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<mods:title id="B16D05F8DEB2A3116FCE879180DFF668">The genus Cephaloleia Chevrolat, 1836 (Coleoptera, Chrysomelidae, Cassidinae)</mods:title>
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<mods:namePart id="57E7EF97F81B75FC3D8D3DF9BF990AB3">Staines, Charles L.</mods:namePart>
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<mods:namePart id="3A314339346635EFAA18336B66D12884">Garcia-Robledo, Carlos</mods:namePart>
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<mods:identifier id="B3FD782F369C086A1329353F23F71FED" type="DOI">http://dx.doi.org/10.3897/zookeys.436.5766</mods:identifier>
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<treatment id="9A4280E11DA8769D10258A80230CFF90" ID-GBIF-Taxon="152055251" LSID="urn:lsid:plazi:treatment:9A4280E11DA8769D10258A80230CFF90" httpUri="http://treatment.plazi.org/id/9A4280E11DA8769D10258A80230CFF90" lastPageId="219" lastPageNumber="220" pageId="218" pageNumber="219" scope_family="Chrysomelidae" scope_order="Coleoptera" scope_subFamily="Cassidinae">
<subSubSection id="EADB9D0DAF761AAA82E893BD9B62D3B1" pageId="218" pageNumber="219" type="multiple">
<paragraph id="3010C12D44331A56377364D22962047D" pageId="218" pageNumber="219">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
</subSubSection>
<subSubSection id="3F3E0DCC5A49A1BE70C48A0A5FF1D291" pageId="218" pageNumber="219" type="nomenclature">
<paragraph id="010D41D49189AAEA0F1409152D7B6536" pageId="218" pageNumber="219">
<taxonomicName id="54F33FB391A5B26FC62860420A091EB7" ID-CoL="SDFZ" LSID="http://species-id.net/wiki/Cephaloleia_princeps" authority="Baly, 1858" authorityName="Baly" authorityYear="1858" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia princeps" order="Coleoptera" pageId="218" pageNumber="219" phylum="Arthropoda" rank="species" species="princeps">Cephaloleia princeps Baly, 1858</taxonomicName>
Fig. 215
</paragraph>
</subSubSection>
<subSubSection id="18F39A454641DE04BA397934AB4B3B45" pageId="218" pageNumber="219" type="reference_group">
<paragraph id="0C0D46ABE4C7126F1D61830D541D2083" pageId="218" pageNumber="219">
<taxonomicName id="A19457FE93BC1ED8604BCC10AD7C8222" genus="Cephalolia" lsidName="Cephalolia princeps" pageId="218" pageNumber="219" rank="species" species="princeps">Cephalolia princeps</taxonomicName>
<bibRefCitation id="26528E6DE03C0A80DAE437E842C93D90" author="Baly, JS" journalOrPublisher="London" pageId="284" pageNumber="285" title="Catalogue of Hispidae in the collection of the British Museum. Part I" year="1858">Baly 1858</bibRefCitation>
: 45.
<bibRefCitation id="BBD37003A51C48BDCA58DDC74C6A9748" author="Gemminger, M" journalOrPublisher="Williams and Norgate, London" pageId="287" pageNumber="288" pagination="3479 - 3822" title="Catalogus Coleopterum hucusque descriptorum, synonymicus et systematicus" year="1876">Gemminger and Harold 1876</bibRefCitation>
: 3602 (catalog);
<bibRefCitation id="21AA2D8A89A29CC94788BE02B555F210" author="Donckier, H" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="286" pageNumber="287" pagination="540 - 615" title="Catalogue systematique des Hispides." volume="68" year="1899">Donckier 1899</bibRefCitation>
: 550 (catalog);
<bibRefCitation id="865DB833CF21E7B807B36F17DA101ED4" author="Weise, J" journalOrPublisher="Archiv fuer Naturgeschite" pageId="297" pageNumber="298" pagination="67 - 127" title="Beitrag zur Kenntnis der amerikanischer Hispinen." volume="76" year="1910">Weise 1910</bibRefCitation>
: 84 (noted),
<bibRefCitation id="B50BE522129B8D78DC593EF346289213" author="Weise, J" journalOrPublisher="W. Junk" pageId="297" pageNumber="298" pagination="1 - 94" title="Coleopterorum Catalogus, Chrysomelidae: Hispinae" year="1911 a">1911a</bibRefCitation>
: 9 (catalog),
<bibRefCitation id="34C1BBC6E7495DE227478599F50F0BDC" author="Weise, J" editor="Wytsman, P" journalOrPublisher="Genera Insectorum, Brussels" pageId="297" pageNumber="298" pagination="1 - 123" title="Coleoptera Phytophaga fam. Chrysomelidae, subfam. Hispinae." volume="fasc. 125" year="1911 b">1911b</bibRefCitation>
: 10 (catalog); Uhmann 1932c: 261 (noted),
<bibRefCitation id="2E8C4C016F202EECB7E2261ADB827F62" author="Uhmann, E" journalOrPublisher="Revista de Entomologia Rio de Janiero" pageId="295" pageNumber="296" pagination="481 - 485" title="Schluessel der mir bekannten Cephaloleia-Arten (Col. Chrys.). 65. Beitrag zur Kenntnis der Hispinen." volume="6" year="1936 b">1936b</bibRefCitation>
: 110 (noted), 1936f: 481 (key), 1953d: 47 (faunal list).
</paragraph>
<paragraph id="2F34A92E564194DEC69F7ADCBE514726" pageId="218" pageNumber="219">
<taxonomicName id="19B3E6F3D66865AC03B42A3C9BA6F2B2" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia princeps" order="Coleoptera" pageId="218" pageNumber="219" phylum="Arthropoda" rank="species" species="princeps">Cephaloleia princeps</taxonomicName>
Baly.
<bibRefCitation id="DF5D34EBA747F455EDF5A07E2BBB3AE7" author="Waterhouse, CO" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="297" pageNumber="298" pagination="260 - 269" title="On the Coleopterous insects belonging to the family Hispidae collected by Mr. Buckley in Ecuador." volume="1881" year="1881">Waterhouse 1881</bibRefCitation>
: 261 (distribution);
<bibRefCitation id="D2BA762300D63162E6F02C163E03E9E4" author="Uhmann, E" journalOrPublisher="Arbeiten ueber Morphologische und Taxonomische Entomologie aus Berlin-Dahlem" pageId="295" pageNumber="296" pagination="93 - 102" title="Der Borstenkegel und das Pygidium bei Cephalolia-Arten. (Coleoptera: Chrysomelidae). 96. Beitrag zur Kenntnis der Hispinen." volume="9" year="1942">Uhmann 1942</bibRefCitation>
b: 110 (pygidium),
<bibRefCitation id="AA1CF16A149A1D71B936F943F7D0C379" author="Uhmann, E" journalOrPublisher="Beitraege zur Entomologie" pageId="296" pageNumber="297" pagination="35 - 42" title="Sechs neue Hispinen aus Suedamerika (Coleoptera: Chrysomelidae). 181. Beitrag zur Kenntnis der Hispinae." volume="7" year="1957 b">1957b</bibRefCitation>
: 24 (catalog);
<bibRefCitation id="86DDF775A2DF3AD585D569FD395EBB39" author="Staines, CL" journalOrPublisher="Beitraege zur Entomologie" pageId="293" pageNumber="294" pagination="489 - 530" title="Joseph Sugar Baly: The man and his entomological works." volume="49" year="1999">Staines and Staines 1999</bibRefCitation>
: 524 (Baly species list);
<bibRefCitation id="965A9BDB90E1EB534E118736AE600516" author="McKenna, DD" journalOrPublisher="Proceedings of the National Academy of Sciences" pageId="290" pageNumber="291" pagination="10947 - 10951" title="Tropical forests are both evolutionary cradles and museums of leaf beetle diversity." url="10.1073/pnas.0602712103" volume="103" year="2006">McKenna and Farrell 2006</bibRefCitation>
: 10949 (phylogeny).
</paragraph>
</subSubSection>
<subSubSection id="21A1DDDDA4F5E6E38E4488764EFACD93" lastPageId="219" lastPageNumber="220" pageId="218" pageNumber="219" type="description">
<paragraph id="20CDDE0CCD9638939EFDAEBE5631A692" pageId="218" pageNumber="219">Description.</paragraph>
<paragraph id="A720FDBA98746E79D14DEFCFF2C9F5AA" lastPageId="219" lastPageNumber="220" pageId="218" pageNumber="219">
Large; elongate; subparallel; subdepressed; shining; head, pronotum, scutellum, venter, and legs reddish; eyes and antennae black, elytra greenish-black.
<pageBreakToken id="BCC60545BE6FE9B67B482EB837A5F118" pageId="219" pageNumber="220" start="start">Head</pageBreakToken>
: vertex finely punctate, with small fovea; frons not projecting; slightly depressed between eyes. Antenna: as long as head and pronotum combined; slender; antennomere 1 obovate, incrassate, longer than 2; 2 transverse, triangular in male; 3 elongate, triangular in male, subequal to 1; 4-5 elongate, subequal in length, each shorter than 3; 4 triangular in male; 6-10 transverse, subequal in length; 11 2
<normalizedToken id="B65FFF100437B78035363AEEB0859179" originalValue="×">x</normalizedToken>
length 10, broadly rounded at apex; 1-5 punctate, glabrous; 6-11 setose. Pronotum: subquadrate; lateral margin dilated, straight for basal
<normalizedToken id="27F72BDA7DB5C37953B037196AD23FDE" originalValue="¾">3/4</normalizedToken>
, then rounding to anterior angle, canaliculate; anterior angle acute, slightly produced; posterior angle acute; anterior margin curved anteriorly; disc subconvex; surface deeply punctate, more dense laterally, sparse on disc; basal impression absent; pronotal length 2.3-2.7 mm; pronotal width 3.2-3.6 mm. Scutellum: pentagonal; impunctate. Elytron: lateral margin straight, smooth, narrowly margined; apex rounded, apical margin thickened; sutural angle without tooth; humerus rounded, not produced; slightly constricted behind humerus; faint carina present behind humerus along puncture row 7; disc flattened along suture; finely punctate-striate, rows converge and unite apically; pygidium finely punctate, rounded at apex; elytral length 8.4-9.0 mm; elytral width 4.0-4.4 mm. Venter: pro-, meso-, and metasterna impunctate; abdominal sterna punctate, each puncture with pale seta; suture between sterna 1 and 2 complete; last sternite with apical margin emarginate medially in male, sinuate in female. Leg: slender; punctate; tibia with fringe of setae on inner margin of apex. Total length: 11.0-11.7 mm.
</paragraph>
<caption id="7B54AE32E9887B9269BBFEBD108DF846" pageId="219" pageNumber="220">
<paragraph id="81F82E2019C9BCA86C1A2D93406C54DE" pageId="219" pageNumber="220">
Figures 215-223. Habitus. 215
<taxonomicName id="165D9F8A4E1ACB2BEF9B91EFAE08B17B" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia princeps" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="princeps">Cephaloleia princeps</taxonomicName>
216
<taxonomicName id="8744656356115D37DE87DB688425D2E4" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia proxima" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="proxima">Cephaloleia proxima</taxonomicName>
217
<taxonomicName id="A5CFE681F1B6544138C660AFEA2360E4" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia pulchella" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="pulchella">Cephaloleia pulchella</taxonomicName>
218
<taxonomicName id="6A833DDDA4313E8E1A9329175A0DCBAD" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia punctatissima" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="punctatissima">Cephaloleia punctatissima</taxonomicName>
219
<taxonomicName id="1AFABCA9DE7B73CA266DAE78D91C9AD9" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia puncticollis" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="puncticollis">Cephaloleia puncticollis</taxonomicName>
220
<taxonomicName id="854F5B3966F6B9AB873C145D97ABB2F6" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia quadrilineata" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="quadrilineata">Cephaloleia quadrilineata</taxonomicName>
221
<taxonomicName id="263575C123009DAEFFA2D191BDA95EAD" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia quinquemaculata" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="quinquemaculata">Cephaloleia quinquemaculata</taxonomicName>
222
<taxonomicName id="4ECD71E2A77BF0E0BB8A08F268732875" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia recondita" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="recondita">Cephaloleia recondita</taxonomicName>
223
<taxonomicName id="9F272A12354012C3D8E20EA71F2AD1E5" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia reventazonica" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="reventazonica">Cephaloleia reventazonica</taxonomicName>
. Scale bars equal 3 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="6A793BD30694F845DC92D69F4278F58F" pageId="219" pageNumber="220" type="diagnosis">
<paragraph id="DA1095C412841BC5889E49693CCD2075" pageId="219" pageNumber="220">Diagnosis.</paragraph>
<paragraph id="E3EDE279AB3AC338C883BC5D780F7012" pageId="219" pageNumber="220">
This species is similar to
<taxonomicName id="6A18AD3B9C4C4952A8C60A1D5DBC01F6" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia abdominalis" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="abdominalis">Cephaloleia abdominalis</taxonomicName>
,
<taxonomicName id="17AC5D87ADF156D38EB20868E67FC1F3" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia amazona" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="amazona">Cephaloleia amazona</taxonomicName>
,
<taxonomicName id="D0916672B23938E02554583F7E4263E4" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia steinhauseni" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="steinhauseni">Cephaloleia steinhauseni</taxonomicName>
,
<taxonomicName id="C8EE46D48132C16BB4A1CD6141A79604" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia susanae" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="susanae">Cephaloleia susanae</taxonomicName>
sp. n., and
<taxonomicName id="994A375288FB5EBDD7D6EA18D9FDB8D7" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia teutonica" order="Coleoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="teutonica">Cephaloleia teutonica</taxonomicName>
. It can be distinguished by the vertex of the head having a medial fovea and by the faint carina behind the humerus along puncture row 7 of the elytra.
</paragraph>
</subSubSection>
<subSubSection id="E468A333D6D95704C8092C617CA8A9AF" pageId="219" pageNumber="220" type="distribution">
<paragraph id="63D1C90FBE355504AFF3295BD2FE6CBB" pageId="219" pageNumber="220">Distribution.</paragraph>
<paragraph id="7016204E828EB2CFBDE5B419CF92C3B1" pageId="219" pageNumber="220">Colombia, Ecuador, Peru.</paragraph>
</subSubSection>
<subSubSection id="59C3607FA9682F9AE8FC8CBCB5FCFE0B" pageId="219" pageNumber="220" type="type material examined">
<paragraph id="28064780D929FDA29E7E54E7DA0C8FD3" pageId="219" pageNumber="220">Type material examined.</paragraph>
<paragraph id="92107B7869E45526CC74C689DB783C0C" pageId="219" pageNumber="220">Holotype male: Peru, Fraser [handwritten label]/ Saunders Coll. [printed label]/ Cephalolia princeps Baly, Peru [blue handwritten label] (BMNH).</paragraph>
</subSubSection>
<subSubSection id="482125AC0D25F974FE7169F262FDDFA4" pageId="219" pageNumber="220" type="specimens examined">
<paragraph id="E14C232431A85D0B81954849A5EC2A06" pageId="219" pageNumber="220">Specimens examined.</paragraph>
<paragraph id="02F4E81E080451D4C4BB57463538D034" pageId="219" pageNumber="220">
No label data (USNM). Colombia: Meta- Villavicencio, May 1946 (USNM). Ecuador: Morona Santiago- Macas (USNM). Imbabura- Cachabe to Paramba, February 1897 (USNM). Napo- Limonchcha, 7 June 1977 (USNM); Pununo, 20 August 1997 (CDFA, USNM);Limonchocha Reserve, 215 m, 10 August 1997 (CDFA); Sacha Lodge, 270 m, 22 March 1999 (SEMC, USNM), 3-13 April 1994, 23 March 1999 (SEMC); Shushufindi, 100 m, 11 August 1997 (CDFA); Sta. Cecilia, 340 m, 8 June- 1 August 1968 (SEMC). Orellana- 1 km S Onkone Gare Camp, Reserva Etnica Waorani, 216.3 m, 23 January 2006 (USNM); Yasuni area, 36 km S Pompeya, 15 August 1997 (USNM);
<normalizedToken id="BF89A71343D35C911B41F8EC96F4B7E0" originalValue="Estación">Estacion</normalizedToken>
Cientifica Yasuni, 215 m, 5-10 November 1999 (EGRC). Pichincha- above Chimba, 3000', August 1897 (USNM); Limonocha, 300 m, 31 March 1975 (EGRC); 10.6 km W Mindo, Mindo Road, 1375 m, 28 March 1999 (SEMC). Peru: no further data (ISNB). Total: 44.
</paragraph>
</subSubSection>
</treatment>
</document>