263 lines
37 KiB
XML
263 lines
37 KiB
XML
<document id="FAFBB1385838D799E0438E59E3CD09F5" ID-ISSN="11755334" ID-ZooBank="2412CB4F-4D29-4988-80C1-205D16767678" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629824082200" checkinUser="felipe" docAuthor="Borkent, Christopher J. & Wheeler, Terry A." docDate="2012" docId="CA3487C862249D54EECCFBF2FA8CECEF" docLanguage="en" docName="zt03549p117.pdf" docOrigin="Zootaxa 3549" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Leptomorphus waodani Borkent 2012, new species" docType="treatment" docVersion="3" lastPageNumber="71" masterDocId="360DFFB062619D12EE5BFF91FFD9EA01" masterDocTitle="Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549" masterLastPageNumber="117" masterPageNumber="1" pageNumber="70" updateTime="1699117448841" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C5E8CD98CF917B35045B3B1C8E8341E3">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
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<mods:namePart id="B61515DC9D32532DAB075B91D696B30A">Borkent, Christopher J.</mods:namePart>
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<mods:namePart id="523805C10C751087BCD98764033DC4D6">Wheeler, Terry A.</mods:namePart>
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<mods:title id="CBF2734E62A06037ACC39A5F4B15674D">Zootaxa</mods:title>
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<mods:date id="A58C861E485D6059E2AE36885FF3DD09">2012</mods:date>
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<mods:number id="75A28E1382247D05A2167B86FC355FB6">3549</mods:number>
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<treatment id="CA3487C862249D54EECCFBF2FA8CECEF" ID-DOI="http://doi.org/10.5281/zenodo.5258373" ID-Zenodo-Dep="5258373" LSID="urn:lsid:plazi:treatment:CA3487C862249D54EECCFBF2FA8CECEF" httpUri="http://treatment.plazi.org/id/CA3487C862249D54EECCFBF2FA8CECEF" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
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<paragraph id="422236DE62249D57EECCFBF2FD34EE7C" blockId="69.[151,749,1123,1182]" box="[151,749,1123,1150]" pageId="69" pageNumber="70">
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<heading id="196A81B262249D57EECCFBF2FD34EE7C" bold="true" box="[151,749,1123,1150]" fontSize="11" level="1" pageId="69" pageNumber="70" reason="1">
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<emphasis id="70E9EACC62249D57EECCFBF2FD34EE7C" bold="true" box="[151,749,1123,1150]" pageId="69" pageNumber="70">
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37.
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<taxonomicName id="859D4D5D62249D57EE98FBF5FD89EE7C" ID-CoL="8CJ4M" authority="Borkent" authorityName="Borkent" authorityYear="2012" box="[195,592,1123,1150]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="waodani" status="sp. nov.">
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<emphasis id="70E9EACC62249D57EE98FBF5FE38EE7F" bold="true" box="[195,481,1124,1150]" italics="true" pageId="69" pageNumber="70">Leptomorphus waodani</emphasis>
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Borkent
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</taxonomicName>
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,
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<taxonomicNameLabel id="6BDA57B762249D57EC06FBF2FD34EE7C" box="[605,749,1123,1149]" pageId="69" pageNumber="70" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A87655562249D57EECCFB17FD8DEE9F" box="[151,596,1158,1182]" pageId="69" pageNumber="70" type="description">
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<paragraph id="422236DE62249D57EECCFB17FD8DEE9F" blockId="69.[151,749,1123,1182]" box="[151,596,1158,1182]" pageId="69" pageNumber="70">
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(
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<figureCitation id="DAA62A5B62249D57EEC5FB17FEC0EE9F" box="[158,281,1158,1182]" captionStart="FIGURES 38–41" captionStartId="91.[151,264,1941,1964]" captionTargetBox="[153,1434,258,1918]" captionTargetId="figure-40@91.[151,1436,255,1920]" captionTargetPageId="91" captionText="FIGURES 38–41. Leptomorphus habitus. 38. L. titiwangsensis, paratype female. 39. L. waodani, holotype male. 40. L. tagbanua, holotype male. 41. L. walkeri, male." pageId="69" pageNumber="70">Figures 39</figureCitation>
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,
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<figureCitation id="DAA62A5B62249D57EF7EFB16FE9AEE9F" box="[293,323,1159,1182]" captionStart="FIGURES 70–77" captionStartId="94.[151,264,1209,1232]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure-136@94.[151,1436,193,1188]" captionTargetPageId="94" captionText="FIGURES 70–77. Leptomorphus wings. 70. L. quadrimaculatus, ventral. 71. L. stigmatus holotype male, dorsal. 72. L. subcaeruleus, ventral. 73. L. tabatius holotype male, dorsal. 74. L. tagbanua holotype male, ventral. 75. L. titiwangsensis paratype female, dorsal. 76. L. walkeri, ventral. 77. L. waodani holotype male, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="69" pageNumber="70">77</figureCitation>
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,
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<figureCitation id="DAA62A5B62249D57EF14FB17FEB4EE9F" box="[335,365,1158,1182]" captionStart="FIGURES 82–90" captionStartId="96.[151,263,1444,1467]" captionTargetBox="[151,1436,211,1413]" captionTargetId="figure-84@96.[151,1436,211,1413]" captionTargetPageId="96" captionText="FIGURES 82–90. Leptomorphus morphology. 82. Head of L. brandiae, anterolateral, showing swollen palpus segment 5 covered in fine white trichia. 83–84. Male midfemur, anterior, showing apicoventral process (arrow). 83. L. nebulosus. 84. L. subcaeruleus. 85–90. Male foretibia, posterior, showing presence (arrow) or absence of dense comb of ventral bristles. 85. L. forcipatus. 86. L. magnificus. 87. L. mandelai. 88. L. tagbanua. 89. L. walkeri. 90. L. waodani." pageId="69" pageNumber="70">90</figureCitation>
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,
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<figureCitation id="DAA62A5B62249D57EF22FB17FE7CEE9F" box="[377,421,1158,1182]" captionStart="FIGURES 121–124" captionStartId="105.[151,264,1775,1798]" captionTargetBox="[176,1407,186,1696]" captionTargetId="figure-118@105.[161,1446,161,1741]" captionTargetPageId="105" captionText="FIGURES 121–124. Leptomorphus male genitalia. 121. L. talyshensis (reprinted with permission from Zaitzev & Ševčík 2002), A. gonocoxite and gonostylus, ventral. B. dorsal. 122. L. titiwangsensis, ventral. 123. L. walkeri, ventral. 124. L. waodani, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="69" pageNumber="70">124</figureCitation>
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,
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<figureCitation id="DAA62A5B62249D57EFEAFB17FE04EE9F" box="[433,477,1158,1182]" captionStart="FIGURE 142–143" captionStartId="110.[151,249,1784,1807]" captionTargetBox="[151,1436,193,1763]" captionTargetId="figure-57@110.[151,1436,193,1763]" captionTargetPageId="110" captionText="FIGURE 142–143. Distribution of Neotropical Leptomorphus. 142. L. amorimi (blue circle), L. crassipilus (red square), L. fasciculatus (black triangle), L. neivai (red star). 143. L. brandiae (purple triangle), L. eberhardi (blue circle), L. femoratus (red star), L. waodani (black square)." pageId="69" pageNumber="70">143</figureCitation>
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,
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<figureCitation id="DAA62A5B62249D57EFB2FB17FDCCEE9F" box="[489,533,1158,1182]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values>50% are shown in bold below the branches." pageId="69" pageNumber="70">150</figureCitation>
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,
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<figureCitation id="DAA62A5B62249D57EC7AFB17FD95EE9F" box="[545,588,1158,1182]" captionStart="FIGURE 155" captionStartId="116.[151,250,858,881]" captionTargetBox="[151,1425,187,838]" captionTargetId="figure-58@116.[151,1436,159,885]" captionTargetPageId="116" captionText="FIGURE 155. One of three equally parsimonious trees showing the relationships between species in the L. furcatus species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="69" pageNumber="70">155</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="0A87655562249D57EECCFB5FFE71EC08" pageId="69" pageNumber="70" type="diagnosis">
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<paragraph id="422236DE62249D57EECCFB5FFD54EF77" blockId="69.[151,1437,1230,2013]" pageId="69" pageNumber="70">
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<emphasis id="70E9EACC62249D57EECCFB5FFEEDEEE6" bold="true" box="[151,308,1230,1255]" pageId="69" pageNumber="70">DIAGNOSIS</emphasis>
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: The only extant species of
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<taxonomicName id="859D4D5D62249D57EC35FB5FFCC9EEE7" authorityName=": Edwards" authorityYear="1925" box="[622,784,1230,1254]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">
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<emphasis id="70E9EACC62249D57EC35FB5FFCC9EEE7" box="[622,784,1230,1254]" italics="true" pageId="69" pageNumber="70">Leptomorphus</emphasis>
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</taxonomicName>
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with the following combination of characters: scutum and scutellum evenly covered with fine trichia; male foretibia with a dense row (comb) of short anteroventral bristles (
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<figureCitation id="DAA62A5B62249D57EEC4FA87FF20EF2F" box="[159,249,1302,1326]" captionStart="FIGURES 82–90" captionStartId="96.[151,263,1444,1467]" captionTargetBox="[151,1436,211,1413]" captionTargetId="figure-84@96.[151,1436,211,1413]" captionTargetPageId="96" captionText="FIGURES 82–90. Leptomorphus morphology. 82. Head of L. brandiae, anterolateral, showing swollen palpus segment 5 covered in fine white trichia. 83–84. Male midfemur, anterior, showing apicoventral process (arrow). 83. L. nebulosus. 84. L. subcaeruleus. 85–90. Male foretibia, posterior, showing presence (arrow) or absence of dense comb of ventral bristles. 85. L. forcipatus. 86. L. magnificus. 87. L. mandelai. 88. L. tagbanua. 89. L. walkeri. 90. L. waodani." pageId="69" pageNumber="70">Fig. 90</figureCitation>
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); palp segment 5 with thick covering of fine, white, setulae; male genitalia with tergite 9 tapering posteriorly to a point, with short pointed processes laterally at 2/3 of length towards apex; gonocoxite with large gonocoxal lobe on medial margin (
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<figureCitation id="DAA62A5B62249D57EC45FACFFD59EF77" box="[542,640,1374,1398]" captionStart="FIGURES 121–124" captionStartId="105.[151,264,1775,1798]" captionTargetBox="[176,1407,186,1696]" captionTargetId="figure-118@105.[161,1446,161,1741]" captionTargetPageId="105" captionText="FIGURES 121–124. Leptomorphus male genitalia. 121. L. talyshensis (reprinted with permission from Zaitzev & Ševčík 2002), A. gonocoxite and gonostylus, ventral. B. dorsal. 122. L. titiwangsensis, ventral. 123. L. walkeri, ventral. 124. L. waodani, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="69" pageNumber="70">Fig. 124</figureCitation>
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).
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</paragraph>
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<paragraph id="422236DE62249D57EE9CFA13FE71EC08" blockId="69.[151,1437,1230,2013]" pageId="69" pageNumber="70">
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This species is distinguished from most other Neotropical species missing R
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<subScript id="DE19349B62249D57EA10FA03FB8AEFA1" attach="left" box="[1099,1107,1426,1440]" fontSize="6" pageId="69" pageNumber="70">4</subScript>
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by mostly yellow abdomen (tergites 6, 7 brown) and having the scutellum lighter brown than the scutum or mediotergite (
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<figureCitation id="DAA62A5B62249D57EA96FA38FAFAEFC0" box="[1229,1315,1449,1473]" captionStart="FIGURES 38–41" captionStartId="91.[151,264,1941,1964]" captionTargetBox="[153,1434,258,1918]" captionTargetId="figure-40@91.[151,1436,255,1920]" captionTargetPageId="91" captionText="FIGURES 38–41. Leptomorphus habitus. 38. L. titiwangsensis, paratype female. 39. L. waodani, holotype male. 40. L. tagbanua, holotype male. 41. L. walkeri, male." pageId="69" pageNumber="70">Fig. 39</figureCitation>
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). It can be distinguished from
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<taxonomicName id="859D4D5D62249D57EF2EFA5FFE2AEFE4" authorityName="Borkent" authorityYear="2012" box="[373,499,1486,1509]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="amorimi">
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<emphasis id="70E9EACC62249D57EF2EFA5FFE2AEFE4" box="[373,499,1486,1509]" italics="true" pageId="69" pageNumber="70">L. amorimi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="859D4D5D62249D57EC75FA5FFD0DEFE7" box="[558,724,1486,1510]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="fasciculatus">
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<emphasis id="70E9EACC62249D57EC75FA5FFD0DEFE7" box="[558,724,1486,1510]" italics="true" pageId="69" pageNumber="70">L. fasciculatus</emphasis>
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</taxonomicName>
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by the posteromedial margin of tergite 9 being pointed rather than concave (
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<figureCitation id="DAA62A5B62249D57EF62FA60FE42EC08" box="[313,411,1521,1545]" captionStart="FIGURES 121–124" captionStartId="105.[151,264,1775,1798]" captionTargetBox="[176,1407,186,1696]" captionTargetId="figure-118@105.[161,1446,161,1741]" captionTargetPageId="105" captionText="FIGURES 121–124. Leptomorphus male genitalia. 121. L. talyshensis (reprinted with permission from Zaitzev & Ševčík 2002), A. gonocoxite and gonostylus, ventral. B. dorsal. 122. L. titiwangsensis, ventral. 123. L. walkeri, ventral. 124. L. waodani, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="69" pageNumber="70">Fig. 124</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="0A87655562249D54EE9CF984FE1BEFAB" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70" type="description">
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<paragraph id="422236DE62249D54EE9CF984FDFCEF63" blockId="69.[151,1437,1230,2013]" lastBlockId="70.[151,1437,151,1774]" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
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<emphasis id="70E9EACC62249D57EE9CF984FE5FEC2F" bold="true" box="[199,390,1557,1582]" pageId="69" pageNumber="70">DESCRIPTION</emphasis>
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:
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<emphasis id="70E9EACC62249D57EFCCF984FE16EC2C" box="[407,463,1557,1581]" italics="true" pageId="69" pageNumber="70">Male</emphasis>
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. (
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<figureCitation id="DAA62A5B62249D57EFB3F984FDE3EC2C" box="[488,570,1557,1581]" captionStart="FIGURES 38–41" captionStartId="91.[151,264,1941,1964]" captionTargetBox="[153,1434,258,1918]" captionTargetId="figure-40@91.[151,1436,255,1920]" captionTargetPageId="91" captionText="FIGURES 38–41. Leptomorphus habitus. 38. L. titiwangsensis, paratype female. 39. L. waodani, holotype male. 40. L. tagbanua, holotype male. 41. L. walkeri, male." pageId="69" pageNumber="70">Fig. 39</figureCitation>
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||
)
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<emphasis id="70E9EACC62249D57EC10F987FD53EC2F" bold="true" box="[587,650,1558,1582]" pageId="69" pageNumber="70">Head</emphasis>
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: brown spot medially from ocelli to antennal base, remainder yellow, somewhat dorsoventrally compressed in anterior view. Antenna dark brown; scape yellow, with brown setae in short row on apicodorsal margin and thick patch covering apicoventral process, remainder bare, anterobasal patch of setulae present; pedicel yellow, with 1–2 large bristles, few setae on apicodorsal margin, patch of fine setae apicoventrally; flagellomere 1 with tapered base yellow remainder dark brown; flagellomere 6 1.3X as long as broad. Clypeus yellow, slightly laterally compressed oval; bristles on clypeus yellow, 4–6 strong bristles on ventral margin directed ventrally, remaining setae directed medioventrally, clypeus 2X as long as face. Face yellow; shape a slightly wider than tall triangle, bare. Frons brown; bare, frontal furrow running 1/2 distance from dorsal margin towards ventral margin, frontal cleft more than 1X diameter of median ocellus anterior of median ocellus. Palpus yellow; segment 1 hidden behind eye, segments increasing in length, segment 5 2X length and 1.5X width of segment 4 with even width from base to apex and covered in fine white setulae, segment 3 without distinct lateral patch of fine setae. Labellum yellow. Eye with very few, short inter-ommatidial setulae scattered on surface. Occiput yellow with appressed, anteriorly directed setae. Three ocelli, in a straight line, space between ocelli less than diameter of laterals, lateral ocelli 2.5X their own diameter from eye margin, ocellar triangle dark brown.
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<emphasis id="70E9EACC62279D54ED3DFF09FC64EAB1" bold="true" box="[870,957,152,176]" pageId="70" pageNumber="71">Thorax</emphasis>
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: Length
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(n = 1). Dark brown with yellow lateral spots. Scutum dark brown/black with blue-green specks, pair of yellow anterolateral spots,
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<specimenCount id="549BFD5762279D54EB2FFF2DFF26EAF6" pageId="70" pageNumber="71" type="generic">one specimen</specimenCount>
|
||
with lateral margins brown and remainder yellow; surface of scutum covered with trichia; acrostichal setae absent; single dorsocentral seta present anteriorly; two thirds row of lateral setae present; patch of setae on scutum at wing base present. Scutellum dark brown with covering of fine trichia; bristles absent. Prescutum anterior half yellow, posterior half brown. Mediotergite dark brown with 6 bristles on posterolateral corners, and covering of appressed trichia. Laterotergite brown, with covering of appressed trichia; anterior margin of laterotergite abutting katepisternum. Anepimeron brown with posterodorsal corner yellow. Anepisternum brown. Katepisternum brown with posterodorsal corner yellow. Antepronotum and proepisternum brown. Margin of anterior and posterior spiracles yellow with yellow trichia. Metepisternum yellow. Anapleural suture with anterior portion slightly curved dorsally. Halter stem yellow, knob light to dark brown.
|
||
<emphasis id="70E9EACC62279D54ECC5FD91FD0CE819" bold="true" box="[670,725,512,536]" pageId="70" pageNumber="71">Legs</emphasis>
|
||
: principally yellow; hind coxa with brown spot along length of anterior surface; extreme anteroapical corner yellow on forefemur, on other femora dark brown. Midfemur without apical spine-like process. Tibia with covering of brown macrotrichia, foretibia with comb of short setae along length of anteroventral surface (
|
||
<figureCitation id="DAA62A5B62279D54EFF7FDFDFE26E885" box="[428,511,620,644]" captionStart="FIGURES 82–90" captionStartId="96.[151,263,1444,1467]" captionTargetBox="[151,1436,211,1413]" captionTargetId="figure-84@96.[151,1436,211,1413]" captionTargetPageId="96" captionText="FIGURES 82–90. Leptomorphus morphology. 82. Head of L. brandiae, anterolateral, showing swollen palpus segment 5 covered in fine white trichia. 83–84. Male midfemur, anterior, showing apicoventral process (arrow). 83. L. nebulosus. 84. L. subcaeruleus. 85–90. Male foretibia, posterior, showing presence (arrow) or absence of dense comb of ventral bristles. 85. L. forcipatus. 86. L. magnificus. 87. L. mandelai. 88. L. tagbanua. 89. L. walkeri. 90. L. waodani." pageId="70" pageNumber="71">Fig. 90</figureCitation>
|
||
), tibial spurs brown, foretibial spur length 2X apical thickness of foretibia, midtibia with strong, dorsal, bare patch of even thickness for 2/3 of its length, placed basally, shortest midtibial spur 0.8X length of longest, longest midtibial spur 4X apical thickness of midtibia, shortest hind tibial spur subequal to length of longest, longest hind tibial spur 4X apical thickness of hind tibia. Foreleg first tarsomere 1.7X length of foretibia.
|
||
<emphasis id="70E9EACC62279D54EECCFD6AFF0EE915" bold="true" box="[151,215,763,788]" pageId="70" pageNumber="71">Wing</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62279D54EEB3FD6DFEE5E915" box="[232,316,764,788]" captionStart="FIGURES 82–90" captionStartId="96.[151,263,1444,1467]" captionTargetBox="[151,1436,211,1413]" captionTargetId="figure-84@96.[151,1436,211,1413]" captionTargetPageId="96" captionText="FIGURES 82–90. Leptomorphus morphology. 82. Head of L. brandiae, anterolateral, showing swollen palpus segment 5 covered in fine white trichia. 83–84. Male midfemur, anterior, showing apicoventral process (arrow). 83. L. nebulosus. 84. L. subcaeruleus. 85–90. Male foretibia, posterior, showing presence (arrow) or absence of dense comb of ventral bristles. 85. L. forcipatus. 86. L. magnificus. 87. L. mandelai. 88. L. tagbanua. 89. L. walkeri. 90. L. waodani." pageId="70" pageNumber="71">Fig. 90</figureCitation>
|
||
): Length
|
||
<quantity id="85659B3B62279D54EFFCFD6DFE24E912" box="[423,509,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="70" pageNumber="71" unit="mm" value="6.7">6.7 mm</quantity>
|
||
(n = 1). Hyaline; apical macula dark brown but fading towards apex and posterior margin, running from anterior to posterior wing margin, beginning halfway along R
|
||
<subScript id="DE19349B62279D54EA6FFCBEFBE2E93C" attach="right" box="[1076,1083,815,829]" fontSize="6" pageId="70" pageNumber="71">5</subScript>
|
||
; medial macula extending from R
|
||
<subScript id="DE19349B62279D54EEF2FCC7FF68E965" attach="left" box="[169,177,854,868]" fontSize="6" pageId="70" pageNumber="71">1</subScript>
|
||
to stem of M
|
||
<subScript id="DE19349B62279D54EF15FCC7FEBCE965" attach="left" box="[334,357,854,868]" fontSize="6" pageId="70" pageNumber="71">1+2</subScript>
|
||
. Macrotrichia in all cells, though absent from posterobasal margin of cell a, and sparse in basal cells. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with a few short setae. Vein sc-r present, apical end joining R within its own length prior to origin of Rs. R
|
||
<subScript id="DE19349B62279D54EA30FC31FBAAE9AF" attach="none" box="[1131,1139,928,942]" fontSize="6" pageId="70" pageNumber="71">4</subScript>
|
||
absent. R
|
||
<subScript id="DE19349B62279D54EA87FC31FB3DE9AF" attach="left" box="[1244,1252,928,942]" fontSize="6" pageId="70" pageNumber="71">5</subScript>
|
||
slightly concave for entire length. M
|
||
<subScript id="DE19349B62279D54EF2AFC56FEA0E9D4" attach="left" box="[369,377,967,981]" fontSize="6" pageId="70" pageNumber="71">1</subScript>
|
||
reaching apex before R
|
||
<subScript id="DE19349B62279D54ECD8FC56FD52E9D4" attach="left" box="[643,651,967,981]" fontSize="6" pageId="70" pageNumber="71">5</subScript>
|
||
, apices of M veins reaching wing margin. M
|
||
<subScript id="DE19349B62279D54EADAFC56FB51E9D4" attach="left" box="[1153,1160,967,981]" fontSize="6" pageId="70" pageNumber="71">4</subScript>
|
||
-CuA fork arising before origin of r-m. A
|
||
<subScript id="DE19349B62279D54EF13FC7FFE89E9FD" attach="left" box="[328,336,1006,1020]" fontSize="6" pageId="70" pageNumber="71">2</subScript>
|
||
absent.
|
||
<emphasis id="70E9EACC62279D54EFF7FC4FFDC6E9F6" bold="true" box="[428,543,990,1015]" pageId="70" pageNumber="71">Abdomen</emphasis>
|
||
: Tergites principally yellow, T1 and 6–7 dark brown, T5 with posterior 1/2 dark brown. Tergite 8 smaller than all other abdominal sclerites, without bristles.
|
||
<emphasis id="70E9EACC62279D54ED87FB94FB91EE1F" bold="true" box="[988,1096,1029,1054]" pageId="70" pageNumber="71">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62279D54EA0CFB97FB61EE1F" box="[1111,1208,1030,1054]" captionStart="FIGURES 121–124" captionStartId="105.[151,264,1775,1798]" captionTargetBox="[176,1407,186,1696]" captionTargetId="figure-118@105.[161,1446,161,1741]" captionTargetPageId="105" captionText="FIGURES 121–124. Leptomorphus male genitalia. 121. L. talyshensis (reprinted with permission from Zaitzev & Ševčík 2002), A. gonocoxite and gonostylus, ventral. B. dorsal. 122. L. titiwangsensis, ventral. 123. L. walkeri, ventral. 124. L. waodani, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="70" pageNumber="71">Fig. 124</figureCitation>
|
||
): mostly brown with apex yellow. Sternite 9 sclerotized, thin, posteriorly tapered sclerite, almost as long as aedeagus, not reaching gonocoxal margin. Tergite 9 longer than wide, rounded basally, tapering into long point at apex with small point 2/3 way to apex. Gonocoxite placed centrally on T9, medial margins not reaching median line, apicolateral margin a short point, apicoventral margin extended into long (2X remainder of gonocoxite) tapering point, bearing gonostylus centrally. Gonostylus a single broad-based lobe tapering to a point apically and slightly curved dorsally with setae on basal half, gonocoxite III associated with dorsal margin of gonostylus but not fused to it. Aedeagus ~1.2X length of S9, tapering to middle and then swelling to apex which ends in a pointed lobe on each lateral corner and a medial rounded bump, apodemes sclerotized and 1/4 total length. Parameres a simple thin taper with a swollen base, apodemes ~1/2 length of parameres.
|
||
</paragraph>
|
||
<paragraph id="422236DE62279D54EE9CFAFFFE47EF87" blockId="70.[151,1437,151,1774]" box="[199,414,1390,1414]" pageId="70" pageNumber="71">
|
||
<emphasis id="70E9EACC62279D54EE9CFAFFFEC2EF87" box="[199,283,1390,1414]" italics="true" pageId="70" pageNumber="71">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph id="422236DE62279D54EE9CFA02FE1BEFAB" blockId="70.[151,1437,151,1774]" box="[199,450,1426,1450]" pageId="70" pageNumber="71">
|
||
<emphasis id="70E9EACC62279D54EE9CFA02FEE7EFAB" box="[199,318,1427,1450]" italics="true" pageId="70" pageNumber="71">Immatures</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562279D54EE9CFA24FE16EFCF" box="[199,463,1461,1486]" pageId="70" pageNumber="71" type="biology_ecology">
|
||
<paragraph id="422236DE62279D54EE9CFA24FE16EFCF" blockId="70.[151,1437,151,1774]" box="[199,463,1461,1486]" pageId="70" pageNumber="71">
|
||
<emphasis id="70E9EACC62279D54EE9CFA24FE93EFCF" bold="true" box="[199,330,1461,1486]" pageId="70" pageNumber="71">BIOLOGY</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562279D54EE9CFA48FD2DEFF3" box="[199,756,1497,1522]" pageId="70" pageNumber="71" type="distribution">
|
||
<paragraph id="422236DE62279D54EE9CFA48FD2DEFF3" blockId="70.[151,1437,151,1774]" box="[199,756,1497,1522]" pageId="70" pageNumber="71">
|
||
<emphasis id="70E9EACC62279D54EE9CFA48FE4FEFF3" bold="true" box="[199,406,1497,1522]" pageId="70" pageNumber="71">DISTRIBUTION</emphasis>
|
||
:
|
||
<collectingCountry id="3A8A764E62279D54EFFFFA4BFDD8EFF3" box="[420,513,1498,1522]" name="Ecuador" pageId="70" pageNumber="71">Ecuador</collectingCountry>
|
||
(
|
||
<figureCitation id="DAA62A5B62279D54EC4AFA4BFDABEFF3" box="[529,626,1498,1522]" captionStart="FIGURE 142–143" captionStartId="110.[151,249,1784,1807]" captionTargetBox="[151,1436,193,1763]" captionTargetId="figure-57@110.[151,1436,193,1763]" captionTargetPageId="110" captionText="FIGURE 142–143. Distribution of Neotropical Leptomorphus. 142. L. amorimi (blue circle), L. crassipilus (red square), L. fasciculatus (black triangle), L. neivai (red star). 143. L. brandiae (purple triangle), L. eberhardi (blue circle), L. femoratus (red star), L. waodani (black square)." pageId="70" pageNumber="71">Fig. 143</figureCitation>
|
||
),
|
||
<quantity id="85659B3B62279D54ECDCFA4BFD29EFF3" box="[647,752,1498,1522]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="70" pageNumber="71" unit="m" value="215.0">215 masl</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562279D54EE9CFA6CFECAEC5F" pageId="70" pageNumber="71" type="etymology">
|
||
<paragraph id="422236DE62279D54EE9CFA6CFECAEC5F" blockId="70.[151,1437,151,1774]" pageId="70" pageNumber="71">
|
||
<emphasis id="70E9EACC62279D54EE9CFA6CFE58EC17" bold="true" box="[199,385,1533,1558]" pageId="70" pageNumber="71">ETYMOLOGY</emphasis>
|
||
:
|
||
<materialsCitation id="F2F53C8362279D54EFCBFA6FFED6EC5F" country="Ecuador" county="Amazonian" location="This" municipality="Waodani" pageId="70" pageNumber="71" specimenCount="1" typeStatus="holotype">
|
||
<location id="4742600562279D54EFCBFA6FFE1BEC17" LSID="urn:lsid:plazi:treatment:CA3487C862249D54EECCFBF2FA8CECEF:4742600562279D54EFCBFA6FFE1BEC17" box="[400,450,1534,1558]" country="Ecuador" county="Amazonian" municipality="Waodani" name="This" pageId="70" pageNumber="71">This</location>
|
||
species is named in honour of the
|
||
<collectingMunicipality id="A246ACA462279D54ED0AFA6FFC6DEC17" box="[849,948,1534,1558]" pageId="70" pageNumber="71">Waodani</collectingMunicipality>
|
||
(also spelled Waorani or Huaorani) people of
|
||
<collectingCounty id="AB434E5262279D54EEE1F9B3FEE2EC3B" box="[186,315,1570,1594]" pageId="70" pageNumber="71">Amazonian</collectingCounty>
|
||
<collectingCountry id="3A8A764E62279D54EF1DF9B3FE7CEC3B" box="[326,421,1570,1594]" name="Ecuador" pageId="70" pageNumber="71">Ecuador</collectingCountry>
|
||
, as the
|
||
<typeStatus id="9D26887C62279D54EC5EF9B3FDBEEC3B" box="[517,615,1570,1594]" pageId="70" pageNumber="71" type="holotype">holotype</typeStatus>
|
||
was collected in their territories. The name is proposed as a noun in apposition
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562279D54EE9CF9F8FA8CECEF" pageId="70" pageNumber="71" type="materials_examined">
|
||
<paragraph id="422236DE62279D54EE9CF9F8FA8CECEF" blockId="70.[151,1437,151,1774]" pageId="70" pageNumber="71">
|
||
<materialsCitation id="F2F53C8362279D54EE9CF9F8FA8CECEF" collectingDate="1996-06-21" collectionCode="MATERIAL, T, USNM" collectorName="S. Onkone Gare Cmp & L. Erwin & Det. C. J. Borkent" country="Ecuador" determinerName="Det. C. J. Borkent" elevation="216" latitude="-0.6572222" location="Waorani" longLatPrecision="21" longitude="-76.45305" pageId="70" pageNumber="71" specimenCode="MAXUS # 1551" specimenCount="1" specimenCount-male="1" stateProvince="Orellana" typeStatus="holotype">
|
||
<emphasis id="70E9EACC62279D54EE9CF9F8FDDFEC83" bold="true" box="[199,518,1641,1666]" pageId="70" pageNumber="71">
|
||
<collectionCode id="248CAE1B62279D54EE9CF9F8FE86EC83" box="[199,351,1641,1666]" pageId="70" pageNumber="71">MATERIAL</collectionCode>
|
||
EXAMINED
|
||
</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC62279D54EC40F9FBFD53EC83" box="[539,650,1642,1666]" italics="true" pageId="70" pageNumber="71">
|
||
<typeStatus id="9D26887C62279D54EC40F9FBFD5DEC83" box="[539,644,1642,1666]" pageId="70" pageNumber="71" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
here designated, adult male, pinned with genitalia in vial on pin, labelled “
|
||
<collectingCountry id="3A8A764E62279D54EF50F91FFE4EECA7" box="[267,407,1678,1702]" name="Ecuador" pageId="70" pageNumber="71">ECUADOR</collectingCountry>
|
||
:
|
||
<collectingRegion id="8059F83C62279D54EFF2F91FFDD5ECA7" box="[425,524,1678,1702]" country="Ecuador" name="Orellana" pageId="70" pageNumber="71">Orellana</collectingRegion>
|
||
, Res. Ethnica/
|
||
<location id="4742600562279D54EC97F91FFCF4ECA7" LSID="urn:lsid:plazi:treatment:CA3487C862249D54EECCFBF2FA8CECEF:4742600562279D54EC97F91FFCF4ECA7" box="[716,813,1678,1702]" country="Ecuador" latitude="-0.6572222" longLatPrecision="21" longitude="-76.45305" name="Waorani" pageId="70" pageNumber="71" stateProvince="Orellana">Waorani</location>
|
||
,
|
||
<quantity id="85659B3B62279D54ED1BF91FFC59ECA7" box="[832,896,1678,1702]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="70" pageNumber="71" unit="km" value="1.0">1 km</quantity>
|
||
<collectorName id="EF68530862279D54EDD4F91FFB41ECA7" box="[911,1176,1678,1702]" pageId="70" pageNumber="71">S. Onkone Gare Cmp</collectorName>
|
||
/ Trans. Ent.
|
||
<quantity id="85659B3B62279D54EB1BF91FFA41ECA7" box="[1344,1432,1678,1702]" metricMagnitude="2" metricUnit="m" metricValue="2.163" pageId="70" pageNumber="71" unit="m" value="216.3">
|
||
<elevation id="C9B0D1ED62279D54EB1BF91FFA41ECA7" box="[1344,1432,1678,1702]" metricMagnitude="2" metricUnit="m" metricValue="2.163" pageId="70" pageNumber="71" unit="m" value="216.3">216.3m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="27A9501962279D54EECCF923FECEECCB" box="[151,279,1714,1738]" degrees="00" direction="south" minutes="39" orientation="latitude" pageId="70" pageNumber="71" precision="15" seconds="26" value="-0.6572222">00°39'26"S</geoCoordinate>
|
||
,/
|
||
<geoCoordinate id="27A9501962279D54EF77F923FE1AECCB" box="[300,451,1714,1738]" degrees="076" direction="west" minutes="27" orientation="longitude" pageId="70" pageNumber="71" precision="15" seconds="11" value="-76.45305">076°27'11"W</geoCoordinate>
|
||
,
|
||
<date id="3623101E62279D54EF95F923FD91ECCB" box="[462,584,1714,1738]" pageId="70" pageNumber="71" value="1996-06-21">
|
||
<collectingDate id="2667E9F662279D54EF95F923FD91ECCB" box="[462,584,1714,1738]" pageId="70" pageNumber="71" value="1996-06-21">21.vi.1996</collectingDate>
|
||
</date>
|
||
, fogging/ terre firme forest,
|
||
<collectionCode id="248CAE1B62279D54EDDEF923FC4EECC8" box="[901,919,1714,1737]" name="Tavera, Department of Geology and Geophysics" pageId="70" pageNumber="71">T</collectionCode>
|
||
.
|
||
<collectorName id="EF68530862279D54EDF9F923FBDCECCB" box="[930,1029,1714,1738]" pageId="70" pageNumber="71">L. Erwin</collectorName>
|
||
<emphasis id="70E9EACC62279D54EA57F922FB9DECCB" box="[1036,1092,1714,1738]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
|
||
/ Trans. 4 Sta. 1 Proj.
|
||
<specimenCode id="123B9EA562279D54EB6CF923FF3EECEF" collectionCode="MAXUS" pageId="70" pageNumber="71">MAXUS # 1551</specimenCode>
|
||
;
|
||
<typeStatus id="9D26887C62279D54EEA8F947FE5EECEF" box="[243,391,1750,1774]" pageId="70" pageNumber="71" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="549BFD5762279D54EFD6F947FE7BECEC" box="[397,418,1750,1773]" pageId="70" pageNumber="71" type="male">♂</specimenCount>
|
||
/
|
||
<taxonomicName id="859D4D5D62279D54EFEAF947FD63ECEF" authorityName="Borkent" authorityYear="2012" box="[433,698,1750,1774]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="waodani">
|
||
<emphasis id="70E9EACC62279D54EFEAF947FD63ECEF" box="[433,698,1750,1774]" italics="true" pageId="70" pageNumber="71">Leptomorphus waodani</emphasis>
|
||
</taxonomicName>
|
||
/ Borkent,
|
||
<taxonomicNameLabel id="6BDA57B762279D54ED63F947FC66ECEF" box="[824,959,1750,1774]" pageId="70" pageNumber="71" rank="species">new species</taxonomicNameLabel>
|
||
/
|
||
<determinerName id="B7FEDA1A62279D54ED8FF947FB4EECEF" box="[980,1175,1750,1774]" pageId="70" pageNumber="71">
|
||
<collectorName id="EF68530862279D54ED8FF947FB4EECEF" box="[980,1175,1750,1774]" pageId="70" pageNumber="71">Det. C.J. Borkent</collectorName>
|
||
</determinerName>
|
||
2012” [
|
||
<collectionCode id="248CAE1B62279D54EAAAF947FA92ECEF" box="[1265,1355,1750,1774]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="70" pageNumber="71" type="Museum">USNM</collectionCode>
|
||
].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |