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<document id="BCB6176C49AE02DB02AC7FF3EF4A7F9B" ID-CLB-Dataset="7262" ID-DOI="10.11646/zootaxa.4954.1.1" ID-GBIF-Dataset="64eab05b-4539-4be2-bb06-7af4369949e0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4690775" ID-ZooBank="A8D2AA60-562C-4F98-8000-D792F1E40C87" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618451905616" checkinUser="plazi" docAuthor="Zahniser, James N." docDate="2021" docId="DE47C351FFFCC15DFF67DBB3FD897841" docLanguage="en" docName="zootaxa.4954.1.1.pdf" docOrigin="Zootaxa 4954 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tenucephalus ecuadorensis Zahniser 2021, n. sp." docType="treatment" docUuid="E84EC615-8CC7-4BEC-BBFA-5AD804BD6542" docUuidSource="ZooBank" docVersion="7" lastPageNumber="104" masterDocId="227EBB29FF9AC13AFFF0DE5DFFF67B54" masterDocTitle="Revision of the New World leafhopper tribe Faltalini (Hemiptera: Cicadellidae: Deltocephalinae) and the evolution of brachyptery" masterLastPageNumber="160" masterPageNumber="1" pageNumber="103" updateTime="1698936753110" updateUser="ExternalLinkService">
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<mods:title id="D21F22C90A5D7158767E630B06CC4D1E">Revision of the New World leafhopper tribe Faltalini (Hemiptera: Cicadellidae: Deltocephalinae) and the evolution of brachyptery</mods:title>
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<mods:namePart id="5EBDAF73B0A27D72BBD102BAA46E020D">Zahniser, James N.</mods:namePart>
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<mods:title id="042979AB8A8DC5B30D140155DBFAD5D8">Zootaxa</mods:title>
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<mods:date id="4C2177865D14FB5B619900EA89420A54">2021</mods:date>
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<mods:number id="EC7191E0868B963935BFB5ADE472F77A">2021-04-13</mods:number>
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<mods:number id="5F6C25A94B56D46078C2DD89438E253C">4954</mods:number>
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<mods:classification id="7B0B729B3BABF1A9278FD9B168168419">journal article</mods:classification>
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<mods:identifier id="DE693CD40366617C59BE230E01C516F9" type="DOI">10.11646/zootaxa.4954.1.1</mods:identifier>
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<treatment id="DE47C351FFFCC15DFF67DBB3FD897841" ID-DOI="http://doi.org/10.5281/zenodo.4701206" ID-GBIF-Taxon="181198302" ID-Zenodo-Dep="4701206" LSID="urn:lsid:zoobank.org:act:E84EC615-8CC7-4BEC-BBFA-5AD804BD6542" httpUri="http://treatment.plazi.org/id/DE47C351FFFCC15DFF67DBB3FD897841" lastPageId="103" lastPageNumber="104" pageId="102" pageNumber="103">
<subSubSection id="1EF421CCFFFCC15CFF67DBB3FDD97D5D" box="[151,559,1518,1545]" pageId="102" pageNumber="103" type="nomenclature">
<paragraph id="56517247FFFCC15CFF67DBB3FDD97D5D" blockId="102.[151,559,1518,1581]" box="[151,559,1518,1545]" pageId="102" pageNumber="103">
<heading id="0D19C52BFFFCC15CFF67DBB3FDD97D5D" bold="true" box="[151,559,1518,1545]" fontSize="11" level="2" pageId="102" pageNumber="103" reason="4">
<emphasis id="649AAE55FFFCC15CFF67DBB3FDD97D5D" bold="true" box="[151,559,1518,1545]" pageId="102" pageNumber="103">
<taxonomicName id="91EE09C4FFFCC15CFF67DBB3FE137D5D" ID-CoL="8KN7N" authority="Zahniser, 2021" authorityName="Zahniser" authorityYear="2021" box="[151,485,1518,1545]" class="Insecta" family="Cicadellidae" genus="Tenucephalus" kingdom="Animalia" order="Hemiptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="ecuadorensis" status="sp. nov.">
<emphasis id="649AAE55FFFCC15CFF67DBB3FE137D5D" bold="true" box="[151,485,1518,1545]" italics="true" pageId="102" pageNumber="103">Tenucephalus ecuadorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FA9132EFFFCC15CFE1CDBB2FDD97D5D" box="[492,559,1519,1545]" pageId="102" pageNumber="103" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1EF421CCFFFCC15CFF67D84EFED87D79" box="[151,302,1555,1581]" pageId="102" pageNumber="103" type="description">
<paragraph id="56517247FFFCC15CFF67D84EFED87D79" blockId="102.[151,559,1518,1581]" box="[151,302,1555,1581]" pageId="102" pageNumber="103">
(
<figureCitation id="CED56EC2FFFCC15CFF6FD84EFF0B7D79" box="[159,253,1555,1581]" captionStart="FIGUIRE 63" captionStartId="101.[151,259,1943,1967]" captionTargetBox="[168,1419,181,1917]" captionTargetId="figure-16@101.[167,1421,181,1920]" captionTargetPageId="101" captionText="FIGUIRE 63. Tenucephalus ecuadorensis n. sp. (AB, DH, J) standard views. (Q) aedeagus, ventral view with paired symmetrical processes of aedeagus-connective and phragma removed from image, apex of aedeagus at bottom of figure." figureDoi="http://doi.org/10.5281/zenodo.4690971" httpUri="https://zenodo.org/record/4690971/files/figure.png" pageId="102" pageNumber="103">Figs. 63</figureCitation>
,
<figureCitation id="CED56EC2FFFCC15CFEF9D84EFED07D79" box="[265,294,1555,1581]" captionStart="FIGUIRE 84" captionStartId="131.[151,259,1973,1997]" captionTargetBox="[247,1341,740,1949]" captionTargetId="figure-219@131.[247,1341,740,1949]" captionTargetPageId="131" captionText="FIGUIRE 84. Distribution of Tenucephalus spp. and Virganana danasensis in northwestern South America." figureDoi="http://doi.org/10.5281/zenodo.4691025" httpUri="https://zenodo.org/record/4691025/files/figure.png" pageId="102" pageNumber="103">84</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1EF421CCFFFCC15DFF67D807FED77AA1" lastPageId="103" lastPageNumber="104" pageId="102" pageNumber="103" type="diagnosis">
<paragraph id="56517247FFFCC15CFF67D807FCE17DB5" blockId="102.[151,1437,1626,2013]" pageId="102" pageNumber="103">
<emphasis id="649AAE55FFFCC15CFF67D807FEE47D20" bold="true" box="[151,274,1626,1652]" pageId="102" pageNumber="103">Diagnosis.</emphasis>
<taxonomicName id="91EE09C4FFFCC15CFEEAD806FE3D7D21" authorityName="Zahniser" authorityYear="2021" box="[282,459,1627,1653]" class="Insecta" family="Cicadellidae" genus="Tenucephalus" kingdom="Animalia" order="Hemiptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="649AAE55FFFCC15CFEEAD806FE3D7D21" box="[282,459,1627,1653]" italics="true" pageId="102" pageNumber="103">T. ecuadorensis</emphasis>
</taxonomicName>
can be distinguished from other species of the genus by a combination of the typical color pattern but transverse white band on anterior margin of head without thin brown or black line above and below, style apophysis thick, foot-like, and apex pointed and directed laterally, and aedeagus robust with dorsal surface roughened by numerous minute anteriorly-inclined teeth.
</paragraph>
<paragraph id="56517247FFFCC15CFF37D8B6FCB57C51" blockId="102.[151,1437,1626,2013]" box="[199,835,1771,1797]" pageId="102" pageNumber="103">
<emphasis id="649AAE55FFFCC15CFF37D8B6FEFD7C51" bold="true" box="[199,267,1771,1797]" pageId="102" pageNumber="103">Body.</emphasis>
Male,
<quantity id="9116DFA2FFFCC15CFEA9D8B6FE157C51" box="[345,483,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="102" pageNumber="103" unit="mm" value="4.05" valueMax="4.2" valueMin="3.9">3.94.2 mm</quantity>
. Female,
<quantity id="9116DFA2FFFCC15CFDBED8B6FD527C51" box="[590,676,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="102" pageNumber="103" unit="mm" value="4.9">4.9 mm</quantity>
to wing apex.
</paragraph>
<paragraph id="56517247FFFCC15CFF37D953FE397C19" blockId="102.[151,1437,1626,2013]" pageId="102" pageNumber="103">
<emphasis id="649AAE55FFFCC15CFF37D953FEE67C7C" bold="true" box="[199,272,1806,1832]" pageId="102" pageNumber="103">Color.</emphasis>
With typical color pattern but transverse white band on anterior margin of head without thin brown or black line above and below.
</paragraph>
<paragraph id="56517247FFFCC15DFF37D90AFC757ADD" blockId="102.[151,1437,1626,2013]" lastBlockId="103.[151,1437,150,789]" lastPageId="103" lastPageNumber="104" pageId="102" pageNumber="103">
<emphasis id="649AAE55FFFCC15CFF37D90AFEFD7C25" bold="true" box="[199,267,1879,1905]" pageId="102" pageNumber="103">Male.</emphasis>
Pygofer length about 2.5x height at midlength; basal half broad; dorsally incised at midlength; apical half lobe-like; ventral margin gently excavated near base; with ~20 long macrosetae on apical lobe; with many short to medium-length fine setae above and below membranous cleft and ventrally near apex; with pair of thick processes arising at base of dorsal incision at midlength, paralleling dorsal margin of pygofer for 4/5 their length and bent ventrad and attenuated at apical 1/5. Valve width about 2x median length; not produced anteriorly. Subgenital plate somewhat truncated apically, trapezoidal; with single row of 67 macrosetae near lateral margin; with many fine setae laterally. Style preapical lobe long, rectangular, apical angle rounded; apophysis thick, foot-like, apex pointed and directed laterally. Connective-aedeagus with symmetrical pair of processes fused near aedeagal base, processes relatively straight in lateral view, slightly undulate, sharply pointed at apex. Aedeagus robust, curving dorsad; with dorsal surface roughened by numerous minute anteriorly-inclined teeth. Phragma lobes with numerous distinct moderately long setae throughout. Segment X sclerotized laterally.
</paragraph>
<paragraph id="56517247FFFDC15DFF37DFCEFED77AA1" blockId="103.[151,1437,150,789]" pageId="103" pageNumber="104">
<emphasis id="649AAE55FFFDC15DFF37DFCEFED37AF9" bold="true" box="[199,293,403,429]" pageId="103" pageNumber="104">Female.</emphasis>
Sternite VII roughly rectangular; width less than 2x median length; posterior margin broadly incised and slightly produced medially. Sternite VIII membranous; arcuate wrinkled tissue slightly exceeding anterior margin of stVII.
</paragraph>
</subSubSection>
<subSubSection id="1EF421CCFFFDC15DFF37DFA3FB327935" pageId="103" pageNumber="104" type="materials_examined">
<paragraph id="56517247FFFDC15DFF37DFA3FB327935" blockId="103.[151,1437,150,789]" pageId="103" pageNumber="104">
<emphasis id="649AAE55FFFDC15DFF37DFA3FE42794C" bold="true" box="[199,436,510,536]" pageId="103" pageNumber="104">Material examined.</emphasis>
<materialsCitation id="E686781AFFFDC15DFE4EDFA2FE117969" ID-GBIF-Occurrence="3080386379" collectingDate="1976-07-22" collectingDateMax="1976-07-28" collectingDateMin="1976-07-22" collectionCode="USNM" country="Ecuador" county="Jeffrey Cohen" location="Santo Domingo" municipality="Malaise" pageId="103" pageNumber="104" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="8955CCE5FFFDC15DFE4EDFA2FDA0794D" box="[446,598,511,537]" pageId="103" pageNumber="104" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="40E8B9CEFFFDC15DFD94DFA2FD7E794D" box="[612,648,511,537]" pageId="103" pageNumber="104" type="male">1♂</specimenCount>
,
<collectingCountry id="2EF932D7FFFDC15DFD68DFA2FCD4794D" box="[664,802,511,537]" name="Ecuador" pageId="103" pageNumber="104">ECUADOR</collectingCountry>
, Pich.,
<location id="5331249CFFFDC15DFC8BDFA2FBD9794D" LSID="urn:lsid:plazi:treatment:DE47C351FFFCC15DFF67DBB3FD897841:5331249CFFFDC15DFC8BDFA2FBD9794D" box="[891,1071,511,537]" country="Ecuador" county="Jeffrey Cohen" municipality="Malaise" name="Santo Domingo" pageId="103" pageNumber="104">Santo Domingo</location>
(47KmS),
<collectingMunicipality id="B635E83DFFFDC15DFB43DFA2FAF8794D" box="[1203,1294,511,537]" pageId="103" pageNumber="104">Malaise</collectingMunicipality>
,
<date id="22505487FFFDC15DFAEDDFA2FF247969" pageId="103" pageNumber="104" value="1976-07-22" valueMax="1976-07-28" valueMin="1976-07-22">
<collectingDate id="3214AD6FFFFDC15DFAEDDFA2FF247969" pageId="103" pageNumber="104" value="1976-07-22" valueMax="1976-07-28" valueMin="1976-07-22">2228-VII- 1976</collectingDate>
</date>
,
<collectingCounty id="BF300ACBFFFDC15DFF2DDC7EFE8F7969" box="[221,377,547,573]" pageId="103" pageNumber="104">Jeffrey Cohen</collectingCounty>
[
<collectionCode id="30FFEA82FFFDC15DFE76DC7FFE167969" box="[390,480,546,573]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="103" pageNumber="104" type="Museum">USNM</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="E686781AFFFDC15DFE00DC7EFC347969" ID-GBIF-Occurrence="3080386441" box="[496,962,547,573]" collectingDate="1976-07-22" collectingDateMax="1976-07-28" collectingDateMin="1976-07-22" collectionCode="USNM" country="Ecuador" county="Jeffrey Cohen" location="Santo Domingo" municipality="Malaise" pageId="103" pageNumber="104" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="8955CCE5FFFDC15DFE00DC7EFD797969" box="[496,655,547,573]" pageId="103" pageNumber="104" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="40E8B9CEFFFDC15DFD69DC7EFD487969" box="[665,702,547,573]" pageId="103" pageNumber="104" type="male">3♂</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="E686781AFFFDC15DFC3CDC7EFF247935" ID-GBIF-Occurrence="3080386385" collectingDate="1976-07-29" collectingDateMax="1976-07-28" collectingDateMin="1976-07-22" collectionCode="USNM" country="Ecuador" county="Jeffrey Cohen" location="Santo Domingo" municipality="Malaise" pageId="103" pageNumber="104" specimenCount="4" specimenCount-male="4" typeStatus="paratype">
<specimenCount id="40E8B9CEFFFDC15DFC3CDC7EFC077969" box="[972,1009,547,573]" pageId="103" pageNumber="104" type="male">4♂</specimenCount>
, same data as holotype except
<date id="22505487FFFDC15DFAB7DC7EFF247935" pageId="103" pageNumber="104" value="1976-07-29">
<collectingDate id="3214AD6FFFFDC15DFAB7DC7EFF247935" pageId="103" pageNumber="104" value="1976-07-29">29-VII- 1976</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="E686781AFFFDC15DFF2DDC1AFB327935" ID-GBIF-Occurrence="3080386396" box="[221,1220,582,609]" collectingDate="1958-03-20" collectionCode="USNM" collectorName="R. W. Hodges." country="Ecuador" location="Sapayo Grande" pageId="103" pageNumber="104" specimenCount="1" specimenCount-female="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCount id="40E8B9CEFFFDC15DFF2DDC1AFEF67935" box="[221,256,583,609]" pageId="103" pageNumber="104" type="female">1♀</specimenCount>
,
<collectingCountry id="2EF932D7FFFDC15DFEFEDC1AFE6E7935" box="[270,408,583,609]" name="Ecuador" pageId="103" pageNumber="104">ECUADOR</collectingCountry>
,
<location id="5331249CFFFDC15DFE54DC1AFDA47935" LSID="urn:lsid:plazi:treatment:DE47C351FFFCC15DFF67DBB3FD897841:5331249CFFFDC15DFE54DC1AFDA47935" box="[420,594,583,609]" country="Ecuador" name="Sapayo Grande" pageId="103" pageNumber="104" stateProvince="Esmeraldas">Sapayo Grande</location>
,
<collectingRegion id="942ABCA5FFFDC15DFDADDC1AFCD57935" box="[605,803,583,609]" country="Ecuador" name="Esmeraldas" pageId="103" pageNumber="104">Esmeraldas Prov.</collectingRegion>
,
<date id="22505487FFFDC15DFCDDDC1BFC457935" box="[813,947,582,609]" pageId="103" pageNumber="104" value="1958-03-20">
<collectingDate id="3214AD6FFFFDC15DFCDDDC1BFC457935" box="[813,947,582,609]" pageId="103" pageNumber="104" value="1958-03-20">20-III-1958</collectingDate>
</date>
,
<collectorName id="FB1B1791FFFDC15DFC4EDC1AFBAD7935" box="[958,1115,583,609]" pageId="103" pageNumber="104">R.W. Hodges.</collectorName>
[
<collectionCode id="30FFEA82FFFDC15DFB98DC1BFB497935" box="[1128,1215,582,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="103" pageNumber="104" type="Museum">USNM</collectionCode>
]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1EF421CCFFFDC15DFF37DC36FC5279D1" box="[199,932,619,645]" pageId="103" pageNumber="104" type="etymology">
<paragraph id="56517247FFFDC15DFF37DC36FC5279D1" blockId="103.[151,1437,150,789]" box="[199,932,619,645]" pageId="103" pageNumber="104">
<emphasis id="649AAE55FFFDC15DFF37DC36FEB879D1" bold="true" box="[199,334,619,645]" pageId="103" pageNumber="104">Etymology.</emphasis>
The species name highlights its presence in
<collectingCountry id="2EF932D7FFFDC15DFCB1DC36FC5679D1" box="[833,928,619,645]" name="Ecuador" pageId="103" pageNumber="104">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1EF421CCFFFDC15DFF37DCD3FB647999" pageId="103" pageNumber="104" type="distribution">
<paragraph id="56517247FFFDC15DFF37DCD3FB647999" blockId="103.[151,1437,150,789]" pageId="103" pageNumber="104">
<emphasis id="649AAE55FFFDC15DFF37DCD3FE9779FC" bold="true" box="[199,353,654,680]" pageId="103" pageNumber="104">Distribution.</emphasis>
The
<typeStatus id="8955CCE5FFFDC15DFE6CDCD2FE3A79FD" box="[412,460,655,681]" pageId="103" pageNumber="104">type</typeStatus>
locality is interpreted from the labels to be in
<collectingRegion id="942ABCA5FFFDC15DFC2EDCD2FB4C79FD" box="[990,1210,655,681]" country="Ecuador" name="Pichincha" pageId="103" pageNumber="104">Pichincha Province</collectingRegion>
,
<quantity id="9116DFA2FFFDC15DFB37DCD2FAF979FD" box="[1223,1295,655,681]" metricMagnitude="4" metricUnit="m" metricValue="4.7" pageId="103" pageNumber="104" unit="km" value="47.0">47 km</quantity>
north of the city of Santo Domingo. So far, it is the only species of
<taxonomicName id="91EE09C4FFFDC15DFD0EDCEEFC617999" box="[766,919,691,717]" class="Insecta" family="Cicadellidae" genus="Tenucephalus" kingdom="Animalia" order="Hemiptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="649AAE55FFFDC15DFD0EDCEEFC617999" box="[766,919,691,717]" italics="true" pageId="103" pageNumber="104">Tenucephalus</emphasis>
</taxonomicName>
known from
<collectingCountry id="2EF932D7FFFDC15DFBDFDCEEFB797999" box="[1071,1167,691,717]" name="Ecuador" pageId="103" pageNumber="104">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1EF421CCFFFDC15DFF37DC8AFD897841" pageId="103" pageNumber="104" type="discussion">
<paragraph id="56517247FFFDC15DFF37DC8AFD897841" blockId="103.[151,1437,150,789]" pageId="103" pageNumber="104">
<emphasis id="649AAE55FFFDC15DFF37DC8AFECD79A5" bold="true" box="[199,315,727,753]" pageId="103" pageNumber="104">Remarks.</emphasis>
The association made here of the female from a separate locality with males from the
<typeStatus id="8955CCE5FFFDC15DFAE0DC8AFAB679A5" box="[1296,1344,727,753]" pageId="103" pageNumber="104">type</typeStatus>
locality should be confirmed with more collections.
</paragraph>
</subSubSection>
</treatment>
</document>