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<document ID-DOI="http://doi.org/10.5281/zenodo.6870843" ID-ISBN="978-84-16728-08-4" ID-Zenodo-Dep="6870843" approvalRequired="304" approvalRequired_for_illustrations="45" approvalRequired_for_matCits="75" approvalRequired_for_taxonomicNames="40" approvalRequired_for_textStreams="75" approvalRequired_for_treatments="69" checkinTime="1658335596803" checkinUser="felipe" docAuthor="Russell A. Mittermeier &amp; Don E. Wilson" docDate="2018" docId="3D474A54A01C8770FFF2AEC81586FE95" docLanguage="en" docName="hbmw_8_Soricidae_0332.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 8 Insectivores, Sloths and Colugos, Barcelona: Lynx Edicions" docTitle="Anourosorex squamipes Milne-Edwards 1872" docType="treatment" docVersion="3" lastPageNumber="447" masterDocId="C17E322CA0288744FF8DAB47125EFFF9" masterDocTitle="Soricidae" masterLastPageNumber="551" masterPageNumber="332" pageNumber="447" updateTime="1658350298165" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Soricidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2018</mods:dateIssued>
<mods:dateOther type="pubDate">2018-07-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 8 Insectivores, Sloths and Colugos</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>332</mods:start>
<mods:end>551</mods:end>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6870843</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-08-4</mods:identifier>
<mods:identifier type="Zenodo-Dep">6870843</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6869890" ID-Zenodo-Dep="6869890" LSID="urn:lsid:plazi:treatment:3D474A54A01C8770FFF2AEC81586FE95" httpUri="http://treatment.plazi.org/id/3D474A54A01C8770FFF2AEC81586FE95" lastPageNumber="447" pageId="52" pageNumber="447">
<heading pageId="52" pageNumber="447">
<subSubSection box="[127,206,1423,1469]" pageId="52" pageNumber="447" type="multiple">
<paragraph blockId="52.[122,1140,1423,1550]" box="[127,206,1423,1469]" pageId="52" pageNumber="447">
<figureCitation box="[127,206,1423,1469]" captionStart="Plate 18: Soricidae" captionStartId="48.[122,151,3054,3079]" captionTargetBox="[12,2744,15,3636]" captionTargetPageId="47" captionText="136. Elliots Short-tailed Shrew (Blarina hulophaga), 137. Northern Short-tailed Shrew (Blarina brevicaudus), 138. Southern Short-tailed Shrew (Blarina carolinensis), 139. Everglades Short-tailed Shrew (Blarina peninsulae), 140. Shermans Short-tailed Shrew (Blarina shermani), 141. Sichuan Short-tailed Shrew (Blarinella quadraticauda), 142. Burmese Short-tailed Shrew (Blarinella wardi), 143. Indochinese Short-tailed Shrew (Blarinella griselda), 144. Chinese Mole Shrew (Anourosorex squamaipes), 145. Taiwanese Mole Shrew (Anowrosorex yamashinai), 146. Assam Mole Shrew (Anowrosorex assamensis), 147. Giant Mole Shrew (Anourosorex schmid), 148. Desert Gray Shrew (Notiosorex crawfordi), 149. Cockrums Gray Shrew (Notiosorex cockrumi), 150. Large-eared Gray Shrew (Notiosorex evolis), 151. Villas Gray Shrew (Notiosorex villa), 152. Mexican Shrew (Megasorex gigas), 153. Taiwanese Brown-toothed (Shrew Epusoriculusfumidus), 154. Arboreal Brown-toothed Shrew (Episoriculus macrurus), 155. Bailey's Brown-toothed Shrew (Episoriculus baileyi), 156. Long-tailed Brown-toothed Shrew (Episoriculus leucops), 157. Hodgsons Brown-toothed Shrew (Episoriculus caudatus), 158. Sichuan Brown-toothed Shrew (Episoriculus sacratus), 159. Hidden Brown-toothed Shrew (Episoriculus umbrinus), 160. Nepalese Brown-toothed Shrew (Episoriculus soluensis), 161. Himalayan Shrew (Soriculus nigrescens), 162. De Wintons Brown-toothed Shrew (Chodsigoa hypsibia), 163. Pygmy Brown-toothed Shrew (Chodsigoa parva), 164. Smiths Brown-toothed Shrew (Chodsigoa smithii), 165. Salenskis Brown-toothed Shrew (Chodsigoa salenskii), 166. Dusky Brown-toothed Shrew (Chodsigoa furva), 167. Lesser Taiwanese Brown-toothed Shrew (Chodsigoa sodalis), 168. Van Sungs Brown-toothed Shrew (Chodsigoa caovansunga), 169. Hoffmanns Brown-toothed Shrew (Chodsigoa hoffmanni), 170. Lowes Brown-toothed Shrew (Chodsigoa parca), 171. Bornean Water Shrew (Chimarrogale phaeura), 172. Sumatran Water Shrew (Chimarrogale sumatrana), 173. Malayan Water Shrew (Chimarrogale hantu), 174. Chinese Water Shrew (Chimarrogale styani), 175. Himalayan Water Shrew (Chimarrogale himalayica), 176. Leanders Water Shrew (Chimarrogale leander), 177. Japanese Water Shrew (Chimarrogale platycephala), 178. Elegant Water Shrew (Nectogale elegans), 179. Mediterranean Water Shrew (Neomys anomalus), 180. Transcaucasian Water (Shrew Neomysteres), 181. Eurasian Water Shrew (Neomys fodiens)" figureDoi="http://doi.org/10.5281/zenodo.6871913" httpUri="https://zenodo.org/record/6871913/files/figure.png" pageId="52" pageNumber="447">144.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[224,654,1423,1469]" pageId="52" pageNumber="447" type="vernacular_names">
<paragraph blockId="52.[122,1140,1423,1550]" box="[224,654,1423,1469]" pageId="52" pageNumber="447">
<vernacularName box="[224,654,1423,1469]" pageId="52" pageNumber="447">Chinese Mole Shrew</vernacularName>
</paragraph>
</subSubSection>
<subSubSection box="[721,1139,1423,1469]" pageId="52" pageNumber="447" type="nomenclature">
<paragraph blockId="52.[122,1140,1423,1550]" box="[721,1139,1423,1469]" pageId="52" pageNumber="447">
<taxonomicName authorityName="Milne-Edwards" authorityYear="1872" box="[721,1139,1423,1469]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="genus" species="squamipes">
<emphasis box="[721,1139,1423,1469]" italics="true" pageId="52" pageNumber="447">Anourosorex squamaipes</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="447" type="vernacular_names">
<paragraph blockId="52.[122,1140,1423,1550]" box="[124,1072,1487,1508]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[124,199,1487,1508]" pageId="52" pageNumber="447">French:</emphasis>
<vernacularName box="[209,389,1487,1508]" pageId="52" pageNumber="447">Musaraigne taupe</vernacularName>
/
<emphasis bold="true" box="[409,500,1487,1508]" pageId="52" pageNumber="447">German:</emphasis>
<vernacularName box="[510,707,1487,1508]" pageId="52" pageNumber="447">Maulwurfspitzmaus</vernacularName>
/
<emphasis bold="true" box="[728,819,1487,1508]" pageId="52" pageNumber="447">Spanish:</emphasis>
<vernacularName box="[829,1072,1487,1508]" pageId="52" pageNumber="447">Musarana topo de China</vernacularName>
</paragraph>
<paragraph blockId="52.[122,1140,1423,1550]" box="[123,1061,1527,1548]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[123,370,1527,1548]" pageId="52" pageNumber="447">Other common names:</emphasis>
<vernacularName box="[379,653,1527,1548]" pageId="52" pageNumber="447">Chinese Short-tailed Shrew</vernacularName>
,
<vernacularName box="[667,788,1527,1548]" pageId="52" pageNumber="447">Mole Shrew</vernacularName>
,
<vernacularName box="[801,1061,1527,1548]" pageId="52" pageNumber="447">Sichuan Burrowing Shrew</vernacularName>
</paragraph>
</subSubSection>
</heading>
<subSubSection pageId="52" pageNumber="447" type="reference_group">
<paragraph blockId="52.[736,1329,1597,2020]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[736,891,1597,1626]" pageId="52" pageNumber="447">Taxonomy.</emphasis>
<taxonomicName authorityName="Milne-Edwards" authorityYear="1872" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="squamipes">Anourosorex squamipes Milne- Edwards, 1872</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="447" type="materials_examined">
<paragraph blockId="52.[736,1329,1597,2020]" pageId="52" pageNumber="447">
<materialsCitation country="China" location="probably Moupin (= Baox- ing)" pageId="52" pageNumber="447" specimenCount="1" stateProvince="Sichuan">
<location LSID="urn:lsid:plazi:treatment:3D474A54A01C8770FFF2AEC81586FE95:B031AD99A01C8770FC31AD271142F950" country="China" name="probably Moupin (= Baox- ing)" pageId="52" pageNumber="447" stateProvince="Sichuan">probably Moupin (= Baox- ing)</location>
,
<collectingRegion box="[814,928,1680,1705]" country="China" name="Sichuan" pageId="52" pageNumber="447">Sichuan</collectingRegion>
,
<collectingCountry box="[945,1031,1680,1705]" name="China" pageId="52" pageNumber="447">China</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="447" type="discussion">
<paragraph blockId="52.[736,1329,1597,2020]" pageId="52" pageNumber="447">
<taxonomicName authorityName="Milne-Edwards" authorityYear="1872" box="[737,1032,1715,1744]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="squamipes">Anourosorex squamipes</taxonomicName>
is the type species of the genus, which was considered mono- typic until recently.
<taxonomicName authorityName="Milne-Edwards" authorityYear="1872" box="[1029,1329,1794,1823]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="squamipes">Anourosorex squamipes</taxonomicName>
used to include
<taxonomicName authorityName="Anderson" authorityYear="1875" box="[991,1128,1838,1863]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="assamensis">assamensis</taxonomicName>
,
<taxonomicName box="[1153,1250,1838,1863]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="schmidi">schmidi</taxonomicName>
, and
<taxonomicName authorityName="Kuroda" authorityYear="1935" box="[736,886,1869,1902]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="yamashinai">yamashinai</taxonomicName>
as subspecies, all of which are currently recognized as full species based on karyotypic differences between
<taxonomicName authorityName="Kuroda" authorityYear="1935" box="[736,886,1991,2020]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="yamashinai">yamashinai</taxonomicName>
and
<taxonomicName authorityName="Milne-Edwards" authorityYear="1872" box="[981,1110,1991,2020]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="squamipes">squamipes</taxonomicName>
and morpho-
</paragraph>
<paragraph blockId="52.[119,1330,2027,3478]" pageId="52" pageNumber="447">
logical differences among
<taxonomicName authorityName="Anderson" authorityYear="1875" box="[502,639,2027,2060]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="assamensis">assamensis</taxonomicName>
,
<taxonomicName authorityName="Milne-Edwards" authorityYear="1872" box="[656,786,2027,2060]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="squamipes">squamipes</taxonomicName>
, and
<taxonomicName authorityName="Kuroda" authorityYear="1935" box="[865,1016,2027,2060]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="yamashinai">yamashinai</taxonomicName>
. The forms capito and capnias are considered synonyms, both of which were collected in Yunnan, China.
<taxonomicName authorityName="Milne-Edwards" authorityYear="1872" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="squamipes">Anourosorex squamipes</taxonomicName>
is sister to A.
<taxonomicName authorityName="Kuroda" authorityYear="1935" box="[586,737,2110,2139]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="yamashinai">yamashinai</taxonomicName>
. Specimens from Mizoram and Assam in India that were assigned to A.
<taxonomicName authorityName="Milne-Edwards" authorityYear="1872" box="[554,683,2145,2178]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="squamipes">squamipes</taxonomicName>
are obviously A.
<taxonomicName authorityName="Anderson" authorityYear="1875" box="[926,1063,2145,2178]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="assamensis">assamensis</taxonomicName>
, which means that A.
<taxonomicName authorityName="Milne-Edwards" authorityYear="1872" box="[163,292,2185,2218]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="squamipes">squamipes</taxonomicName>
is not distributed in India. It is unclear where the distributional boundary is, especially between A.
<taxonomicName authorityName="Anderson" authorityYear="1875" box="[470,607,2224,2257]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="assamensis">assamensis</taxonomicName>
and A.
<taxonomicName authorityName="Milne-Edwards" authorityYear="1872" box="[716,846,2224,2257]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="52" pageNumber="447" phylum="Chordata" rank="species" species="squamipes">squamipes</taxonomicName>
. Monotypic.
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="447" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6871180" ID-Zenodo-Dep="6871180" httpUri="https://zenodo.org/record/6871180/files/figure.png" inLine="true" pageId="52" pageNumber="447" targetBox="[122,713,1601,2015]" targetPageId="52">
<paragraph blockId="52.[119,1330,2027,3478]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[124,300,2263,2296]" pageId="52" pageNumber="447">Distribution.</emphasis>
Widely distributed in C &amp; S China (from S Gansu S to Guangdong), adjacent N &amp; E Myanmar, N Laos, N Vietnam, and N Thailand.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="52" pageNumber="447" type="description">
<paragraph blockId="52.[119,1330,2027,3478]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[124,377,2342,2375]" pageId="52" pageNumber="447">Descriptive notes.</emphasis>
Head-body 74-110 mm, tail 8-19 mm, hindfoot 11-16 mm. No specific data are available for body weight. Condylo-incisive lengths are 23-3-26-5 mm, and upper tooth rows are 10-7-12:6 mm. The Chinese Mole Shrew is medium-sized, characterized by small eyes and short tail. It resembles a shrew mole in external appearance but with smaller claws. Body fur is dense and lax. Dorsal pelage is black or dark grayish brown, and ventral pelage is slightly paler. Eyes are small. External ears are completely reduced, and only slim openings can be traced. Forefeet have lengthened claws. Tail is short and dark brown. It has a two upper unicuspids, and first is longer than second. Upper P*and M' are quadrangle, and M* is usually triangular. Upper M? is much reduced. Cusps of teeth are unpigmented. It has very strong smell. Dental formula for all species of Anourosorexis13/2, C0/0,P 1/1, M 3/3 (x2) = 26. Chromosomal complement has 2n = 48 and FNa = 92.
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="447" type="biology_ecology">
<paragraph blockId="52.[119,1330,2027,3478]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[125,236,2823,2848]" pageId="52" pageNumber="447">Habitat.</emphasis>
Various habitats including broad-leaved, conifer, bamboo forests, subalpine shrubs, and grassland at elevations of 300-4000 m. The Chinese Mole Shrew is well adapted to anthropogenic habitats including farms, plantations, gardens, and parks.
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="447" type="food_feeding">
<paragraph blockId="52.[119,1330,2027,3478]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[125,391,2937,2966]" pageId="52" pageNumber="447">Food and Feeding.</emphasis>
The Chinese Mole Shrew is insectivorous, foraging on a wide variety of invertebrates. Examination of stomach contents found earthworms, insects, insect larvae, spiders, and frogs. Captive individuals ate raw meat, eels, fish, and occasionally oatmeal.
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="447" type="breeding">
<paragraph blockId="52.[119,1330,2027,3478]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[126,260,3091,3124]" pageId="52" pageNumber="447">Breeding.</emphasis>
Breeding of the Chinese Mole Shrew in south-western China occurs in April-October. Litters have 3-5 young.
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="447" type="activity">
<paragraph blockId="52.[119,1330,2027,3478]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[124,361,3169,3202]" pageId="52" pageNumber="447">Activity patterns.</emphasis>
Most Chinese Mole Shrews were trapped during dusk and at night. It is terrestrial and semi-fossorial, moving on the ground and frequently using tunnels.
</paragraph>
</subSubSection>
<subSubSection box="[126,1064,3248,3281]" pageId="52" pageNumber="447" type="biology_ecology">
<paragraph blockId="52.[119,1330,2027,3478]" box="[126,1064,3248,3281]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[126,826,3248,3281]" pageId="52" pageNumber="447">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="447" type="conservation">
<paragraph blockId="52.[119,1330,2027,3478]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[126,478,3291,3320]" pageId="52" pageNumber="447">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. The Chinese Mole Shrew is among the most common species of shrews in China. It is considered a pest in Sichuan, because populations can be large in farmland. It is known to be a host of hantavirus. Population size is highly variable but unlikely to be declining overall.
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="447" type="bibRefCitation_list">
<paragraph blockId="52.[1395,2600,298,366]" pageId="52" pageNumber="447">
<emphasis bold="true" box="[1396,1549,298,323]" pageId="52" pageNumber="447">Bibliography.</emphasis>
Hutterer (1985, 2005), Kawada et al. (2014), Mandal &amp; Das (1969), Mandal et al. (1995), Motokawa &amp; Lin Liangkong (2002), Motokawa et al. (2004).
</paragraph>
</subSubSection>
</treatment>
</document>