis id="B94BEAB3FA5AFFD1FF50E5065B1F1F1A" box="[196,321,1304,1329]" italics="true" pageId="1" pageNumber="62">Abdomen:</emphasis>
Tergum I whitish with wide anteromedian blackish mark. Terga IIVI whitish translucent bearing 3 pairs of sub apical marks: median, sublateral and lateral. Terga VIIX orange brown. Terga VIIIX possessing one lateral sub apical blackish mark; anterior and posterior margins dark brown, except for anterior margin of tergum IX. All terga possessing a blackish dash on lateral margins (
<figureCitation id="13042A24FA5AFFD1FBEAE5965E931F89" box="[1150,1229,1416,1442]" captionStart="FIGURES 1 5" captionStartId="2.[151,269,1744,1768]" captionTargetBox="[159,1434,211,1722]" captionTargetId="figure@2.[151,1436,194,1723]" captionTargetPageId="2" captionText="FIGURES 1 5. Thraulodes jones sp. nov. Male imago. 1. Fore leg. 2. Forewing. 3. Hind wing. 4 — Abdominal terga II— X. 5. Genitalia (scale: 1.0 mm)." httpUri="https://zenodo.org/record/194896/files/figure.png" pageId="1" pageNumber="62">Fig. 4</figureCitation>
). Sterna whitish translucent. Sterna IVII with pair of median, sub apical and apical-lateral blackish spots. Sterna VIIIIX without marks.
</paragraph>
<paragraph id="8B8036A1FA5AFFD1FF50E5E65B421CE7" blockId="1.[151,1437,631,2002]" pageId="1" pageNumber="62">
<emphasis id="B94BEAB3FA5AFFD1FF50E5E65B611C3A" box="[196,319,1528,1553]" italics="true" pageId="1" pageNumber="62">Genitalia:</emphasis>
Styliger plate whitish bearing very subtle grooves, posterior median projection present with rounded apex. Forceps whitish, first segment with distal half washed with gray, second segment with rounded internal distal projection. Penes long and slender, without lateral pouch. Inner margin sclerotized; apicolateral area with acute projection directed externally and posterioly; subapical spines long and projected internally and posteriorly (
<figureCitation id="13042A24FA5AFFD1FEF2E6935BED1C8C" box="[358,435,1677,1703]" captionStart="FIGURES 1 5" captionStartId="2.[151,269,1744,1768]" captionTargetBox="[159,1434,211,1722]" captionTargetId="figure@2.[151,1436,194,1723]" captionTargetPageId="2" captionText="FIGURES 1 5. Thraulodes jones sp. nov. Male imago. 1. Fore leg. 2. Forewing. 3. Hind wing. 4 — Abdominal terga II— X. 5. Genitalia (scale: 1.0 mm)." httpUri="https://zenodo.org/record/194896/files/figure.png" pageId="1" pageNumber="62">Fig. 5</figureCitation>
). Caudal filaments whitish, with wide and narrow dark brown bands irregularly distributed.
</paragraph>
<paragraph id="8B8036A1FA5AFFD1FF50E6C65B901DAC" blockId="1.[151,1437,631,2002]" pageId="1" pageNumber="62">
<emphasis id="B94BEAB3FA5AFFD1FF50E6C65B161CDA" box="[196,328,1752,1777]" italics="true" pageId="1" pageNumber="62">Variations:</emphasis>
Body measurements were smaller in some specimens, about 10.0 mm body length; 11.0 mm forewing; and 2.0 mm hind wing;
<typeStatus id="54848803FA5AFFD1FDA7E6E358F91D3C" box="[563,679,1789,1815]" pageId="1" pageNumber="62" type="paratype">Paratypes</typeStatus>
showed scape whitish and flagella shaded with dark brown. In one male
<typeStatus id="54848803FA5AFFD1FE9BE73C5B251D17" box="[271,379,1826,1852]" pageId="1" pageNumber="62" type="paratype">paratype</typeStatus>
, the triangular mark at base of fore femora was absent and only 4 cross-veins basal to bullae were found. The dashes on lateral margins of abdominal terga were less conspicuous at first abdominal segments on one
<typeStatus id="54848803FA5AFFD1FEF5E7735B941DAC" box="[353,458,1901,1927]" pageId="1" pageNumber="62" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="8B8036A1FA5AFFD1FF52E78C59A61DF9" blockId="1.[151,1437,631,2002]" pageId="1" pageNumber="62">
<emphasis id="B94BEAB3FA5AFFD1FF52E78C5BDD1D87" box="[198,387,1938,1964]" italics="true" pageId="1" pageNumber="62">
<emphasis id="B94BEAB3FA5AFFD1FF52E78C5B271D87" bold="true" box="[198,377,1938,1964]" italics="true" pageId="1" pageNumber="62">Female Imago</emphasis>
:
</emphasis>
Length: body 10.011.0 mm; forewing
<quantity id="4CC79B44FA5AFFD1FCF7E78C5E4F1D87" box="[867,1041,1938,1964]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" metricValueMax="1.27" metricValueMin="1.15" pageId="1" pageNumber="62" unit="mm" value="12.1" valueMax="12.7" valueMin="11.5">11.512.7 mm</quantity>
; hind wing 1.42.0 mm. General coloration light brown, abdomen shaded with dark brown. Wings hyaline.
</paragraph>
<caption id="DF406629FA59FFD2FF03E6CE5BB71D23" httpUri="https://zenodo.org/record/194896/files/figure.png" pageId="2" pageNumber="63" targetBox="[159,1434,211,1722]" targetPageId="2">
<paragraph id="8B8036A1FA59FFD2FF03E6CE5BB71D23" blockId="2.[151,1436,1744,1800]" pageId="2" pageNumber="63">
<emphasis id="B94BEAB3FA59FFD2FF03E6CE5B1C1CC3" bold="true" box="[151,322,1744,1768]" pageId="2" pageNumber="63">FIGURES 15.</emphasis>
<taxonomicName id="4C3F4D22FA59FFD2FEDCE6CF5BA11CC3" box="[328,511,1745,1768]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="jones" status="sp. nov.">
<emphasis id="B94BEAB3FA59FFD2FEDCE6CF5BA11CC3" box="[328,511,1745,1768]" italics="true" pageId="2" pageNumber="63">Thraulodes jones</emphasis>
</taxonomicName>
<emphasis id="B94BEAB3FA59FFD2FD92E6CE58041CC3" bold="true" box="[518,602,1744,1768]" pageId="2" pageNumber="63">
<taxonomicNameLabel id="A27857C8FA59FFD2FD92E6CE58041CC3" box="[518,602,1744,1768]" pageId="2" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male imago. 1. Fore leg. 2. Forewing. 3. Hind wing. 4—Abdominal terga II— X. 5. Genitalia (scale: 1.0 mm).
</paragraph>
</caption>
<paragraph id="8B8036A1FA59FFD2FF50E72F5B7B1DBE" blockId="2.[151,1437,1841,2016]" pageId="2" pageNumber="63">
<emphasis id="B94BEAB3FA59FFD2FF50E72F5B511D61" box="[196,271,1841,1866]" italics="true" pageId="2" pageNumber="63">Head:</emphasis>
Area between eyes and ocelli shaded with dark brown. Eyes black. Ocelli white surrounded by black. Anterior margin of head yellowish. Scape yellowish, pedicel dark brown and flagellum shaded with dark brown.
</paragraph>
<paragraph id="8B8036A1FA59FFD3FF50E7BF58D51AFC" blockId="2.[151,1437,1841,2016]" lastBlockId="3.[151,1436,152,327]" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
<emphasis id="B94BEAB3FA59FFD2FF50E7BF5B791D91" box="[196,295,1953,1978]" italics="true" pageId="2" pageNumber="63">Thorax:</emphasis>
Pronotum yellowish; anterior and posterior margins of pronotum dark brown; median mark shaded with dark brown from which 2 pairs of marks arise, one on each side of pronotum. General color of mesonotum and metanotum as in male. Prosternum and metasternum whitish. Mesosternum dark brown with two median longitudinal light brown lines.
</paragraph>
<paragraph id="8B8036A1FA58FFD3FF50E0FC585A1AD7" blockId="3.[151,1436,152,327]" box="[196,516,226,252]" pageId="3" pageNumber="64">
<emphasis id="B94BEAB3FA58FFD3FF50E0FC5B5B1AD0" box="[196,261,226,251]" italics="true" pageId="3" pageNumber="64">Legs:</emphasis>
Legs similar to male.
</paragraph>
<paragraph id="8B8036A1FA58FFD3FF50E11658741B6C" blockId="3.[151,1436,152,327]" pageId="3" pageNumber="64">
<emphasis id="B94BEAB3FA58FFD3FF50E1165B4B1B0A" box="[196,277,264,289]" italics="true" pageId="3" pageNumber="64">Wings:</emphasis>
Wings similar to male, except for having 34 cross-veins basal to bullae and C, Sc and R1 area of forewing not whitish toward apex.
</paragraph>
<caption id="DF406629FA58FFD3FF03E7445B1F1DF9" httpUri="https://zenodo.org/record/194897/files/figure.png" pageId="3" pageNumber="64" targetBox="[151,1435,399,1858]" targetPageId="3">
<paragraph id="8B8036A1FA58FFD3FF03E7445B1F1DF9" blockId="3.[151,1437,1882,2002]" pageId="3" pageNumber="64">
<emphasis id="B94BEAB3FA58FFD3FF03E7445B111D59" bold="true" box="[151,335,1882,1906]" pageId="3" pageNumber="64">FIGURES 614.</emphasis>
<taxonomicName id="4C3F4D22FA58FFD3FEC1E74558531D59" box="[341,525,1883,1906]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="jones" status="sp. nov.">
<emphasis id="B94BEAB3FA58FFD3FEC1E74558531D59" box="[341,525,1883,1906]" italics="true" pageId="3" pageNumber="64">Thraulodes jones</emphasis>
</taxonomicName>
<emphasis id="B94BEAB3FA58FFD3FD87E74458361D59" bold="true" box="[531,616,1882,1906]" pageId="3" pageNumber="64">
<taxonomicNameLabel id="A27857C8FA58FFD3FD87E74458361D59" box="[531,616,1882,1906]" pageId="3" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Nymph. 6. Labrum d.v. 7. Detail of denticles on labrum d.v. 8. Left mandible d.v. 9. Right mandible d.v. 10. Detail of right mandible d.v. 11. Detail of left mandible d.v. 12. Maxilla d.v. 13. Hypopharynx d.v. 14. Labium (left: v.v.; right: d.v.). d.v. - dorsal view, v.v—ventral view (scale: 0.5mm, except for Figs. 1011, 0.1mm).
</paragraph>
</caption>
<caption id="DF406629FA5FFFD4FF03E78958781DC5" httpUri="https://zenodo.org/record/194898/files/figure.png" pageId="4" pageNumber="65" targetBox="[189,1407,201,1919]" targetPageId="4">
<paragraph id="8B8036A1FA5FFFD4FF03E78958781DC5" blockId="4.[151,1436,1943,2030]" pageId="4" pageNumber="65">
<emphasis id="B94BEAB3FA5FFFD4FF03E7895B381D84" bold="true" box="[151,358,1943,1967]" pageId="4" pageNumber="65">FIGURES 1520.</emphasis>
<taxonomicName id="4C3F4D22FA5FFFD4FEF9E78958711D85" box="[365,559,1943,1966]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="jones" status="sp. nov.">
<emphasis id="B94BEAB3FA5FFFD4FEF9E78958711D85" box="[365,559,1943,1966]" italics="true" pageId="4" pageNumber="65">Thraulodes jones</emphasis>
</taxonomicName>
<emphasis id="B94BEAB3FA5FFFD4FDA3E78958CC1D84" bold="true" box="[567,658,1943,1967]" pageId="4" pageNumber="65">
<taxonomicNameLabel id="A27857C8FA5FFFD4FDA3E78958CC1D84" box="[567,658,1943,1967]" pageId="4" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Nymph. 15. Fore leg. 16. Tarsal claw. 17. Mid leg. 18. Hind leg. 19. Abdominal terga. 20. Gill. (scale: Figs. 15, 17, 18, 20, 1.0 mm; Fig. 16, 0.1 mm; Fig. 19, 2.5 mm)(Drawings of legs and abdomen from a non mature nymph).
</paragraph>
</caption>
<paragraph id="8B8036A1FA5EFFD5FF50E0865E071BB9" blockId="5.[151,1438,152,2007]" pageId="5" pageNumber="66">
<emphasis id="B94BEAB3FA5EFFD5FF50E0865B1C1A9A" box="[196,322,152,177]" italics="true" pageId="5" pageNumber="66">Abdomen:</emphasis>
Terga IIII shaded with dark brown; tergum IV also shaded except for yellowish median triangular mark extending from middle to posterior margin; terga VVI yellowish shaded with dark brown on anterior area; tergum VII yellowish with anterior and posterior margins shaded with dark brown; tergum VIII yellowish, shaded with dark brown on anterior margin; terga IXX yellowish; terga IIIVI with a pair of sub apical dark brown marks; all terga possessing a dark brown dash on lateral margins. Sterna IVI with a pair of apico-lateral dark brown marks, remaining sterna yellowish. Apex of female sternum cleft, V-shaped. Caudal filaments whitish, with wide and narrow dark brown bands irregularly distributed.
</paragraph>
<paragraph id="8B8036A1FA5EFFD5FF50E1835EAF180C" blockId="5.[151,1438,152,2007]" pageId="5" pageNumber="66">
<emphasis id="B94BEAB3FA5EFFD5FF50E1835B181B9D" box="[196,326,413,438]" italics="true" pageId="5" pageNumber="66">Variations:</emphasis>
Some females showed terga VIIIX more heavily shaded with dark brown. Females collected at Macaé de Cima, Nova Friburgo, were found to be slightly bigger: body 11.512.0 mm; forewing: 15.0
<quantity id="4CC79B44FA5EFFD5FAFDE1DC5A921829" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="5" pageNumber="66" unit="mm" value="16.2">16.2 mm</quantity>
; hind wing: 2.0
<quantity id="4CC79B44FA5EFFD5FE3DE1F658531829" box="[425,525,488,514]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="66" unit="mm" value="2.1">2.1 mm</quantity>
. They also showed a color pattern more intense than described above, prosternum with dark brown median mark and abdominal sterna with color pattern as in males.
</paragraph>
<paragraph id="8B8036A1FA5EFFD5FF50E22C58851859" blockId="5.[151,1438,152,2007]" pageId="5" pageNumber="66">
<emphasis id="B94BEAB3FA5EFFD5FF50E22C5BBD1860" box="[196,483,562,588]" italics="true" pageId="5" pageNumber="66">
<emphasis id="B94BEAB3FA5EFFD5FF50E22C5BDC1867" bold="true" box="[196,386,562,588]" italics="true" pageId="5" pageNumber="66">Mature Nymph</emphasis>
(male):
</emphasis>
Length: body: 10.011.0 mm; caudal filaments: more than 13.0 mm (broken). General color dark brown with light brown areas.
</paragraph>
<paragraph id="8B8036A1FA5EFFD5FF50E2635E8A1FA9" blockId="5.[151,1438,152,2007]" pageId="5" pageNumber="66">
<emphasis id="B94BEAB3FA5EFFD5FF50E2635B4F18BD" box="[196,273,637,662]" italics="true" pageId="5" pageNumber="66">Head:</emphasis>
Ocelli black; area surrounding ocelli light brown. Eyes black; turbinated portion dark brown reddish. Antennae about 2.62.7 times the size of the head, scape and pedicel dark brown, flagellum light brown. Labrum brown with basal lateral area light brown, wider than clypeus; its width about 3 times its length; presence of apical and sub-apical rows of whitish long setae (
<figureCitation id="13042A24FA5EFFD5FC4DE2F35E7C192C" box="[985,1058,749,775]" captionStart="FIGURES 6 14" captionStartId="3.[151,270,1882,1906]" captionTargetBox="[151,1435,399,1858]" captionTargetId="figure@3.[151,1436,383,1858]" captionTargetPageId="3" captionText="FIGURES 6 14. Thraulodes jones sp. nov. Nymph. 6. Labrum d. v. 7. Detail of denticles on labrum d. v. 8. Left mandible d. v. 9. Right mandible d. v. 10. Detail of right mandible d. v. 11. Detail of left mandible d. v. 12. Maxilla d. v. 13. Hypopharynx d. v. 14. Labium (left: v. v.; right: d. v.). d. v. - dorsal view, v. v—ventral view (scale: 0.5 mm, except for Figs. 10 11, 0.1 mm)." httpUri="https://zenodo.org/record/194897/files/figure.png" pageId="5" pageNumber="66">Fig. 6</figureCitation>
); anterior margin straight; with five very subtle emarginations (
<figureCitation id="13042A24FA5EFFD5FD9DE30C580E1907" box="[521,592,786,812]" captionStart="FIGURES 6 14" captionStartId="3.[151,270,1882,1906]" captionTargetBox="[151,1435,399,1858]" captionTargetId="figure@3.[151,1436,383,1858]" captionTargetPageId="3" captionText="FIGURES 6 14. Thraulodes jones sp. nov. Nymph. 6. Labrum d. v. 7. Detail of denticles on labrum d. v. 8. Left mandible d. v. 9. Right mandible d. v. 10. Detail of right mandible d. v. 11. Detail of left mandible d. v. 12. Maxilla d. v. 13. Hypopharynx d. v. 14. Labium (left: v. v.; right: d. v.). d. v. - dorsal view, v. v—ventral view (scale: 0.5 mm, except for Figs. 10 11, 0.1 mm)." httpUri="https://zenodo.org/record/194897/files/figure.png" pageId="5" pageNumber="66">Fig. 7</figureCitation>
); lateral margins rounded. Mandibles with row of whitish setae on outer margin extending to level of outer incisor; setae long on distal half and short on basal half; outer margin projected; inner margin of outer incisor and both margins of inner incisor serrate; molars with serrate ridges; prosthecae well developed. Incisors, outer margin and basal third of mandibles brown; apical third whitish; molars and prosthecae light brown (
<figureCitation id="13042A24FA5EFFD5FDA9E3B658F119E9" box="[573,687,936,962]" captionStart="FIGURES 6 14" captionStartId="3.[151,270,1882,1906]" captionTargetBox="[151,1435,399,1858]" captionTargetId="figure@3.[151,1436,383,1858]" captionTargetPageId="3" captionText="FIGURES 6 14. Thraulodes jones sp. nov. Nymph. 6. Labrum d. v. 7. Detail of denticles on labrum d. v. 8. Left mandible d. v. 9. Right mandible d. v. 10. Detail of right mandible d. v. 11. Detail of left mandible d. v. 12. Maxilla d. v. 13. Hypopharynx d. v. 14. Labium (left: v. v.; right: d. v.). d. v. - dorsal view, v. v—ventral view (scale: 0.5 mm, except for Figs. 10 11, 0.1 mm)." httpUri="https://zenodo.org/record/194897/files/figure.png" pageId="5" pageNumber="66">Figs. 89</figureCitation>
). Right mandible outer incisor with three apical denticles; inner incisor with two denticles (
<figureCitation id="13042A24FA5EFFD5FE48E3D3586819CC" box="[476,566,973,999]" captionStart="FIGURES 6 14" captionStartId="3.[151,270,1882,1906]" captionTargetBox="[151,1435,399,1858]" captionTargetId="figure@3.[151,1436,383,1858]" captionTargetPageId="3" captionText="FIGURES 6 14. Thraulodes jones sp. nov. Nymph. 6. Labrum d. v. 7. Detail of denticles on labrum d. v. 8. Left mandible d. v. 9. Right mandible d. v. 10. Detail of right mandible d. v. 11. Detail of left mandible d. v. 12. Maxilla d. v. 13. Hypopharynx d. v. 14. Labium (left: v. v.; right: d. v.). d. v. - dorsal view, v. v—ventral view (scale: 0.5 mm, except for Figs. 10 11, 0.1 mm)." httpUri="https://zenodo.org/record/194897/files/figure.png" pageId="5" pageNumber="66">Fig. 10</figureCitation>
); setae near molars light brown. Left mandible possessing 3 denticles on inner and outer incisors (
<figureCitation id="13042A24FA5EFFD5FE2DE3EC58501E27" box="[441,526,1010,1036]" captionStart="FIGURES 6 14" captionStartId="3.[151,270,1882,1906]" captionTargetBox="[151,1435,399,1858]" captionTargetId="figure@3.[151,1436,383,1858]" captionTargetPageId="3" captionText="FIGURES 6 14. Thraulodes jones sp. nov. Nymph. 6. Labrum d. v. 7. Detail of denticles on labrum d. v. 8. Left mandible d. v. 9. Right mandible d. v. 10. Detail of right mandible d. v. 11. Detail of left mandible d. v. 12. Maxilla d. v. 13. Hypopharynx d. v. 14. Labium (left: v. v.; right: d. v.). d. v. - dorsal view, v. v—ventral view (scale: 0.5 mm, except for Figs. 10 11, 0.1 mm)." httpUri="https://zenodo.org/record/194897/files/figure.png" pageId="5" pageNumber="66">Fig. 11</figureCitation>
). Maxilla whitish, flattened and rectangular; apical margin brown with row of apical long brown setae, simple and pectinated, and sub-apical interrupted row of shorter but thicker setae; inner margin bordered by long light brown setae, culminating on single pectinated setae; outer margin brown, dark brown basal to palpi insertion; posterior margin light brown, with light brown setae; palpi brown with outer margin bordered by long whitish setae, denser on segment III; inner margin of segment III and apical third of segment II with row of long setae (
<figureCitation id="13042A24FA5EFFD5FD19E4B358BA1EEC" box="[653,740,1197,1223]" captionStart="FIGURES 6 14" captionStartId="3.[151,270,1882,1906]" captionTargetBox="[151,1435,399,1858]" captionTargetId="figure@3.[151,1436,383,1858]" captionTargetPageId="3" captionText="FIGURES 6 14. Thraulodes jones sp. nov. Nymph. 6. Labrum d. v. 7. Detail of denticles on labrum d. v. 8. Left mandible d. v. 9. Right mandible d. v. 10. Detail of right mandible d. v. 11. Detail of left mandible d. v. 12. Maxilla d. v. 13. Hypopharynx d. v. 14. Labium (left: v. v.; right: d. v.). d. v. - dorsal view, v. v—ventral view (scale: 0.5 mm, except for Figs. 10 11, 0.1 mm)." httpUri="https://zenodo.org/record/194897/files/figure.png" pageId="5" pageNumber="66">Fig. 12</figureCitation>
). Hypopharynx whitish with light brown setae; superlingua bearing long setae on apical margin and at base (
<figureCitation id="13042A24FA5EFFD5FD7DE4CC591D1EC7" box="[745,835,1234,1260]" captionStart="FIGURES 6 14" captionStartId="3.[151,270,1882,1906]" captionTargetBox="[151,1435,399,1858]" captionTargetId="figure@3.[151,1436,383,1858]" captionTargetPageId="3" captionText="FIGURES 6 14. Thraulodes jones sp. nov. Nymph. 6. Labrum d. v. 7. Detail of denticles on labrum d. v. 8. Left mandible d. v. 9. Right mandible d. v. 10. Detail of right mandible d. v. 11. Detail of left mandible d. v. 12. Maxilla d. v. 13. Hypopharynx d. v. 14. Labium (left: v. v.; right: d. v.). d. v. - dorsal view, v. v—ventral view (scale: 0.5 mm, except for Figs. 10 11, 0.1 mm)." httpUri="https://zenodo.org/record/194897/files/figure.png" pageId="5" pageNumber="66">Fig. 13</figureCitation>
); apex of lingua ventrally covered by short setae; outer margin more densely setose so that the region appears to be somewhat darkened. Labium yellowish white; densely covered by long light brown setae on glossa and apical third of paraglossa; first segment of palpi with scattered whitish setae on inner and outer margins; second segment with denser setae at apical third, outer margin with long setae and inner margin bearing short, spine-like setae (
<figureCitation id="13042A24FA5EFFD5FBFBE5765E991FA9" box="[1135,1223,1384,1410]" captionStart="FIGURES 6 14" captionStartId="3.[151,270,1882,1906]" captionTargetBox="[151,1435,399,1858]" captionTargetId="figure@3.[151,1436,383,1858]" captionTargetPageId="3" captionText="FIGURES 6 14. Thraulodes jones sp. nov. Nymph. 6. Labrum d. v. 7. Detail of denticles on labrum d. v. 8. Left mandible d. v. 9. Right mandible d. v. 10. Detail of right mandible d. v. 11. Detail of left mandible d. v. 12. Maxilla d. v. 13. Hypopharynx d. v. 14. Labium (left: v. v.; right: d. v.). d. v. - dorsal view, v. v—ventral view (scale: 0.5 mm, except for Figs. 10 11, 0.1 mm)." httpUri="https://zenodo.org/record/194897/files/figure.png" pageId="5" pageNumber="66">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8B8036A1FA5EFFD5FF50E5935F571C3C" blockId="5.[151,1438,152,2007]" pageId="5" pageNumber="66">
<emphasis id="B94BEAB3FA5EFFD5FF50E5935B791F8D" box="[196,295,1421,1446]" italics="true" pageId="5" pageNumber="66">Thorax:</emphasis>
Pronotum brown with several scattered light brown spots; lateral margins light brown with sublateral dark brown stripe, posterior margin with light brown border not reaching lateral margins. General coloration of mesonotum dark brown, scattered with several brown spots; two brown longitudinal lines one on each side of median suture. Wing pads light brown, veins dark brown. Thoracic sterna yellowish.
</paragraph>
<paragraph id="8B8036A1FA5EFFD6FF50E63D59EC1AD7" blockId="5.[151,1438,152,2007]" lastBlockId="6.[151,1437,152,1648]" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66">
<emphasis id="B94BEAB3FA5EFFD5FF50E63D5B581C17" box="[196,262,1571,1596]" italics="true" pageId="5" pageNumber="66">Legs:</emphasis>
Femora of all legs brown with light brown marks, tibia, tarsi and claw light brown. Foreleg (
<figureCitation id="13042A24FA5EFFD5FAF8E63C5AE61C49" captionStart="FIGURES 15 20" captionStartId="4.[151,273,1943,1967]" captionTargetBox="[189,1407,201,1919]" captionTargetId="figure@4.[183,1407,194,1919]" captionTargetPageId="4" captionText="FIGURES 15 20. Thraulodes jones sp. nov. Nymph. 15. Fore leg. 16. Tarsal claw. 17. Mid leg. 18. Hind leg. 19. Abdominal terga. 20. Gill. (scale: Figs. 15, 17, 18, 20, 1.0 mm; Fig. 16, 0.1 mm; Fig. 19, 2.5 mm) (Drawings of legs and abdomen from a non mature nymph)." httpUri="https://zenodo.org/record/194898/files/figure.png" pageId="5" pageNumber="66">Fig. 15</figureCitation>
): coxa with wide dark brown margin, outer margin bordered by long simple setae; femora dark brown with one basal, one sub-median and one sub-apical light brown marks; basal mark lighter, with inner grayish spot; outer margin with row of simple long setae and fewer shorter clavate setae, inner margin with few long simple setae, not as long as those of outer margin; margins and dorsal area of femora presenting short bristles; outer margin and dorsal region of tibia with row of simple setae, not as long as those on femora; inner margin with row of spines; apical half of tibia and tarsus somewhat shaded with brown; tarsus bearing simple setae. Tarsal claws with seven denticles, apical denticle over twice longer than subapical denticle (
<figureCitation id="13042A24FA5EFFD5FB1BE7365EB91D69" box="[1167,1255,1832,1858]" captionStart="FIGURES 15 20" captionStartId="4.[151,273,1943,1967]" captionTargetBox="[189,1407,201,1919]" captionTargetId="figure@4.[183,1407,194,1919]" captionTargetPageId="4" captionText="FIGURES 15 20. Thraulodes jones sp. nov. Nymph. 15. Fore leg. 16. Tarsal claw. 17. Mid leg. 18. Hind leg. 19. Abdominal terga. 20. Gill. (scale: Figs. 15, 17, 18, 20, 1.0 mm; Fig. 16, 0.1 mm; Fig. 19, 2.5 mm) (Drawings of legs and abdomen from a non mature nymph)." httpUri="https://zenodo.org/record/194898/files/figure.png" pageId="5" pageNumber="66">Fig. 16</figureCitation>
). Mid leg (
<figureCitation id="13042A24FA5EFFD5FAF8E7365AE61D4C" captionStart="FIGURES 15 20" captionStartId="4.[151,273,1943,1967]" captionTargetBox="[189,1407,201,1919]" captionTargetId="figure@4.[183,1407,194,1919]" captionTargetPageId="4" captionText="FIGURES 15 20. Thraulodes jones sp. nov. Nymph. 15. Fore leg. 16. Tarsal claw. 17. Mid leg. 18. Hind leg. 19. Abdominal terga. 20. Gill. (scale: Figs. 15, 17, 18, 20, 1.0 mm; Fig. 16, 0.1 mm; Fig. 19, 2.5 mm) (Drawings of legs and abdomen from a non mature nymph)." httpUri="https://zenodo.org/record/194898/files/figure.png" pageId="5" pageNumber="66">Fig. 17</figureCitation>
): similar to foreleg except femora with one basal and one sub apical light brown mark and inner margin with row of strong short spines; outer margin of tibia with fringe of long simple setae; inner margin without setae.
<collectingCountry id="F3287631FA5EFFD5FF75E7865B421D99" box="[225,284,1944,1970]" name="India" pageId="5" pageNumber="66">Hind</collectingCountry>
leg (
<figureCitation id="13042A24FA5EFFD5FECDE7865BEC1D99" box="[345,434,1944,1970]" captionStart="FIGURES 15 20" captionStartId="4.[151,273,1943,1967]" captionTargetBox="[189,1407,201,1919]" captionTargetId="figure@4.[183,1407,194,1919]" captionTargetPageId="4" captionText="FIGURES 15 20. Thraulodes jones sp. nov. Nymph. 15. Fore leg. 16. Tarsal claw. 17. Mid leg. 18. Hind leg. 19. Abdominal terga. 20. Gill. (scale: Figs. 15, 17, 18, 20, 1.0 mm; Fig. 16, 0.1 mm; Fig. 19, 2.5 mm) (Drawings of legs and abdomen from a non mature nymph)." httpUri="https://zenodo.org/record/194898/files/figure.png" pageId="5" pageNumber="66">Fig. 18</figureCitation>
): similar to foreleg except basal mark of femora less conspicuous than in other legs, reduced to a line; inner margin with row of strong short spines, slightly longer than those of mid leg; outer margin of tibia fringed with long simple setae, short bristles and few clavate setae, inner margin with row of short bristles and row of long simple setae, although not as long as those of femora; outer margin of tarsus with row of simple long setae, some clavate setae and short bristles;
</paragraph>
<paragraph id="8B8036A1FA5DFFD6FF50E116590C1B9C" blockId="6.[151,1437,152,1648]" pageId="6" pageNumber="67">
<emphasis id="B94BEAB3FA5DFFD6FF50E1165B601B0A" box="[196,318,264,289]" italics="true" pageId="6" pageNumber="67">Abdomen:</emphasis>
Terga IVII mainly dark brown, postero-lateral areas light brown; terga VIIIX light brown, somewhat yellowish. Tergum X with median mark and lateral margins dark brown (
<figureCitation id="13042A24FA5DFFD6FBFDE1335E9E1B6C" box="[1129,1216,301,327]" captionStart="FIGURES 15 20" captionStartId="4.[151,273,1943,1967]" captionTargetBox="[189,1407,201,1919]" captionTargetId="figure@4.[183,1407,194,1919]" captionTargetPageId="4" captionText="FIGURES 15 20. Thraulodes jones sp. nov. Nymph. 15. Fore leg. 16. Tarsal claw. 17. Mid leg. 18. Hind leg. 19. Abdominal terga. 20. Gill. (scale: Figs. 15, 17, 18, 20, 1.0 mm; Fig. 16, 0.1 mm; Fig. 19, 2.5 mm) (Drawings of legs and abdomen from a non mature nymph)." httpUri="https://zenodo.org/record/194898/files/figure.png" pageId="6" pageNumber="67">Fig. 19</figureCitation>
). Caudal filaments brownish. Posterior margin of terga and caudal filaments with row of spines, longer on caudal filaments. Gills purplish gray, with symmetrical lamellae; main trachea strongly pigmented but branches are visible although weakly marked; gills broad, tapering toward apex (
<figureCitation id="13042A24FA5DFFD6FD78E183591A1B9C" box="[748,836,413,439]" captionStart="FIGURES 15 20" captionStartId="4.[151,273,1943,1967]" captionTargetBox="[189,1407,201,1919]" captionTargetId="figure@4.[183,1407,194,1919]" captionTargetPageId="4" captionText="FIGURES 15 20. Thraulodes jones sp. nov. Nymph. 15. Fore leg. 16. Tarsal claw. 17. Mid leg. 18. Hind leg. 19. Abdominal terga. 20. Gill. (scale: Figs. 15, 17, 18, 20, 1.0 mm; Fig. 16, 0.1 mm; Fig. 19, 2.5 mm) (Drawings of legs and abdomen from a non mature nymph)." httpUri="https://zenodo.org/record/194898/files/figure.png" pageId="6" pageNumber="67">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="8B8036A1FA5DFFD6FF50E1DC59221867" blockId="6.[151,1437,152,1648]" pageId="6" pageNumber="67">
<emphasis id="B94BEAB3FA5DFFD6FF50E1DC5B101BF0" box="[196,334,450,475]" italics="true" pageId="6" pageNumber="67">Variations:</emphasis>
Body length was higher on female mature nymphs, reaching 12.0 mm. Some specimens showed different abdominal color pattern, with dark brown anterior margin on tergum VIII and on tergum IX, this coloration was interrupted on median region. Aside that, subtle variations on color intensity were found, with specimens overall darker or lighter on their color patterns.
</paragraph>
</subSubSection>
<subSubSection id="C325652AFA5DFFD6FF50E24958D41F9D" pageId="6" pageNumber="67" type="materials_examined">
<paragraph id="8B8036A1FA5DFFD6FF50E2495F411983" blockId="6.[151,1437,152,1648]" pageId="6" pageNumber="67">
<emphasis id="B94BEAB3FA5DFFD6FF50E2495BDD185A" bold="true" box="[196,387,599,625]" pageId="6" pageNumber="67">
<typeStatus id="54848803FA5DFFD6FF50E2495B5C185A" box="[196,258,599,625]" pageId="6" pageNumber="67">Type</typeStatus>
material:
</emphasis>
<typeStatus id="54848803FA5DFFD6FE1AE246585B1859" box="[398,517,600,626]" pageId="6" pageNumber="67" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3287631FA5DFFD6FD85E246583E1859" box="[529,608,600,626]" name="Brazil" pageId="6" pageNumber="67">Brazil</collectingCountry>
, Rio de Janeiro: Nova Friburgo, Lumiar, Rio Boa Vista, Cachoeira Indiana
<taxonomicName id="4C3F4D22FA5DFFD6FF62E29E5B6418B1" box="[246,314,640,666]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="68" phylum="Arthropoda" rank="species" species="jones" status="sp. nov.">Jones</taxonomicName>
,
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,
<geoCoordinate id="EE0B5066FA5DFFD6FE0AE29E586F18B1" box="[414,561,637,666]" direction="south" orientation="latitude" pageId="6" pageNumber="67" precision="1" value="-22.317251">22o19'02.1&quot;S</geoCoordinate>
/
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,
<date id="FF811061FA5DFFD6FD78E29E593218B1" box="[748,876,640,666]" pageId="6" pageNumber="67" value="2008-11-15">15.xi.2008</date>
, Gonçalves, I.C.
<emphasis id="B94BEAB3FA5DFFD6FBA1E29E5E3F18B2" box="[1077,1121,640,665]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
<specimenCount id="9D39FD28FA5DFFD6FBFCE29E5EE918B1" box="[1128,1207,640,666]" pageId="6" pageNumber="67" type="male">1 male</specimenCount>
imago (
<collectionCode id="ED2EAE64FA5DFFD6FA83E29E5F3C18B1" box="[1303,1378,640,666]" pageId="6" pageNumber="67">DZRJ</collectionCode>
800); Alotype: Nova Friburgo, Lumiar, tributário de primeira ordem do Córrego Santa Margarida,
<quantity id="4CC79B44FA5DFFD6FAD0E2BB5FC61894" box="[1348,1432,677,703]" metricMagnitude="2" metricUnit="m" metricValue="8.44" pageId="6" pageNumber="67" unit="m" value="844.0">844 m</quantity>
,
<geoCoordinate id="EE0B5066FA5DFFD6FF03E2D35B7518CC" box="[151,299,715,743]" direction="south" orientation="latitude" pageId="6" pageNumber="67" precision="1" value="-22.343195">22o20'35.5&quot;S</geoCoordinate>
/
<geoCoordinate id="EE0B5066FA5DFFD6FEA0E2D35B8618CC" box="[308,472,715,743]" direction="west" orientation="longitude" pageId="6" pageNumber="67" precision="1" value="-42.3">42o18'00.0&quot;W</geoCoordinate>
,
<date id="FF811061FA5DFFD6FE70E2D3583A18CC" box="[484,612,717,743]" pageId="6" pageNumber="67" value="2008-11-16">16.xi.2008</date>
, Gonçalves, I.C.
<emphasis id="B94BEAB3FA5DFFD6FCB9E2D0590618CC" box="[813,856,718,743]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
(Light trap)
<specimenCount id="9D39FD28FA5DFFD6FC79E2D35E0D18CC" box="[1005,1107,717,743]" pageId="6" pageNumber="67" type="female">1 female</specimenCount>
imago (alotype;
<collectionCode id="ED2EAE64FA5DFFD6FA8DE2D35F3A18CC" box="[1305,1380,717,743]" pageId="6" pageNumber="67">DZRJ</collectionCode>
807).
<typeStatus id="54848803FA5DFFD6FF03E2ED5B501926" box="[151,270,755,781]" pageId="6" pageNumber="67" type="paratype">Paratypes</typeStatus>
: same data, 2 nymphs (
<collectionCode id="ED2EAE64FA5DFFD6FD8AE2ED58D81926" box="[542,646,755,781]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="67">MZUSP</collectionCode>
); same data, 1 nymph (
<collectionCode id="ED2EAE64FA5DFFD6FC00E2ED59811926" box="[916,991,755,781]" pageId="6" pageNumber="67">DZRJ</collectionCode>
802); same locality and date, Alecrim, V.P.
<emphasis id="B94BEAB3FA5DFFD6FF58E3075AA91919" box="[204,247,793,818]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
6 nymphs (
<collectionCode id="ED2EAE64FA5DFFD6FE17E3065B911919" box="[387,463,792,818]" pageId="6" pageNumber="67">DZRJ</collectionCode>
803); same data, 1 nymph (
<collectionCode id="ED2EAE64FA5DFFD6FC99E30659061919" box="[781,856,792,818]" pageId="6" pageNumber="67">DZRJ</collectionCode>
804); Nova Friburgo, Lumiar, Rio Boa Vista,
<quantity id="4CC79B44FA5DFFD6FAE4E3065AEE1970" metricMagnitude="2" metricUnit="m" metricValue="5.83" pageId="6" pageNumber="67" unit="m" value="583.0">583 m</quantity>
,
<geoCoordinate id="EE0B5066FA5DFFD6FF29E35F5B071970" box="[189,345,829,859]" direction="south" orientation="latitude" pageId="6" pageNumber="67" precision="1" value="-22.399416">22o23' 57.9&quot;S</geoCoordinate>
/
<geoCoordinate id="EE0B5066FA5DFFD6FEF7E35F58561970" box="[355,520,829,859]" direction="west" orientation="longitude" pageId="6" pageNumber="67" precision="1" value="-42.32072">42o19'14.6&quot;W</geoCoordinate>
,
<date id="FF811061FA5DFFD6FD81E35F58C81970" box="[533,662,833,859]" pageId="6" pageNumber="67" value="2008-11-15">15.xi.2008</date>
, Alecrim, V.P.
<emphasis id="B94BEAB3FA5DFFD6FCD3E35F592C1971" box="[839,882,833,858]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
1 nymph (
<collectionCode id="ED2EAE64FA5DFFD6FC66E35F5E621970" box="[1010,1084,833,859]" pageId="6" pageNumber="67">DZRJ</collectionCode>
805); Nova Friburgo, Lumiar, first order tributary of Córrego Santa Margarida,
<quantity id="4CC79B44FA5DFFD6FD4BE377597319A8" box="[735,813,873,899]" metricMagnitude="2" metricUnit="m" metricValue="8.44" pageId="6" pageNumber="67" unit="m" value="844.0">844 m</quantity>
,
<geoCoordinate id="EE0B5066FA5DFFD6FCAEE377599019A8" box="[826,974,870,899]" direction="south" orientation="latitude" pageId="6" pageNumber="67" precision="1" value="-22.343195">22o20'35.5&quot;S</geoCoordinate>
/
<geoCoordinate id="EE0B5066FA5DFFD6FC4CE3775E2019A8" box="[984,1150,870,899]" direction="west" orientation="longitude" pageId="6" pageNumber="67" precision="1" value="-42.3">42o18'00.0&quot;W</geoCoordinate>
,
<date id="FF811061FA5DFFD6FB1EE3775F5519A8" box="[1162,1291,873,899]" pageId="6" pageNumber="67" value="2008-11-16">16.xi.2008</date>
, Gonçalves, I.C.
<emphasis id="B94BEAB3FA5DFFD6FF5EE3905AAB198C" box="[202,245,910,935]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
(Light trap),
<specimenCount id="9D39FD28FA5DFFD6FE07E3905BBC1983" box="[403,482,910,936]" pageId="6" pageNumber="67" type="male">1 male</specimenCount>
and
<specimenCount id="9D39FD28FA5DFFD6FD8FE39058DF1983" box="[539,641,910,936]" pageId="6" pageNumber="67" type="female">3 female</specimenCount>
imagos (
<collectionCode id="ED2EAE64FA5DFFD6FD78E390590B1983" box="[748,853,910,936]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="67">MZUSP</collectionCode>
); same data,
<specimenCount id="9D39FD28FA5DFFD6FC7FE3905E651983" box="[1003,1083,910,936]" pageId="6" pageNumber="67" type="male">1 male</specimenCount>
imago (
<collectionCode id="ED2EAE64FA5DFFD6FB0EE3905EBB1983" box="[1178,1253,910,936]" pageId="6" pageNumber="67">DZRJ</collectionCode>
806);
</paragraph>
<paragraph id="8B8036A1FA5DFFD6FF50E3AD58D41F9D" blockId="6.[151,1437,152,1648]" pageId="6" pageNumber="67">
<emphasis id="B94BEAB3FA5DFFD6FF50E3AD5B9419E6" bold="true" box="[196,458,947,973]" pageId="6" pageNumber="67">Additional material</emphasis>
:
<emphasis id="B94BEAB3FA5DFFD6FE4FE3AD595719E6" bold="true" box="[475,777,947,973]" pageId="6" pageNumber="67">
<collectingCountry id="F3287631FA5DFFD6FE4FE3AD587019E6" box="[475,558,947,973]" name="Brazil" pageId="6" pageNumber="67">Brazil</collectingCountry>
, Rio de Janeiro:
</emphasis>
Nova Friburgo, Macaé de Cima, Rio Macaé,
<quantity id="4CC79B44FA5DFFD6FAD2E3AD5FC619E6" box="[1350,1432,947,973]" metricMagnitude="2" metricUnit="m" metricValue="9.35" pageId="6" pageNumber="67" unit="m" value="935.0">935 m</quantity>
,
<geoCoordinate id="EE0B5066FA5DFFD6FF03E3C25B4B19DD" box="[151,277,985,1014]" direction="south" orientation="latitude" pageId="6" pageNumber="67" precision="15" value="-22.412777">22o24'46&quot;S</geoCoordinate>
/
<geoCoordinate id="EE0B5066FA5DFFD6FE8AE3C25B9D19DD" box="[286,451,985,1014]" direction="west" orientation="longitude" pageId="6" pageNumber="67" precision="1" value="-42.521168">42o31'16.2&quot;W</geoCoordinate>
,
<date id="FF811061FA5DFFD6FE5BE3C2581119DD" box="[463,591,988,1014]" pageId="6" pageNumber="67" value="2009-09-12">12.ix.2009</date>
, Alecrim, V.P.
<emphasis id="B94BEAB3FA5DFFD6FD6AE3C2597719DE" box="[766,809,988,1013]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
,
<specimenCount id="9D39FD28FA5DFFD6FCACE3C259C019DD" box="[824,926,988,1014]" pageId="6" pageNumber="67" type="female">4 female</specimenCount>
imagos (
<collectionCode id="ED2EAE64FA5DFFD6FB9DE3C25E0A19DD" box="[1033,1108,988,1014]" pageId="6" pageNumber="67">DZRJ</collectionCode>
812); same data, Gonçalves, I.C.
<emphasis id="B94BEAB3FA5DFFD6FF58E41F5AA71E31" box="[204,249,1025,1050]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
, 1 nymph (
<collectionCode id="ED2EAE64FA5DFFD6FE11E41F5BAE1E30" box="[389,496,1025,1051]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="67">MZUSP</collectionCode>
); same locality,
<date id="FF811061FA5DFFD6FD20E41F59691E30" box="[692,823,1025,1051]" pageId="6" pageNumber="67" value="2009-09-13">13.ix.2009</date>
, Gonçalves, I.C.
<emphasis id="B94BEAB3FA5DFFD6FB9DE41F5E681E31" box="[1033,1078,1025,1050]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
,
<specimenCount id="9D39FD28FA5DFFD6FBD2E41F5EEE1E30" box="[1094,1200,1025,1051]" pageId="6" pageNumber="67" type="female">1 female</specimenCount>
imago (
<collectionCode id="ED2EAE64FA5DFFD6FA80E41F5F3F1E30" box="[1300,1377,1025,1051]" pageId="6" pageNumber="67">DZRJ</collectionCode>
809); same data,
<specimenCount id="9D39FD28FA5DFFD6FEB7E4385BD11E6B" box="[291,399,1062,1088]" pageId="6" pageNumber="67" type="female">1 female</specimenCount>
subimago (
<collectionCode id="ED2EAE64FA5DFFD6FD8BE43858301E6B" box="[543,622,1062,1088]" pageId="6" pageNumber="67">DZRJ</collectionCode>
810); same data, 5 nymphs (
<collectionCode id="ED2EAE64FA5DFFD6FC5EE4385E441E6B" box="[970,1050,1062,1088]" pageId="6" pageNumber="67">DZRJ</collectionCode>
811); same locality,
<date id="FF811061FA5DFFD6FA85E4385FC91E6B" box="[1297,1431,1062,1088]" pageId="6" pageNumber="67" value="2009-09-14">14.ix.2009</date>
, Alecrim, V.P.
<emphasis id="B94BEAB3FA5DFFD6FED2E4525B2A1E4E" box="[326,372,1100,1125]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
,
<specimenCount id="9D39FD28FA5DFFD6FE11E4525BAF1E4D" box="[389,497,1100,1126]" pageId="6" pageNumber="67" type="female">3 female</specimenCount>
imagos (
<collectionCode id="ED2EAE64FA5DFFD6FDF1E452588D1E4D" box="[613,723,1100,1126]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="67">MZUSP</collectionCode>
); same locality,
<date id="FF811061FA5DFFD6FC0AE4525E7B1E4D" box="[926,1061,1100,1126]" pageId="6" pageNumber="67" value="2008-09-18">18.ix.2008</date>
, Alecrim, V.P.
<emphasis id="B94BEAB3FA5DFFD6FB70E4525F4C1E4E" box="[1252,1298,1100,1125]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
, 4 nymphs (
<collectionCode id="ED2EAE64FA5DFFD6FF0BE46F5AB21EA0" box="[159,236,1137,1163]" pageId="6" pageNumber="67">DZRJ</collectionCode>
808); Nova Friburgo, Macaé de Cima, Rio Macaé,
<quantity id="4CC79B44FA5DFFD6FCDAE46F59C31EA0" box="[846,925,1137,1163]" metricMagnitude="2" metricUnit="m" metricValue="9.77" pageId="6" pageNumber="67" unit="m" value="977.0">977 m</quantity>
,
<geoCoordinate id="EE0B5066FA5DFFD6FC3EE46F5E151EA0" box="[938,1099,1137,1163]" direction="south" orientation="latitude" pageId="6" pageNumber="67" precision="1" value="-22.425165">22º2530.6S</geoCoordinate>
/
<geoCoordinate id="EE0B5066FA5DFFD6FBC1E46F5F571EA0" box="[1109,1289,1137,1163]" direction="west" orientation="longitude" pageId="6" pageNumber="67" precision="1" value="-42.533527">42º3200.7W</geoCoordinate>
,
<date id="FF811061FA5DFFD6FA81E46F5FC91EA0" box="[1301,1431,1137,1163]" pageId="6" pageNumber="67" value="2009-09-13">13.ix.2009</date>
, Gonçalves, I.C.
<emphasis id="B94BEAB3FA5DFFD6FEC0E4895BDE1E9B" box="[340,384,1175,1200]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
, 3 nymphs (
<collectionCode id="ED2EAE64FA5DFFD6FD85E48858241E9B" box="[529,634,1174,1200]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="67">MZUSP</collectionCode>
); Teresópolis, Parque Nacional da Serra dos Órgãos, Rio Paquequer,
<quantity id="4CC79B44FA5DFFD6FF03E4A25ABB1EFD" box="[151,229,1212,1238]" metricMagnitude="2" metricUnit="m" metricValue="9.33" pageId="6" pageNumber="67" unit="m" value="933.0">933 m</quantity>
,
<geoCoordinate id="EE0B5066FA5DFFD6FF66E4A25BD71EFD" box="[242,393,1212,1238]" direction="south" orientation="latitude" pageId="6" pageNumber="67" precision="1" value="-22.45226">22°27'8.13&quot;S</geoCoordinate>
/
<geoCoordinate id="EE0B5066FA5DFFD6FE07E4A258131EFD" box="[403,589,1212,1238]" direction="west" orientation="longitude" pageId="6" pageNumber="67" precision="1" value="-42.992157">42°59'31.77&quot;W</geoCoordinate>
,
<date id="FF811061FA5DFFD6FDCDE4A2588E1EFD" box="[601,720,1212,1238]" pageId="6" pageNumber="67" value="2008-05-10">10.v.2008</date>
, Henriques-Oliveira, A.L., Santos, A.P.M. &amp; Ferreira-Jr., N.
<emphasis id="B94BEAB3FA5DFFD6FF03E4FF5AE31ED1" box="[151,189,1249,1274]" italics="true" pageId="6" pageNumber="67">leg</emphasis>
. (light trap),
<specimenCount id="9D39FD28FA5DFFD6FEF0E4FF5BE61ED0" box="[356,440,1249,1275]" pageId="6" pageNumber="67" type="male">1 male</specimenCount>
subimago (
<collectionCode id="ED2EAE64FA5DFFD6FDDDE4FF58C61ED0" box="[585,664,1249,1275]" pageId="6" pageNumber="67">DZRJ</collectionCode>
816); Macaé, Sana, Córrego do Colégio,
<quantity id="4CC79B44FA5DFFD6FB06E4FF5EBD1ED0" box="[1170,1251,1249,1275]" metricMagnitude="2" metricUnit="m" metricValue="2.94" pageId="6" pageNumber="67" unit="m" value="294.0">294 m</quantity>
,
<geoCoordinate id="EE0B5066FA5DFFD6FB65E4FF5FCD1ED0" box="[1265,1427,1249,1275]" direction="south" orientation="latitude" pageId="6" pageNumber="67" precision="1" value="-22.339834">22°20'23.4&quot;S</geoCoordinate>
/
<geoCoordinate id="EE0B5066FA5DFFD6FF03E5185B1F1F0B" box="[151,321,1286,1312]" direction="west" orientation="longitude" pageId="6" pageNumber="67" precision="1" value="-42.20375">42°12'13.5&quot;W</geoCoordinate>
,
<date id="FF811061FA5DFFD6FEDAE5185B941F0B" box="[334,458,1286,1312]" pageId="6" pageNumber="67" value="2009-02-19">19.ii.2009</date>
, Gonçalves, I.C.
<emphasis id="B94BEAB3FA5DFFD6FD08E51958961F0B" box="[668,712,1287,1312]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
(light trap), one female subimago (
<collectionCode id="ED2EAE64FA5DFFD6FBE1E5185E9C1F0B" box="[1141,1218,1286,1312]" pageId="6" pageNumber="67">DZRJ</collectionCode>
818); same data,
<specimenCount id="9D39FD28FA5DFFD6FA19E5185A8C1F6D" pageId="6" pageNumber="67" type="male">1 male</specimenCount>
and
<specimenCount id="9D39FD28FA5DFFD6FE9BE5325B241F6D" box="[271,378,1324,1350]" pageId="6" pageNumber="67" type="female">2 female</specimenCount>
imagos (
<collectionCode id="ED2EAE64FA5DFFD6FE78E53258651F6D" box="[492,571,1324,1350]" pageId="6" pageNumber="67">DZRJ</collectionCode>
819);
<emphasis id="B94BEAB3FA5DFFD6FDEBE53559551F6E" bold="true" box="[639,779,1323,1349]" pageId="6" pageNumber="67">São Paulo:</emphasis>
Ubatuba, Parque Estadual da Serra do Mar Núcleo Picinguaba, Rio da Fazenda,
<quantity id="4CC79B44FA5DFFD6FE63E54F58691F40" box="[503,567,1361,1387]" metricMagnitude="1" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="67" unit="m" value="41.0">41 m</quantity>
,
<geoCoordinate id="EE0B5066FA5DFFD6FDD0E54F58801F40" box="[580,734,1361,1387]" direction="south" orientation="latitude" pageId="6" pageNumber="67" precision="1" value="-23.337973">23°20'16.7&quot;S</geoCoordinate>
/
<geoCoordinate id="EE0B5066FA5DFFD6FD7CE54F59CB1F40" box="[744,917,1361,1387]" direction="west" orientation="longitude" pageId="6" pageNumber="67" precision="1" value="-44.83636">44°50'10.9&quot;W</geoCoordinate>
,
<date id="FF811061FA5DFFD6FC35E54F5E421F40" box="[929,1052,1361,1387]" pageId="6" pageNumber="67" value="2005-10-02">02.x.2005</date>
, Santos, A.P.M. &amp; Dumas, L.L.
<emphasis id="B94BEAB3FA5DFFD6FF03E5695A9D1FBB" box="[151,195,1399,1424]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
(Light trap),
<specimenCount id="9D39FD28FA5DFFD6FEFEE5685BE31FBB" box="[362,445,1398,1424]" pageId="6" pageNumber="67" type="male">1 male</specimenCount>
and
<specimenCount id="9D39FD28FA5DFFD6FE6DE568583C1FBB" box="[505,610,1398,1424]" pageId="6" pageNumber="67" type="female">2 female</specimenCount>
subimagos (
<collectionCode id="ED2EAE64FA5DFFD6FD6AE56859131FBB" box="[766,845,1398,1424]" pageId="6" pageNumber="67">DZRJ</collectionCode>
821); same locality and date, Santos, A.P.M.
<emphasis id="B94BEAB3FA5DFFD6FAFBE5695FC21FBB" box="[1391,1436,1399,1424]" italics="true" pageId="6" pageNumber="67">leg.</emphasis>
(Light trap),
<specimenCount id="9D39FD28FA5DFFD6FEB9E5825B221F9D" box="[301,380,1436,1462]" pageId="6" pageNumber="67" type="male">1 male</specimenCount>
subimago (
<collectionCode id="ED2EAE64FA5DFFD6FD91E582580E1F9D" box="[517,592,1436,1462]" pageId="6" pageNumber="67">DZRJ</collectionCode>
822).
</paragraph>
</subSubSection>
<subSubSection id="C325652AFA5DFFD6FF52E5DE58181C60" pageId="6" pageNumber="67" type="etymology">
<paragraph id="8B8036A1FA5DFFD6FF52E5DE5B591C2B" blockId="6.[151,1437,152,1648]" pageId="6" pageNumber="67">
<emphasis id="B94BEAB3FA5DFFD6FF52E5DE5B131FF1" bold="true" box="[198,333,1472,1498]" pageId="6" pageNumber="67">Etymology</emphasis>
: The name
<taxonomicName id="4C3F4D22FA5DFFD6FE73E5DF587B1FF1" box="[487,549,1473,1498]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="jones">
<emphasis id="B94BEAB3FA5DFFD6FE73E5DF587B1FF1" box="[487,549,1473,1498]" italics="true" pageId="6" pageNumber="67">jones</emphasis>
</taxonomicName>
is a reference to the
<typeStatus id="54848803FA5DFFD6FCA2E5DF59371FF0" box="[822,873,1473,1499]" pageId="6" pageNumber="67">type</typeStatus>
locality, the Boa Vista river at Indiana
<taxonomicName id="4C3F4D22FA5DFFD6FACFE5DF5FC21FF0" box="[1371,1436,1473,1499]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="68" phylum="Arthropoda" rank="species" species="jones" status="sp. nov.">Jones</taxonomicName>
waterfall.
</paragraph>
<paragraph id="8B8036A1FA5DFFD6FF52E61558181C60" blockId="6.[151,1437,152,1648]" pageId="6" pageNumber="67">
<emphasis id="B94BEAB3FA5DFFD6FF52E6155BB81C0E" bold="true" box="[198,486,1547,1573]" pageId="6" pageNumber="67">Life cycle associations:</emphasis>
Nymphs and adults were associated from one male nymph (
<typeStatus id="54848803FA5DFFD6FB2FE6125F751C0D" box="[1211,1323,1548,1574]" pageId="6" pageNumber="67" type="holotype">Holotype</typeStatus>
) and two female nymphs reared to adult stage.
</paragraph>
</subSubSection>
<subSubSection id="C325652AFA5DFFD6FF52E6485EFB1C5B" box="[198,1189,1622,1648]" pageId="6" pageNumber="67" type="biology_ecology">
<paragraph id="8B8036A1FA5DFFD6FF52E6485EFB1C5B" blockId="6.[151,1437,152,1648]" box="[198,1189,1622,1648]" pageId="6" pageNumber="67">
<emphasis id="B94BEAB3FA5DFFD6FF52E6485B7A1C5B" bold="true" box="[198,292,1622,1648]" pageId="6" pageNumber="67">Biology</emphasis>
: Nymphs were found on rocky stream bottoms in areas of moderate current.
</paragraph>
</subSubSection>
</treatment>
</document>