292 lines
46 KiB
XML
292 lines
46 KiB
XML
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<mods:titleInfo id="CCF352F5A35796A53DD58C6E95BCF147">
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<mods:title id="222182674CC1279F8DDACDE09CAA70EF">Harvestmen (Arachnida: Opiliones) in Eocene Rovno amber (Ukraine)</mods:title>
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</mods:titleInfo>
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<mods:name id="46509DF83D110F53648FD82A0AA3D047" type="personal">
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<mods:roleTerm id="3C3C8D95DC0DC2C0A837AB39832B31AB">Author</mods:roleTerm>
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<mods:namePart id="8F7078A51C5CF3EDE276516FA70EE91B">Mitov, Plamen G.</mods:namePart>
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<mods:roleTerm id="6416DB61698358E964AFF2409FFBEADC">Author</mods:roleTerm>
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<mods:namePart id="B7C042A8FD7C95E54E096915E0D647E0">Perkovsky, Evgeny E.</mods:namePart>
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<mods:namePart id="AAFC67D39BCC59446DBA2D40EE3BFBA9">Dunlop, Jason A.</mods:namePart>
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<mods:nameIdentifier id="CEF3B5DB603A20E3CE6DE3F5C4DD2310" type="email">jason.dunlop@mfn.berlin</mods:nameIdentifier>
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<mods:title id="80FA34854E9CB5C16AEE5C98C3FAD91E">Zootaxa</mods:title>
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<mods:date id="DF2D92789CA7F6206D28FAF6A0C2EED3">2021</mods:date>
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<mods:number id="E54A26EDF532D22F49F4BF0FE1030CED">2021-06-10</mods:number>
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<mods:number id="EA96AD11CC0F05846B96F2E3DCC77AB7">4984</mods:number>
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<treatment id="038106260C63717918C0FF40FD9F405A" ID-DOI="http://doi.org/10.5281/zenodo.4928218" ID-GBIF-Taxon="184587116" ID-Zenodo-Dep="4928218" LSID="urn:lsid:plazi:treatment:038106260C63717918C0FF40FD9F405A" httpUri="http://treatment.plazi.org/id/038106260C63717918C0FF40FD9F405A" lastPageId="14" lastPageNumber="57" pageId="10" pageNumber="53">
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<subSubSection id="C332E4BB0C63717D18C0FF40FDC342D6" box="[151,547,151,178]" pageId="10" pageNumber="53" type="nomenclature">
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<paragraph id="8B97B7300C63717D18C0FF40FDC342D6" blockId="10.[151,547,151,213]" box="[151,547,151,178]" pageId="10" pageNumber="53">
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<heading id="D0DF005C0C63717D18C0FF40FDC342D6" bold="true" box="[151,547,151,178]" fontSize="11" level="1" pageId="10" pageNumber="53" reason="1">
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<emphasis id="B95C6B220C63717D18C0FF40FDC342D6" bold="true" box="[151,547,151,178]" pageId="10" pageNumber="53">
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<taxonomicName id="4C28CCB30C63717D18C0FF40FE3942D6" ID-CoL="9X5HZ" authority="Mitov & Perkovsky & Dunlop, 2021" authorityName="Mitov & Perkovsky & Dunlop" authorityYear="2021" box="[151,473,151,178]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="53" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
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<emphasis id="B95C6B220C63717D18C0FF40FE3942D6" bold="true" box="[151,473,151,178]" italics="true" pageId="10" pageNumber="53">Metaphalangium martensi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26FD6590C63717D19B7FF40FDC342D6" box="[480,547,151,177]" pageId="10" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C332E4BB0C63717D18C0FF6CFEBA42B2" box="[151,346,187,213]" pageId="10" pageNumber="53" type="description">
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<paragraph id="8B97B7300C63717D18C0FF6CFEBA42B2" blockId="10.[151,547,151,213]" box="[151,346,187,213]" pageId="10" pageNumber="53">
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<figureCitation id="1313ABB50C63717D18C0FF6CFEE842B2" box="[151,264,187,213]" captionStart-0="FIGURES 9–10" captionStart-1="FIGURES 11–12" captionStartId-0="11.[151,264,1930,1954]" captionStartId-1="12.[151,264,1808,1832]" captionTargetBox-0="[282,1308,237,1904]" captionTargetBox-1="[223,1364,181,1782]" captionTargetId-0="figure-54@11.[279,1308,237,1907]" captionTargetId-1="figure-18@12.[223,1364,181,1785]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 9–10. Metaphalangium martensi sp. n., holotype, SIZK K-3197, from Rovno amber (Klesov, Pugach quarry, Ukraine), late Eocene (ca. 36–37 Ma). 9 Dorso-lateral overview of the complete animal. 10 Detail of body in dorso-lateral view. Scale bars: 2 mm." captionText-1="FIGURES 11–12. Metaphalangium spp., detail of the anterior prosoma and pedipalps. 11 Metaphalangium martensi sp. n. (holotype, SIZK K-3197, from Rovno amber, Klesov, Pugach quarry, Ukraine; late Eocene). Note the palpal patella with small and blunt medio-distal prominence (black arrow) with stout spine-like setae and mesial brush of stout spine-like setae on palpal tibia (white arrow), ventral view. 12 Comparative image of the extant species Metaphalangium cirtanum (C. L. Koch) (juvenile from Albania: Zvërnec), white arrows indicate palpal patella and tibia, mesially covered with stout spine-like setae, dorso-lateral view. Scale bars: 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4928504" figureDoi-1="http://doi.org/10.5281/zenodo.4928506" httpUri-0="https://zenodo.org/record/4928504/files/figure.png" httpUri-1="https://zenodo.org/record/4928506/files/figure.png" pageId="10" pageNumber="53">Figs 9–11</figureCitation>
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,
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<figureCitation id="1313ABB50C63717D1944FF6CFEBA42B2" box="[275,346,187,213]" captionStart="FIGURES 13–14" captionStartId="13.[151,264,1885,1909]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-18@13.[151,1435,181,1861]" captionTargetPageId="13" captionText="FIGURES 13–14. Metaphalangium martensi sp. n., paratype, SIZK K-1254, from Rovno amber (Klesov, Pugach quarry, Ukraine), late Eocene. 13 Lateral overview of the complete animal (white arrow points the pedipalps). 14 Detail of pedipalps, lateral. Scale bars: 13: 1 mm; 14: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4928510" httpUri="https://zenodo.org/record/4928510/files/figure.png" pageId="10" pageNumber="53">13–14</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C332E4BB0C63717D18C0FED4FBD54302" pageId="10" pageNumber="53" type="materials_examined">
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<paragraph id="8B97B7300C63717D18C0FED4FBD54302" blockId="10.[151,1437,259,2014]" pageId="10" pageNumber="53">
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<emphasis id="B95C6B220C63717D18C0FED4FE3D437A" bold="true" box="[151,477,259,285]" pageId="10" pageNumber="53">
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Type material.
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<collectingCountry id="F33FF7A00C63717D1918FED4FE33437A" box="[335,467,259,285]" name="Ukraine" pageId="10" pageNumber="53">UKRAINE</collectingCountry>
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:
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</emphasis>
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<emphasis id="B95C6B220C63717D19B1FED4FDB4437A" box="[486,596,259,285]" italics="true" pageId="10" pageNumber="53">
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<typeStatus id="549309920C63717D19B1FED4FDB0437A" box="[486,592,259,285]" pageId="10" pageNumber="53" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED392FF50C63717D1A0BFED4FD7A437A" box="[604,666,259,285]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="10" pageNumber="53">SIZK</collectionCode>
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K-3197,
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<collectingRegion id="49EC79D20C63717D1B5EFED3FC46437A" box="[777,934,259,286]" country="Ukraine" name="Rivne" pageId="10" pageNumber="53">Rovno region</collectingRegion>
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, Sarny District, Klesov, Pugach quarry; late Eocene. Syninclusions:
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<collectionCode id="ED392FF50C63717D19FDFEF0FE084326" box="[426,488,295,321]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="10" pageNumber="53">SIZK</collectionCode>
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K-3197, stellate hairs/trichomes;
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<taxonomicName id="4C28CCB30C63717D1B25FEF0FC2A4326" box="[882,970,295,321]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="53" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
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:
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<taxonomicName id="4C28CCB30C63717D1B8FFEF0FB894326" box="[984,1129,295,321]" class="Insecta" family="Rhagionidae" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="53" phylum="Arthropoda" rank="family">Rhagionidae</taxonomicName>
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.
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<emphasis id="B95C6B220C63717D1C20FEF0FB0D4326" bold="true" box="[1143,1261,295,321]" pageId="10" pageNumber="53">
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<typeStatus id="549309920C63717D1C20FEF0FB084326" box="[1143,1256,295,321]" pageId="10" pageNumber="53" type="paratype">Paratype</typeStatus>
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:
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</emphasis>
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<collectionCode id="ED392FF50C63717D1CA0FEF0FAD54326" box="[1271,1333,295,321]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="10" pageNumber="53">SIZK</collectionCode>
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K-1254, with same data as for
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<typeStatus id="549309920C63717D19DBFE9CFE114302" box="[396,497,331,357]" pageId="10" pageNumber="53" type="holotype">holotype</typeStatus>
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, but syninclusions absent. Both from
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<collectingRegion id="49EC79D20C63717D1BC0FE9BFC024301" box="[919,994,332,358]" country="Ukraine" name="Rivne" pageId="10" pageNumber="53">Rovno</collectingRegion>
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amber.
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</paragraph>
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</subSubSection>
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<subSubSection id="C332E4BB0C63717D1890FEB8FB3343C9" pageId="10" pageNumber="53" type="etymology">
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<paragraph id="8B97B7300C63717D1890FEB8FB3343C9" blockId="10.[151,1437,259,2014]" pageId="10" pageNumber="53">
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<emphasis id="B95C6B220C63717D1890FEB8FEAE43EE" bold="true" box="[199,334,367,393]" pageId="10" pageNumber="53">Etymology.</emphasis>
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In honour of Prof. Jochen Martens (Mainz,
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<collectingCountry id="F33FF7A00C63717D1B6EFEB8FC4343EE" box="[825,931,367,393]" name="Germany" pageId="10" pageNumber="53">Germany</collectingCountry>
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) in recognition of his extensive contributions to arachnology/opilionology, and on the occasion of his 80th birthday; name in the genitive case.
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</paragraph>
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</subSubSection>
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<subSubSection id="C332E4BB0C63717D1890FE60FCD94006" pageId="10" pageNumber="53" type="diagnosis">
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<paragraph id="8B97B7300C63717D1890FE60FCD94006" blockId="10.[151,1437,259,2014]" pageId="10" pageNumber="53">
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<emphasis id="B95C6B220C63717D1890FE60FEDA43B6" bold="true" box="[199,314,439,465]" pageId="10" pageNumber="53">Diagnosis</emphasis>
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. Easily distinguished from all other fossil phalangiid species by the unique combination of: (1) palpal patella with a small and blunt medio-distal prominence, (2) palpal patella and tibia mesially covered with not densely spaced stout spine-like setae, typically arranged brush-like in 4–5 rows, (3) absence of a distal apophysis on the palpal tibia, (4) the ocular tubercle (ellipsoidal, normal in size) is up to twice its length from the frontal margin of the carapace, and (5) having thin and relatively long legs.
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</paragraph>
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</subSubSection>
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<subSubSection id="C332E4BB0C63717D1890FDBCFEE145F2" pageId="10" pageNumber="53" type="description">
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<paragraph id="8B97B7300C63717D1890FDBCFA994139" blockId="10.[151,1437,259,2014]" pageId="10" pageNumber="53">
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<emphasis id="B95C6B220C63717D1890FDBCFEB940E2" bold="true" box="[199,345,619,645]" pageId="10" pageNumber="53">Description.</emphasis>
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The
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<typeStatus id="549309920C63717D19C2FDBCFE1A40E2" box="[405,506,619,645]" pageId="10" pageNumber="53" type="holotype">holotype</typeStatus>
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, SIZK K-3197, is an almost complete specimen (
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<figureCitation id="1313ABB50C63717D1C7CFDBCFB4140E2" box="[1067,1185,619,645]" captionStart="FIGURES 9–10" captionStartId="11.[151,264,1930,1954]" captionTargetBox="[282,1308,237,1904]" captionTargetId="figure-54@11.[279,1308,237,1907]" captionTargetPageId="11" captionText="FIGURES 9–10. Metaphalangium martensi sp. n., holotype, SIZK K-3197, from Rovno amber (Klesov, Pugach quarry, Ukraine), late Eocene (ca. 36–37 Ma). 9 Dorso-lateral overview of the complete animal. 10 Detail of body in dorso-lateral view. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4928504" httpUri="https://zenodo.org/record/4928504/files/figure.png" pageId="10" pageNumber="53">Figs 9–10</figureCitation>
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), possibly hinting at a juvenile/immature, best seen in dorsal view, but missing the ventral surface of the body (together with the leg coxae, and parts of the pedipalpal coxae and femora). Body, partly dorsally hidden in a gas bubble, and by a white emulsion anterior to the ocular tubercle. Body ovoid, light colored (colouration in amber – pale gray-yellow) with a silverish shine; length ca. 1.75; width of prosoma 1.5; width of opisthosoma 1.65. Ocular tubercle broader than long: length 0.175, and width 0.30, with 2–3 tubercles on the eye rings; diameter of each lens 0.15. Distance from ocular tubercle to front of prosoma 0.325. Ocular tubercle up to two times of its length from the frontal margin of the carapace.
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</paragraph>
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<paragraph id="8B97B7300C63717D1890FCB0FCA7461A" blockId="10.[151,1437,259,2014]" pageId="10" pageNumber="53">
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Pedipalps (with sawn-off coxae and femurs; the latter have only their distal parts preserved, with a length of 0.25), with article lengths of: Pa: 0.2; Ti: 0.32; Ta: 0.65; the claw, 0.07. On the preserved distal parts of the femurs there are spine-like setae apically and ventrally, ordered roughly in two rows. Palpal patella with a small and blunt medio-distal prominence; palpal patella and tibia mesially covered with not densely spaced stout spine-like setae, specifically brush-like, ordered in 4–5 rows (
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<figureCitation id="1313ABB50C63717D1ACBFC20FD114676" box="[668,753,1015,1041]" captionStart="FIGURES 11–12" captionStartId="12.[151,264,1808,1832]" captionTargetBox="[223,1364,181,1782]" captionTargetId="figure-18@12.[223,1364,181,1785]" captionTargetPageId="12" captionText="FIGURES 11–12. Metaphalangium spp., detail of the anterior prosoma and pedipalps. 11 Metaphalangium martensi sp. n. (holotype, SIZK K-3197, from Rovno amber, Klesov, Pugach quarry, Ukraine; late Eocene). Note the palpal patella with small and blunt medio-distal prominence (black arrow) with stout spine-like setae and mesial brush of stout spine-like setae on palpal tibia (white arrow), ventral view. 12 Comparative image of the extant species Metaphalangium cirtanum (C. L. Koch) (juvenile from Albania: Zvërnec), white arrows indicate palpal patella and tibia, mesially covered with stout spine-like setae, dorso-lateral view. Scale bars: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4928506" httpUri="https://zenodo.org/record/4928506/files/figure.png" pageId="10" pageNumber="53">Fig. 11</figureCitation>
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: arrows). These spine-like setae are 0.05–0.07 high and the distance between them is 0.03–0.04. On the medial surface of the patella they number 16–18, while on the medial side of the tibia they are about 26–28. Similar spine-like setae are scattered on the surface of the palpal tarsus as well, along with thinner setae; tarsus ends in a claw (
|
||
<figureCitation id="1313ABB50C63717D1ABFFBB4FCD9461A" box="[744,825,1123,1149]" captionStart="FIGURES 11–12" captionStartId="12.[151,264,1808,1832]" captionTargetBox="[223,1364,181,1782]" captionTargetId="figure-18@12.[223,1364,181,1785]" captionTargetPageId="12" captionText="FIGURES 11–12. Metaphalangium spp., detail of the anterior prosoma and pedipalps. 11 Metaphalangium martensi sp. n. (holotype, SIZK K-3197, from Rovno amber, Klesov, Pugach quarry, Ukraine; late Eocene). Note the palpal patella with small and blunt medio-distal prominence (black arrow) with stout spine-like setae and mesial brush of stout spine-like setae on palpal tibia (white arrow), ventral view. 12 Comparative image of the extant species Metaphalangium cirtanum (C. L. Koch) (juvenile from Albania: Zvërnec), white arrows indicate palpal patella and tibia, mesially covered with stout spine-like setae, dorso-lateral view. Scale bars: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4928506" httpUri="https://zenodo.org/record/4928506/files/figure.png" pageId="10" pageNumber="53">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B97B7300C63717D1890FB5FFE164756" blockId="10.[151,1437,259,2014]" pageId="10" pageNumber="53">
|
||
Legs relatively complete (
|
||
<figureCitation id="1313ABB50C63717D19BEFB50FDBC46C6" box="[489,604,1159,1185]" captionStart="FIGURES 9–10" captionStartId="11.[151,264,1930,1954]" captionTargetBox="[282,1308,237,1904]" captionTargetId="figure-54@11.[279,1308,237,1907]" captionTargetPageId="11" captionText="FIGURES 9–10. Metaphalangium martensi sp. n., holotype, SIZK K-3197, from Rovno amber (Klesov, Pugach quarry, Ukraine), late Eocene (ca. 36–37 Ma). 9 Dorso-lateral overview of the complete animal. 10 Detail of body in dorso-lateral view. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4928504" httpUri="https://zenodo.org/record/4928504/files/figure.png" pageId="10" pageNumber="53">Figs 9–10</figureCitation>
|
||
), all elongate and slender, and when preserved at their full length ending in a single claw. Leg article length: Leg I (Fe: 1.85, Pa: 0.5, Ti: 1.6, Mt: 1.75, Ta: 2.85–3.0); Leg II (Fe: 3.7, Pa: 0.5–0.6, Ti: 3.5, Mt: 3.25, Ta: 7.75); Leg III (Fe: 1.9, Pa: 0.35; full length of tibia, metatarsus, and tarsus not preserved [i.e., sawn-off]); Leg IV (Fe: 2.75, Pa: 0.55, Ti: 2.0, Mt: 2.85; full length of tarsus not preserved [partially sawn-off]). Index of leg length (IL) = 1.23.
|
||
</paragraph>
|
||
<paragraph id="8B97B7300C63717D1890FAECFDDD4782" blockId="10.[151,1437,259,2014]" pageId="10" pageNumber="53">The leg segments are cylindrical; femurs with longitudinal rows of acute inclined tubercles; thinnest are the femurs of the second pair of legs, thickest those of the first and third pairs. Their patellae bear dorsally and ventrally sparse setae and dorso-distally two tubercles each. Tibia distally slightly dorso-ventrally flattened, with sparse setae laterally and with more setae on the ventral side. Metatarsus with few setae on all sides. Tarsi with short, fine and numerous setae on their ventral sides.</paragraph>
|
||
<paragraph id="8B97B7300C63717D1890FA38FEE145F2" blockId="10.[151,1437,259,2014]" pageId="10" pageNumber="53">
|
||
The
|
||
<typeStatus id="549309920C63717D18ACFA27FEBE446D" box="[251,350,1520,1546]" pageId="10" pageNumber="53" type="paratype">paratype</typeStatus>
|
||
, SIZK K-1254, is enclosed in very dark amber (
|
||
<figureCitation id="1313ABB50C63717D1BD5FA38FC36446E" box="[898,982,1519,1545]" captionStart="FIGURES 13–14" captionStartId="13.[151,264,1885,1909]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-18@13.[151,1435,181,1861]" captionTargetPageId="13" captionText="FIGURES 13–14. Metaphalangium martensi sp. n., paratype, SIZK K-1254, from Rovno amber (Klesov, Pugach quarry, Ukraine), late Eocene. 13 Lateral overview of the complete animal (white arrow points the pedipalps). 14 Detail of pedipalps, lateral. Scale bars: 13: 1 mm; 14: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4928510" httpUri="https://zenodo.org/record/4928510/files/figure.png" pageId="10" pageNumber="53">Fig. 13</figureCitation>
|
||
). During processing of the amber piece, the ventral body surface was sawn off with parts of the legs. Body length 1.62; prosoma width 1.25, abdomen width 1.25, possibly also hinting at a juvenile/immature. The body dorsally, the ocular tubercle, the chelicerae, the pedipalps and parts of the legs are covered with a white emulsion. Therefore, only part of the pedipalpal and leg segments can be seen and measured. Pedipalps (
|
||
<figureCitation id="1313ABB50C63717D1AE9F9A8FCF244FE" box="[702,786,1663,1689]" captionStart="FIGURES 13–14" captionStartId="13.[151,264,1885,1909]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-18@13.[151,1435,181,1861]" captionTargetPageId="13" captionText="FIGURES 13–14. Metaphalangium martensi sp. n., paratype, SIZK K-1254, from Rovno amber (Klesov, Pugach quarry, Ukraine), late Eocene. 13 Lateral overview of the complete animal (white arrow points the pedipalps). 14 Detail of pedipalps, lateral. Scale bars: 13: 1 mm; 14: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4928510" httpUri="https://zenodo.org/record/4928510/files/figure.png" pageId="10" pageNumber="53">Fig. 14</figureCitation>
|
||
), with article lengths of: Fe: 0.35–0.4, Pa: 0.21, Ti: 0.37, Ta: 0.62. Palpal femurs dorso-apically and ventrally with stout spine-like setae, palpal patella and tibia medially also with stout spine-like setae. Palpal patella and tibia dorsally with setae; palpal tarsus with many setae; tarsus ends with a claw. Palps covered with a white emulsion, thus features of the setae appear much thicker than they would have been in life – compare with the
|
||
<typeStatus id="549309920C63717D1ADBF8D8FD0E454E" box="[652,750,1807,1833]" pageId="10" pageNumber="53" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="1313ABB50C63717D1AA9F8D8FCB0454E" box="[766,848,1807,1833]" captionStart="FIGURES 11–12" captionStartId="12.[151,264,1808,1832]" captionTargetBox="[223,1364,181,1782]" captionTargetId="figure-18@12.[223,1364,181,1785]" captionTargetPageId="12" captionText="FIGURES 11–12. Metaphalangium spp., detail of the anterior prosoma and pedipalps. 11 Metaphalangium martensi sp. n. (holotype, SIZK K-3197, from Rovno amber, Klesov, Pugach quarry, Ukraine; late Eocene). Note the palpal patella with small and blunt medio-distal prominence (black arrow) with stout spine-like setae and mesial brush of stout spine-like setae on palpal tibia (white arrow), ventral view. 12 Comparative image of the extant species Metaphalangium cirtanum (C. L. Koch) (juvenile from Albania: Zvërnec), white arrows indicate palpal patella and tibia, mesially covered with stout spine-like setae, dorso-lateral view. Scale bars: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4928506" httpUri="https://zenodo.org/record/4928506/files/figure.png" pageId="10" pageNumber="53">Fig. 11</figureCitation>
|
||
). Leg article length: Leg I (Fe: ca. 1.5, Pa: 0.37, Ti: 1.25, Mt: 1.75, Ta: 1.125); Leg II (Fe: 2.75, Pa: 0.37); Leg III (Fe: 1.3, Pa: 0.36); Leg IV: femur is sawn off, the rest of it 2.0 mm long. Index of leg length (IL) = 1.2. The remaining leg segments are sawn off or cannot be measured correctly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332E4BB0C6371791890F848FD9F405A" lastPageId="14" lastPageNumber="57" pageId="10" pageNumber="53" type="discussion">
|
||
<paragraph id="8B97B7300C63717C1890F848FB3C42B2" blockId="10.[151,1437,259,2014]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="54" pageId="10" pageNumber="53">
|
||
<emphasis id="B95C6B220C63717D1890F848FEDB45DE" bold="true" box="[199,315,1951,1977]" pageId="10" pageNumber="53">Remarks.</emphasis>
|
||
The key characters of this new fossil
|
||
<taxonomicName id="4C28CCB30C63717D1AB4F848FC1A45DE" box="[739,1018,1951,1977]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="53" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B95C6B220C63717D1AB4F848FC4245DE" box="[739,930,1951,1977]" italics="true" pageId="10" pageNumber="53">Metaphalangium</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
are the small mesal projection on the palpal patella, in combination with the characteristically arranged stout spine-like setae along the entire mesal surface of palpal patella and tibia (
|
||
<figureCitation id="1313ABB50C62717C19A0FF40FDAD42D6" box="[503,589,151,177]" captionStart="FIGURES 11–12" captionStartId="12.[151,264,1808,1832]" captionTargetBox="[223,1364,181,1782]" captionTargetId="figure-18@12.[223,1364,181,1785]" captionTargetPageId="12" captionText="FIGURES 11–12. Metaphalangium spp., detail of the anterior prosoma and pedipalps. 11 Metaphalangium martensi sp. n. (holotype, SIZK K-3197, from Rovno amber, Klesov, Pugach quarry, Ukraine; late Eocene). Note the palpal patella with small and blunt medio-distal prominence (black arrow) with stout spine-like setae and mesial brush of stout spine-like setae on palpal tibia (white arrow), ventral view. 12 Comparative image of the extant species Metaphalangium cirtanum (C. L. Koch) (juvenile from Albania: Zvërnec), white arrows indicate palpal patella and tibia, mesially covered with stout spine-like setae, dorso-lateral view. Scale bars: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4928506" httpUri="https://zenodo.org/record/4928506/files/figure.png" pageId="11" pageNumber="54">Figs 11</figureCitation>
|
||
,
|
||
<figureCitation id="1313ABB50C62717C1A0FFF40FD9742D6" box="[600,631,151,177]" captionStart="FIGURES 13–14" captionStartId="13.[151,264,1885,1909]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-18@13.[151,1435,181,1861]" captionTargetPageId="13" captionText="FIGURES 13–14. Metaphalangium martensi sp. n., paratype, SIZK K-1254, from Rovno amber (Klesov, Pugach quarry, Ukraine), late Eocene. 13 Lateral overview of the complete animal (white arrow points the pedipalps). 14 Detail of pedipalps, lateral. Scale bars: 13: 1 mm; 14: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4928510" httpUri="https://zenodo.org/record/4928510/files/figure.png" pageId="11" pageNumber="54">14</figureCitation>
|
||
). However, these stout spine-like setae (in the
|
||
<typeStatus id="549309920C62717C1C28FF40FB0142D6" box="[1151,1249,151,177]" pageId="11" pageNumber="54" type="holotype">holotype</typeStatus>
|
||
at least;
|
||
<figureCitation id="1313ABB50C62717C1D12FF40FA7342D6" box="[1349,1427,151,177]" captionStart="FIGURES 11–12" captionStartId="12.[151,264,1808,1832]" captionTargetBox="[223,1364,181,1782]" captionTargetId="figure-18@12.[223,1364,181,1785]" captionTargetPageId="12" captionText="FIGURES 11–12. Metaphalangium spp., detail of the anterior prosoma and pedipalps. 11 Metaphalangium martensi sp. n. (holotype, SIZK K-3197, from Rovno amber, Klesov, Pugach quarry, Ukraine; late Eocene). Note the palpal patella with small and blunt medio-distal prominence (black arrow) with stout spine-like setae and mesial brush of stout spine-like setae on palpal tibia (white arrow), ventral view. 12 Comparative image of the extant species Metaphalangium cirtanum (C. L. Koch) (juvenile from Albania: Zvërnec), white arrows indicate palpal patella and tibia, mesially covered with stout spine-like setae, dorso-lateral view. Scale bars: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4928506" httpUri="https://zenodo.org/record/4928506/files/figure.png" pageId="11" pageNumber="54">Fig. 11</figureCitation>
|
||
) are much thicker than those in extant forms; see the juvenile
|
||
<taxonomicName id="4C28CCB30C62717C1B16FF6CFB8C42B3" box="[833,1132,187,213]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="54" phylum="Arthropoda" rank="species" species="cirtanum">
|
||
<emphasis id="B95C6B220C62717C1B16FF6CFB8C42B3" box="[833,1132,187,213]" italics="true" pageId="11" pageNumber="54">Metaphalangium cirtanum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1313ABB50C62717C1C2BFF6CFB2F42B2" box="[1148,1231,187,213]" captionStart="FIGURES 11–12" captionStartId="12.[151,264,1808,1832]" captionTargetBox="[223,1364,181,1782]" captionTargetId="figure-18@12.[223,1364,181,1785]" captionTargetPageId="12" captionText="FIGURES 11–12. Metaphalangium spp., detail of the anterior prosoma and pedipalps. 11 Metaphalangium martensi sp. n. (holotype, SIZK K-3197, from Rovno amber, Klesov, Pugach quarry, Ukraine; late Eocene). Note the palpal patella with small and blunt medio-distal prominence (black arrow) with stout spine-like setae and mesial brush of stout spine-like setae on palpal tibia (white arrow), ventral view. 12 Comparative image of the extant species Metaphalangium cirtanum (C. L. Koch) (juvenile from Albania: Zvërnec), white arrows indicate palpal patella and tibia, mesially covered with stout spine-like setae, dorso-lateral view. Scale bars: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4928506" httpUri="https://zenodo.org/record/4928506/files/figure.png" pageId="11" pageNumber="54">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF57E7B80C62717C18C0F85DFEAE458D" ID-DOI="http://doi.org/10.5281/zenodo.4928504" ID-Zenodo-Dep="4928504" httpUri="https://zenodo.org/record/4928504/files/figure.png" pageId="11" pageNumber="54" startId="11.[151,264,1930,1954]" targetBox="[282,1308,237,1904]" targetPageId="11">
|
||
<paragraph id="8B97B7300C62717C18C0F85DFEAE458D" blockId="11.[151,1437,1930,2027]" pageId="11" pageNumber="54">
|
||
<emphasis id="B95C6B220C62717C18C0F85DFEAC45C4" bold="true" box="[151,332,1930,1955]" pageId="11" pageNumber="54">FIGURES 9–10.</emphasis>
|
||
<taxonomicName id="4C28CCB30C62717C1901F85CFD8745C5" authority="Mitov & Perkovsky & Dunlop, 2021" authorityName="Mitov & Perkovsky & Dunlop" authorityYear="2021" box="[342,615,1931,1955]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="54" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B95C6B220C62717C1901F85CFD8745C5" box="[342,615,1931,1955]" italics="true" pageId="11" pageNumber="54">Metaphalangium martensi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C6B220C62717C1A25F85CFD4F45C4" bold="true" box="[626,687,1931,1955]" pageId="11" pageNumber="54">
|
||
<taxonomicNameLabel id="A26FD6590C62717C1A25F85CFD4F45C4" box="[626,687,1931,1955]" pageId="11" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, SIZK K-3197, from Rovno amber (Klesov, Pugach quarry, Ukraine), late Eocene (ca. 36–37 Ma). 9 Dorso-lateral overview of the complete animal. 10 Detail of body in dorso-lateral view. Scale bars: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF57E7B80C65717B18C0F8C7FE7D45BB" ID-DOI="http://doi.org/10.5281/zenodo.4928506" ID-Zenodo-Dep="4928506" httpUri="https://zenodo.org/record/4928506/files/figure.png" pageId="12" pageNumber="55" startId="12.[151,264,1808,1832]" targetBox="[223,1364,181,1782]" targetPageId="12">
|
||
<paragraph id="8B97B7300C65717B18C0F8C7FE7D45BB" blockId="12.[151,1436,1808,2013]" pageId="12" pageNumber="55">
|
||
<emphasis id="B95C6B220C65717B18C0F8C7FEB5454F" bold="true" box="[151,341,1808,1832]" pageId="12" pageNumber="55">FIGURES 11–12.</emphasis>
|
||
<taxonomicName id="4C28CCB30C65717B1909F8C6FDA0454F" box="[350,576,1809,1833]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B95C6B220C65717B1909F8C6FDEB454E" box="[350,523,1809,1833]" italics="true" pageId="12" pageNumber="55">Metaphalangium</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, detail of the anterior prosoma and pedipalps. 11
|
||
<taxonomicName id="4C28CCB30C65717B1C1EF8C6FAB8454F" authority="Mitov & Perkovsky & Dunlop, 2021" authorityName="Mitov & Perkovsky & Dunlop" authorityYear="2021" box="[1097,1368,1809,1833]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B95C6B220C65717B1C1EF8C6FAB8454F" box="[1097,1368,1809,1833]" italics="true" pageId="12" pageNumber="55">Metaphalangium martensi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C6B220C65717B1D36F8C6FA7C454E" bold="true" box="[1377,1436,1809,1833]" pageId="12" pageNumber="55">
|
||
<taxonomicNameLabel id="A26FD6590C65717B1D36F8C6FA7C454E" box="[1377,1436,1809,1833]" pageId="12" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype, SIZK K-3197, from Rovno amber, Klesov, Pugach quarry, Ukraine; late Eocene). Note the palpal patella with small and blunt medio-distal prominence (black arrow) with stout spine-like setae and mesial brush of stout spine-like setae on palpal tibia (white arrow), ventral view. 12 Comparative image of the extant species
|
||
<taxonomicName id="4C28CCB30C65717B1BF0F8AAFADC45F2" authority="(C. L. Koch)" baseAuthorityName="C. L. Koch" box="[935,1340,1917,1941]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="cirtanum">
|
||
<emphasis id="B95C6B220C65717B1BF0F8AAFB5545F3" box="[935,1205,1917,1941]" italics="true" pageId="12" pageNumber="55">Metaphalangium cirtanum</emphasis>
|
||
(C. L. Koch)
|
||
</taxonomicName>
|
||
(juvenile from Albania: Zvërnec), white arrows indicate palpal patella and tibia, mesially covered with stout spine-like setae, dorso-lateral view. Scale bars: 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF57E7B80C64717A18C0F88AFDA245DA" ID-DOI="http://doi.org/10.5281/zenodo.4928510" ID-Zenodo-Dep="4928510" httpUri="https://zenodo.org/record/4928510/files/figure.png" pageId="13" pageNumber="56" startId="13.[151,264,1885,1909]" targetBox="[151,1435,181,1859]" targetPageId="13">
|
||
<paragraph id="8B97B7300C64717A18C0F88AFDA245DA" blockId="13.[151,1437,1885,1981]" pageId="13" pageNumber="56">
|
||
<emphasis id="B95C6B220C64717A18C0F88AFEB84512" bold="true" box="[151,344,1885,1909]" pageId="13" pageNumber="56">FIGURES 13–14.</emphasis>
|
||
<taxonomicName id="4C28CCB30C64717A1936F88AFD924512" authority="Mitov & Perkovsky & Dunlop, 2021" authorityName="Mitov & Perkovsky & Dunlop" authorityYear="2021" box="[353,626,1885,1909]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="56" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B95C6B220C64717A1936F88AFD924512" box="[353,626,1885,1909]" italics="true" pageId="13" pageNumber="56">Metaphalangium martensi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C6B220C64717A1A2CF88AFD584512" bold="true" box="[635,696,1885,1909]" pageId="13" pageNumber="56">
|
||
<taxonomicNameLabel id="A26FD6590C64717A1A2CF88AFD584512" box="[635,696,1885,1909]" pageId="13" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, SIZK K-1254, from Rovno amber (Klesov, Pugach quarry, Ukraine), late Eocene. 13 Lateral overview of the complete animal (white arrow points the pedipalps). 14 Detail of pedipalps, lateral. Scale bars: 13: 1 mm; 14: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B97B7300C6771791890FF40FD9F405A" blockId="14.[151,1437,151,573]" pageId="14" pageNumber="57">
|
||
When apophyses are present in phalangiid harvestmen, they can be more prominent in immature instars and become less well expressed in adults (
|
||
<bibRefCitation id="EFB9CAC10C6771791A6CFF6CFCE142B2" author="Mitov, P. G. & Dunlop, J. A. & Penney, D." box="[571,769,186,213]" pageId="14" pageNumber="57" pagination="37 - 42" refId="ref18760" refString="Mitov, P. G., Dunlop, J. A. & Penney, D. (2015) A new species of Lacinius in amber (Arachnida: Opiliones). Fossil Record, 18, 37 - 42. https: // doi. org / 10.5194 / fr- 18 - 37 - 2015" type="journal article" year="2015">
|
||
Mitov
|
||
<emphasis id="B95C6B220C6771791AD0FF6CFD5F42B2" box="[647,703,187,213]" italics="true" pageId="14" pageNumber="57">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
: p. 41; PGM, pers. obs.) and representatives of
|
||
<taxonomicName id="4C28CCB30C6771791D5BFF6CFF2F429F" authorityName="Roewer" authorityYear="1911" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C6B220C6771791D5BFF6CFF2F429F" italics="true" pageId="14" pageNumber="57">Metaphalangium</emphasis>
|
||
</taxonomicName>
|
||
are no exception. In extant members of this genus only the adults (both sexes) of
|
||
<taxonomicName id="4C28CCB30C6771791C38FF08FEE1437A" authority="(Roewer)" baseAuthorityName="Roewer" baseAuthorityYear="1956" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="57" phylum="Arthropoda" rank="species" species="corsicum">
|
||
<emphasis id="B95C6B220C6771791C38FF08FA7D429F" box="[1135,1437,223,249]" italics="true" pageId="14" pageNumber="57">Metaphalangium corsicum</emphasis>
|
||
(Roewer)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C28CCB30C677179196DFED4FD16437A" authority="(Roewer)" baseAuthorityName="Roewer" baseAuthorityYear="1956" box="[314,758,259,285]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="57" phylum="Arthropoda" rank="species" species="lusitanicum">
|
||
<emphasis id="B95C6B220C677179196DFED4FD64437A" box="[314,644,259,285]" italics="true" pageId="14" pageNumber="57">Metaphalangium lusitanicum</emphasis>
|
||
(Roewer)
|
||
</taxonomicName>
|
||
have short palpal apophyses. Of the studied representatives of the genus (see Material and methods), in adults of
|
||
<taxonomicName id="4C28CCB30C6771791AA2FEF0FC614327" box="[757,897,295,320]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="57" phylum="Arthropoda" rank="species" species="cirtanum">
|
||
<emphasis id="B95C6B220C6771791AA2FEF0FC614327" box="[757,897,295,320]" italics="true" pageId="14" pageNumber="57">M. cirtanum</emphasis>
|
||
</taxonomicName>
|
||
(females) and
|
||
<taxonomicName id="4C28CCB30C6771791C7CFEF0FB404326" baseAuthorityName="Nosek" baseAuthorityYear="1905" box="[1067,1184,295,321]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="57" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B95C6B220C6771791C7CFEF0FB404326" box="[1067,1184,295,321]" italics="true" pageId="14" pageNumber="57">M. strandi</emphasis>
|
||
</taxonomicName>
|
||
(both sexes), only the palpal patella bears a mesially pronounced rounded protrusion at the distal end, while in juvenile
|
||
<taxonomicName id="4C28CCB30C6771791CB0FE9CFA924303" box="[1255,1394,331,356]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="57" phylum="Arthropoda" rank="species" species="cirtanum">
|
||
<emphasis id="B95C6B220C6771791CB0FE9CFA924303" box="[1255,1394,331,356]" italics="true" pageId="14" pageNumber="57">M. cirtanum</emphasis>
|
||
</taxonomicName>
|
||
the pedipalpal patellae bear clearly formed apophyses (
|
||
<figureCitation id="1313ABB50C6771791AB5FEB8FCD743EE" box="[738,823,367,393]" captionStart="FIGURES 11–12" captionStartId="12.[151,264,1808,1832]" captionTargetBox="[223,1364,181,1782]" captionTargetId="figure-18@12.[223,1364,181,1785]" captionTargetPageId="12" captionText="FIGURES 11–12. Metaphalangium spp., detail of the anterior prosoma and pedipalps. 11 Metaphalangium martensi sp. n. (holotype, SIZK K-3197, from Rovno amber, Klesov, Pugach quarry, Ukraine; late Eocene). Note the palpal patella with small and blunt medio-distal prominence (black arrow) with stout spine-like setae and mesial brush of stout spine-like setae on palpal tibia (white arrow), ventral view. 12 Comparative image of the extant species Metaphalangium cirtanum (C. L. Koch) (juvenile from Albania: Zvërnec), white arrows indicate palpal patella and tibia, mesially covered with stout spine-like setae, dorso-lateral view. Scale bars: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4928506" httpUri="https://zenodo.org/record/4928506/files/figure.png" pageId="14" pageNumber="57">Fig. 12</figureCitation>
|
||
). Conversely, in immature specimens of
|
||
<taxonomicName id="4C28CCB30C6771791D5BFEB8FEDB43CB" authority="Mitov & Perkovsky & Dunlop, 2021" authorityName="Mitov & Perkovsky & Dunlop" authorityYear="2021" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="57" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B95C6B220C6771791D5BFEB8FEDB43CB" italics="true" pageId="14" pageNumber="57">Metaphalangium martensi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C6B220C6771791912FE44FE6843CA" bold="true" box="[325,392,403,429]" pageId="14" pageNumber="57">
|
||
<taxonomicNameLabel id="A26FD6590C6771791912FE44FE6843CA" box="[325,392,403,429]" pageId="14" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
such apophyses are missing—the palpal patella bears only a small and blunt medio-distal prominence (
|
||
<figureCitation id="1313ABB50C677179197BFE60FE6043B6" box="[300,384,439,465]" captionStart="FIGURES 11–12" captionStartId="12.[151,264,1808,1832]" captionTargetBox="[223,1364,181,1782]" captionTargetId="figure-18@12.[223,1364,181,1785]" captionTargetPageId="12" captionText="FIGURES 11–12. Metaphalangium spp., detail of the anterior prosoma and pedipalps. 11 Metaphalangium martensi sp. n. (holotype, SIZK K-3197, from Rovno amber, Klesov, Pugach quarry, Ukraine; late Eocene). Note the palpal patella with small and blunt medio-distal prominence (black arrow) with stout spine-like setae and mesial brush of stout spine-like setae on palpal tibia (white arrow), ventral view. 12 Comparative image of the extant species Metaphalangium cirtanum (C. L. Koch) (juvenile from Albania: Zvërnec), white arrows indicate palpal patella and tibia, mesially covered with stout spine-like setae, dorso-lateral view. Scale bars: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4928506" httpUri="https://zenodo.org/record/4928506/files/figure.png" pageId="14" pageNumber="57">Fig. 11</figureCitation>
|
||
: black arrow). Differences can also be seen in the morphology of the legs: in both immature fossil specimens they are somewhat more slender than in extant juvenile
|
||
<taxonomicName id="4C28CCB30C6771791BB3FE0CFB434392" authorityName="Roewer" authorityYear="1911" box="[996,1187,475,501]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95C6B220C6771791BB3FE0CFB434392" box="[996,1187,475,501]" italics="true" pageId="14" pageNumber="57">Metaphalangium</emphasis>
|
||
</taxonomicName>
|
||
, and the index of leg length (IL) in the smallest studied juvenile
|
||
<taxonomicName id="4C28CCB30C6771791A29FE28FCE8407F" box="[638,776,511,536]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="57" phylum="Arthropoda" rank="species" species="cirtanum">
|
||
<emphasis id="B95C6B220C6771791A29FE28FCE8407F" box="[638,776,511,536]" italics="true" pageId="14" pageNumber="57">M. cirtanum</emphasis>
|
||
</taxonomicName>
|
||
(with L: 3.3) is 1.06., while in
|
||
<taxonomicName id="4C28CCB30C6771791C3DFE28FB12407F" authority="Mitov & Perkovsky & Dunlop, 2021" authorityName="Mitov & Perkovsky & Dunlop" authorityYear="2021" box="[1130,1266,511,536]" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="57" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B95C6B220C6771791C3DFE28FB12407F" box="[1130,1266,511,536]" italics="true" pageId="14" pageNumber="57">M. martensi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95C6B220C6771791CADFE28FADA407E" bold="true" box="[1274,1338,511,537]" pageId="14" pageNumber="57">
|
||
<taxonomicNameLabel id="A26FD6590C6771791CADFE28FADA407E" box="[1274,1338,511,537]" pageId="14" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(with L: 1.62–1.75), IL varies between 1.2 and 1.23.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |