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<document id="F35774ECA8F82B1F0327F21CACF87483" ID-CLB-Dataset="7262" ID-DOI="10.11646/zootaxa.4954.1.1" ID-GBIF-Dataset="64eab05b-4539-4be2-bb06-7af4369949e0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4690775" ID-ZooBank="A8D2AA60-562C-4F98-8000-D792F1E40C87" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618451905616" checkinUser="plazi" docAuthor="Zahniser, James N." docDate="2021" docId="DE47C351FFBAC11BFF67DB5FFA6B7CBA" docLanguage="en" docName="zootaxa.4954.1.1.pdf" docOrigin="Zootaxa 4954 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Bonamus Oman 1937" docType="treatment" docVersion="5" lastPageNumber="34" masterDocId="227EBB29FF9AC13AFFF0DE5DFFF67B54" masterDocTitle="Revision of the New World leafhopper tribe Faltalini (Hemiptera: Cicadellidae: Deltocephalinae) and the evolution of brachyptery" masterLastPageNumber="160" masterPageNumber="1" pageNumber="33" updateTime="1698936753110" updateUser="ExternalLinkService">
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<mods:title id="CD612C43653CF02C6AAFCF7BF3F78C89">Revision of the New World leafhopper tribe Faltalini (Hemiptera: Cicadellidae: Deltocephalinae) and the evolution of brachyptery</mods:title>
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<taxonomicName id="91EE09C4FFBAC11AFF67DB5FFEFD7E48" ID-CoL="8HDYS" authorityName="Oman" authorityYear="1937" box="[151,267,1282,1308]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="649AAE55FFBAC11AFF67DB5FFEFD7E48" bold="true" box="[151,267,1282,1308]" italics="true" pageId="32" pageNumber="33">Bonamus</emphasis>
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<collectingCountry id="2EF932D7FFBAC11AFEE2DB5CFE967E4F" box="[274,352,1281,1307]" name="Oman" pageId="32" pageNumber="33">Oman</collectingCountry>
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<typeStatus id="8955CCE5FFBAC11AFF67DB1BFF3F7E09" box="[151,201,1350,1373]" pageId="32" pageNumber="33">Type</typeStatus>
species:
<taxonomicName id="91EE09C4FFBAC11AFED6DB1BFD947E0A" authority="Oman, 1938" authorityName="Oman" authorityYear="1938" box="[294,610,1350,1374]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="649AAE55FFBAC11AFED6DB1BFE2D7E0A" box="[294,475,1350,1374]" italics="true" pageId="32" pageNumber="33">Bonamus lineatus</emphasis>
<bibRefCitation id="327F0FB6FFBAC11AFE12DB1BFD947E0A" author="Oman, P. W." box="[482,610,1350,1374]" pageId="32" pageNumber="33" pagination="343 - 420" refId="ref80707" refString="Oman, P. W. (1938) A generic revision of American Bythoscopinae and South American Jassinae. University of Kansas Science Bulletin, 37, 343 - 420." type="journal article" year="1938">
<collectingCountry id="2EF932D7FFBAC11AFE12DB1BFDD47E0A" box="[482,546,1350,1374]" name="Oman" pageId="32" pageNumber="33">Oman</collectingCountry>
, 1938
</bibRefCitation>
</taxonomicName>
; designated.
</paragraph>
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<taxonomicName id="91EE09C4FFBAC11AFF67DB38FE407E28" ID-CoL="8HDYS" authority="Oman, 1938: 365" authorityName="Oman" authorityPageNumber="365" authorityYear="1938" box="[151,438,1380,1404]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="649AAE55FFBAC11AFF67DB38FF037E28" box="[151,245,1381,1404]" italics="true" pageId="32" pageNumber="33">Bonamus</emphasis>
<treatmentCitation id="D74F5456FFBAC11AFF0FDB39FE407E28" author="Oman, P. W." box="[255,438,1380,1404]" page="365" pageId="32" pageNumber="33" year="1938">
<bibRefCitation id="327F0FB6FFBAC11AFF0FDB39FE407E28" author="Oman, P. W." box="[255,438,1380,1404]" pageId="32" pageNumber="33" pagination="343 - 420" refId="ref80707" refString="Oman, P. W. (1938) A generic revision of American Bythoscopinae and South American Jassinae. University of Kansas Science Bulletin, 37, 343 - 420." type="journal article" year="1938">
<collectingCountry id="2EF932D7FFBAC11AFF0FDB39FEC97E28" box="[255,319,1380,1404]" name="Oman" pageId="32" pageNumber="33">Oman</collectingCountry>
, 1938: 365
</bibRefCitation>
</treatmentCitation>
</taxonomicName>
[original description, illustration, morphology, new species];
<treatmentCitation id="D74F5456FFBAC11AFBCEDB38FB037E28" author="Evans, J. W." box="[1086,1269,1380,1404]" page="140" pageId="32" pageNumber="33" year="1947">
<bibRefCitation id="327F0FB6FFBAC11AFBCEDB38FB037E28" author="Evans, J. W." box="[1086,1269,1380,1404]" pageId="32" pageNumber="33" pagination="105 - 271" refId="ref79346" refString="Evans, J. W. (1947) A natural classification of leaf-hoppers (Jassoidea, Homoptera). Part 3: Jassidae. Transactions of the Royal Entomological Society of London, 98, 105 - 271. https: // doi. org / 10.1111 / j. 1365 - 2311.1947. tb 01054. x" type="journal article" year="1947">Evans, 1947: 140</bibRefCitation>
</treatmentCitation>
[classification];
<treatmentCitation id="D74F5456FFBAC11AFF37DBDEFE5C7ECF" author="Linnavuori, R." box="[199,426,1411,1435]" page="138" pageId="32" pageNumber="33" year="1957">
<bibRefCitation id="327F0FB6FFBAC11AFF37DBDEFE5C7ECF" author="Linnavuori, R." box="[199,426,1411,1435]" pageId="32" pageNumber="33" pagination="133 - 143" refId="ref79921" refString="Linnavuori, R. (1957) The neotropical Hecalinae (Hom. Cicadellidae). Suomen Hyonteistieteellinen Aikakauskirja, 23 (3), 133 - 143." type="journal article" year="1957">Linnavuori, 1957: 138</bibRefCitation>
</treatmentCitation>
[description];
<treatmentCitation id="D74F5456FFBAC11AFDCEDBDEFCCC7ECF" author="Linnavuori, R." box="[574,826,1411,1435]" page="63" pageId="32" pageNumber="33" year="1959">
<bibRefCitation id="327F0FB6FFBAC11AFDCEDBDEFCCC7ECF" author="Linnavuori, R." box="[574,826,1411,1435]" pageId="32" pageNumber="33" pagination="1 - 370" refId="ref79950" refString="Linnavuori, R. (1959) Revision of the Neotropical Deltocephalinae and some related subfamilies (Homoptera). Annales Zoologici Societatis Zoologicae-Botanicae Fennicae ' Vanamo', 20, 1 - 370." type="journal article" year="1959">Linnavuori, 1959: 63–64</bibRefCitation>
</treatmentCitation>
[key];
<treatmentCitation id="D74F5456FFBAC11AFC73DBDEFB5C7ECF" author="Linnavuori, R. &amp; Heller, F." box="[899,1194,1411,1435]" page="5" pageId="32" pageNumber="33" year="1962">
<bibRefCitation id="327F0FB6FFBAC11AFC73DBDEFB5C7ECF" author="Linnavuori, R. &amp; Heller, F." box="[899,1194,1411,1435]" pageId="32" pageNumber="33" pagination="1 - 14" refId="ref80185" refString="Linnavuori, R. &amp; Heller, F. (1962) Beitrag zur Cicadelliden-Fauna von Peru. Entomologische Ergebnisse der Stuttgarter Anden- Expedition 1957 - Nr. 1. Stuttgarter Beitraege zur Naturkunde aus dem Staatlichen Museum fur Naturkunde in Stuttgart, 67, 1 - 14." type="journal article" year="1962">Linnavuori &amp; Heller, 1962: 5</bibRefCitation>
</treatmentCitation>
[new species];
<treatmentCitation id="D74F5456FFBAC11AFAB7DBDEFED77EEE" author="Metcalf, Z. P." page="60" pageId="32" pageNumber="33" year="1963">
<bibRefCitation id="327F0FB6FFBAC11AFAB7DBDEFED77EEE" author="Metcalf, Z. P." pageId="32" pageNumber="33" refId="ref80364" refString="Metcalf, Z. P. (1963) General Catalogue of the Homoptera. Fascicle VI. Cicdelloidea. Part 9. Hecalidae. U. S. Department of Agriculture, Agricultural Research Service, Washington, D. C., 123 pp." type="book" year="1963">Metcalf, 1963: 60</bibRefCitation>
</treatmentCitation>
[catalogue];
<treatmentCitation id="D74F5456FFBAC11AFE59DBFFFD897EEE" author="Hamilton, K. G. A." box="[425,639,1442,1466]" page="437" pageId="32" pageNumber="33" year="2000">
<bibRefCitation id="327F0FB6FFBAC11AFE59DBFFFD897EEE" author="Hamilton, K. G. A." box="[425,639,1442,1466]" pageId="32" pageNumber="33" pagination="429 - 503" refId="ref79635" refString="Hamilton, K. G. A. (2000) Five genera of New-World &quot; Shovel-Headed &quot; and &quot; Spoon-Bill &quot; leafhoppers (Hemiptera: Cicadellidae: Dorycephalini and Hecalini). Canadian Entomologist, 132 (4), 429 - 503. https: // doi. org / 10.4039 / Ent 132429 - 4" type="journal article" year="2000">Hamilton, 2000: 437</bibRefCitation>
</treatmentCitation>
, 453 [key; morphology; classification];
<treatmentCitation id="D74F5456FFBAC11AFBE9DBFFFB4A7EEE" author="Zanol, K. M. R." box="[1049,1212,1442,1466]" page="97" pageId="32" pageNumber="33" year="2006">
<bibRefCitation id="327F0FB6FFBAC11AFBE9DBFFFB4A7EEE" author="Zanol, K. M. R." box="[1049,1212,1442,1466]" pageId="32" pageNumber="33" pagination="89 - 161" refId="ref82441" refString="Zanol, K. M. R. (2006) Catalogue of the neotropical (including north of Mexico) Deltocephalinae (Hemiptera, Cicadellidae). Part I - Athysanini and Deltocephalini excluded Acta Biologica Paranaense, Curitiba, 35 (3 - 4), 89 - 161. https: // doi. org / 10.5380 / abpr. v 35 i 0.6869" type="journal article" year="2006">Zanol, 2006: 97</bibRefCitation>
</treatmentCitation>
[catalogue];
<treatmentCitation id="D74F5456FFBAC11AFAB4DBFFFE2A7E8C" author="Zahniser, J. N. &amp; Dietrich, C. H." page="507" pageId="32" pageNumber="33" year="2010">
<bibRefCitation id="327F0FB6FFBAC11AFAB4DBFFFE2A7E8C" author="Zahniser, J. N. &amp; Dietrich, C. H." pageId="32" pageNumber="33" pagination="489 - 511" refId="ref82181" refString="Zahniser, J. N. &amp; Dietrich, C. H. (2010) Phylogeny of the leafhopper subfamily Deltocephalinae (Hemiptera: Cicadellidae) based on molecular and morphological data with a revised family-group classification. Systematic Entomology, 35 (3), 489 - 511. https: // doi. org / 10.1111 / j. 1365 - 3113.2010.00522. x" type="journal article" year="2010">Zahniser &amp; Dietrich, 2010: 507–508</bibRefCitation>
</treatmentCitation>
[classification; morphology];
<treatmentCitation id="D74F5456FFBAC11AFCE2DB9DFBBE7E8C" author="Zahniser, J. N. &amp; Dietrich, C. H." box="[786,1096,1472,1496]" page="83" pageId="32" pageNumber="33" year="2013">
<bibRefCitation id="327F0FB6FFBAC11AFCE2DB9DFBBE7E8C" author="Zahniser, J. N. &amp; Dietrich, C. H." box="[786,1096,1472,1496]" pageId="32" pageNumber="33" pagination="1 - 211" refId="ref82248" refString="Zahniser, J. N. &amp; Dietrich, C. H. (2013) A review of the tribes of Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). European Journal of Taxonomy, 45, 1 - 211. https: // doi. org / 10.5852 / ejt. 2013.45" type="journal article" year="2013">Zahniser &amp; Dietrich, 2013: 83</bibRefCitation>
</treatmentCitation>
, 85 [classification; morphology];
<bibRefCitation id="327F0FB6FFBAC11AFF37DB82FEA87EA3" author="Zahniser, J. N." box="[199,350,1503,1527]" pageId="32" pageNumber="33" refId="ref82139" refString="Zahniser, J. N. (2007) An online interactive key and searchable database of Deltocephalinae (Hemiptera: Cicadellidae). Available from: http: // zahniser. speciesfile. org / (accessed 20 June 2020)" type="url" year="2007">Zahniser, 2007</bibRefCitation>
[online catalogue];
<bibRefCitation id="327F0FB6FFBAC11AFDD9DB82FCEC7EA3" author="Freytag, P. H. &amp; Gaiani, M." box="[553,794,1503,1527]" pageId="32" pageNumber="33" refId="ref79405" refString="Freytag, P. H. &amp; Gaiani, M. (2017) Neotropical Leafhoppers. Available from: http: // neotropicalcicadellidae. myspecies. info / (accessed 25 January 2020)" type="url" year="2017">Freytag &amp; Gaiani, 2017</bibRefCitation>
[online catalogue]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1EF421CCFFBAC11AFF67D878FAF57DA7" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="56517247FFBAC11AFF67D878FAF57DA7" blockId="32.[151,1437,1573,2032]" pageId="32" pageNumber="33">
<emphasis id="649AAE55FFBAC11AFF67D878FEE47D6B" bold="true" box="[151,274,1573,1599]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
<taxonomicName id="91EE09C4FFBAC11AFEE7D87BFE897D6B" authorityName="Oman" authorityYear="1937" box="[279,383,1574,1599]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="649AAE55FFBAC11AFEE7D87BFE897D6B" box="[279,383,1574,1599]" italics="true" pageId="32" pageNumber="33">Bonamus</emphasis>
</taxonomicName>
can be distinguished from other genera of the tribe by the macropterous males and females, the ocelli close to the eyes, anterior margin of head with numerous transverse carinae, discal portion of crown smooth, shiny, margin of pronotum much shorter than greatest width of eye, dorsum with distinct longitudinal orange stripes, subgenital plates with numerous long macrosetae arranged across entire plate, pygofer produced ventroapically into a process, pygofer with an internal lobelike process, connective articulated with aedeagus, ovipositor extending very far beyond pygofer apex, and first valvula dorsal sculpturing maculose and dorsoventrally elongated.
</paragraph>
</subSubSection>
<subSubSection id="1EF421CCFFBAC11BFF37D8A0FC8A78F1" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="56517247FFBAC11BFF37D8A0FAAA7A15" blockId="32.[151,1437,1573,2032]" lastBlockId="33.[151,1437,151,1473]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="649AAE55FFBAC11AFF37D8A0FEFD7C43" bold="true" box="[199,267,1789,1815]" pageId="32" pageNumber="33">Body.</emphasis>
Male,
<quantity id="9116DFA2FFBAC11AFEAED8A0FE1D7C4C" box="[350,491,1789,1816]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="7.2" metricValueMin="5.1" pageId="32" pageNumber="33" unit="mm" value="6.15" valueMax="7.2" valueMin="5.1">5.1–7.2 mm</quantity>
. Female,
<quantity id="9116DFA2FFBAC11AFDAAD8A0FD127C4C" box="[602,740,1789,1816]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="8.6" metricValueMin="5.5" pageId="32" pageNumber="33" unit="mm" value="7.05" valueMax="8.6" valueMin="5.5">5.5–8.6 mm</quantity>
to wing tip, 6.0–
<quantity id="9116DFA2FFBAC11AFC5DD8A0FBF37C4C" box="[941,1029,1789,1816]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="32" pageNumber="33" unit="mm" value="9.7">9.7 mm</quantity>
to ovipositor tip. Body length 4.5x width of pronotum (male), 4.5–5.0x (female). Anterior margin of head with numerous transverse carinae. Crown length 0.6x (male) and 0.8x (female) interocular width; texture mostly smooth with faint longitudinal striae; slightly depressed. Ocelli 0.5–1.0x their own diameter distant from eye; on anterior margin or slightly dorsad. Frontoclypeus texture shagreen; flat to concave, anteriorly abruptly angled dorsad to anterior margin of head. Antennal sockets situated close upper margin of face. Pronotum carinate laterally; lateral margin shorter than greatest width of eye; texture smooth. Scutellum texture smooth. Protrochanter without distinct setae. Profemur row AV with ~8–14 very short, stout setae; intercalary row with 9–10 thin setae; AM1 present; dorsally with pair of apical setae; row PD with distinct row of 3–4 macrosetae and some smaller setae. Protibia row AD with apical seta; row PD with 8–20 or more macrosetae. Mesofemur row AV with numerous short, stout setae. Mesotibia row AD with 6–8 macrosetae; row PD with 6–10 macrosetae. Metafemur apical setae 2+2+1. Metatibia gradually arched throughout length. Metatarsomere I long, slender; apex with row of 3 or 4 platellae, flanked on each side with tapered seta. Macropterous.
</paragraph>
<paragraph id="56517247FFBBC11BFF37DF17FDA07A85" blockId="33.[151,1437,151,1473]" pageId="33" pageNumber="34">
<emphasis id="649AAE55FFBBC11BFF37DF17FEE67A30" bold="true" box="[199,272,330,356]" pageId="33" pageNumber="34">Color.</emphasis>
General color whitish to yellowish or pale greenish. Head with three longitudinal orange stripes continuing onto pronotum and scutellum. Pronotum with additional pair of lateral orange stripes. Forewing with orange stripes continuing from pronotum and scutellum; with orange stripes in other cells of corium extending nearly to apical cells. Ventrally pale to yellowish.
</paragraph>
<paragraph id="56517247FFBBC11BFF37DF86FB3479A5" blockId="33.[151,1437,151,1473]" pageId="33" pageNumber="34">
<emphasis id="649AAE55FFBBC11BFF37DF86FEFD7AA1" bold="true" box="[199,267,475,501]" pageId="33" pageNumber="34">Male.</emphasis>
Pygofer incised dorsally nearly 2/3 its length to base; ventrally produced to large quadrate or broad spinelike process; without macrosetae. Subgenital plates relatively long, triangular; apices bluntly angled or rounded; with numerous long macrosetae across entire ventral surface. Valve long, triangular; length 0.7–0.8x width. Connective short; Y-shaped. Style broadly bilobed at base; preapical lobe bluntly quadrate. Aedeagus base about half as tall as shaft; shaft arising ventrally from base; curving dorsad; simple or with pair of apical processes; gonopore apical or on posterior margin; articulated with connective. Phragma sclerotized as setose lobe or pair of bulbous lobes articulating with dorsal rim of aedeagus; with an internal membranous process short and digitate (
<taxonomicName id="91EE09C4FFBBC11BFAECDCEEFA657999" box="[1308,1427,691,717]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="649AAE55FFBBC11BFAECDCEEFA657999" box="[1308,1427,691,717]" italics="true" pageId="33" pageNumber="34">B. lineatus</emphasis>
</taxonomicName>
) or elongate and lobelike (
<taxonomicName id="91EE09C4FFBBC11BFE45DC8AFDD879A5" authorityName="Zahniser" authorityYear="2021" box="[437,558,727,753]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="jussaral">
<emphasis id="649AAE55FFBBC11BFE45DC8AFDD879A5" box="[437,558,727,753]" italics="true" pageId="33" pageNumber="34">B. jussaral</emphasis>
</taxonomicName>
). Segment X large, long, sclerotized dorsally and laterally.
</paragraph>
<paragraph id="56517247FFBBC11BFF37DCA6FC8A78F1" blockId="33.[151,1437,151,1473]" pageId="33" pageNumber="34">
<emphasis id="649AAE55FFBBC11BFF37DCA6FED37841" bold="true" box="[199,293,763,789]" pageId="33" pageNumber="34">Female.</emphasis>
Pygofer with numerous macrosetae. Ovipositor elongate; extending very far beyond pygofer. Shape of sternite VII variable. First valvula slender, straight; dorsal sculptured area with dorsoventrally elongated maculalike texture, becoming more granular toward apex, submarginal with broad unsculptured band along dorsal margin; VSA present, distinctly delimited, sculpturing similar dorsal sculpturing. Second valvula lanceolate, narrow; without dorsal teeth. Gonoplac with distinct row of 12–16 macrosetae.
</paragraph>
</subSubSection>
<subSubSection id="1EF421CCFFBBC11BFF37DDF3FCA57F45" pageId="33" pageNumber="34" type="distribution">
<paragraph id="56517247FFBBC11BFF37DDF3FCA57F45" blockId="33.[151,1437,151,1473]" pageId="33" pageNumber="34">
<emphasis id="649AAE55FFBBC11BFF37DDF3FE97789C" bold="true" box="[199,353,942,968]" pageId="33" pageNumber="34">Distribution.</emphasis>
<collectingCountry id="2EF932D7FFBBC11BFE97DDF2FE2F789D" box="[359,473,943,969]" name="Argentina" pageId="33" pageNumber="34">Argentina</collectingCountry>
,
<collectingCountry id="2EF932D7FFBBC11BFE16DDF2FDD8789D" box="[486,558,943,969]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
. One undescribed species, apparently a member of this genus, is known from a single female from
<collectingCountry id="2EF932D7FFBBC11BFE7DDD8EFE3578B9" box="[397,451,979,1005]" name="Peru" pageId="33" pageNumber="34">Peru</collectingCountry>
,
<collectingRegion id="942ABCA5FFBBC11BFE20DD8EFDAC78B9" box="[464,602,979,1005]" country="Peru" name="Cusco" pageId="33" pageNumber="34">Cusco Prov.</collectingRegion>
(specimen at USNM). If confirmed by association with a male, it would substantially extend the known geographic range of the genus.
</paragraph>
</subSubSection>
<subSubSection id="1EF421CCFFBBC11BFF37DA46FE147DB5" pageId="33" pageNumber="34" type="discussion">
<paragraph id="56517247FFBBC11BFF37DA46FE057F91" blockId="33.[151,1437,151,1473]" pageId="33" pageNumber="34">
<emphasis id="649AAE55FFBBC11BFF37DA46FECD7F61" bold="true" box="[199,315,1051,1077]" pageId="33" pageNumber="34">Remarks.</emphasis>
<taxonomicName id="91EE09C4FFBBC11BFEB4DA46FCFD7F61" authority="Linnavuori &amp; Heller" authorityName="Linnavuori &amp; Heller" baseAuthorityName="Zahniser" baseAuthorityYear="2021" box="[324,779,1051,1077]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="amabilis">
<emphasis id="649AAE55FFBBC11BFEB4DA46FDE07F61" box="[324,534,1051,1077]" italics="true" pageId="33" pageNumber="34">Bonamus amabilis</emphasis>
Linnavuori &amp; Heller
</taxonomicName>
is transferred to
<taxonomicName id="91EE09C4FFBBC11BFC22DA46FB9D7F61" box="[978,1131,1051,1077]" class="Insecta" family="Cicadellidae" genus="Tenucephalus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="649AAE55FFBBC11BFC22DA46FB9D7F61" box="[978,1131,1051,1077]" italics="true" pageId="33" pageNumber="34">Tenucephalus</emphasis>
</taxonomicName>
here. The species was described based on a female specimen and possesses the –
<taxonomicName id="91EE09C4FFBBC11BFCEEDA62FC5E7F0C" authorityName="Zahniser" authorityYear="2021" box="[798,936,1087,1112]" class="Insecta" family="Cicadellidae" genus="Tenucephalus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="interstinctus">
<emphasis id="649AAE55FFBBC11BFCEEDA62FC5E7F0C" box="[798,936,1087,1112]" italics="true" pageId="33" pageNumber="34">interstinctus</emphasis>
</taxonomicName>
color pattern present in some other species of
<taxonomicName id="91EE09C4FFBBC11BFF46DA3EFEB97F29" box="[182,335,1123,1149]" class="Insecta" family="Cicadellidae" genus="Tenucephalus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="649AAE55FFBBC11BFF46DA3EFEB97F29" box="[182,335,1123,1149]" italics="true" pageId="33" pageNumber="34">Tenucephalus</emphasis>
</taxonomicName>
. It is significantly smaller (
<quantity id="9116DFA2FFBBC11BFD77DA3EFD167F29" box="[647,736,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="33" pageNumber="34" unit="mm" value="4.7">4.7 mm</quantity>
) than the females currently placed in
<taxonomicName id="91EE09C4FFBBC11BFB7BDA3EFB057F28" authorityName="Oman" authorityYear="1937" box="[1163,1267,1123,1148]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="649AAE55FFBBC11BFB7BDA3EFB057F28" box="[1163,1267,1123,1148]" italics="true" pageId="33" pageNumber="34">Bonamus</emphasis>
</taxonomicName>
(
<quantity id="9116DFA2FFBBC11BFAF3DA3FFA667F29" box="[1283,1424,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="8.6" metricValueMin="5.5" pageId="33" pageNumber="34" unit="mm" value="7.05" valueMax="8.6" valueMin="5.5">5.5–8.6 mm</quantity>
).
<bibRefCitation id="327F0FB6FFBBC11BFF67DADAFE3D7FF6" author="Linnavuori, R. &amp; Heller, F." box="[151,459,1159,1186]" pageId="33" pageNumber="34" pagination="1 - 14" refId="ref80185" refString="Linnavuori, R. &amp; Heller, F. (1962) Beitrag zur Cicadelliden-Fauna von Peru. Entomologische Ergebnisse der Stuttgarter Anden- Expedition 1957 - Nr. 1. Stuttgarter Beitraege zur Naturkunde aus dem Staatlichen Museum fur Naturkunde in Stuttgart, 67, 1 - 14." type="journal article" year="1962">Linnavuori &amp; Heller (1962)</bibRefCitation>
indicated an uncertain generic placement of the species by placing a question mark after the genus name in the heading.
</paragraph>
<paragraph id="56517247FFBBC11BFF37DA92FD7D7E59" blockId="33.[151,1437,151,1473]" pageId="33" pageNumber="34">
The unique internal membranous process of the pygofer (
<figureCitation id="CED56EC2FFBBC11BFCA7DA92FC5C7FBD" box="[855,938,1231,1257]" captionStart="FIGUIRE 19" captionStartId="37.[151,259,1570,1594]" captionTargetBox="[166,1425,204,1516]" captionTargetId="figure-16@37.[152,1436,201,1524]" captionTargetPageId="37" captionText="FIGUIRE 19. Bonamus lineatus, male pygofer with internal genitalia, subgenital plates, and valve removed. (Q) dorsal view. (R) ventral view. (S) detail of Q. (T) detail of R." figureDoi="http://doi.org/10.5281/zenodo.4690829" httpUri="https://zenodo.org/record/4690829/files/figure.png" pageId="33" pageNumber="34">Fig. 19</figureCitation>
) is an extension of the phragma which also forms the setose bulbous lobes in this genus.
</paragraph>
<paragraph id="56517247FFBBC11BFF37DB4AFCE97E95" blockId="33.[151,1437,151,1473]" pageId="33" pageNumber="34">
The genus was resolved as sister to
<taxonomicName id="91EE09C4FFBBC11BFDA8DB4AFD077E65" box="[600,753,1303,1329]" class="Insecta" family="Cicadellidae" genus="Tenucephalus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="649AAE55FFBBC11BFDA8DB4AFD077E65" box="[600,753,1303,1329]" italics="true" pageId="33" pageNumber="34">Tenucephalus</emphasis>
</taxonomicName>
, and the two genera together form a highly supported monophyletic clade which is sister to the remaining
<taxonomicName id="91EE09C4FFBBC11BFD52DB66FCF57E01" authorityName=": Linnavuori &amp; DeLong" authorityYear="1977" box="[674,771,1339,1365]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Faltalini">Faltalini</taxonomicName>
. The two genera share characters of the ovipositor which is unusually long and protruding very far beyond the pygofer, the first valvula dorsal sculpturing pattern maculose and dorsoventrally elongated, and gonoplac with macrosetae. Males of the genera share the the phragma forming setose bulbous lobes articulating dorsally with base of aedeagus.
</paragraph>
<paragraph id="56517247FFBBC11BFF67DBB2FE927D5C" blockId="33.[151,356,1518,1545]" box="[151,356,1518,1545]" pageId="33" pageNumber="34">
<emphasis id="649AAE55FFBBC11BFF67DBB2FE927D5C" bold="true" box="[151,356,1518,1545]" pageId="33" pageNumber="34">Included species:</emphasis>
</paragraph>
<paragraph id="56517247FFBBC11BFF67D86AFE607D06" blockId="33.[151,605,1591,1654]" box="[151,406,1591,1618]" pageId="33" pageNumber="34">
<heading id="0D19C52BFFBBC11BFF67D86AFE607D06" box="[151,406,1591,1618]" fontSize="11" level="2" pageId="33" pageNumber="34" reason="5">
<taxonomicName id="91EE09C4FFBBC11BFF67D86AFF077D05" authority="Zahniser, 2021" authorityName="Zahniser" authorityYear="2021" box="[151,241,1591,1617]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="jussaral" status="sp. nov.">
<emphasis id="649AAE55FFBBC11BFF67D86AFF077D05" box="[151,241,1591,1617]" italics="true" pageId="33" pageNumber="34">jussaral</emphasis>
</taxonomicName>
<emphasis id="649AAE55FFBBC11BFF08D86AFEC17D05" bold="true" box="[248,311,1591,1617]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="7FA9132EFFBBC11BFF08D86AFEC17D05" box="[248,311,1591,1617]" pageId="33" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="2EF932D7FFBBC11BFEB6D86AFE7B7D06" box="[326,397,1591,1618]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
)
</heading>
</paragraph>
<paragraph id="56517247FFBBC11BFF67D806FDAB7D22" blockId="33.[151,605,1591,1654]" box="[151,605,1627,1654]" pageId="33" pageNumber="34">
<taxonomicName id="91EE09C4FFBBC11BFF67D806FE777D21" authority="Oman, 1938" authorityName="Oman" authorityYear="1938" box="[151,385,1627,1653]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="649AAE55FFBBC11BFF67D806FF067D21" box="[151,240,1627,1653]" italics="true" pageId="33" pageNumber="34">lineatus</emphasis>
<bibRefCitation id="327F0FB6FFBBC11BFF07D806FE777D21" author="Oman, P. W." box="[247,385,1627,1653]" pageId="33" pageNumber="34" pagination="343 - 420" refId="ref80707" refString="Oman, P. W. (1938) A generic revision of American Bythoscopinae and South American Jassinae. University of Kansas Science Bulletin, 37, 343 - 420." type="journal article" year="1938">
<collectingCountry id="2EF932D7FFBBC11BFF07D806FEC87D21" box="[247,318,1627,1653]" name="Oman" pageId="33" pageNumber="34">Oman</collectingCountry>
, 1938
</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="2EF932D7FFBBC11BFE60D806FDF27D21" box="[400,516,1627,1653]" name="Argentina" pageId="33" pageNumber="34">Argentina</collectingCountry>
,
<collectingCountry id="2EF932D7FFBBC11BFDFFD806FDA37D22" box="[527,597,1627,1654]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
)
</paragraph>
<paragraph id="56517247FFBBC11BFF67D89AFE147DB5" blockId="33.[151,482,1734,1761]" box="[151,482,1734,1761]" pageId="33" pageNumber="34">
<emphasis id="649AAE55FFBBC11BFF67D89AFE147DB5" bold="true" box="[151,482,1734,1761]" pageId="33" pageNumber="34">
Key to species of
<taxonomicName id="91EE09C4FFBBC11BFE9ED89AFE147DB5" authorityName="Oman" authorityYear="1937" box="[366,482,1735,1761]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="649AAE55FFBBC11BFE9ED89AFE147DB5" bold="true" box="[366,482,1735,1761]" italics="true" pageId="33" pageNumber="34">Bonamus</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="1EF421CCFFBBC11BFF67D951FA6B7CBA" pageId="33" pageNumber="34" type="key">
<keyStep id="ED1A6AE2FFBBC11BFF67D951FA6B7CBA" pageId="33" pageNumber="34">
<key id="227F2576FFBBC11BFF67D951FA6B7CBA" pageId="33" pageNumber="34">
<paragraph id="56517247FFBBC11BFF67D951FA6A7C2D" blockId="33.[151,1437,1803,2031]" pageId="33" pageNumber="34">
<keyLead id="ED1FD172FFBBC11BFF67D951FA6A7C2D" pageId="33" pageNumber="34">
1. Smaller (length of male
<quantity id="9116DFA2FFBBC11BFE3CD956FDB47C75" box="[460,578,1803,1825]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.4" metricValueMin="5.1" pageId="33" pageNumber="34" unit="mm" value="5.25" valueMax="5.4" valueMin="5.1">5.1–5.4 mm</quantity>
, female 6.0–
<quantity id="9116DFA2FFBBC11BFD4ED951FCFF7C75" box="[702,777,1804,1826]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="33" pageNumber="34" unit="mm" value="6.7">6.7 mm</quantity>
) (17A,B); male pygofer ventral process with several large and small blunt teeth ventrally (17D); phragma ventrolateral membranous process short, digitate, reaching at most to midlength of segment X (19Q–T); aedeagus without apical processes (17H,I); female sternite VII posterior margin deeply excavated medially (17J)........................................................................................
<taxonomicName id="91EE09C4FFBBC11BFAC6D939FA6A7C2D" ID-CoL="MH49" box="[1334,1436,1892,1913]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="649AAE55FFBBC11BFAC6D939FA6A7C2D" box="[1334,1436,1892,1913]" italics="true" pageId="33" pageNumber="34">B. lineatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="56517247FFBBC11BFF67D9DCFA6B7CBA" blockId="33.[151,1437,1803,2031]" pageId="33" pageNumber="34">
<keyLead id="ED1FD172FFBBC11BFF67D9DCFA6B7CBA" pageId="33" pageNumber="34">
1’. Larger (length of male
<quantity id="9116DFA2FFBBC11BFE4DD9DCFDFE7CC2" box="[445,520,1921,1942]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="33" pageNumber="34" unit="mm" value="7.2">7.2 mm</quantity>
, female
<quantity id="9116DFA2FFBBC11BFDA8D9DCFD547CC2" box="[600,674,1921,1942]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="33" pageNumber="34" unit="mm" value="9.7">9.7 mm</quantity>
) (16A1,A2,B); male pygofer process without ventral teeth, ventral side rounded to base (16D); phragma ventrolateral membranous process much larger, wider, reaching nearly to apex of segment X (16D,E); aedeagus with pair of apical branched processes (16H,I); female sternite VII posterior margin not excavated medially (16J).......................................................................................
<taxonomicName id="91EE09C4FFBBC11BFB08D987FA967CBA" ID-CoL="8KL68" authority="Zahniser, 2021" authorityName="Zahniser" authorityYear="2021" box="[1272,1376,2009,2031]" class="Insecta" family="Cicadellidae" genus="Bonamus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="jussaral" status="sp. nov.">
<emphasis id="649AAE55FFBBC11BFB08D987FA967CBA" box="[1272,1376,2009,2031]" italics="true" pageId="33" pageNumber="34">B. jussaral</emphasis>
</taxonomicName>
<emphasis id="649AAE55FFBBC11BFA96D984FA6B7CBA" bold="true" box="[1382,1437,2009,2030]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="7FA9132EFFBBC11BFA96D984FA6B7CBA" box="[1382,1437,2009,2030]" pageId="33" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</key>
</keyStep>
</subSubSection>
</treatment>
</document>
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