303 lines
63 KiB
XML
303 lines
63 KiB
XML
<document id="285963234D58250EF35295621BE8E2FA" ID-CLB-Dataset="8807" ID-DOI="10.11646/zootaxa.4822.2.1" ID-GBIF-Dataset="73a6f164-2334-466a-aee0-d39129bdf7e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401369" ID-ZooBank="E3B5713F-ADE8-4075-9ABF-8F6DE9D3A88E" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609335057997" checkinUser="plazi" docAuthor="Miranda, Gil Felipe Gonçalves, Skevington, Jeffrey H. & Marshall, Stephen A." docDate="2020" docId="A51D5F67FFEC3A6DFF50FD9EFCE39FF2" docLanguage="en" docName="zootaxa.4822.2.1.pdf" docOrigin="Zootaxa 4822 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fragosa Miranda & Skevington & Marshall 2020, gen. nov." docType="treatment" docUuid="59261D3C-F2FC-46FC-B54E-E3B205B56290" docUuidSource="ZooBank" docVersion="8" lastPageNumber="154" masterDocId="5924271FFFEE3A6EFFC7FFBDFFE39F0B" masterDocTitle="New generic concepts for orphaned lineages formerly treated as part of the genus Ocyptamus Macquart, 1834 (Diptera, Syrphidae)" masterLastPageNumber="174" masterPageNumber="151" pageNumber="153" updateTime="1698849646355" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4CB0FD0AF97CAAF076F416257F5F643C">New generic concepts for orphaned lineages formerly treated as part of the genus Ocyptamus Macquart, 1834 (Diptera, Syrphidae)</mods:title>
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<mods:name id="7AEC2AC6706D98C68788FD7B54C9490F" type="personal">
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<mods:namePart id="AA2E665D51DBC5DCD1E5ECB8AAB0E5B3">Miranda, Gil Felipe Gonçalves</mods:namePart>
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<mods:nameIdentifier id="467EED1382ABCFDC23A21CC3D838DD80" type="email">gilfgm@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="C12AE7B6C30D26D43CB9716E51C8494D">Author</mods:roleTerm>
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<mods:namePart id="1BB30BBF353E995BFC9F64178C079B64">Skevington, Jeffrey H.</mods:namePart>
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<mods:nameIdentifier id="2DFDCB48DA28C25E864FA2B657373F61" type="ORCID">0000-0001-7919-2639</mods:nameIdentifier>
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<mods:affiliation id="7BFF1C2533619B316EF27484C756058E">jhskevington @ gmail. com; https: // orcid. org / 0000 - 0002 - 1445 - 9870</mods:affiliation>
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<mods:nameIdentifier id="CBDA09EAE7B52BE170FCBEBADC40FAEA" type="email">gilfgm@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B04B0BC0BC4EB25F3C483D6A1B04E6D0">Marshall, Stephen A.</mods:namePart>
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<mods:nameIdentifier id="C5DBBD93422828D5D0847272708E969E" type="ORCID">0000-0001-7919-2639</mods:nameIdentifier>
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<mods:affiliation id="1BFA6CEDA502AF7170E4ED15A6C409DE">University of Guelph Insect Collection and Insect Systematics Laboratory - School of Environmental Sciences - University of Guelph. Guelph, Ontario, Canada.</mods:affiliation>
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<mods:nameIdentifier id="A81382E570ED22975770943E00014815" type="email">gilfgm@gmail.com</mods:nameIdentifier>
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<mods:title id="C074BA8EE2E0A85CE655F7EE135C8F15">Zootaxa</mods:title>
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<mods:part id="B3D836B3B41F6A19A62FC3C700010441">
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<mods:date id="0B6F3E774AA466B699B6EB4D8E758CB2">2020</mods:date>
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<mods:number id="7F33DC590B334A2576973143741D7828">2020-08-05</mods:number>
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<treatment id="A51D5F67FFEC3A6DFF50FD9EFCE39FF2" ID-DOI="http://doi.org/10.5281/zenodo.4449971" ID-GBIF-Taxon="177008123" ID-Zenodo-Dep="4449971" LSID="urn:lsid:zoobank.org:act:59261D3C-F2FC-46FC-B54E-E3B205B56290" httpUri="http://treatment.plazi.org/id/A51D5F67FFEC3A6DFF50FD9EFCE39FF2" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153">
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<subSubSection id="65AEBDFAFFEC3A6CFF50FD9EFDDC9D36" box="[151,575,547,573]" pageId="2" pageNumber="153" type="nomenclature">
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<paragraph id="2D0BEE71FFEC3A6CFF50FD9EFDDC9D36" blockId="2.[151,962,547,646]" box="[151,575,547,573]" pageId="2" pageNumber="153">
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<emphasis id="1FC03263FFEC3A6CFF50FD9EFDDC9D36" bold="true" box="[151,575,547,573]" pageId="2" pageNumber="153">
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Genus
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<taxonomicName id="EAB495F2FFEC3A6CFF29FD9EFE299D36" ID-CoL="89NNT" ID-ENA="2929881" authority="Miranda" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" box="[238,458,547,573]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="1FC03263FFEC3A6CFF29FD9EFEB09D36" bold="true" box="[238,339,547,573]" italics="true" pageId="2" pageNumber="153">Fragosa</emphasis>
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Miranda
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</taxonomicName>
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<taxonomicNameLabel id="04F38F18FFEC3A6CFE16FD9EFDDC9D36" box="[465,575,547,573]" pageId="2" pageNumber="153" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="65AEBDFAFFEC3A6CFF50FDFAFD459C0B" pageId="2" pageNumber="153" type="description">
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<paragraph id="2D0BEE71FFEC3A6CFF50FDFAFDC89D69" blockId="2.[151,962,547,646]" box="[151,555,583,610]" pageId="2" pageNumber="153">
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<figureCitation id="B58FF2F4FFEC3A6CFF50FDFAFEE49D6A" box="[151,263,583,609]" captionStart="FIGURES 1–11" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="2" pageNumber="153">Figs 1–11</figureCitation>
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,
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<figureCitation id="B58FF2F4FFEC3A6CFED4FDFAFED29D6A" box="[275,305,583,609]" captionStart="FIGURES 61–67. 61" captionStartId="15.[151,264,1624,1648]" captionTargetBox="[201,1387,181,1594]" captionTargetId="figure-18@15.[199,1387,181,1600]" captionTargetPageId="15" captionText="FIGURES 61–67. 61: Pseudoscaeva diversifasciata (Knab) (CNC_Diptera040088), female, katatergite, lateral. 62: Styxia eblis Hull (debu01088838), male, pleuron, lateral. 63: Hybobathus norina (Curran) (JSS25233), male, hypandrium, lateral. 64: Orphnabaccha aff. calda (CNC_Diptera210099), male, hypandrium, lateral. 65: Mimocalla erebus (Hull) (CNC_Diptera209355), male, epandrium, lateral, photo previously published in Mengual et al. (2018). 66: Fragosa argentina (Curran) (CNC_Diptera174232), female, tergum 1, dorsal. 67: Maiana callida (Hine) (CNC_Diptera105396), male, tergum 1, dorsal. c: cercus; ca: calypter; d: distiphallus; db: dorsal bridge; e: epandrium; h: hypandrium; k: katatergite; l: lobe; n: lateral notch; p: postgonite; ps: posterior spiracle; s: surstylus; t1: tergum 1." figureDoi="http://doi.org/10.5281/zenodo.4401383" httpUri="https://zenodo.org/record/4401383/files/figure.png" pageId="2" pageNumber="153">66</figureCitation>
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,
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<figureCitation id="B58FF2F4FFEC3A6CFEFAFDFAFE659D6A" box="[317,390,583,609]" captionStart="FIGURES 68–77. 68" captionStartId="16.[151,264,1424,1448]" captionTargetBox="[211,1375,205,1398]" captionTargetId="figure-19@16.[205,1381,181,1400]" captionTargetPageId="16" captionText="FIGURES 68–77. 68: Maiana callida (Hine) (CNC_Diptera105396), male, abdomen, ventral. 69: Maiana pumila (Austen) (JSS32861), male, terminalia, oblique fronto-ventral. 70–73: Hybobathus pola (Curran). 70: female terminalia (CNC_Diptera 161361), apex, dorsal. 71–72: male terminalia (CNC_Diptera 161366). 71: hypandrium, ventral. 72: terminalia, lateral. 73: holotype Baccha pola Curran (AMNH), male, habitus, dorsal, by AMNH. 74: Fragosa aurora (Hull) (holotype Baccha aurora Hull, AMNH), male, habitus, dorsal, by AMNH. 75: Nuntianus variegatus (Macquart) (holotype Baccha variegata Macquart, MNHNP, ED8265), female, habitus, dorsal, by MNHNP. 76: Fragosa virgilio (Hull) (holotype Baccha virgilio Hull, CNC, CNC_Diptera189124), male, habitus, dorsal, by CNC. 77: Victoriana oblonga (Walker) (holotype Baccha oblonga Walker, BMNH, NHMUK 010369937), female, habitus, dorsal, copyright of The Trustees of the Natural History Museum, London. c: cercus; e: epiproct; h: hypandrium; l: lobe; n: lateral notch; p: postgonite; ph: phallus; s: surstylus; s4 and 5: sternum 4 and 5; t7: tergum 7." figureDoi="http://doi.org/10.5281/zenodo.4401385" httpUri="https://zenodo.org/record/4401385/files/figure.png" pageId="2" pageNumber="153">74, 76</figureCitation>
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,
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<figureCitation id="B58FF2F4FFEC3A6CFE56FDFAFE399D6A" box="[401,474,583,609]" captionStart="FIGURES 78–88. 78" captionStartId="17.[151,264,1665,1689]" captionTargetBox="[301,1287,192,1634]" captionTargetId="figure-18@17.[294,1294,181,1642]" captionTargetPageId="17" captionText="FIGURES 78–88. 78: Fragosa harlequina (Hull) (holotype Baccha harlequina Hull, AMNH), male, thorax, lateral, by AMNH (line drawing represents outline of the alula). 79: Fragosa argentina (Curran) (CNC_Diptera174232), thorax, postero-ventral (line drawing represents outline of the incomplete post-metacoxal bridge). 80–81: Fragosa virgilio (Hull) (holotype Baccha virgilio Hull, CNC, CNC_Diptera189124), male. 80: habitus, lateral, by CNC. 81: head, oblique frontal, by CNC. 82: Fragosa titania (Hull) (JSS25229), female, head, dorsal. 83: Fragosa stenogaster (Williston) (debu01088840), male, head, dorsal. 84–85: Hypocritanus fascipennis (Wiedemann), female, head. 84: frontal, by Ximo Mengual (http://syrphidae.myspecies.info/ taxonomy/term/879). 85: lateral (CNC_Diptera501). 86–87: Nuntianus cecrops (Hull) (INPA-DIP000246), female, head. 86: dorsal. 87: lateral. 88: Victoriana aff. melanorrhina (DEBU), female, head, lateral. a: alula; ai: antennal insertion; f: frons; ft: frontal triangle; lem: lateral eye margin; mt: metafemur; o: ocellar triangle; oc: occiput; pmt: post-metacoxal bridge; t: facial tubercle." figureDoi="http://doi.org/10.5281/zenodo.4401387" httpUri="https://zenodo.org/record/4401387/files/figure.png" pageId="2" pageNumber="153">78–83</figureCitation>
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,
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<figureCitation id="B58FF2F4FFEC3A6CFE22FDFAFDC89D69" box="[485,555,583,610]" captionStart="FIGURES 89–102. 89" captionStartId="18.[151,264,1641,1665]" captionTargetBox="[209,1377,189,1607]" captionTargetId="figure-19@18.[203,1383,181,1617]" captionTargetPageId="18" captionText="FIGURES 89–102. 89: Victoriana sativa (Curran) (holotype Baccha sativa Curran, AMNH), habitus, dorsal, by AMNH. 90– 91: Victoriana laudabilis (Williston) (holotype Baccha laudabilis Williston, BMNH, NHMUK010369870), copyright of The Trustees of the Natural History Museum, London. 90: base of abdomen and metafemur, dorsal. 91: base of wing, dorsal (line drawing represents outline of the alula). 92–95: Fragosa sp. (species group tenuis, CNC_Diptera196012), male. 92: hypandrium, lateral. 93: surstylus, ventro-lateral. 94: epandrium, lateral. 95: epandrium, ventral (line drawing represents outline of the subepandrial sclerite). 96–97: Nuntianus abata (Curran) (CNC_Diptera209361), male, hypandrium. 96: lateral. 97: ventral. 98: Syrphus conjunctus Wiedemann (currently incertae sedis, CNC_Diptera209347), male, hypandrium, lateral. 99: Victoriana aff. melanorrhina (debu00279441), male, hypandrium, ventral. 100: Victoriana aff. melanorrhina (DEBU), female, terminalia, dorsal. 101: Victoriana parvicornis (Loew) (CNC_Diptera237922), terminalia, dorsal. 102: Maiana pumila (Austen) (debu00178088), female, terminalia, dorsal. a: alula; c: cercus; d: distiphallus; e: epiproct; l: lobe; p: baso-posterior pile; t7 and 8: tergum 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401389" httpUri="https://zenodo.org/record/4401389/files/figure.png" pageId="2" pageNumber="153">92–95</figureCitation>
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</paragraph>
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<paragraph id="2D0BEE71FFEC3A6CFF50FDD6FC219D8D" blockId="2.[151,962,547,646]" box="[151,962,619,646]" pageId="2" pageNumber="153">
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<uri id="5925E273FFEC3A6CFF50FDD6FC219D8D" box="[151,962,619,646]" pageId="2" pageNumber="153">
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urn:lsid:zoobank.org:act:
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<uuid id="5912D4A4FFEC3A6CFE61FDD6FC219D8D" box="[422,962,619,646]" pageId="2" pageNumber="153">59261D3C-F2FC-46FC-B54E-E3B205B56290</uuid>
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</uri>
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</paragraph>
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<paragraph id="2D0BEE71FFEC3A6CFF50FD16FD9C9DC9" blockId="2.[151,1319,682,799]" box="[151,639,682,707]" pageId="2" pageNumber="153">
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<typeStatus id="F20F50D3FFEC3A6CFF50FD16FF2A9DC8" box="[151,201,683,707]" pageId="2" pageNumber="153">Type</typeStatus>
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species:
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<taxonomicName id="EAB495F2FFEC3A6CFEE1FD17FD9C9DC9" authority="Schiner, 1868" authorityName="Schiner" authorityYear="1868" box="[294,639,682,706]" class="Insecta" family="Syrphidae" genus="Baccha" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="rugosifrons">
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<emphasis id="1FC03263FFEC3A6CFEE1FD17FE0E9DC9" box="[294,493,682,706]" italics="true" pageId="2" pageNumber="153">Baccha rugosifrons</emphasis>
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<bibRefCitation id="49259380FFEC3A6CFE34FD17FD9C9DC9" author="Schiner, I. R." box="[499,639,682,706]" pageId="2" pageNumber="153" refId="ref17881" refString="Schiner, I. R. (1868) Diptera. In: Wullerstorf-Urbair, B. von (in charge), Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf-Urbair. Vol. 2. Band 1. Abtheilung B. Kaiserlich-konigliche Hof- und Staatsdruckerei in Commission bei Karl Gerold's Sohn, Wien, pp. i-vi + 1 - 388, 4 pls." type="journal volume" year="1868">Schiner, 1868</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="2D0BEE71FFEC3A6CFF50FD74FD779DEA" blockId="2.[151,1319,682,799]" box="[151,660,713,737]" pageId="2" pageNumber="153">
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<taxonomicName id="EAB495F2FFEC3A6CFF50FD74FE979DEA" authorityName="Loew" authorityYear="1863" box="[151,372,713,737]" class="Insecta" family="Syrphidae" genus="Baccha" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="obscuricornis">
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<emphasis id="1FC03263FFEC3A6CFF50FD74FE979DEA" box="[151,372,713,737]" italics="true" pageId="2" pageNumber="153">Baccha obscuricornis</emphasis>
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</taxonomicName>
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species group.
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<bibRefCitation id="49259380FFEC3A6CFDD5FD74FD779DEA" author="Hull, F. M." box="[530,660,713,737]" pageId="2" pageNumber="153" pagination="89 - 285" refId="ref16669" refString="Hull, F. M. (1949 a) The genus Baccha from the New World. Entomologica Americana, 27, 89 - 285, 47 pls. [1947]" type="journal article" year="1949">Hull (1949a)</bibRefCitation>
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</paragraph>
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<paragraph id="2D0BEE71FFEC3A6CFF50FD55FD459C0B" blockId="2.[151,1319,682,799]" box="[151,678,744,768]" pageId="2" pageNumber="153">
|
||
<taxonomicName id="EAB495F2FFEC3A6CFF50FD55FEDB9DF4" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" baseAuthorityName="Hull" baseAuthorityYear="1941" box="[151,312,744,768]" class="Insecta" family="Syrphidae" genus="Baccha" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="victoria">
|
||
<emphasis id="1FC03263FFEC3A6CFF50FD55FEDB9DF4" box="[151,312,744,768]" italics="true" pageId="2" pageNumber="153">Baccha victoria</emphasis>
|
||
</taxonomicName>
|
||
species group, in part.
|
||
<bibRefCitation id="49259380FFEC3A6CFDE3FD55FD459C0B" author="Hull, F. M." box="[548,678,744,768]" pageId="2" pageNumber="153" pagination="89 - 285" refId="ref16669" refString="Hull, F. M. (1949 a) The genus Baccha from the New World. Entomologica Americana, 27, 89 - 285, 47 pls. [1947]" type="journal article" year="1949">Hull (1949a)</bibRefCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65AEBDFAFFEC3A6CFF50FCBAFAC49C15" box="[151,1319,774,799]" pageId="2" pageNumber="153" type="reference_group">
|
||
<paragraph id="2D0BEE71FFEC3A6CFF50FCBAFAC49C15" blockId="2.[151,1319,682,799]" box="[151,1319,774,799]" pageId="2" pageNumber="153">
|
||
<taxonomicName id="EAB495F2FFEC3A6CFF50FCBAFE679C15" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" baseAuthorityName="Williston" baseAuthorityYear="1888" box="[151,388,775,798]" class="Insecta" family="Syrphidae" genus="Ocyptamus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="stenogaster">
|
||
<emphasis id="1FC03263FFEC3A6CFF50FCBAFE679C15" box="[151,388,775,798]" italics="true" pageId="2" pageNumber="153">Ocyptamus stenogaster</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
<bibRefCitation id="49259380FFEC3A6CFDE6FCBBFD389C15" author="Thompson, F. C." box="[545,731,774,798]" pageId="2" pageNumber="153" pagination="1 - 200" refId="ref18053" refString="Thompson, F. C. (1981) The flower flies of the West Indies (Diptera: Syrphidae). Memoirs of the Entomological Society of Washington, 9, 1 - 200." type="journal article" year="1981">Thompson (1981)</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="49259380FFEC3A6CFD22FCBBFC5F9C15" author="Mengual, X. & Stahls, G. & Rojo, S." box="[741,956,774,798]" pageId="2" pageNumber="153" pagination="191 - 205" refId="ref17150" refString="Mengual, X., Stahls, G. & Rojo, S. (2012) Is the mega-diverse genus Ocyptamus (Diptera, Syrphidae) monophyletic? Evidence from molecular characters including the secondary structure of 28 S rRNA. Molecular Phylogenetics and Evolution, 62, 191 - 205. https: // doi. org / 10.1016 / j. ympev. 2011.09.014" type="journal article" year="2012">
|
||
Mengual
|
||
<emphasis id="1FC03263FFEC3A6CFC82FCBAFC919C15" box="[837,882,774,798]" italics="true" pageId="2" pageNumber="153">et al</emphasis>
|
||
. (2012
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="49259380FFEC3A6CFC01FCBBFC1E9C15" author="Mengual, X. & Miranda, G. F. G. & Thompson, F. C." box="[966,1021,774,798]" pageId="2" pageNumber="153" pagination="1 - 44" refId="ref17217" refString="Mengual, X., Miranda, G. F. G. & Thompson, F. C. (2018) Unraveling Ocyptamus and the Baccha legacy (Diptera: Syrphidae): redefinition of groups and new species descriptions. Zootaxa, 4461 (1), 1 - 44. https: // doi. org / 10.11646 / zootaxa. 4461.1.1" type="journal article" year="2018">2018</bibRefCitation>
|
||
);
|
||
<bibRefCitation id="49259380FFEC3A6CFBC8FCBBFB029C15" author="Miranda, G. F. G. & Marshall, S. A. & Skevington, J. H." box="[1039,1249,774,798]" pageId="2" pageNumber="153" pagination="1 - 154" refId="ref17439" refString="Miranda, G. F. G., Marshall, S. A. & Skevington, J. H. (2014) Revision of the genus Pelecinobaccha Shannon, description of Relictanum gen. nov. and redescription of Atylobaccha flukiella (Curran, 1941). Zootaxa, 3819 (1), 1 - 154. https: // doi. org / 10.11646 / zootaxa. 3819.1.1" type="journal article" year="2014">
|
||
Miranda
|
||
<emphasis id="1FC03263FFEC3A6CFBACFCBAFB749C15" box="[1131,1175,774,798]" italics="true" pageId="2" pageNumber="153">et al</emphasis>
|
||
. (2014
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="49259380FFEC3A6CFB2BFCBBFAC39C15" author="Miranda, G. F. G. & Skevington, J. H. & Marshall, S. A. & Kelso, S." box="[1260,1312,774,798]" pageId="2" pageNumber="153" pagination="161 - 176" refId="ref17512" refString="Miranda, G. F. G., Skevington, J. H., Marshall, S. A. & Kelso, S. (2016) The genus Ocyptamus Macquart (Diptera: Syrphidae): A molecular phylogenetic analysis. Arthropod Systematics and Phylogeny, 74, 161 - 176." type="journal article" year="2016">2016</bibRefCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65AEBDFAFFEC3A6CFF50FCF1FDA09A54" pageId="2" pageNumber="153" type="description">
|
||
<paragraph id="2D0BEE71FFEC3A6CFF50FCF1FDA09A54" blockId="2.[151,1437,844,2023]" pageId="2" pageNumber="153">
|
||
<emphasis id="1FC03263FFEC3A6CFF50FCF1FECA9C6D" bold="true" box="[151,297,844,870]" pageId="2" pageNumber="153">Description.</emphasis>
|
||
Delicate-looking flies.
|
||
<emphasis id="1FC03263FFEC3A6CFDF6FCF1FD8C9C6C" bold="true" box="[561,623,844,871]" italics="true" pageId="2" pageNumber="153">Head</emphasis>
|
||
. Face very narrow to narrow (around 1/4 to less than 1/3 of head width). Frons usually ~1/3 of head width. Antennal insertions confluent. Female ocellar triangle adjacent or separated by approximately one ocellus-width from lateral eye margin. Dorsal occiput usually with two rows of pile (sometimes with one row).
|
||
<emphasis id="1FC03263FFEC3A6CFE8BFC05FE419CD8" bold="true" box="[332,418,952,979]" italics="true" pageId="2" pageNumber="153">Thorax</emphasis>
|
||
. Scutum without distinct anterior row of pile. Anterior anepisternum pilose. Katatergite with short microtrichia that gives the sclerite a ‘velvet’ appearance. Metaepisternum pilose. Metasternum bare. Upper calypter margin either bare, with pile shorter than pile on the ventral calypter margin, or with longer darker pile. Metafemur with normal pile.
|
||
<emphasis id="1FC03263FFEC3A6CFDD9FB99FDB89B35" bold="true" box="[542,603,1060,1086]" italics="true" pageId="2" pageNumber="153">Wing</emphasis>
|
||
. Alula often absent (some species with a trace of alula, e.g.
|
||
<taxonomicName id="EAB495F2FFEC3A6CFAC2FB98FA7F9B35" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" baseAuthorityName="Hull" baseAuthorityYear="1948" box="[1285,1436,1060,1086]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="harlequina">
|
||
<emphasis id="1FC03263FFEC3A6CFAC2FB98FA7F9B35" box="[1285,1436,1060,1086]" italics="true" pageId="2" pageNumber="153">F. harlequina</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B58FF2F4FFEC3A6CFF67FBF5FF129B68" box="[160,241,1096,1123]" captionStart="FIGURES 78–88. 78" captionStartId="17.[151,264,1665,1689]" captionTargetBox="[301,1287,192,1634]" captionTargetId="figure-18@17.[294,1294,181,1642]" captionTargetPageId="17" captionText="FIGURES 78–88. 78: Fragosa harlequina (Hull) (holotype Baccha harlequina Hull, AMNH), male, thorax, lateral, by AMNH (line drawing represents outline of the alula). 79: Fragosa argentina (Curran) (CNC_Diptera174232), thorax, postero-ventral (line drawing represents outline of the incomplete post-metacoxal bridge). 80–81: Fragosa virgilio (Hull) (holotype Baccha virgilio Hull, CNC, CNC_Diptera189124), male. 80: habitus, lateral, by CNC. 81: head, oblique frontal, by CNC. 82: Fragosa titania (Hull) (JSS25229), female, head, dorsal. 83: Fragosa stenogaster (Williston) (debu01088840), male, head, dorsal. 84–85: Hypocritanus fascipennis (Wiedemann), female, head. 84: frontal, by Ximo Mengual (http://syrphidae.myspecies.info/ taxonomy/term/879). 85: lateral (CNC_Diptera501). 86–87: Nuntianus cecrops (Hull) (INPA-DIP000246), female, head. 86: dorsal. 87: lateral. 88: Victoriana aff. melanorrhina (DEBU), female, head, lateral. a: alula; ai: antennal insertion; f: frons; ft: frontal triangle; lem: lateral eye margin; mt: metafemur; o: ocellar triangle; oc: occiput; pmt: post-metacoxal bridge; t: facial tubercle." figureDoi="http://doi.org/10.5281/zenodo.4401387" httpUri="https://zenodo.org/record/4401387/files/figure.png" pageId="2" pageNumber="153">Fig. 78</figureCitation>
|
||
) and
|
||
<taxonomicName id="EAB495F2FFEC3A6CFEF6FBF4FE6B9B69" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" baseAuthorityName="Curran" baseAuthorityYear="1941" box="[305,392,1097,1122]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="mara">
|
||
<emphasis id="1FC03263FFEC3A6CFEF6FBF4FE6B9B69" box="[305,392,1097,1122]" italics="true" pageId="2" pageNumber="153">F. mara</emphasis>
|
||
</taxonomicName>
|
||
). Male wing light brown, usually bare on small basal areas of cells c, r and bm; female wing pigmentation and microtrichosity variable; anal lobe narrowed basally.
|
||
<emphasis id="1FC03263FFEC3A6CFC68FBD1FBFE9B8C" bold="true" box="[943,1053,1132,1159]" italics="true" pageId="2" pageNumber="153">Abdomen</emphasis>
|
||
. Abdomen slender; tergum 1 crescent-shaped with lateral extremities directed laterally; terga usually only with quadrangular pale maculae on the baso-lateral corners of terga 3 and 4 (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFDF2FB09FD999BC4" box="[565,634,1204,1231]" captionStart="FIGURES 1–11" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="2" pageNumber="153">Fig. 2</figureCitation>
|
||
).
|
||
<emphasis id="1FC03263FFEC3A6CFD49FB09FCEE9BC4" bold="true" box="[654,781,1204,1231]" italics="true" pageId="2" pageNumber="153">Terminalia</emphasis>
|
||
. *Female tergum 7 quadrangular (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFB4AFB09FB339BC4" box="[1165,1232,1204,1231]" captionStart="FIGURES 1–11" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="2" pageNumber="153">Fig. 8</figureCitation>
|
||
); female tergum 8 rectangular with concave basal and apical margins; *epiproct partially divided into a pair of separate sclerites connected to each other by a basal narrow strip or more extensively fused, each with baso-lateral apodemes extending into the abdominal cavity (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFE07FA9DFDE09A30" box="[448,515,1312,1339]" captionStart="FIGURES 1–11" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="2" pageNumber="153">Fig. 8</figureCitation>
|
||
). Male basiphallus teardrop-shaped (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFC59FA9DFC0D9A30" box="[926,1006,1312,1339]" captionStart="FIGURES 1–11" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="2" pageNumber="153">Fig. 11</figureCitation>
|
||
), distiphallus membranous with dorsal sclerotized triangular region (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFE25FAF9FDD59A54" box="[482,566,1348,1375]" captionStart="FIGURES 1–11" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="2" pageNumber="153">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65AEBDFAFFEC3A6CFF00FAD4FEE19AAD" pageId="2" pageNumber="153" type="etymology">
|
||
<paragraph id="2D0BEE71FFEC3A6CFF00FAD4FEE19AAD" blockId="2.[151,1437,844,2023]" pageId="2" pageNumber="153">
|
||
<emphasis id="1FC03263FFEC3A6CFF00FAD4FEAD9A88" bold="true" box="[199,334,1385,1411]" pageId="2" pageNumber="153">Etymology.</emphasis>
|
||
The name is a reference to the ‘brittle’ (Latin), delicate, aspect of these flies. It is to be treated as feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65AEBDFAFFEC3A6DFF00FA0DFCE39FF2" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153" type="discussion">
|
||
<paragraph id="2D0BEE71FFEC3A6CFF00FA0DFC4F99E1" blockId="2.[151,1437,844,2023]" pageId="2" pageNumber="153">
|
||
<emphasis id="1FC03263FFEC3A6CFF00FA0DFEAB9AC1" bold="true" box="[199,328,1456,1482]" pageId="2" pageNumber="153">Comments</emphasis>
|
||
. This genus, equivalent to the former
|
||
<taxonomicName id="EAB495F2FFEC3A6CFD2DFA0CFC0D9AC1" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" baseAuthorityName="Williston" baseAuthorityYear="1888" box="[746,1006,1457,1483]" class="Insecta" family="Syrphidae" genus="Ocyptamus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="stenogaster">
|
||
<emphasis id="1FC03263FFEC3A6CFD2DFA0CFC0D9AC1" box="[746,1006,1457,1483]" italics="true" pageId="2" pageNumber="153">Ocyptamus stenogaster</emphasis>
|
||
</taxonomicName>
|
||
species group, is made up of relatively delicate flies (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFEF4FA69FE399AE4" box="[307,474,1492,1519]" captionStart="FIGURES 1–11" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="2" pageNumber="153">Figs 1, 2 and 6</figureCitation>
|
||
) superficially similar to
|
||
<taxonomicName id="EAB495F2FFEC3A6CFD2BFA69FCA39AE5" authorityName="Fabricius" authorityYear="1805" box="[748,832,1492,1518]" class="Insecta" family="Syrphidae" genus="Baccha" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FC03263FFEC3A6CFD2BFA69FCA39AE5" box="[748,832,1492,1518]" italics="true" pageId="2" pageNumber="153">Baccha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EAB495F2FFEC3A6CFC89FA69FC059AE5" authorityName="Hull" authorityYear="1949" box="[846,998,1492,1518]" class="Insecta" family="Syrphidae" genus="Leucopodella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FC03263FFEC3A6CFC89FA69FC059AE5" box="[846,998,1492,1518]" italics="true" pageId="2" pageNumber="153">Leucopodella</emphasis>
|
||
</taxonomicName>
|
||
, and the species of the
|
||
<taxonomicName id="EAB495F2FFEC3A6CFB2EFA69FEFA9919" authorityName="Schiner" authorityYear="1868" class="Insecta" family="Syrphidae" genus="Pelecinobaccha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="brevipennis">
|
||
<emphasis id="1FC03263FFEC3A6CFB2EFA69FEFA9919" italics="true" pageId="2" pageNumber="153">Pelecinobaccha brevipennis</emphasis>
|
||
</taxonomicName>
|
||
group. Most species lack an alula (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFD74FA45FD1A9918" box="[691,761,1528,1555]" captionStart="FIGURES 1–11" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="2" pageNumber="153">Fig. 3</figureCitation>
|
||
), or have it reduced to a barely discernible trace (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFAFEFA45FA6C9918" box="[1337,1423,1528,1555]" captionStart="FIGURES 78–88. 78" captionStartId="17.[151,264,1665,1689]" captionTargetBox="[301,1287,192,1634]" captionTargetId="figure-18@17.[294,1294,181,1642]" captionTargetPageId="17" captionText="FIGURES 78–88. 78: Fragosa harlequina (Hull) (holotype Baccha harlequina Hull, AMNH), male, thorax, lateral, by AMNH (line drawing represents outline of the alula). 79: Fragosa argentina (Curran) (CNC_Diptera174232), thorax, postero-ventral (line drawing represents outline of the incomplete post-metacoxal bridge). 80–81: Fragosa virgilio (Hull) (holotype Baccha virgilio Hull, CNC, CNC_Diptera189124), male. 80: habitus, lateral, by CNC. 81: head, oblique frontal, by CNC. 82: Fragosa titania (Hull) (JSS25229), female, head, dorsal. 83: Fragosa stenogaster (Williston) (debu01088840), male, head, dorsal. 84–85: Hypocritanus fascipennis (Wiedemann), female, head. 84: frontal, by Ximo Mengual (http://syrphidae.myspecies.info/ taxonomy/term/879). 85: lateral (CNC_Diptera501). 86–87: Nuntianus cecrops (Hull) (INPA-DIP000246), female, head. 86: dorsal. 87: lateral. 88: Victoriana aff. melanorrhina (DEBU), female, head, lateral. a: alula; ai: antennal insertion; f: frons; ft: frontal triangle; lem: lateral eye margin; mt: metafemur; o: ocellar triangle; oc: occiput; pmt: post-metacoxal bridge; t: facial tubercle." figureDoi="http://doi.org/10.5281/zenodo.4401387" httpUri="https://zenodo.org/record/4401387/files/figure.png" pageId="2" pageNumber="153">Fig. 78</figureCitation>
|
||
), and have a reduced anal lobe, very long and narrow abdominal segments, a crescent-shaped tergum 1 with lateral extremities directed laterally (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFE21F9FDFDDA9950" box="[486,569,1600,1627]" captionStart="FIGURES 61–67. 61" captionStartId="15.[151,264,1624,1648]" captionTargetBox="[201,1387,181,1594]" captionTargetId="figure-18@15.[199,1387,181,1600]" captionTargetPageId="15" captionText="FIGURES 61–67. 61: Pseudoscaeva diversifasciata (Knab) (CNC_Diptera040088), female, katatergite, lateral. 62: Styxia eblis Hull (debu01088838), male, pleuron, lateral. 63: Hybobathus norina (Curran) (JSS25233), male, hypandrium, lateral. 64: Orphnabaccha aff. calda (CNC_Diptera210099), male, hypandrium, lateral. 65: Mimocalla erebus (Hull) (CNC_Diptera209355), male, epandrium, lateral, photo previously published in Mengual et al. (2018). 66: Fragosa argentina (Curran) (CNC_Diptera174232), female, tergum 1, dorsal. 67: Maiana callida (Hine) (CNC_Diptera105396), male, tergum 1, dorsal. c: cercus; ca: calypter; d: distiphallus; db: dorsal bridge; e: epandrium; h: hypandrium; k: katatergite; l: lobe; n: lateral notch; p: postgonite; ps: posterior spiracle; s: surstylus; t1: tergum 1." figureDoi="http://doi.org/10.5281/zenodo.4401383" httpUri="https://zenodo.org/record/4401383/files/figure.png" pageId="2" pageNumber="153">Fig. 66</figureCitation>
|
||
), and (usually) quadrangular pale maculae on the baso-lateral corners of terga 3 and 4 (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFF3DF9D9FEDC9974" box="[250,319,1636,1663]" captionStart="FIGURES 1–11" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="2" pageNumber="153">Fig. 2</figureCitation>
|
||
). They usually have a very long and narrow 2
|
||
<superScript id="DAC14339FFEC3A6CFC85F9D9FCB19979" attach="left" box="[834,850,1636,1650]" fontSize="6" pageId="2" pageNumber="153">nd</superScript>
|
||
abdominal segment (over 8 times longer than wide) (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFF58F935FEE199A8" box="[159,258,1672,1699]" captionStart="FIGURES 1–11" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="2" pageNumber="153">Figs 1, 2</figureCitation>
|
||
) with a naturally arched tergum giving the segment a cylindrical appearance. The unique condition of the female epiproct (see description above) is distinctive and provides strong evidence for the monophyly of the genus. Facial coloration seems to vary among specimens of the same species.
|
||
</paragraph>
|
||
<paragraph id="2D0BEE71FFEC3A6CFF00F948FAEE98ED" blockId="2.[151,1437,844,2023]" pageId="2" pageNumber="153">
|
||
<taxonomicName id="EAB495F2FFEC3A6CFF00F948FEC59805" authority="Miranda & Skevington & Marshall, 2020" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" box="[199,294,1781,1806]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="1FC03263FFEC3A6CFF00F948FEC59805" box="[199,294,1781,1806]" italics="true" pageId="2" pageNumber="153">Fragosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FC03263FFEC3A6CFEEBF948FE779804" bold="true" box="[300,404,1781,1807]" pageId="2" pageNumber="153">
|
||
<taxonomicNameLabel id="04F38F18FFEC3A6CFEEBF948FE779804" box="[300,404,1781,1807]" pageId="2" pageNumber="153" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="EAB495F2FFEC3A6CFE0EF948FD829805" authority="Miranda & Skevington & Marshall, 2020" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" box="[457,609,1781,1806]" class="Insecta" family="Syrphidae" genus="Hypocritanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="1FC03263FFEC3A6CFE0EF948FD829805" box="[457,609,1781,1806]" italics="true" pageId="2" pageNumber="153">Hypocritanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FC03263FFEC3A6CFDA0F948FD2C9804" bold="true" box="[615,719,1781,1807]" pageId="2" pageNumber="153">
|
||
<taxonomicNameLabel id="04F38F18FFEC3A6CFDA0F948FD2C9804" box="[615,719,1781,1807]" pageId="2" pageNumber="153" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
share a dorsally sclerotized female cercus with setulae restricted dorsally (
|
||
<figureCitation id="B58FF2F4FFEC3A6CFEC3F8A5FEB39838" box="[260,336,1816,1843]" captionStart="FIGURES 1–11" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 1–11. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 9–11. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="2" pageNumber="153">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="B58FF2F4FFEC3A6CFEA6F8A4FE4E9838" box="[353,429,1816,1843]" captionStart="FIGURES 12–22. 12–15" captionStartId="6.[151,264,1700,1724]" captionTargetBox="[259,1328,566,1667]" captionTargetId="figure-200@6.[248,1339,561,1677]" captionTargetPageId="6" captionText="FIGURES 12–22. 12–15. Hypocritanus Miranda gen. nov.. 12: Hypocritanus fascipennis (Wiedemann), female, habitus, dorsal view, by Steve Marshall. 13: Hypocritanus fascipennis (Wiedemann) (debu01088980), male, epandrium, ventral view. 14–15. Hypocritanus fascipennis (Wiedemann) (debu00119165), female, terminalia. 14: dorsal view. 15: lateral view. 16–22. Maiana Miranda gen. nov. 16: Maiana pumila (Austen), female, habitus, dorsal view, by Steve Marshall. 17: Maiana sp. (INPA- DIP000172), male, head, dorsal view. 18: Maiana pumila (Austen) (JSS32861), male, wing. 19: Maiana pumila (Austen) (debu00178088), female, scutellum and tergum 1, dorsal view. 20–22. Maiana pumila (Austen) (JSS32861), male, terminalia. 20: epandrium, lateral view. 21: epandrium, ventral view. 22: hypandrium, lateral view. c: cercus; e: epiproct; h: hypandrium; p: postgonite; ph: phallus; s: surstylus; se: subepandrial sclerite; l: lobe; le: lobular expansion; lem: lateral eye margin; o: ocellar triangle; pem: posterior eye margin; t1: tergum 1." figureDoi="http://doi.org/10.5281/zenodo.4401373" httpUri="https://zenodo.org/record/4401373/files/figure.png" pageId="2" pageNumber="153">14, 15</figureCitation>
|
||
), however, we consider this condition to be a plesiomorphy since recent phylogenetic analyses always keep them as separate taxa (
|
||
<bibRefCitation id="49259380FFEC3A6CFD56F881FC98985C" author="Mengual, X. & Stahls, G. & Rojo, S." box="[657,891,1852,1879]" pageId="2" pageNumber="153" pagination="191 - 205" refId="ref17150" refString="Mengual, X., Stahls, G. & Rojo, S. (2012) Is the mega-diverse genus Ocyptamus (Diptera, Syrphidae) monophyletic? Evidence from molecular characters including the secondary structure of 28 S rRNA. Molecular Phylogenetics and Evolution, 62, 191 - 205. https: // doi. org / 10.1016 / j. ympev. 2011.09.014" type="journal article" year="2012">
|
||
Mengual
|
||
<emphasis id="1FC03263FFEC3A6CFD39F880FCD3985D" box="[766,816,1852,1878]" italics="true" pageId="2" pageNumber="153">et al</emphasis>
|
||
. 2012
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="49259380FFEC3A6CFC40F881FB88985C" author="Miranda, G. F. G. & Skevington, J. H. & Marshall, S. A. & Kelso, S." box="[903,1131,1852,1879]" pageId="2" pageNumber="153" pagination="161 - 176" refId="ref17512" refString="Miranda, G. F. G., Skevington, J. H., Marshall, S. A. & Kelso, S. (2016) The genus Ocyptamus Macquart (Diptera: Syrphidae): A molecular phylogenetic analysis. Arthropod Systematics and Phylogeny, 74, 161 - 176." type="journal article" year="2016">
|
||
<taxonomicName id="EAB495F2FFEC3A6CFC40F881FC05985D" authority="Miranda & Skevington & Marshall, 2020" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" box="[903,998,1852,1878]" class="Arachnida" family="Araneidae" genus="Miranda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">Miranda</taxonomicName>
|
||
<emphasis id="1FC03263FFEC3A6CFC29F880FBC3985D" box="[1006,1056,1852,1878]" italics="true" pageId="2" pageNumber="153">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
). Furthermore, a single genus including both taxa would be hard to diagnose, while keeping them separated makes both genera distinct and diagnosable with their own unique characters. Both taxa prey on
|
||
<taxonomicName id="EAB495F2FFEC3A6CFCA8F839FB569894" authority="(Hemiptera)" baseAuthorityName="Hemiptera" box="[879,1205,1924,1951]" class="Insecta" family="Pseudococcidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="family">Pseudococcidae (Hemiptera)</taxonomicName>
|
||
, and we hypothesize that the Neotropical
|
||
<taxonomicName id="EAB495F2FFEC3A6CFEBCF814FE3998C9" authority="Miranda & Skevington & Marshall, 2020" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" box="[379,474,1961,1986]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="1FC03263FFEC3A6CFEBCF814FE3998C9" box="[379,474,1961,1986]" italics="true" pageId="2" pageNumber="153">Fragosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FC03263FFEC3A6CFE27F814FDAB98C8" bold="true" box="[480,584,1961,1987]" pageId="2" pageNumber="153">
|
||
<taxonomicNameLabel id="04F38F18FFEC3A6CFE27F814FDAB98C8" box="[480,584,1961,1987]" pageId="2" pageNumber="153" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and the Nearctic
|
||
<taxonomicName id="EAB495F2FFEC3A6CFCCCF814FC4098C9" authority="Miranda & Skevington & Marshall, 2020" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" box="[779,931,1961,1986]" class="Insecta" family="Syrphidae" genus="Hypocritanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="1FC03263FFEC3A6CFCCCF814FC4098C9" box="[779,931,1961,1986]" italics="true" pageId="2" pageNumber="153">Hypocritanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FC03263FFEC3A6CFC6DF814FBF198C8" bold="true" box="[938,1042,1961,1987]" pageId="2" pageNumber="153">
|
||
<taxonomicNameLabel id="04F38F18FFEC3A6CFC6DF814FBF198C8" box="[938,1042,1961,1987]" pageId="2" pageNumber="153" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
probably evolved independently to occupy the same ecological niche, explaining the absence of the other in each biogeographical region.
|
||
</paragraph>
|
||
<paragraph id="2D0BEE71FFED3A6DFF00FF2AFCE39FF2" blockId="3.[151,1437,151,249]" pageId="3" pageNumber="154">
|
||
Based on morphological characters observed, and the sister group relationship of
|
||
<taxonomicName id="EAB495F2FFED3A6DFB9BFF2AFB229FBB" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" baseAuthorityName="Hull" baseAuthorityYear="1943" box="[1116,1217,151,176]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="titania">
|
||
<emphasis id="1FC03263FFED3A6DFB9BFF2AFB229FBB" box="[1116,1217,151,176]" italics="true" pageId="3" pageNumber="154">F. titania</emphasis>
|
||
</taxonomicName>
|
||
with the remaining
|
||
<taxonomicName id="EAB495F2FFED3A6DFF50FF06FF159FDF" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" box="[151,246,187,212]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FC03263FFED3A6DFF50FF06FF159FDF" box="[151,246,187,212]" italics="true" pageId="3" pageNumber="154">Fragosa</emphasis>
|
||
</taxonomicName>
|
||
in a previous phylogenetic analysis (
|
||
<bibRefCitation id="49259380FFED3A6DFD58FF06FC659FDD" author="Miranda, G. F. G. & Skevington, J. H. & Marshall, S. A. & Kelso, S." box="[671,902,187,214]" pageId="3" pageNumber="154" pagination="161 - 176" refId="ref17512" refString="Miranda, G. F. G., Skevington, J. H., Marshall, S. A. & Kelso, S. (2016) The genus Ocyptamus Macquart (Diptera: Syrphidae): A molecular phylogenetic analysis. Arthropod Systematics and Phylogeny, 74, 161 - 176." type="journal article" year="2016">
|
||
<taxonomicName id="EAB495F2FFED3A6DFD58FF06FCE39FDE" authority="Miranda & Skevington & Marshall, 2020" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" box="[671,768,187,213]" class="Arachnida" family="Araneidae" genus="Miranda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">Miranda</taxonomicName>
|
||
<emphasis id="1FC03263FFED3A6DFCCFFF06FCA19FDE" box="[776,834,187,213]" italics="true" pageId="3" pageNumber="154">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
), we propose two species groups for the genus
|
||
<taxonomicName id="EAB495F2FFED3A6DFF50FF62FF159FF3" authority="Miranda & Skevington & Marshall, 2020" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" box="[151,246,223,248]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="1FC03263FFED3A6DFF50FF62FF159FF3" box="[151,246,223,248]" italics="true" pageId="3" pageNumber="154">Fragosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FC03263FFED3A6DFF3AFF62FE859FF2" bold="true" box="[253,358,223,249]" pageId="3" pageNumber="154">
|
||
<taxonomicNameLabel id="04F38F18FFED3A6DFF3AFF62FE859FF2" box="[253,358,223,249]" pageId="3" pageNumber="154" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
:
|
||
<taxonomicName id="EAB495F2FFED3A6DFEB2FF62FDF39FF2" authorityName="Schiner" authorityYear="1868" box="[373,528,223,249]" class="Insecta" family="Syrphidae" genus="Baccha" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="rugosifrons">
|
||
<emphasis id="1FC03263FFED3A6DFEB2FF62FDF39FF2" box="[373,528,223,249]" italics="true" pageId="3" pageNumber="154">F. rugosifrons</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EAB495F2FFED3A6DFD80FF62FD469FF3" authorityName="Miranda & Skevington & Marshall" authorityYear="2020" baseAuthorityName="Walker" baseAuthorityYear="1852" box="[583,677,223,248]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="tenuis">
|
||
<emphasis id="1FC03263FFED3A6DFD80FF62FD469FF3" box="[583,677,223,248]" italics="true" pageId="3" pageNumber="154">F. tenuis</emphasis>
|
||
</taxonomicName>
|
||
groups.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |