164 lines
18 KiB
XML
164 lines
18 KiB
XML
<document ID-ISSN="2201-4349" approvalRequired="134" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="24" approvalRequired_for_textStreams="101" approvalRequired_for_treatments="8" checkinTime="1629578325228" checkinUser="felipe" docAuthor="Moulds, M. S." docDate="2003" docId="03811E7D0911FF9E6254FCD2FBA88DA6" docLanguage="en" docName="RecAustMus.55.3.245-304.pdf" docOrigin="Records of the Australian Museum 55" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.1:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="1" docTitle="Aleeta Moulds 2003, n.gen." docType="treatment" docVersion="3" lastPageNumber="263" masterDocId="FFB866050903FF8C6338FFD8FFDA8950" masterDocTitle="An Appraisal of the Cicadas of the Genus Abricta Stål and Allied Genera (Hemiptera: Auchenorrhyncha: Cicadidae)" masterLastPageNumber="304" masterPageNumber="245" pageNumber="263" updateTime="1676039802306" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>An Appraisal of the Cicadas of the Genus Abricta Stål and Allied Genera (Hemiptera: Auchenorrhyncha: Cicadidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moulds, M. S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2003</mods:date>
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<mods:detail type="volume">
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<mods:number>55</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>245</mods:start>
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<mods:end>304</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="ISSN">2201-4349</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5239822" ID-Zenodo-Dep="5239822" LSID="urn:lsid:plazi:treatment:03811E7D0911FF9E6254FCD2FBA88DA6" httpUri="http://treatment.plazi.org/id/03811E7D0911FF9E6254FCD2FBA88DA6" lastPageNumber="263" pageId="18" pageNumber="263">
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<subSubSection box="[364,596,778,802]" pageId="18" pageNumber="263" type="nomenclature">
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<paragraph blockId="18.[364,596,778,802]" box="[364,596,778,802]" pageId="18" pageNumber="263">
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<heading box="[364,596,778,802]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="263" reason="2">
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<emphasis bold="true" box="[364,596,778,802]" pageId="18" pageNumber="263">
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Genus
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<taxonomicName ID-CoL="8HCFF" ID-ENA="1218667" authority="Moulds, 2003" authorityName="Moulds" authorityYear="2003" box="[445,518,778,802]" class="Insecta" family="Cicadidae" genus="Aleeta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="genus" status="n.gen.">
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<emphasis bold="true" box="[445,518,778,802]" italics="true" pageId="18" pageNumber="263">Aleeta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[525,596,778,802]" pageId="18" pageNumber="263" rank="genus">n.gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="263" type="type_taxon">
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<paragraph blockId="18.[166,681,830,854]" box="[166,681,830,854]" pageId="18" pageNumber="263">
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<emphasis bold="true" box="[166,308,830,854]" pageId="18" pageNumber="263">
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<typeStatus box="[166,222,830,854]" pageId="18" pageNumber="263">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName authority="Germar, 1834" authorityName="Germar" authorityYear="1834" box="[322,676,830,853]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="species" species="curvicosta">
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<emphasis box="[322,519,830,853]" italics="true" pageId="18" pageNumber="263">Cicada curvicosta</emphasis>
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<bibRefCitation author="Germar, E" box="[525,676,830,853]" pageId="18" pageNumber="263" pagination="49 - 82" refId="ref61046" refString="Germar, E. F., 1834. Observations sur plusieurs espsces du genre Cicada, Latr. Revue Entomologique, Silbermann 2: 49 - 82, pls." type="journal article" year="1834">Germar, 1834</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="18.[166,587,882,906]" box="[166,587,882,906]" pageId="18" pageNumber="263">
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<emphasis bold="true" box="[166,352,882,906]" pageId="18" pageNumber="263">Included species</emphasis>
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.
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<taxonomicName authority="(Germar)" baseAuthorityName="Germar" box="[363,583,882,905]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="species" species="curvicosta">
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<emphasis box="[363,475,882,905]" italics="true" pageId="18" pageNumber="263">curvicosta</emphasis>
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(Germar)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="263" type="etymology">
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<paragraph blockId="18.[166,794,934,1074]" pageId="18" pageNumber="263">
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<emphasis bold="true" box="[166,290,934,958]" pageId="18" pageNumber="263">Etymology</emphasis>
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. Derived from the Greek
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<emphasis box="[568,635,934,957]" italics="true" pageId="18" pageNumber="263">aleton</emphasis>
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meaning flour or meal and pertaining to the flour-like “dusting” partly covering the body of
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<taxonomicName box="[396,539,993,1016]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="species" species="curvicosta">
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<emphasis box="[396,539,993,1016]" italics="true" pageId="18" pageNumber="263">A. curvicosta</emphasis>
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</taxonomicName>
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, resulting from the fine silver body pubescence. This pubescence is easily abraded and is often substantially lacking on older adults.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="263" type="diagnosis">
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<paragraph blockId="18.[166,795,1103,2006]" pageId="18" pageNumber="263">
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<emphasis bold="true" box="[166,287,1103,1127]" pageId="18" pageNumber="263">Diagnosis</emphasis>
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. Postclypeus slightly produced. Rostrum reaching or just passing bases of hind coxae. Width of head including eyes equal to, or slightly wider than, anterior pronotum and equal to, or narrower than, mesonotum; width of abdomen as wide as thorax. Pronotal collar narrow with lateral angles ampliate and rounded; rudimentary and essentially confluent with adjoining sclerites anterior of lateral angles. Wings hyaline. Fore wing with infuscation at bases of apical cells 2 and 3; costal margin ampliate to node, the maximum dilation greater than width of costal vein; basal cell usually with translucent pigmentation. Hind wing with plaga along much of vein 3A plus inner margin of anal lobe and along length of vein 2A; plaga bordered by black infuscation that is partly expanded along wing margin at distal end of 2A. Tymbals (
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<figureCitation box="[576,656,1514,1537]" captionStart="Figs" captionStartId="9.[794,842,976,997]" captionTargetBox="[794,1388,151,961]" captionTargetId="figure-737@9.[794,1388,151,961]" captionTargetPageId="9" captionText="Figs. 18–21. Tymbals of Trismarcha, Abricta, Aleeta and Chrysolasia species.(18) Trismarcha umbrosa, Boukoko, Republic of Central Africa; (19) Abricta brunnea, Mauritius; (20) Aleeta curvicosta, Kuranda, northern Qld; (21) Chrysolasia guatemalena, Purula, Guatemala." pageId="18" pageNumber="263">Fig. 20</figureCitation>
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) with 11–13 long ribs (usually 12) evenly spaced but slightly converging dorsally; basal plate small. Male opercula tending tear-drop in shape, weakly angled inwards before midpoint; nearly flat; not quite meeting; extending laterally beyond lateral margins of abdomen and distally a little beyond tympanal cavities. Newly emerged individuals bear fine silver pubescence over much of body and along basal fore wing veins.
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</paragraph>
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<paragraph blockId="18.[166,795,1103,2006]" lastBlockId="18.[839,1467,153,440]" pageId="18" pageNumber="263">
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Male genitalia (
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<figureCitation box="[362,487,1778,1801]" captionStart="Figs" captionStartId="19.[121,166,1120,1141]" captionTargetBox="[121,1428,151,1080]" captionTargetId="figure-466@19.[121,1428,151,1080]" captionTargetPageId="19" captionText="Figs. 42–49. Male genitalia of Abricta, Abroma, Aleeta and Tryella species showing pygofer with uncus and aedeagus in right lateral view (left) and ventral view (right): (42–43) Abricta ferruginosa, Mauritius; (44–45) Abroma guerinii, Madagascar; (46–47) Aleeta curvicosta, genitalia prep.AB32; and (48–49) Tryella noctua, genitalia prep.AB21." pageId="18" pageNumber="263">Figs. 46, 47</figureCitation>
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) with apical spine of pygofer large, long and pointed; upper pygofer lobes much shorter than apical spine, tending bilobed; pygofer basal lobes divided into primary outer and secondary inner lobes, the outer lobe in lateral view substantially webbed to inner lobe. Uncal lobes widely separated, beak-like, downturned; no lateral process at base of uncus. Aedeagus in lateral view straight or gently curved for most of its length, basal plate a fused pair of nearly circular discs; conjunctival claws directed ventrally, claw-like with no associated sclerotization, flattened, broad in lateral view, narrow in ventral view, apex simple and pointed; a small sublateral rounded flange either side of theca near distal end; palearis absent.
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</paragraph>
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<paragraph blockId="18.[839,1467,153,440]" pageId="18" pageNumber="263">
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Female abdominal segment 9 long, clearly more than half as long as abdominal remainder, nearly conical; ovipositor (
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<figureCitation box="[962,1024,358,381]" captionStart="Figs" captionStartId="6.[166,211,1158,1179]" captionTargetBox="[166,794,151,1138]" captionTargetId="figure-692@6.[166,794,151,1138]" captionTargetPageId="6" captionText="Figs. 7–14. Distal end of female ovipositor in lateral view: (7) Trismarcha umbrosa, Boukoko, Republic of Central Africa; (8) Aleeta curvicosta, Kuranda, northern Qld; (9) Tryella burnsi, Kuranda, northern Qld; (10) T. castanea, Waterhouse R., NT; (11) T. kauma, East Haydon, northern Qld; (12) T. lachlani, “York Downs”, northern Qld; (13) T. rubra, Waterhouse R., NT; (14) T. stalkeri, De Grey R., WA." pageId="18" pageNumber="263">Fig. 8</figureCitation>
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) long with strong dorsal downward slope in lateral view; ovipositor sheath terminating about level with, or slightly extending beyond, apex of dorsal beak.
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</paragraph>
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<paragraph blockId="18.[839,1467,468,785]" pageId="18" pageNumber="263">
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<emphasis bold="true" box="[839,1116,468,492]" pageId="18" pageNumber="263">Distinguishing features</emphasis>
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. Male genitalia show features unique within the
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<taxonomicName authorityName="Stal and Allied" authorityYear="1866" box="[1045,1126,498,521]" class="Insecta" family="Cicadidae" genus="Abricta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis box="[1045,1126,498,521]" italics="true" pageId="18" pageNumber="263">Abricta</emphasis>
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</taxonomicName>
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complex of genera: the beaklike uncal lobes and the partially bifurcate upper pygofer lobes do not occur elsewhere in the
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<taxonomicName authorityName="Stal and Allied" authorityYear="1866" box="[1216,1297,557,580]" class="Insecta" family="Cicadidae" genus="Abricta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis box="[1216,1297,557,580]" italics="true" pageId="18" pageNumber="263">Abricta</emphasis>
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</taxonomicName>
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complex, while the pair of sublateral flanges near the distal end of the theca and the laterally flattened conjunctival claws are unknown in any other
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<taxonomicName authorityName="Latreille" authorityYear="1802" box="[978,1103,645,668]" pageId="18" pageNumber="303" rank="superFamily" superFamily="Cicadoidea">Cicadoidea</taxonomicName>
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. The broad male opercula, which extend clearly beyond the lateral margins of the abdomen, are also unique within the
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<taxonomicName authorityName="Stal and Allied" authorityYear="1866" box="[1124,1205,703,726]" class="Insecta" family="Cicadidae" genus="Abricta" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis box="[1124,1205,703,726]" italics="true" pageId="18" pageNumber="263">Abricta</emphasis>
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</taxonomicName>
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complex of genera. The strongly ampliate fore wing costa is shared with
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<taxonomicName authorityName="Moulds" authorityYear="2003" box="[1351,1427,733,756]" class="Insecta" family="Cicadidae" genus="Tryella" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis box="[1351,1427,733,756]" italics="true" pageId="18" pageNumber="263">Tryella</emphasis>
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</taxonomicName>
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but is otherwise only found in a few distantly related genera.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="263" type="discussion">
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<paragraph blockId="18.[839,1467,814,1160]" pageId="18" pageNumber="263">
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<emphasis bold="true" box="[839,944,814,838]" pageId="18" pageNumber="263">Remarks</emphasis>
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. The distinctive nature of this monotypic genus is also reflected in its genetic makeup. Allozyme electrophoresis revealed a high number of fixed differences between
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<taxonomicName box="[937,1080,902,925]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="species" species="curvicosta">
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<emphasis box="[937,1080,902,925]" italics="true" pageId="18" pageNumber="263">A. curvicosta</emphasis>
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</taxonomicName>
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and some allied species examined, reflecting the high genetic dissimilarity and long genetic distance between
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<taxonomicName box="[1058,1218,961,984]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="species" species="curvicosta">
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<emphasis box="[1058,1218,961,984]" italics="true" pageId="18" pageNumber="263">A. curvicosta</emphasis>
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</taxonomicName>
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and
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<taxonomicName box="[1285,1466,961,984]" class="Insecta" family="Cicadidae" genus="Tryella" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis box="[1285,1369,961,984]" italics="true" pageId="18" pageNumber="263">Tryella</emphasis>
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species
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</taxonomicName>
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(Serkowski & Moulds, unpub. data). Further, phylogenetic analyses of both electrophoretic and morphological data confirm the evolutionary distinctiveness of
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<taxonomicName box="[1322,1466,1049,1072]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="species" species="curvicosta">
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<emphasis box="[1322,1466,1049,1072]" italics="true" pageId="18" pageNumber="263">A. curvicosta</emphasis>
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</taxonomicName>
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from its sister group
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<taxonomicName authorityName="Moulds" authorityYear="2003" box="[1060,1136,1078,1101]" class="Insecta" family="Cicadidae" genus="Tryella" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis box="[1060,1136,1078,1101]" italics="true" pageId="18" pageNumber="263">Tryella</emphasis>
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</taxonomicName>
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(Serkowski & Moulds, unpub. data, and
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<figureCitation box="[940,1057,1107,1130]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="7.[121,166,1605,1626]" captionStartId-1="8.[166,202,745,766]" captionTargetBox-0="[121,1270,151,2003]" captionTargetBox-1="[166,1451,151,1281]" captionTargetId-0="figure-146@7.[121,1270,151,1288]" captionTargetId-1="figure-915@8.[192,1452,151,1281]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figs. 15 and 16. Strict consensus trees:(15) derived from the 12 most parsimonious trees generated when all multistate characters were left unordered and using Trismarcha umbrosa as outgroup.Analysis incorporates all species in the Abricta complex of genera except for the genus Abroma which is represented by the type species only. Character state changes shown are those present on all 12 most parsimonious trees. Black bars = non- homoplasious forward change; grey bars = homoplasious forward change; white bars = reversal (whether homoplasious or not). (16) Strict consensus tree derived from the six most parsimonious trees when multistate characters1,11,12,14,18, 19and20 wereordered.Character state changes shown are those present on all three most parsimonious trees.All other data as for Fig. 15." captionText-1="Fig. 17. Preferred tree of the six most parsimonious trees when multistate characters 1, 11, 12, 14, 18, 19 and 20 were ordered.All other data as for Fig. 15." pageId="18" pageNumber="263">Figs. 15–17</figureCitation>
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). See also discussion under
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<emphasis italics="true" pageId="18" pageNumber="263">Rationale for new taxa</emphasis>
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(pp. 254–255).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="263" type="distribution">
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<paragraph blockId="18.[839,1467,1188,1270]" pageId="18" pageNumber="263">
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<emphasis bold="true" box="[839,982,1188,1212]" pageId="18" pageNumber="263">Distribution</emphasis>
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. The single species included in this genus occurs in eastern
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<collectingCountry box="[1035,1134,1218,1241]" name="Australia" pageId="18" pageNumber="263">Australia</collectingCountry>
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from northern
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<collectingRegion box="[1308,1435,1218,1241]" country="Australia" name="Queensland" pageId="18" pageNumber="263">Queensland</collectingRegion>
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to southern
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<collectingRegion box="[939,1135,1247,1270]" country="Australia" name="New South Wales" pageId="18" pageNumber="263">New South Wales</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |