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<document id="5F8CB65D5EAA4F7E67E6F705EE0AC27F" ID-CLB-Dataset="20828" ID-DOI="10.5252/geodiversitas2022v44a4" ID-GBIF-Dataset="7228fba0-fd8b-4e14-8602-a68f0f6c955c" ID-ISSN="1638-9395" ID-Zenodo-Dep="6000901" ID-ZooBank="urn:lsid:zoobank.org:pub:CAF128F4-1301-4620-8BB1-17A668A32723" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643645900571" checkinUser="felipe" docAuthor="Haug, Joachim T., Hörnig, Marie K., Kiesmüller, Christine, Pazinato, Paula G., Baranov, Viktor &amp; Haug, Carolin" docDate="2022" docId="3E1E33385B52EC1FFC9AFA6CFE6CD97B" docLanguage="en" docName="Geodiversitas.44.4.57-73.pdf" docOrigin="Geodiversitas 44 (4)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Gryllobencain patrickmuelleri, n. gen., n. sp." docType="treatment" docUuid="80C207C0-1B2E-44F9-90F8-A53E9229461B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="67" masterDocId="C2274B405B57EC13FFE4FF9EFFC7DE37" masterDocTitle="A 100 - million-year-old ensiferan with unusual mouthparts and comments on the evolution of raptorial appendages within Polyneoptera" masterLastPageNumber="73" masterPageNumber="57" pageNumber="60" updateTime="1699304844753" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="4E981DFECF146ED9E7883E293CBCBFEA">A 100 - million-year-old ensiferan with unusual mouthparts and comments on the evolution of raptorial appendages within Polyneoptera</mods:title>
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<mods:namePart id="EC5D73B5B1A74F8FD705F6075F4FEEF7">Haug, Joachim T.</mods:namePart>
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<mods:namePart id="A96EBE40F168E32D3C42443418FD9DB7">Hörnig, Marie K.</mods:namePart>
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<mods:namePart id="109F09EACED932A16A2B841EFA1128FB">Kiesmüller, Christine</mods:namePart>
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<mods:namePart id="4C20C8CE12E8BECB50883994D8E9B5F3">Pazinato, Paula G.</mods:namePart>
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<mods:namePart id="3E41A975CA7D89A41ED876FC749F63FD">Baranov, Viktor</mods:namePart>
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<mods:namePart id="2336C26E330FFC700038E89358C52B6B">Haug, Carolin</mods:namePart>
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<mods:title id="3BD492E858372AE0704104A0820CE94E">Geodiversitas</mods:title>
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<mods:date id="F0E98ED23443F74A81DFFB104A274BAB">2022</mods:date>
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<mods:number id="0159584BE23EF340A8EEE0B344021842">2022-01-27</mods:number>
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<mods:number id="42B5AED62139ACDD292E314BC79898D6">44</mods:number>
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<treatment id="3E1E33385B52EC1FFC9AFA6CFE6CD97B" ID-DOI="http://doi.org/10.5281/zenodo.6301733" ID-GBIF-Taxon="193253581" ID-Zenodo-Dep="6301733" LSID="urn:lsid:zoobank.org:act:80C207C0-1B2E-44F9-90F8-A53E9229461B" httpUri="http://treatment.plazi.org/id/3E1E33385B52EC1FFC9AFA6CFE6CD97B" lastPageId="12" lastPageNumber="67" pageId="5" pageNumber="60">
<subSubSection id="FEADD1A55B52EC16FC9AFA6CFA99D83A" box="[894,1374,1522,1549]" pageId="5" pageNumber="60" type="nomenclature">
<paragraph id="B608822E5B52EC16FC9AFA6CFA99D83A" blockId="5.[894,1374,1522,1549]" box="[894,1374,1522,1549]" pageId="5" pageNumber="60">
<heading id="ED4035425B52EC16FC9AFA6CFA99D83A" box="[894,1374,1522,1549]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="60" reason="2">
<taxonomicName id="71B7F9AD5B52EC16FC9AFA6CFB03D83A" ID-CoL="9GXTT" authority="Haug, Hörnig, Kiesmüller, Pazinato, Baranov &amp; Haug 2022" box="[894,1220,1522,1549]" class="Insecta" family="Tettigoniidae" genus="Gryllobencain" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="patrickmuelleri" status="n. gen., n. sp.">
<emphasis id="84C35E3C5B52EC16FC9AFA6CFB03D83A" bold="true" box="[894,1220,1522,1549]" italics="true" pageId="5" pageNumber="60">Gryllobencain patrickmuelleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF0E3475B52EC16FB2FFA6DFA99D83A" box="[1227,1374,1523,1549]" pageId="5" pageNumber="60" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</heading>
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urn:lsid:zoobank.org:act:
<uuid id="C211B8FB5B52EC16FBE9F9A4FA68D87B" box="[1037,1455,1594,1612]" pageId="5" pageNumber="60">80C207C0-1B2E-44F9-90F8-A53E9229461B</uuid>
</uri>
</paragraph>
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<subSubSection id="FEADD1A55B52EC16FCC9F91CFC1DD92B" pageId="5" pageNumber="60" type="materials_examined">
<paragraph id="B608822E5B52EC16FCC9F91CFBF4D8AD" blockId="5.[813,1075,1666,1690]" box="[813,1075,1666,1690]" pageId="5" pageNumber="60">
<materialsCitation id="06DF88735B52EC16FCC9F91CFBF4D8AD" ID-GBIF-Occurrence="3461896302" box="[813,1075,1666,1690]" collectionCode="PED" pageId="5" pageNumber="60" specimenCode="PED 0147" specimenCount="1" typeStatus="holotype">
<typeStatus id="690C3C8C5B52EC16FCC9F91CFC50D8AE" box="[813,919,1666,1689]" pageId="5" pageNumber="60" type="holotype">HOLOTYPE</typeStatus>
. —
<specimenCode id="E6112A555B52EC16FC27F91CFBF7D8AD" box="[963,1072,1666,1690]" pageId="5" pageNumber="60">PED 0147</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="B608822E5B52EC16FCC9F928FBEFD8F9" blockId="5.[813,1064,1718,1742]" box="[813,1064,1718,1742]" pageId="5" pageNumber="60">
<materialsCitation id="06DF88735B52EC16FCC9F928FBEFD8F9" ID-GBIF-Occurrence="3461896301" box="[813,1064,1718,1742]" collectionCode="PED" pageId="5" pageNumber="60" specimenCode="PED 0178" specimenCount="1" typeStatus="paratype">
<typeStatus id="690C3C8C5B52EC16FCC9F928FC4BD8FA" box="[813,908,1718,1741]" pageId="5" pageNumber="60" type="paratype">PARATYPE</typeStatus>
. —
<specimenCode id="E6112A555B52EC16FC5CF928FBE3D8F9" box="[952,1060,1718,1742]" pageId="5" pageNumber="60">PED 0178</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="B608822E5B52EC16FCC9F974FC1DD92B" blockId="5.[813,1455,1770,1820]" pageId="5" pageNumber="60">
ADDITIONAL MATERIAL. —
<materialsCitation id="06DF88735B52EC16FBDFF974FC1DD92B" ID-GBIF-Occurrence="3461896303" collectionCode="BUB" pageId="5" pageNumber="60" specimenCode="BUB 3072, 3073" specimenCount="1">
<specimenCode id="E6112A555B52EC16FBDFF974FB6FD935" box="[1083,1192,1770,1794]" pageId="5" pageNumber="60">BUB 3072</specimenCode>
,
<specimenCode id="E6112A555B52EC16FB57F974FB2FD935" box="[1203,1256,1770,1794]" pageId="5" pageNumber="60">3073</specimenCode>
, from the collection of Patrick Müller.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FEADD1A55B52EC16FCC9F8A7FB13D95C" pageId="5" pageNumber="60" type="etymology">
<paragraph id="B608822E5B52EC16FCC9F8A7FB13D95C" blockId="5.[812,1455,1849,1900]" pageId="5" pageNumber="60">
DERIVATIO NOMINIS. — In honour of Patrick Müller and his effort for research, especially on
<collectingCountry id="CEA0C2BE5B52EC16FBCFF8CAFB4CD95B" box="[1067,1163,1876,1900]" name="Myanmar" pageId="5" pageNumber="60">Myanmar</collectingCountry>
amber.
</paragraph>
</subSubSection>
<subSubSection id="FEADD1A55B52EC15FCC9F81AFD09D814" lastPageId="6" lastPageNumber="61" pageId="5" pageNumber="60" type="diagnosis">
<paragraph id="B608822E5B52EC15FCC9F81AFD09D814" blockId="5.[811,1458,1923,2027]" lastBlockId="6.[132,776,1521,1572]" lastPageId="6" lastPageNumber="61" pageId="5" pageNumber="60">DIAGNOSIS. — Specimen with prominent maxillae [maxillulae]; medio-proximal part prominent sickle-shaped; palp prominent with long and strong setae; foreleg and mid leg [thorax appendages 1 and 2] with many long and strong setae mostly medially, on femur and tibia. Tibia additionally with strong disto-median spur-like spine. Cerci prominent with numerous very long and strong setae.</paragraph>
</subSubSection>
<caption id="E2C8D2A65B51EC15FF61FAE9FC2BDB81" ID-DOI="http://doi.org/10.5281/zenodo.6000907" ID-Zenodo-Dep="6000907" httpUri="https://zenodo.org/record/6000907/files/figure.png" pageId="6" pageNumber="61" startId="6.[133,144,1399,1417]" targetBox="[151,1437,215,1353]" targetPageId="6">
<paragraph id="B608822E5B51EC15FF61FAE9FC2BDB81" blockId="6.[133,1456,1398,1462]" pageId="6" pageNumber="61">
FIG. 2. — Holotype of
<taxonomicName id="71B7F9AD5B51EC15FEDFFAE8FDECDBBE" box="[315,555,1398,1417]" class="Insecta" family="Tettigoniidae" genus="Gryllobencain" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="patrickmuelleri" status="n. gen., n. sp.">
<emphasis id="84C35E3C5B51EC15FEDFFAE8FDECDBBE" box="[315,555,1398,1417]" italics="true" pageId="6" pageNumber="61">Gryllobencain patrickmuelleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF0E3475B51EC15FDD5FAE9FD60DBBE" box="[561,679,1399,1417]" pageId="6" pageNumber="61" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, PED 0147, close-up on thorax appendages:
<emphasis id="84C35E3C5B51EC15FBC4FAE9FBEADBBE" bold="true" box="[1056,1069,1399,1417]" pageId="6" pageNumber="61">A</emphasis>
, thorax appendage 1;
<emphasis id="84C35E3C5B51EC15FB0EFAE9FB30DBBE" bold="true" box="[1258,1271,1399,1417]" pageId="6" pageNumber="61">B</emphasis>
, thorax appendage 2;
<emphasis id="84C35E3C5B51EC15FF61FA13FF54DBA8" bold="true" box="[133,147,1421,1439]" pageId="6" pageNumber="61">C</emphasis>
, thorax appendage 3. Abbreviations:
<emphasis id="84C35E3C5B51EC15FE28FA13FE25DBA8" bold="true" box="[460,482,1421,1439]" pageId="6" pageNumber="61">c2</emphasis>
, coxa of thorax appendage 2;
<emphasis id="84C35E3C5B51EC15FD05FA13FD31DBA8" bold="true" box="[737,758,1421,1439]" pageId="6" pageNumber="61">ce</emphasis>
, cercus;
<emphasis id="84C35E3C5B51EC15FCA7FA13FC94DBA8" bold="true" box="[835,851,1421,1439]" pageId="6" pageNumber="61">cl</emphasis>
, claw of thorax appendage;
<emphasis id="84C35E3C5B51EC15FBA5FA13FBA4DBA8" bold="true" box="[1089,1123,1421,1439]" pageId="6" pageNumber="61">f1-3</emphasis>
, femur of thorax appendage 1-3;
<emphasis id="84C35E3C5B51EC15FA99FA13FA6DDBA8" bold="true" box="[1405,1450,1421,1439]" pageId="6" pageNumber="61">ta1-3</emphasis>
, tarsus of thorax appendage 1-3;
<emphasis id="84C35E3C5B51EC15FE72FA3AFE6BDB81" bold="true" box="[406,428,1444,1462]" pageId="6" pageNumber="61">tcl</emphasis>
, tibia claw;
<emphasis id="84C35E3C5B51EC15FDF4FA3AFDF0DB81" bold="true" box="[528,567,1444,1462]" pageId="6" pageNumber="61">ti1-3</emphasis>
, tibia of thorax appendage 1-3. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="FEADD1A55B51EC1FFF60F9CDFE6CD97B" lastPageId="12" lastPageNumber="67" pageId="6" pageNumber="61" type="description">
<paragraph id="B608822E5B51EC15FF60F9CDFED2D85A" blockId="6.[132,775,1619,1804]" box="[132,277,1619,1645]" pageId="6" pageNumber="61">DESCRIPTION</paragraph>
<paragraph id="B608822E5B51EC15FF60F9ECFF13D8BB" blockId="6.[132,775,1619,1804]" box="[132,212,1650,1676]" pageId="6" pageNumber="61">
<emphasis id="84C35E3C5B51EC15FF60F9ECFF13D8BB" box="[132,212,1650,1676]" italics="true" pageId="6" pageNumber="61">General</emphasis>
</paragraph>
<paragraph id="B608822E5B51EC15FF60F90CFEB7D93B" blockId="6.[132,775,1619,1804]" pageId="6" pageNumber="61">
Body organised into head and trunk, the latter subdivided into thorax and abdomen (
<figureCitation id="2E8C9EAB5B51EC15FE57F92CFDECD8FA" box="[435,555,1714,1741]" captionStart="FIG" captionStartId="5.[132,143,1383,1401]" captionTargetBox="[149,1434,216,1340]" captionTargetId="figure-278@5.[149,1434,216,1341]" captionTargetPageId="5" captionText="FIG. 1. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp.,PED 0147:A, overview in dorsal view; B, overview in ventral view; C, close-up on maxillary palp, distal part; D, close-up on thorax appendage 1. Abbreviations: an, antenna; ce, cercus; cl1, claw of thorax appendage 1; f1-2, femur of thorax appendage 1-2; hc, head capsule; mp, maxillary palp; sp, spine-like seta; t1-3, thorax segment 1-3; ta1-3, tarsus of thorax appendage 1-3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, B, 1 mm; C, 0.25 mm; D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000905" httpUri="https://zenodo.org/record/6000905/files/figure.png" pageId="6" pageNumber="61">Figs 1A, B</figureCitation>
;
<figureCitation id="2E8C9EAB5B51EC15FDDFF92CFDB8D8FB" box="[571,639,1714,1740]" captionStart="FIG" captionStartId="8.[132,143,1546,1564]" captionTargetBox="[151,1436,214,1499]" captionTargetId="figure-224@8.[151,1436,214,1500]" captionTargetPageId="8" captionText="FIG. 4. — Paratype of Gryllobencain patrickmuelleri n. gen., n. sp., PED 0178:A, overview in dorso-lateral view; B, detail of thorax appendage 1 in posterior view; C, overview in ventro-lateral view; D, detail of thorax appendage 1 in anterior view. Abbreviations: an, antenna; c1, coxa of thorax appendage 1; ce, cercus; ey, compound eye; f1-3, femur of thorax appendage 1-3; hc, head capsule; mp, maxillary palp; sp, spine-like seta; ta1, 3, tarsus of thorax appendage 1, 3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3; tr1, trochanter of thorax appendage 1. Scale bars: A, C, 1 mm; B, D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000913" httpUri="https://zenodo.org/record/6000913/files/figure.png" pageId="6" pageNumber="61">4A, C</figureCitation>
;
<figureCitation id="2E8C9EAB5B51EC15FD6AF92CFD15D8FB" box="[654,722,1714,1741]" captionStart="FIG" captionStartId="9.[132,143,1546,1564]" captionTargetBox="[149,1435,216,1501]" captionTargetId="figure-212@9.[149,1434,216,1502]" captionTargetPageId="9" captionText="FIG. 5. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3073: A, overview in dorsal view; B, close-up in anterior head area with ocelli; C, overview in ventral view; D, close-up on distal part of thorax appendage 1. In A and C background was desaturated to enhance contrast. Abbreviations: an, antenna; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; mp, maxillary palp; oc, ocelli; ta1, tarsus of thorax appendage 1; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, C, 0.5 mm; B, D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6000917" httpUri="https://zenodo.org/record/6000917/files/figure.png" pageId="6" pageNumber="61">5A, C</figureCitation>
;
<figureCitation id="2E8C9EAB5B51EC15FD06F92CFF5DD8DB" captionStart="FIG" captionStartId="10.[132,143,1543,1561]" captionTargetBox="[153,1439,215,1500]" captionTargetId="figure-180@10.[153,1438,214,1500]" captionTargetPageId="10" captionText="FIG. 6. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3072: A, overview in dorsal view; B, close-up on cercus; C, overview in ventral view; D, close-up on head. Abbreviations: an, antenna; ce, cercus; f1, 3, femur of thorax appendage 1, 3; hc, head capsule; mp, maxillary palp; ti1, 3, tibia of thorax appendage 1, 3. Scale bars: A, C, 1 mm; B, D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000921" httpUri="https://zenodo.org/record/6000921/files/figure.png" pageId="6" pageNumber="61">6A, C</figureCitation>
), presumably with 20 segments, ocular segment plus 19 post-ocular segments.
</paragraph>
<subSection id="DF389BC55B51EC14FF60F8AFFAF8DF05" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61" type="multiple">
<paragraph id="B608822E5B51EC15FF60F8AFFF7BD97C" blockId="6.[132,188,1841,1867]" box="[132,188,1841,1867]" pageId="6" pageNumber="61">
<heading id="ED4035425B51EC15FF60F8AFFF7BD97C" box="[132,188,1841,1867]" fontSize="11" level="2" pageId="6" pageNumber="61" reason="8">
<emphasis id="84C35E3C5B51EC15FF60F8AFFF7BD97C" box="[132,188,1841,1867]" italics="true" pageId="6" pageNumber="61">Head</emphasis>
</heading>
</paragraph>
<paragraph id="B608822E5B51EC15FF60F8CFFD2FD9DB" blockId="6.[129,777,1873,2028]" pageId="6" pageNumber="61">
Ocular segment and post-ocular segments1-5forming distinct capsule (head capsule) (
<figureCitation id="2E8C9EAB5B51EC15FEF7F8EFFEA5D9BB" box="[275,354,1905,1932]" captionStart="FIG" captionStartId="5.[132,143,1383,1401]" captionTargetBox="[149,1434,216,1340]" captionTargetId="figure-278@5.[149,1434,216,1341]" captionTargetPageId="5" captionText="FIG. 1. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp.,PED 0147:A, overview in dorsal view; B, overview in ventral view; C, close-up on maxillary palp, distal part; D, close-up on thorax appendage 1. Abbreviations: an, antenna; ce, cercus; cl1, claw of thorax appendage 1; f1-2, femur of thorax appendage 1-2; hc, head capsule; mp, maxillary palp; sp, spine-like seta; t1-3, thorax segment 1-3; ta1-3, tarsus of thorax appendage 1-3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, B, 1 mm; C, 0.25 mm; D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000905" httpUri="https://zenodo.org/record/6000905/files/figure.png" pageId="6" pageNumber="61">Figs1A</figureCitation>
;
<figureCitation id="2E8C9EAB5B51EC15FE8DF8EFFE4ED9BB" box="[361,393,1905,1932]" captionStart="FIG" captionStartId="8.[132,143,1546,1564]" captionTargetBox="[151,1436,214,1499]" captionTargetId="figure-224@8.[151,1436,214,1500]" captionTargetPageId="8" captionText="FIG. 4. — Paratype of Gryllobencain patrickmuelleri n. gen., n. sp., PED 0178:A, overview in dorso-lateral view; B, detail of thorax appendage 1 in posterior view; C, overview in ventro-lateral view; D, detail of thorax appendage 1 in anterior view. Abbreviations: an, antenna; c1, coxa of thorax appendage 1; ce, cercus; ey, compound eye; f1-3, femur of thorax appendage 1-3; hc, head capsule; mp, maxillary palp; sp, spine-like seta; ta1, 3, tarsus of thorax appendage 1, 3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3; tr1, trochanter of thorax appendage 1. Scale bars: A, C, 1 mm; B, D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000913" httpUri="https://zenodo.org/record/6000913/files/figure.png" pageId="6" pageNumber="61">4A</figureCitation>
;
<figureCitation id="2E8C9EAB5B51EC15FE74F8EFFE77D9BB" box="[400,432,1905,1932]" captionStart="FIG" captionStartId="9.[132,143,1546,1564]" captionTargetBox="[149,1435,216,1501]" captionTargetId="figure-212@9.[149,1434,216,1502]" captionTargetPageId="9" captionText="FIG. 5. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3073: A, overview in dorsal view; B, close-up in anterior head area with ocelli; C, overview in ventral view; D, close-up on distal part of thorax appendage 1. In A and C background was desaturated to enhance contrast. Abbreviations: an, antenna; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; mp, maxillary palp; oc, ocelli; ta1, tarsus of thorax appendage 1; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, C, 0.5 mm; B, D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6000917" httpUri="https://zenodo.org/record/6000917/files/figure.png" pageId="6" pageNumber="61">5A</figureCitation>
;
<figureCitation id="2E8C9EAB5B51EC15FE53F8EFFE1FD9BB" box="[439,472,1905,1932]" captionStart="FIG" captionStartId="10.[132,143,1543,1561]" captionTargetBox="[153,1439,215,1500]" captionTargetId="figure-180@10.[153,1438,214,1500]" captionTargetPageId="10" captionText="FIG. 6. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3072: A, overview in dorsal view; B, close-up on cercus; C, overview in ventral view; D, close-up on head. Abbreviations: an, antenna; ce, cercus; f1, 3, femur of thorax appendage 1, 3; hc, head capsule; mp, maxillary palp; ti1, 3, tibia of thorax appendage 1, 3. Scale bars: A, C, 1 mm; B, D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000921" httpUri="https://zenodo.org/record/6000921/files/figure.png" pageId="6" pageNumber="61">6A</figureCitation>
).Head capsule oval in dorsal(or ventral) view; about 1.6× wider than long. Surface of head capsule dorsallywithprominentY-shapedmoultingsuture(epicranialsuture) (
<figureCitation id="2E8C9EAB5B51EC15FF68F84FFF10D9DC" box="[140,215,2001,2028]" captionStart="FIG" captionStartId="9.[132,143,1546,1564]" captionTargetBox="[149,1435,216,1501]" captionTargetId="figure-212@9.[149,1434,216,1502]" captionTargetPageId="9" captionText="FIG. 5. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3073: A, overview in dorsal view; B, close-up in anterior head area with ocelli; C, overview in ventral view; D, close-up on distal part of thorax appendage 1. In A and C background was desaturated to enhance contrast. Abbreviations: an, antenna; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; mp, maxillary palp; oc, ocelli; ta1, tarsus of thorax appendage 1; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, C, 0.5 mm; B, D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6000917" httpUri="https://zenodo.org/record/6000917/files/figure.png" pageId="6" pageNumber="61">Fig. 5A</figureCitation>
). Lateral edges of head capsule rounded with few setae.
</paragraph>
<paragraph id="B608822E5B51EC15FCA7FA6DFB21D91B" blockId="6.[810,1457,1523,2028]" pageId="6" pageNumber="61">
Ocular segment recognisable by prominent compound eyes laterally projecting from the head capsule; additionally, three ocelli present [median eyes] (
<figureCitation id="2E8C9EAB5B51EC15FB80F9ACFB72D87A" box="[1124,1205,1586,1613]" captionStart="FIG" captionStartId="9.[132,143,1546,1564]" captionTargetBox="[149,1435,216,1501]" captionTargetId="figure-212@9.[149,1434,216,1502]" captionTargetPageId="9" captionText="FIG. 5. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3073: A, overview in dorsal view; B, close-up in anterior head area with ocelli; C, overview in ventral view; D, close-up on distal part of thorax appendage 1. In A and C background was desaturated to enhance contrast. Abbreviations: an, antenna; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; mp, maxillary palp; oc, ocelli; ta1, tarsus of thorax appendage 1; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, C, 0.5 mm; B, D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6000917" httpUri="https://zenodo.org/record/6000917/files/figure.png" pageId="6" pageNumber="61">Fig. 5B</figureCitation>
). Compound eyes with numerous facets; difficult to count, at least 40 rows of facets (
<figureCitation id="2E8C9EAB5B51EC15FCD2F9ECFC6AD8BA" box="[822,941,1650,1677]" captionStart="FIG" captionStartId="11.[132,143,1539,1557]" captionTargetBox="[153,1438,214,1494]" captionTargetId="figure-227@11.[704,982,755,1033]" captionTargetPageId="11" captionText="FIG. 7. — Close-ups on head area in different specimens of Gryllobencain patrickmuelleri n. gen., n. sp. as normal versions (A, C, E) and colour-marked (B, D, F): A-D, PED 0178; A, B, latero-posterior view; C, D, mostly anterior view; elements of maxillary palp difficult to differentiate; E, F, BUB 3073, ventral view. Abbreviations: lp, labial palp; lr, labrum; md, mandible; mmx, median part of maxilla; mp, maxillary palp; mx, maxilla; pmx, proximal part of maxilla. Scale bars: A-D, 0.3 mm; E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6000923" httpUri="https://zenodo.org/record/6000923/files/figure.png" pageId="6" pageNumber="61">Fig. 7A, B</figureCitation>
). Ocelli arranged in a triangle; central ocellus dorsal to lateral ones. Appendage derivative of ocular segment (clypeo-labrum complex) [hypostome-labrum complex] well developed (
<figureCitation id="2E8C9EAB5B51EC15FC3EF94CFBF7D8DB" box="[986,1072,1746,1772]" captionStart="FIG" captionStartId="11.[132,143,1539,1557]" captionTargetBox="[153,1438,214,1494]" captionTargetId="figure-227@11.[704,982,755,1033]" captionTargetPageId="11" captionText="FIG. 7. — Close-ups on head area in different specimens of Gryllobencain patrickmuelleri n. gen., n. sp. as normal versions (A, C, E) and colour-marked (B, D, F): A-D, PED 0178; A, B, latero-posterior view; C, D, mostly anterior view; elements of maxillary palp difficult to differentiate; E, F, BUB 3073, ventral view. Abbreviations: lp, labial palp; lr, labrum; md, mandible; mmx, median part of maxilla; mp, maxillary palp; mx, maxilla; pmx, proximal part of maxilla. Scale bars: A-D, 0.3 mm; E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6000923" httpUri="https://zenodo.org/record/6000923/files/figure.png" pageId="6" pageNumber="61">Fig. 7D</figureCitation>
). Clypeus [hypostome] rectangular, about 2× wider than long. Labrum lobe-like, about as large as clypeus, anterior edge gently rounded.
</paragraph>
<paragraph id="B608822E5B51EC14FCA7F8AFFEBCDBBA" blockId="6.[810,1457,1523,2028]" lastBlockId="7.[130,777,1299,2028]" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61">
Post-ocular segment 1 recognisable by its appendage, antenna [antennula]. Antenna arising from head capsule antero-dorsally; lateral to the clypeus; very long, longer than main body, with numerous elements (antennomeres), more than 40 (
<figureCitation id="2E8C9EAB5B51EC15FAB1F80FFC87D9FB" captionStart="FIG" captionStartId="5.[132,143,1383,1401]" captionTargetBox="[149,1434,216,1340]" captionTargetId="figure-278@5.[149,1434,216,1341]" captionTargetPageId="5" captionText="FIG. 1. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp.,PED 0147:A, overview in dorsal view; B, overview in ventral view; C, close-up on maxillary palp, distal part; D, close-up on thorax appendage 1. Abbreviations: an, antenna; ce, cercus; cl1, claw of thorax appendage 1; f1-2, femur of thorax appendage 1-2; hc, head capsule; mp, maxillary palp; sp, spine-like seta; t1-3, thorax segment 1-3; ta1-3, tarsus of thorax appendage 1-3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, B, 1 mm; C, 0.25 mm; D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000905" httpUri="https://zenodo.org/record/6000905/files/figure.png" pageId="6" pageNumber="61">Figs 1A, B</figureCitation>
;
<figureCitation id="2E8C9EAB5B51EC15FCA9F82FFC56D9FC" box="[845,913,1969,1996]" captionStart="FIG" captionStartId="9.[132,143,1546,1564]" captionTargetBox="[149,1435,216,1501]" captionTargetId="figure-212@9.[149,1434,216,1502]" captionTargetPageId="9" captionText="FIG. 5. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3073: A, overview in dorsal view; B, close-up in anterior head area with ocelli; C, overview in ventral view; D, close-up on distal part of thorax appendage 1. In A and C background was desaturated to enhance contrast. Abbreviations: an, antenna; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; mp, maxillary palp; oc, ocelli; ta1, tarsus of thorax appendage 1; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, C, 0.5 mm; B, D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6000917" httpUri="https://zenodo.org/record/6000917/files/figure.png" pageId="6" pageNumber="61">5A, C</figureCitation>
); some elements are longer and possess a distinct constriction (indication of future subdivision?); length of elements strongly varying; proximal elements with few, but prominent setae. More distal elements with numerous, but shorter setae. Post-ocular segment 2 (intercalary segment) not recognisable externally.
</paragraph>
<caption id="E2C8D2A65B50EC14FF60FB19FEE4DAEA" ID-DOI="http://doi.org/10.5281/zenodo.6000911" ID-Zenodo-Dep="6000911" httpUri="https://zenodo.org/record/6000911/files/figure.png" pageId="7" pageNumber="62" startId="7.[132,143,1159,1177]" targetBox="[132,775,215,1117]" targetPageId="7">
<paragraph id="B608822E5B50EC14FF60FB19FEE4DAEA" blockId="7.[132,777,1159,1245]" pageId="7" pageNumber="62">
FIG. 3. — Holotype of
<taxonomicName id="71B7F9AD5B50EC14FEDFFB19FDEBDAAE" box="[315,556,1159,1177]" class="Insecta" family="Tettigoniidae" genus="Gryllobencain" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="patrickmuelleri" status="n. gen., n. sp.">
<emphasis id="84C35E3C5B50EC14FEDFFB19FDEBDAAE" box="[315,556,1159,1177]" italics="true" pageId="7" pageNumber="62">Gryllobencain patrickmuelleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF0E3475B50EC14FDD5FB19FD60DAAE" box="[561,679,1159,1177]" pageId="7" pageNumber="62" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, PED 0147, close-up on posterior end with thorax appendage 3 and cerci. Abbreviations:
<emphasis id="84C35E3C5B50EC14FF60FB2BFF5EDAF0" bold="true" box="[132,153,1205,1223]" pageId="7" pageNumber="62">ce</emphasis>
, cercus;
<emphasis id="84C35E3C5B50EC14FF00FB2BFF38DAF0" bold="true" box="[228,255,1205,1223]" pageId="7" pageNumber="62">ta3</emphasis>
, tarsus of thorax appendage 3;
<emphasis id="84C35E3C5B50EC14FDE6FB2BFDD0DAF0" bold="true" box="[514,535,1205,1223]" pageId="7" pageNumber="62">ti3</emphasis>
, tibia of thorax appendage 3. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="B608822E5B50EC14FF7FFA0DFEBEDBDA" blockId="7.[130,777,1299,2028]" pageId="7" pageNumber="62">
Post-ocular segment 3 recognisable by its appendage, mandible (
<figureCitation id="2E8C9EAB5B50EC14FF29FA2DFEC9DBFA" box="[205,270,1459,1485]" captionStart="FIG" captionStartId="11.[132,143,1539,1557]" captionTargetBox="[153,1438,214,1494]" captionTargetId="figure-227@11.[704,982,755,1033]" captionTargetPageId="11" captionText="FIG. 7. — Close-ups on head area in different specimens of Gryllobencain patrickmuelleri n. gen., n. sp. as normal versions (A, C, E) and colour-marked (B, D, F): A-D, PED 0178; A, B, latero-posterior view; C, D, mostly anterior view; elements of maxillary palp difficult to differentiate; E, F, BUB 3073, ventral view. Abbreviations: lp, labial palp; lr, labrum; md, mandible; mmx, median part of maxilla; mp, maxillary palp; mx, maxilla; pmx, proximal part of maxilla. Scale bars: A-D, 0.3 mm; E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6000923" httpUri="https://zenodo.org/record/6000923/files/figure.png" pageId="7" pageNumber="62">Fig. 7</figureCitation>
A-D). Mandibles prominent, curved, bearing teeth on inner surface.
</paragraph>
<paragraph id="B608822E5B50EC14FF7FFA6DFEA0D9DC" blockId="7.[130,777,1299,2028]" pageId="7" pageNumber="62">
Post-ocular segment 4 recognisable by its appendage, maxilla [maxillula] (
<figureCitation id="2E8C9EAB5B50EC14FED4F98CFEA9D81A" box="[304,366,1554,1581]" captionStart="FIG" captionStartId="11.[132,143,1539,1557]" captionTargetBox="[153,1438,214,1494]" captionTargetId="figure-227@11.[704,982,755,1033]" captionTargetPageId="11" captionText="FIG. 7. — Close-ups on head area in different specimens of Gryllobencain patrickmuelleri n. gen., n. sp. as normal versions (A, C, E) and colour-marked (B, D, F): A-D, PED 0178; A, B, latero-posterior view; C, D, mostly anterior view; elements of maxillary palp difficult to differentiate; E, F, BUB 3073, ventral view. Abbreviations: lp, labial palp; lr, labrum; md, mandible; mmx, median part of maxilla; mp, maxillary palp; mx, maxilla; pmx, proximal part of maxilla. Scale bars: A-D, 0.3 mm; E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6000923" httpUri="https://zenodo.org/record/6000923/files/figure.png" pageId="7" pageNumber="62">Fig. 7</figureCitation>
A-F). Proximal part of maxilla (cardo) [basipod?] medially drawn out; cardo distally carrying palp [endopod]. Drawn out part (cardo? stipes? both?) prominent, sickle-shaped; laterally with at least three setae. Palp with five elements (
<figureCitation id="2E8C9EAB5B50EC14FEC4F90CFEB0D89B" box="[288,375,1682,1708]" captionStart="FIG" captionStartId="5.[132,143,1383,1401]" captionTargetBox="[149,1434,216,1340]" captionTargetId="figure-278@5.[149,1434,216,1341]" captionTargetPageId="5" captionText="FIG. 1. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp.,PED 0147:A, overview in dorsal view; B, overview in ventral view; C, close-up on maxillary palp, distal part; D, close-up on thorax appendage 1. Abbreviations: an, antenna; ce, cercus; cl1, claw of thorax appendage 1; f1-2, femur of thorax appendage 1-2; hc, head capsule; mp, maxillary palp; sp, spine-like seta; t1-3, thorax segment 1-3; ta1-3, tarsus of thorax appendage 1-3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, B, 1 mm; C, 0.25 mm; D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000905" httpUri="https://zenodo.org/record/6000905/files/figure.png" pageId="7" pageNumber="62">Figs 1C</figureCitation>
;
<figureCitation id="2E8C9EAB5B50EC14FE67F90CFE6ED89A" box="[387,425,1682,1709]" captionStart="FIG" captionStartId="10.[132,143,1543,1561]" captionTargetBox="[153,1439,215,1500]" captionTargetId="figure-180@10.[153,1438,214,1500]" captionTargetPageId="10" captionText="FIG. 6. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3072: A, overview in dorsal view; B, close-up on cercus; C, overview in ventral view; D, close-up on head. Abbreviations: an, antenna; ce, cercus; f1, 3, femur of thorax appendage 1, 3; hc, head capsule; mp, maxillary palp; ti1, 3, tibia of thorax appendage 1, 3. Scale bars: A, C, 1 mm; B, D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000921" httpUri="https://zenodo.org/record/6000921/files/figure.png" pageId="7" pageNumber="62">6D</figureCitation>
;
<figureCitation id="2E8C9EAB5B50EC14FE51F90DFE12D89A" box="[437,469,1683,1709]" captionStart="FIG" captionStartId="11.[132,143,1539,1557]" captionTargetBox="[153,1438,214,1494]" captionTargetId="figure-227@11.[704,982,755,1033]" captionTargetPageId="11" captionText="FIG. 7. — Close-ups on head area in different specimens of Gryllobencain patrickmuelleri n. gen., n. sp. as normal versions (A, C, E) and colour-marked (B, D, F): A-D, PED 0178; A, B, latero-posterior view; C, D, mostly anterior view; elements of maxillary palp difficult to differentiate; E, F, BUB 3073, ventral view. Abbreviations: lp, labial palp; lr, labrum; md, mandible; mmx, median part of maxilla; mp, maxillary palp; mx, maxilla; pmx, proximal part of maxilla. Scale bars: A-D, 0.3 mm; E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6000923" httpUri="https://zenodo.org/record/6000923/files/figure.png" pageId="7" pageNumber="62">7F</figureCitation>
). Element 1 (most proximal one) short, as long as wide (diameter). Element 2 sub-similar to element 1. Element 3 elongate, at least 4.5× longer than wide (diameter); with at least two strong spine-like setae and numerous smaller setae of varying sizes. Element 4 shorter than preceding element, elongate, at least 2.8× longer than wide (diameter); with at least two strong spine-like setae and numerous smaller setae of varying sizes. Element 5 (most distal one) elongate, about 4× longer than wide (diameter); with at least six strong spine-like setae and numerous smaller setae of varying sizes.
</paragraph>
<paragraph id="B608822E5B50EC14FCA7FF49FAF8DF05" blockId="7.[811,1456,215,306]" pageId="7" pageNumber="62">
Post-ocular segment 5 recognisable by its appendage, labium [maxilla] (
<figureCitation id="2E8C9EAB5B50EC14FC73FF69FC39DF25" box="[919,1022,247,274]" captionStart="FIG" captionStartId="11.[132,143,1539,1557]" captionTargetBox="[153,1438,214,1494]" captionTargetId="figure-227@11.[704,982,755,1033]" captionTargetPageId="11" captionText="FIG. 7. — Close-ups on head area in different specimens of Gryllobencain patrickmuelleri n. gen., n. sp. as normal versions (A, C, E) and colour-marked (B, D, F): A-D, PED 0178; A, B, latero-posterior view; C, D, mostly anterior view; elements of maxillary palp difficult to differentiate; E, F, BUB 3073, ventral view. Abbreviations: lp, labial palp; lr, labrum; md, mandible; mmx, median part of maxilla; mp, maxillary palp; mx, maxilla; pmx, proximal part of maxilla. Scale bars: A-D, 0.3 mm; E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6000923" httpUri="https://zenodo.org/record/6000923/files/figure.png" pageId="7" pageNumber="62">Fig. 7E, F</figureCitation>
); only distal parts, palp [endopod], apparent. Palp mostly concealed, with numerous setae.
</paragraph>
</subSection>
<subSection id="DF389BC55B50EC1AFCC9FEC9FBE4D9DB" lastPageId="9" lastPageNumber="64" pageId="7" pageNumber="62" type="multiple">
<paragraph id="B608822E5B50EC14FCC9FEC9FBE6DF46" blockId="7.[813,1057,343,369]" box="[813,1057,343,369]" pageId="7" pageNumber="62">
<heading id="ED4035425B50EC14FCC9FEC9FBE6DF46" box="[813,1057,343,369]" fontSize="11" level="2" pageId="7" pageNumber="62" reason="8">
<emphasis id="84C35E3C5B50EC14FCC9FEC9FBE6DF46" box="[813,1057,343,369]" italics="true" pageId="7" pageNumber="62">Thorax [anterior trunk]</emphasis>
</heading>
</paragraph>
<paragraph id="B608822E5B50EC14FCC9FEE9FB09DCA7" blockId="7.[811,1458,375,2028]" pageId="7" pageNumber="62">
Post-ocular segments 6-8 differentiated from posterior ones, forming a distinct unit, thorax (
<figureCitation id="2E8C9EAB5B50EC14FB73FE08FB29DF86" box="[1175,1262,406,433]" captionStart="FIG" captionStartId="9.[132,143,1546,1564]" captionTargetBox="[149,1435,216,1501]" captionTargetId="figure-212@9.[149,1434,216,1502]" captionTargetPageId="9" captionText="FIG. 5. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3073: A, overview in dorsal view; B, close-up in anterior head area with ocelli; C, overview in ventral view; D, close-up on distal part of thorax appendage 1. In A and C background was desaturated to enhance contrast. Abbreviations: an, antenna; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; mp, maxillary palp; oc, ocelli; ta1, tarsus of thorax appendage 1; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, C, 0.5 mm; B, D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6000917" httpUri="https://zenodo.org/record/6000917/files/figure.png" pageId="7" pageNumber="62">Fig. 5A</figureCitation>
). Tergite of postocular segment 6 (tergite of thorax segment 1; pronotum) about 0.55× narrower than head; about 1.6× wider than long; rectangular-shaped in dorsal view; with six smaller setae medio-anteriorly. Post-ocular segment 6 (thorax segment 1; prothorax) ventrally with a pair of prominent appendages (foreleg) [thoracopod 1] (
<figureCitation id="2E8C9EAB5B50EC14FBDAFDC8FB52DC47" box="[1086,1173,598,624]" captionStart="FIG" captionStartId="5.[132,143,1383,1401]" captionTargetBox="[149,1434,216,1340]" captionTargetId="figure-278@5.[149,1434,216,1341]" captionTargetPageId="5" captionText="FIG. 1. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp.,PED 0147:A, overview in dorsal view; B, overview in ventral view; C, close-up on maxillary palp, distal part; D, close-up on thorax appendage 1. Abbreviations: an, antenna; ce, cercus; cl1, claw of thorax appendage 1; f1-2, femur of thorax appendage 1-2; hc, head capsule; mp, maxillary palp; sp, spine-like seta; t1-3, thorax segment 1-3; ta1-3, tarsus of thorax appendage 1-3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, B, 1 mm; C, 0.25 mm; D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000905" httpUri="https://zenodo.org/record/6000905/files/figure.png" pageId="7" pageNumber="62">Figs1D</figureCitation>
;
<figureCitation id="2E8C9EAB5B50EC14FB44FDC8FB05DC47" box="[1184,1218,598,624]" captionStart="FIG" captionStartId="6.[133,144,1399,1417]" captionTargetBox="[151,1437,215,1353]" captionTargetId="figure-276@6.[151,1436,214,1353]" captionTargetPageId="6" captionText="FIG. 2. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp., PED 0147, close-up on thorax appendages: A, thorax appendage 1; B, thorax appendage 2; C, thorax appendage 3. Abbreviations: c2, coxa of thorax appendage 2; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; ta1-3, tarsus of thorax appendage 1-3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000907" httpUri="https://zenodo.org/record/6000907/files/figure.png" pageId="7" pageNumber="62">2A</figureCitation>
;
<figureCitation id="2E8C9EAB5B50EC14FB28FDC8FAD7DC47" box="[1228,1296,598,624]" captionStart="FIG" captionStartId="8.[132,143,1546,1564]" captionTargetBox="[151,1436,214,1499]" captionTargetId="figure-224@8.[151,1436,214,1500]" captionTargetPageId="8" captionText="FIG. 4. — Paratype of Gryllobencain patrickmuelleri n. gen., n. sp., PED 0178:A, overview in dorso-lateral view; B, detail of thorax appendage 1 in posterior view; C, overview in ventro-lateral view; D, detail of thorax appendage 1 in anterior view. Abbreviations: an, antenna; c1, coxa of thorax appendage 1; ce, cercus; ey, compound eye; f1-3, femur of thorax appendage 1-3; hc, head capsule; mp, maxillary palp; sp, spine-like seta; ta1, 3, tarsus of thorax appendage 1, 3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3; tr1, trochanter of thorax appendage 1. Scale bars: A, C, 1 mm; B, D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000913" httpUri="https://zenodo.org/record/6000913/files/figure.png" pageId="7" pageNumber="62">4B, D</figureCitation>
;
<figureCitation id="2E8C9EAB5B50EC14FAFEFDC8FA86DC47" box="[1306,1345,598,625]" captionStart="FIG" captionStartId="9.[132,143,1546,1564]" captionTargetBox="[149,1435,216,1501]" captionTargetId="figure-212@9.[149,1434,216,1502]" captionTargetPageId="9" captionText="FIG. 5. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3073: A, overview in dorsal view; B, close-up in anterior head area with ocelli; C, overview in ventral view; D, close-up on distal part of thorax appendage 1. In A and C background was desaturated to enhance contrast. Abbreviations: an, antenna; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; mp, maxillary palp; oc, ocelli; ta1, tarsus of thorax appendage 1; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, C, 0.5 mm; B, D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6000917" httpUri="https://zenodo.org/record/6000917/files/figure.png" pageId="7" pageNumber="62">5D</figureCitation>
); basically cylindrical, composed of five elements.
</paragraph>
<paragraph id="B608822E5B50EC14FCA7FD08FC7CDBDA" blockId="7.[811,1458,375,2028]" pageId="7" pageNumber="62">
Element 1 (coxa) [basipod?] about as long (proximal-distal axis) as wide (diameter); with at least three smaller, thin setae posteriorly. Element 2 (trochanter) [endopod element 1?] slightly longer than wide, slightly longer than coxa; with more than nine small, thin setae medially. Element 3 (femur) [endopod element 2?] very elongate; about 10× longer than coxa, similar width at proximal part, tapering towards distally to about 0.48× of maximum width; with more than ten small, thin setae proximo-medially and disto-laterally; also with more than ten strong spine-like setae arranged in two rows (
<figureCitation id="2E8C9EAB5B50EC14FC8BFC4AFC02DDD8" box="[879,965,980,1007]" captionStart="FIG" captionStartId="8.[132,143,1546,1564]" captionTargetBox="[151,1436,214,1499]" captionTargetId="figure-224@8.[151,1436,214,1500]" captionTargetPageId="8" captionText="FIG. 4. — Paratype of Gryllobencain patrickmuelleri n. gen., n. sp., PED 0178:A, overview in dorso-lateral view; B, detail of thorax appendage 1 in posterior view; C, overview in ventro-lateral view; D, detail of thorax appendage 1 in anterior view. Abbreviations: an, antenna; c1, coxa of thorax appendage 1; ce, cercus; ey, compound eye; f1-3, femur of thorax appendage 1-3; hc, head capsule; mp, maxillary palp; sp, spine-like seta; ta1, 3, tarsus of thorax appendage 1, 3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3; tr1, trochanter of thorax appendage 1. Scale bars: A, C, 1 mm; B, D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000913" httpUri="https://zenodo.org/record/6000913/files/figure.png" pageId="7" pageNumber="62">Fig. 4D</figureCitation>
), the majority of which is located proximally, with one exception disto-laterally. Element 4 (tibia) [endopod element 3?] elongate, about 0.75× the length of the femur; about as wide as the femur, though slightly less proximally; covered with smaller setae, more distally than proximally; with three strong spine-like setae disto-medially of varying size and one strong spine-like seta laterally, more towards proximal than distal; with one very strong spine very medio-distally. Element 5 (tarsus) [endopod element 4?] slightly shorter than preceding element, more than 0.8× of the length and about half of the width; with numerous smaller, thin setae throughout its surface; distally with a pair of claws, possible attachment structure between claws not apparent; tarsus subdivided into three sub-elements (tarsomeres) which are of mostly similar length (0.3× of total length) and similar width, most proximal element has three stronger spine-like setae distally.
</paragraph>
<paragraph id="B608822E5B50EC14FCA7FA6DFBE0D8DB" blockId="7.[811,1458,375,2028]" pageId="7" pageNumber="62">
Post-ocular segment 7 (thorax segment 2; mesothorax) about half as long as prothorax and about as wide as the latter, but slightly tapering posteriorly; trapezoid-shaped; with one small seta on each side respectively (antero-laterally), close to the coxa. Ventrally with a pair of prominent appendages (mid leg) [thoracopod 2] (
<figureCitation id="2E8C9EAB5B50EC14FB91F90CFB01D89A" box="[1141,1222,1682,1709]" captionStart="FIG" captionStartId="6.[133,144,1399,1417]" captionTargetBox="[151,1437,215,1353]" captionTargetId="figure-276@6.[151,1436,214,1353]" captionTargetPageId="6" captionText="FIG. 2. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp., PED 0147, close-up on thorax appendages: A, thorax appendage 1; B, thorax appendage 2; C, thorax appendage 3. Abbreviations: c2, coxa of thorax appendage 2; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; ta1-3, tarsus of thorax appendage 1-3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000907" httpUri="https://zenodo.org/record/6000907/files/figure.png" pageId="7" pageNumber="62">Fig. 2B</figureCitation>
); basically cylindrical, composed of five elements; as a whole slightly longer than foreleg [thoracopod 1].
</paragraph>
<paragraph id="B608822E5B50EC1BFCA7F96CFB58D95C" blockId="7.[811,1458,375,2028]" lastBlockId="8.[811,1457,1681,2028]" lastPageId="8" lastPageNumber="63" pageId="7" pageNumber="62">Element 1 (coxa) [basipod?] about as long (proximal-distal axis) as wide (diameter); with two smaller setae medially. Element 2 (trochanter) [endopod element 1?] slightly longer than wide, but as wide as the coxa; with one small, thin seta and two stronger, spine-like setae medio-distally. Element 3 (femur) [endopod element 2?] very elongate; about 10× longer than coxa, almost similar width at proximal part, then widening to 1.5× width at about 30 % of total femur length, then tapering towards distally to about 0.67× of maximum width; with more than ten small, thin setae laterally throughout its surface; also with more than ten strong spine-like setae with varying sizes, the majority of which located medially, with one exception laterally. Element 4 (tibia) [endopod element 3?] elongate, slightly shorter than femur; about as wide as femur, though slightly less so proximally; covered with smaller setae, more distally than proximally; with two strong spine-like setae and more than five smaller spine-like setae disto-medially; also with one strong spine-like seta laterally, more towards proximal than distal, and one very strong spine very medio-distally. Element 5 (tarsus) [endopod element 4?] slightly shorter than tibia, more than 0.8× of the length and about half of the width; with numerous smaller, thin setae throughout; distally with a pair of claws, possible attachment structure between claws not apparent; tarsus subdivided into three sub-elements which are of mostly similar length (0.3× of total length) and similar width.</paragraph>
<caption id="E2C8D2A65B5FEC1BFF60F994FBAAD857" ID-DOI="http://doi.org/10.5281/zenodo.6000913" ID-Zenodo-Dep="6000913" httpUri="https://zenodo.org/record/6000913/files/figure.png" pageId="8" pageNumber="63" startId="8.[132,143,1546,1564]" targetBox="[151,1436,214,1499]" targetPageId="8">
<paragraph id="B608822E5B5FEC1BFF60F994FBAAD857" blockId="8.[132,1457,1546,1632]" pageId="8" pageNumber="63">
FIG. 4. — Paratype of
<taxonomicName id="71B7F9AD5B5FEC1BFED3F994FDE0D82B" box="[311,551,1546,1564]" class="Insecta" family="Tettigoniidae" genus="Gryllobencain" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="patrickmuelleri" status="n. gen., n. sp.">
<emphasis id="84C35E3C5B5FEC1BFED3F994FDE0D82B" box="[311,551,1546,1564]" italics="true" pageId="8" pageNumber="63">Gryllobencain patrickmuelleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF0E3475B5FEC1BFDC8F994FD67D82B" box="[556,672,1546,1564]" pageId="8" pageNumber="63" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, PED 0178:
<emphasis id="84C35E3C5B5FEC1BFCE6F994FCC8D82B" bold="true" box="[770,783,1546,1564]" pageId="8" pageNumber="63">A</emphasis>
, overview in dorso-lateral view;
<emphasis id="84C35E3C5B5FEC1BFBFDF994FBE1D82B" bold="true" box="[1049,1062,1546,1564]" pageId="8" pageNumber="63">B</emphasis>
, detail of thorax appendage 1 in posterior view;
<emphasis id="84C35E3C5B5FEC1BFF60F9BFFF55D804" bold="true" box="[132,146,1569,1587]" pageId="8" pageNumber="63">C</emphasis>
, overview in ventro-lateral view;
<emphasis id="84C35E3C5B5FEC1BFE46F9BFFE77D804" bold="true" box="[418,432,1569,1587]" pageId="8" pageNumber="63">D</emphasis>
, detail of thorax appendage 1 in anterior view. Abbreviations:
<emphasis id="84C35E3C5B5FEC1BFC56F9BFFC0FD804" bold="true" box="[946,968,1569,1587]" pageId="8" pageNumber="63">an</emphasis>
, antenna;
<emphasis id="84C35E3C5B5FEC1BFBC4F9BFFBF3D804" bold="true" box="[1056,1076,1569,1587]" pageId="8" pageNumber="63">c1</emphasis>
, coxa of thorax appendage 1;
<emphasis id="84C35E3C5B5FEC1BFAD0F9BFFA8ED804" bold="true" box="[1332,1353,1569,1587]" pageId="8" pageNumber="63">ce</emphasis>
, cercus;
<emphasis id="84C35E3C5B5FEC1BFA72F9BFFA6DD804" bold="true" box="[1430,1450,1569,1587]" pageId="8" pageNumber="63">ey</emphasis>
, compound eye;
<emphasis id="84C35E3C5B5FEC1BFEEFF9A6FEEBD87D" bold="true" box="[267,300,1592,1610]" pageId="8" pageNumber="63">f1-3</emphasis>
, femur of thorax appendage 1-3;
<emphasis id="84C35E3C5B5FEC1BFDA7F9A6FD9ED87D" bold="true" box="[579,601,1592,1610]" pageId="8" pageNumber="63">hc</emphasis>
, head capsule;
<emphasis id="84C35E3C5B5FEC1BFD38F9A6FD3FD87D" bold="true" box="[732,760,1592,1610]" pageId="8" pageNumber="63">mp</emphasis>
, maxillary palp;
<emphasis id="84C35E3C5B5FEC1BFC9BF9A6FC53D87D" bold="true" box="[895,916,1592,1610]" pageId="8" pageNumber="63">sp</emphasis>
, spine-like seta;
<emphasis id="84C35E3C5B5FEC1BFBC4F9A6FBFCD87D" bold="true" box="[1056,1083,1592,1610]" pageId="8" pageNumber="63">ta1</emphasis>
,
<emphasis id="84C35E3C5B5FEC1BFBA2F9A6FB97D87D" bold="true" box="[1094,1104,1592,1610]" pageId="8" pageNumber="63">3</emphasis>
, tarsus of thorax appendage 1, 3;
<emphasis id="84C35E3C5B5FEC1BFA88F9A6FA45D87D" bold="true" box="[1388,1410,1592,1610]" pageId="8" pageNumber="63">tcl</emphasis>
, tibia claw;
<emphasis id="84C35E3C5B5FEC1BFF51F9D0FF1BD857" bold="true" box="[181,220,1614,1632]" pageId="8" pageNumber="63">ti1-3</emphasis>
, tibia of thorax appendage 1-3;
<emphasis id="84C35E3C5B5FEC1BFE0DF9D0FDC6D857" bold="true" box="[489,513,1614,1632]" pageId="8" pageNumber="63">tr1</emphasis>
, trochanter of thorax appendage 1. Scale bars: A, C, 1 mm; B, D, 0.5 mm.
</paragraph>
</caption>
<paragraph id="B608822E5B5FEC1AFCA7F8EFFDCCD8FB" blockId="8.[811,1457,1681,2028]" lastBlockId="9.[131,777,1682,2027]" lastPageId="9" lastPageNumber="64" pageId="8" pageNumber="63">
Post-ocular segment 8 (thorax segment 3; metathorax) about 1.4× longer than preceding segment and about as wide as that as well; rectangular shaped. Ventrally with a pair of prominent appendages (hind leg) [thoracopod 3] (
<figureCitation id="2E8C9EAB5B5FEC1BFB29F84FFAE1D9DC" box="[1229,1318,2001,2027]" captionStart="FIG" captionStartId="6.[133,144,1399,1417]" captionTargetBox="[151,1437,215,1353]" captionTargetId="figure-276@6.[151,1436,214,1353]" captionTargetPageId="6" captionText="FIG. 2. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp., PED 0147, close-up on thorax appendages: A, thorax appendage 1; B, thorax appendage 2; C, thorax appendage 3. Abbreviations: c2, coxa of thorax appendage 2; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; ta1-3, tarsus of thorax appendage 1-3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000907" httpUri="https://zenodo.org/record/6000907/files/figure.png" pageId="8" pageNumber="63">Figs 2C</figureCitation>
;
<figureCitation id="2E8C9EAB5B5FEC1BFAD5F84FFAF8D9DC" box="[1329,1343,2001,2027]" captionStart="FIG" captionStartId="7.[132,143,1159,1177]" captionTargetBox="[132,775,215,1117]" captionTargetId="figure-720@7.[407,684,485,756]" captionTargetPageId="7" captionText="FIG. 3. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp., PED 0147, close-up on posterior end with thorax appendage 3 and cerci. Abbreviations: ce, cercus; ta3, tarsus of thorax appendage 3; ti3, tibia of thorax appendage 3. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000911" httpUri="https://zenodo.org/record/6000911/files/figure.png" pageId="8" pageNumber="63">3</figureCitation>
); basically cylindrical, composed of supposedly five elements; slightly longer than mid leg [thoracopod 2].
</paragraph>
<caption id="E2C8D2A65B5EEC1AFF60F994FD8FD857" ID-DOI="http://doi.org/10.5281/zenodo.6000917" ID-Zenodo-Dep="6000917" httpUri="https://zenodo.org/record/6000917/files/figure.png" pageId="9" pageNumber="64" startId="9.[132,143,1546,1564]" targetBox="[149,1435,216,1501]" targetPageId="9">
<paragraph id="B608822E5B5EEC1AFF60F994FD8FD857" blockId="9.[132,1457,1545,1632]" pageId="9" pageNumber="64">
FIG. 5. —
<taxonomicName id="71B7F9AD5B5EEC1AFF30F997FE02D82B" box="[212,453,1545,1564]" class="Insecta" family="Tettigoniidae" genus="Gryllobencain" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="patrickmuelleri" status="n. gen., n. sp.">
<emphasis id="84C35E3C5B5EEC1AFF30F997FE02D82B" box="[212,453,1545,1564]" italics="true" pageId="9" pageNumber="64">Gryllobencain patrickmuelleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF0E3475B5EEC1AFE2EF994FDF8D82B" box="[458,575,1546,1564]" pageId="9" pageNumber="64" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, BUB 3073:
<emphasis id="84C35E3C5B5EEC1AFD42F994FD74D82B" bold="true" box="[678,691,1546,1564]" pageId="9" pageNumber="64">A</emphasis>
, overview in dorsal view;
<emphasis id="84C35E3C5B5EEC1AFC6DF994FC51D82B" bold="true" box="[905,918,1546,1564]" pageId="9" pageNumber="64">B</emphasis>
, close-up in anterior head area with ocelli;
<emphasis id="84C35E3C5B5EEC1AFB18F997FACDD82C" bold="true" box="[1276,1290,1545,1563]" pageId="9" pageNumber="64">C</emphasis>
, overview in ventral view;
<emphasis id="84C35E3C5B5EEC1AFF51F9BEFF04D805" bold="true" box="[181,195,1568,1586]" pageId="9" pageNumber="64">D</emphasis>
, close-up on distal part of thorax appendage 1. In A and C background was desaturated to enhance contrast. Abbreviations:
<emphasis id="84C35E3C5B5EEC1AFB07F9BEFB3ED805" bold="true" box="[1251,1273,1568,1586]" pageId="9" pageNumber="64">an</emphasis>
, antenna;
<emphasis id="84C35E3C5B5EEC1AFAB6F9BEFAA0D805" bold="true" box="[1362,1383,1568,1586]" pageId="9" pageNumber="64">ce</emphasis>
, cercus;
<emphasis id="84C35E3C5B5EEC1AFF60F9A9FF53D87E" bold="true" box="[132,148,1591,1609]" pageId="9" pageNumber="64">cl</emphasis>
, claw of thorax appendage;
<emphasis id="84C35E3C5B5EEC1AFE60F9A9FE61D87E" bold="true" box="[388,422,1591,1609]" pageId="9" pageNumber="64">f1-3</emphasis>
, femur of thorax appendage 1-3;
<emphasis id="84C35E3C5B5EEC1AFD26F9A9FD19D87E" bold="true" box="[706,734,1591,1609]" pageId="9" pageNumber="64">mp</emphasis>
, maxillary palp;
<emphasis id="84C35E3C5B5EEC1AFC8CF9A9FCB9D87E" bold="true" box="[872,894,1591,1609]" pageId="9" pageNumber="64">oc</emphasis>
, ocelli;
<emphasis id="84C35E3C5B5EEC1AFC5AF9A9FC1DD87E" bold="true" box="[958,986,1591,1609]" pageId="9" pageNumber="64">ta1</emphasis>
, tarsus of thorax appendage 1;
<emphasis id="84C35E3C5B5EEC1AFB03F9A9FAC9D87E" bold="true" box="[1255,1294,1591,1609]" pageId="9" pageNumber="64">ti1-3</emphasis>
, tibia of thorax appendage 1-3. Scale bars: A, C, 0.5 mm; B, D, 0.1 mm.
</paragraph>
</caption>
<paragraph id="B608822E5B5EEC1AFF7FF94CFBE4D9DB" blockId="9.[131,777,1682,2027]" lastBlockId="9.[811,1457,1682,2028]" pageId="9" pageNumber="64">Element 1 (coxa) [basipod?] not differentiable from next element, i.e. element 2 (trochanter) [endopod element 1?]. Proximal region (coxa + trochanter?) slightly longer than wide. Element 3 (femur) [endopod element 2?] very elongate; about 8× longer than proximal region, similar width as preceding element, but tapering slightly distally; with four strong spine-like setae laterally. Element 4 (tibia) [endopod element 3?] elongate, but about 0.8× shorter than femur; about 0.67× the width of the femur, tapering slightly distally; covered with smaller setae and stronger spine-like setae; no very strong spine very medio-distally, but with one strong spine-like seta on a similar position as on the tibia of fore and mid legs, but shorter than in the latter. Element 5 (tarsus) [endopod element 4?] as long as tibia and slightly less wide than that as well; with numerous smaller, thin setae throughout; distally with a pair of claws, possible attachment structure between claws not apparent; tarsus subdivided into three sub-elements which are of similar width, but with varying length (from proximal to distal: 0.52×, 0.22× and 0.26× of total length).</paragraph>
</subSection>
<caption id="E2C8D2A65B5DEC19FF60F999FE1FD870" ID-DOI="http://doi.org/10.5281/zenodo.6000921" ID-Zenodo-Dep="6000921" httpUri="https://zenodo.org/record/6000921/files/figure.png" pageId="10" pageNumber="65" startId="10.[132,143,1543,1561]" targetBox="[153,1439,215,1500]" targetPageId="10">
<paragraph id="B608822E5B5DEC19FF60F999FE1FD870" blockId="10.[132,1456,1543,1607]" pageId="10" pageNumber="65">
FIG. 6. —
<taxonomicName id="71B7F9AD5B5DEC19FF30F999FE02D82E" box="[212,453,1543,1561]" class="Insecta" family="Tettigoniidae" genus="Gryllobencain" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="patrickmuelleri" status="n. gen., n. sp.">
<emphasis id="84C35E3C5B5DEC19FF30F999FE02D82E" box="[212,453,1543,1561]" italics="true" pageId="10" pageNumber="65">Gryllobencain patrickmuelleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF0E3475B5DEC19FE2EF999FD87D82E" box="[458,576,1543,1561]" pageId="10" pageNumber="65" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, BUB 3072:
<emphasis id="84C35E3C5B5DEC19FD42F999FD74D82E" bold="true" box="[678,691,1543,1561]" pageId="10" pageNumber="65">A</emphasis>
, overview in dorsal view;
<emphasis id="84C35E3C5B5DEC19FC6DF999FC51D82E" bold="true" box="[905,918,1543,1561]" pageId="10" pageNumber="65">B</emphasis>
, close-up on cercus;
<emphasis id="84C35E3C5B5DEC19FBAFF999FB9ED82E" bold="true" box="[1099,1113,1543,1561]" pageId="10" pageNumber="65">C</emphasis>
, overview in ventral view;
<emphasis id="84C35E3C5B5DEC19FAD0F999FA85D82E" bold="true" box="[1332,1346,1543,1561]" pageId="10" pageNumber="65">D</emphasis>
, close-up on head. Abbreviations:
<emphasis id="84C35E3C5B5DEC19FED2F980FE8BD807" bold="true" box="[310,332,1566,1584]" pageId="10" pageNumber="65">an</emphasis>
, antenna;
<emphasis id="84C35E3C5B5DEC19FE40F980FE7ED807" bold="true" box="[420,441,1566,1584]" pageId="10" pageNumber="65">ce</emphasis>
, cercus;
<emphasis id="84C35E3C5B5DEC19FDE3F980FDD0D807" bold="true" box="[519,535,1566,1584]" pageId="10" pageNumber="65">f1</emphasis>
,
<emphasis id="84C35E3C5B5DEC19FDC6F980FDEBD807" bold="true" box="[546,556,1566,1584]" pageId="10" pageNumber="65">3</emphasis>
, femur of thorax appendage 1, 3;
<emphasis id="84C35E3C5B5DEC19FCA8F980FCA5D807" bold="true" box="[844,866,1566,1584]" pageId="10" pageNumber="65">hc</emphasis>
, head capsule;
<emphasis id="84C35E3C5B5DEC19FC03F980FBC4D807" bold="true" box="[999,1027,1566,1584]" pageId="10" pageNumber="65">mp</emphasis>
, maxillary palp;
<emphasis id="84C35E3C5B5DEC19FB69F980FB65D807" bold="true" box="[1165,1186,1566,1584]" pageId="10" pageNumber="65">ti1</emphasis>
,
<emphasis id="84C35E3C5B5DEC19FB49F980FB70D807" bold="true" box="[1197,1207,1566,1584]" pageId="10" pageNumber="65">3</emphasis>
, tibia of thorax appendage 1, 3. Scale bars: A, C, 1 mm; B, D, 0.5 mm.
</paragraph>
</caption>
<subSection id="DF389BC55B5DEC18FF60F90CFB50D9DC" lastPageId="11" lastPageNumber="66" pageId="10" pageNumber="65" type="multiple">
<paragraph id="B608822E5B5DEC19FF60F90CFE50D89B" blockId="10.[132,407,1682,1709]" box="[132,407,1682,1709]" pageId="10" pageNumber="65">
<heading id="ED4035425B5DEC19FF60F90CFE50D89B" box="[132,407,1682,1709]" fontSize="11" level="2" pageId="10" pageNumber="65" reason="8">
<emphasis id="84C35E3C5B5DEC19FF60F90CFE50D89B" box="[132,407,1682,1709]" italics="true" pageId="10" pageNumber="65">Abdomen [posterior trunk]</emphasis>
</heading>
</paragraph>
<paragraph id="B608822E5B5DEC19FF60F92CFD32D93B" blockId="10.[130,776,1714,2028]" pageId="10" pageNumber="65">
Post-ocular segments 9-19 differentiated from further anterior ones, forming distinct unit (abdomen; not corresponding to abdomen in other crustacean groups) (
<figureCitation id="2E8C9EAB5B5DEC19FDC0F96CFD5FD93B" box="[548,664,1778,1804]" captionStart="FIG" captionStartId="5.[132,143,1383,1401]" captionTargetBox="[149,1434,216,1340]" captionTargetId="figure-278@5.[149,1434,216,1341]" captionTargetPageId="5" captionText="FIG. 1. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp.,PED 0147:A, overview in dorsal view; B, overview in ventral view; C, close-up on maxillary palp, distal part; D, close-up on thorax appendage 1. Abbreviations: an, antenna; ce, cercus; cl1, claw of thorax appendage 1; f1-2, femur of thorax appendage 1-2; hc, head capsule; mp, maxillary palp; sp, spine-like seta; t1-3, thorax segment 1-3; ta1-3, tarsus of thorax appendage 1-3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, B, 1 mm; C, 0.25 mm; D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000905" httpUri="https://zenodo.org/record/6000905/files/figure.png" pageId="10" pageNumber="65">Figs 1A, B</figureCitation>
;
<figureCitation id="2E8C9EAB5B5DEC19FD40F96CFD2FD93B" box="[676,744,1778,1805]" captionStart="FIG" captionStartId="9.[132,143,1546,1564]" captionTargetBox="[149,1435,216,1501]" captionTargetId="figure-212@9.[149,1434,216,1502]" captionTargetPageId="9" captionText="FIG. 5. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3073: A, overview in dorsal view; B, close-up in anterior head area with ocelli; C, overview in ventral view; D, close-up on distal part of thorax appendage 1. In A and C background was desaturated to enhance contrast. Abbreviations: an, antenna; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; mp, maxillary palp; oc, ocelli; ta1, tarsus of thorax appendage 1; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, C, 0.5 mm; B, D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6000917" httpUri="https://zenodo.org/record/6000917/files/figure.png" pageId="10" pageNumber="65">5A, C</figureCitation>
).
</paragraph>
<paragraph id="B608822E5B5DEC19FF7FF88CFD82D9BC" blockId="10.[130,776,1714,2028]" pageId="10" pageNumber="65">Only eight distinct units differentiable, first seven probably corresponding to abdominal segments 1 to 7, rectangularshaped. Last unit most likely corresponding to several abdomen segments (8-11?), triangular-shaped.</paragraph>
<paragraph id="B608822E5B5DEC19FF7FF80FFCCFD9DC" blockId="10.[130,776,1714,2028]" pageId="10" pageNumber="65">Post-ocular segment 9 (abdomen segment 1) about 1.4× longer than metathorax; but less wide, about 0.67× of metathorax width; one spine-like seta postero-laterally on each side.</paragraph>
<paragraph id="B608822E5B5DEC19FCA7F92CFBABD91B" blockId="10.[811,1457,1714,2028]" pageId="10" pageNumber="65">Post-ocular segment 10 (abdomen segment 2) similar to preceding segment, but very slightly shorter and slightly wider posteriorly (widening posteriorly); one spine-like seta postero-laterally on each side.</paragraph>
<paragraph id="B608822E5B5DEC19FCA7F8AFFB57D9BC" blockId="10.[811,1457,1714,2028]" pageId="10" pageNumber="65">Post-ocular segment 11 (abdomen segment 3) about half as long, but as wide as the preceding segment; one spine-like seta postero-laterally respectively.</paragraph>
<paragraph id="B608822E5B5DEC19FCA7F80FFA77D9DC" blockId="10.[811,1457,1714,2028]" pageId="10" pageNumber="65">Post-ocular segment 12 (abdomen segment 4) slightly longer and wider than preceding segment; two spine-like setae postero-laterally respectively, one slightly longer than the other.</paragraph>
<caption id="E2C8D2A65B5CEC18FF60F99DFEA1D86E" ID-DOI="http://doi.org/10.5281/zenodo.6000923" ID-Zenodo-Dep="6000923" httpUri="https://zenodo.org/record/6000923/files/figure.png" pageId="11" pageNumber="66" startId="11.[132,143,1539,1557]" targetBox="[153,1438,214,1494]" targetPageId="11">
<paragraph id="B608822E5B5CEC18FF60F99DFEA1D86E" blockId="11.[132,1457,1538,1625]" pageId="11" pageNumber="66">
FIG. 7. — Close-ups on head area in different specimens of
<taxonomicName id="71B7F9AD5B5CEC18FD9EF99CFCACD822" box="[634,875,1538,1557]" class="Insecta" family="Tettigoniidae" genus="Gryllobencain" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="patrickmuelleri" status="n. gen., n. sp.">
<emphasis id="84C35E3C5B5CEC18FD9EF99CFCACD822" box="[634,875,1538,1557]" italics="true" pageId="11" pageNumber="66">Gryllobencain patrickmuelleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF0E3475B5CEC18FC95F99DFC21D822" box="[881,998,1539,1557]" pageId="11" pageNumber="66" rank="species">n. gen., n. sp.</taxonomicNameLabel>
as normal versions (
<emphasis id="84C35E3C5B5CEC18FB77F99DFB67D822" bold="true" box="[1171,1184,1539,1557]" pageId="11" pageNumber="66">A</emphasis>
,
<emphasis id="84C35E3C5B5CEC18FB4FF99CFB7ED823" bold="true" box="[1195,1209,1538,1556]" pageId="11" pageNumber="66">C</emphasis>
,
<emphasis id="84C35E3C5B5CEC18FB20F99DFB17D822" bold="true" box="[1220,1232,1539,1557]" pageId="11" pageNumber="66">E</emphasis>
) and colour-marked (
<emphasis id="84C35E3C5B5CEC18FA60F99DFA56D822" bold="true" box="[1412,1425,1539,1557]" pageId="11" pageNumber="66">B</emphasis>
,
<emphasis id="84C35E3C5B5CEC18FA78F99DFA6DD822" bold="true" box="[1436,1450,1539,1557]" pageId="11" pageNumber="66">D</emphasis>
,
<emphasis id="84C35E3C5B5CEC18FF60F987FF48D81C" bold="true" box="[132,143,1561,1579]" pageId="11" pageNumber="66">F</emphasis>
):
<emphasis id="84C35E3C5B5CEC18FF7BF987FF06D81C" bold="true" box="[159,193,1561,1579]" pageId="11" pageNumber="66">A -D</emphasis>
, PED 0178;
<emphasis id="84C35E3C5B5CEC18FECEF987FEF0D81C" bold="true" box="[298,311,1561,1579]" pageId="11" pageNumber="66">A</emphasis>
,
<emphasis id="84C35E3C5B5CEC18FEA5F987FE89D81C" bold="true" box="[321,334,1561,1579]" pageId="11" pageNumber="66">B</emphasis>
, latero-posterior view;
<emphasis id="84C35E3C5B5CEC18FDEBF987FDDAD81C" bold="true" box="[527,541,1561,1579]" pageId="11" pageNumber="66">C</emphasis>
,
<emphasis id="84C35E3C5B5CEC18FDC3F987FDF2D81C" bold="true" box="[551,565,1561,1579]" pageId="11" pageNumber="66">D</emphasis>
, mostly anterior view; elements of maxillary palp difficult to differentiate;
<emphasis id="84C35E3C5B5CEC18FB73F987FB64D81C" bold="true" box="[1175,1187,1561,1579]" pageId="11" pageNumber="66">E</emphasis>
,
<emphasis id="84C35E3C5B5CEC18FB4AF987FB7ED81C" bold="true" box="[1198,1209,1561,1579]" pageId="11" pageNumber="66">F</emphasis>
, BUB 3073, ventral view. Abbreviations:
<emphasis id="84C35E3C5B5CEC18FF0EF9AEFF3DD875" bold="true" box="[234,250,1584,1602]" pageId="11" pageNumber="66">lp</emphasis>
, labial palp;
<emphasis id="84C35E3C5B5CEC18FE82F9AEFEB5D875" bold="true" box="[358,370,1584,1602]" pageId="11" pageNumber="66">lr</emphasis>
, labrum;
<emphasis id="84C35E3C5B5CEC18FE26F9AEFE19D875" bold="true" box="[450,478,1584,1602]" pageId="11" pageNumber="66">md</emphasis>
, mandible;
<emphasis id="84C35E3C5B5CEC18FDA6F9AEFDA9D875" bold="true" box="[578,622,1584,1602]" pageId="11" pageNumber="66">mmx</emphasis>
, median part of maxilla;
<emphasis id="84C35E3C5B5CEC18FCDAF9AEFC9DD875" bold="true" box="[830,858,1584,1602]" pageId="11" pageNumber="66">mp</emphasis>
, maxillary palp;
<emphasis id="84C35E3C5B5CEC18FC01F9AEFBC7D875" bold="true" box="[997,1024,1584,1602]" pageId="11" pageNumber="66">mx</emphasis>
, maxilla;
<emphasis id="84C35E3C5B5CEC18FBB4F9AEFBB1D875" bold="true" box="[1104,1142,1584,1602]" pageId="11" pageNumber="66">pmx</emphasis>
, proximal part of maxilla. Scale bars: A-D, 0.3 mm; E, F, 0.2 mm.
</paragraph>
</caption>
<paragraph id="B608822E5B5CEC18FF7FF90CFDEED93B" blockId="11.[131,777,1682,2028]" pageId="11" pageNumber="66">Post-ocular segment 13 (abdomen segment 5) similar to preceding segment, but slightly longer and very slightly tapering posteriorly; two spine-like setae postero-laterally on each side, one slightly longer than the other.</paragraph>
<paragraph id="B608822E5B5CEC18FF7FF88CFF28D9BC" blockId="11.[131,777,1682,2028]" pageId="11" pageNumber="66">Post-ocular segment 14 (abdomen segment 6) slightly shorter and less wide than preceding segment, tapering slightly posteriorly; one spine-like seta postero-laterally on each side.</paragraph>
<paragraph id="B608822E5B5CEC18FF7FF80FFD55D9DC" blockId="11.[131,777,1682,2028]" pageId="11" pageNumber="66">Post-ocular segment 15 (abdomen segment 7) similar to preceding segment, but less wide and tapering posteriorly; one spine-like seta postero-laterally on each side.</paragraph>
<paragraph id="B608822E5B5CEC18FCA7F90CFB50D9DC" blockId="11.[810,1457,1682,2028]" pageId="11" pageNumber="66">
Trunk end (post-ocular segments 16-19?; abdomen segments 8-11?) as long as first abdomen segment; anteriorly as wide as preceding segment, but tapering drastically posteriorly. Ventrally with a pair of prominent appendages (cerci) (
<figureCitation id="2E8C9EAB5B5CEC18FAB0F96CFC87D91B" captionStart="FIG" captionStartId="5.[132,143,1383,1401]" captionTargetBox="[149,1434,216,1340]" captionTargetId="figure-278@5.[149,1434,216,1341]" captionTargetPageId="5" captionText="FIG. 1. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp.,PED 0147:A, overview in dorsal view; B, overview in ventral view; C, close-up on maxillary palp, distal part; D, close-up on thorax appendage 1. Abbreviations: an, antenna; ce, cercus; cl1, claw of thorax appendage 1; f1-2, femur of thorax appendage 1-2; hc, head capsule; mp, maxillary palp; sp, spine-like seta; t1-3, thorax segment 1-3; ta1-3, tarsus of thorax appendage 1-3; tcl, tibia claw; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, B, 1 mm; C, 0.25 mm; D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000905" httpUri="https://zenodo.org/record/6000905/files/figure.png" pageId="11" pageNumber="66">Figs 1A, B</figureCitation>
;
<figureCitation id="2E8C9EAB5B5CEC18FCAEF88CFC9FD91B" box="[842,856,1810,1836]" captionStart="FIG" captionStartId="7.[132,143,1159,1177]" captionTargetBox="[132,775,215,1117]" captionTargetId="figure-720@7.[407,684,485,756]" captionTargetPageId="7" captionText="FIG. 3. — Holotype of Gryllobencain patrickmuelleri n. gen., n. sp., PED 0147, close-up on posterior end with thorax appendage 3 and cerci. Abbreviations: ce, cercus; ta3, tarsus of thorax appendage 3; ti3, tibia of thorax appendage 3. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000911" httpUri="https://zenodo.org/record/6000911/files/figure.png" pageId="11" pageNumber="66">3</figureCitation>
;
<figureCitation id="2E8C9EAB5B5CEC18FC81F88FFC6FD91B" box="[869,936,1809,1836]" captionStart="FIG" captionStartId="9.[132,143,1546,1564]" captionTargetBox="[149,1435,216,1501]" captionTargetId="figure-212@9.[149,1434,216,1502]" captionTargetPageId="9" captionText="FIG. 5. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3073: A, overview in dorsal view; B, close-up in anterior head area with ocelli; C, overview in ventral view; D, close-up on distal part of thorax appendage 1. In A and C background was desaturated to enhance contrast. Abbreviations: an, antenna; ce, cercus; cl, claw of thorax appendage; f1-3, femur of thorax appendage 1-3; mp, maxillary palp; oc, ocelli; ta1, tarsus of thorax appendage 1; ti1-3, tibia of thorax appendage 1-3. Scale bars: A, C, 0.5 mm; B, D, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6000917" httpUri="https://zenodo.org/record/6000917/files/figure.png" pageId="11" pageNumber="66">5A, C</figureCitation>
;
<figureCitation id="2E8C9EAB5B5CEC18FC55F88CFC13D91B" box="[945,980,1810,1836]" captionStart="FIG" captionStartId="10.[132,143,1543,1561]" captionTargetBox="[153,1439,215,1500]" captionTargetId="figure-180@10.[153,1438,214,1500]" captionTargetPageId="10" captionText="FIG. 6. — Gryllobencain patrickmuelleri n. gen., n. sp., BUB 3072: A, overview in dorsal view; B, close-up on cercus; C, overview in ventral view; D, close-up on head. Abbreviations: an, antenna; ce, cercus; f1, 3, femur of thorax appendage 1, 3; hc, head capsule; mp, maxillary palp; ti1, 3, tibia of thorax appendage 1, 3. Scale bars: A, C, 1 mm; B, D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6000921" httpUri="https://zenodo.org/record/6000921/files/figure.png" pageId="11" pageNumber="66">6B</figureCitation>
); basically cylindrical, not subdivided; about 0.8× the length of the antenna; as wide as the tibia of the hind leg proximally and tapering slightly until about 70 % of its length, then strongly tapering towards tip; bearing numerous spine-like setae of varying sizes, mostly strong and very long (compared to similar structures elsewhere); also with a few smaller, shorter setae throughout.
</paragraph>
</subSection>
<caption id="E2C8D2A65B5BEC1FFF60FA74FE32D824" ID-DOI="http://doi.org/10.5281/zenodo.6000927" ID-Zenodo-Dep="6000927" httpUri="https://zenodo.org/record/6000927/files/figure.png" pageId="12" pageNumber="67" startId="12.[132,143,1514,1532]" targetBox="[152,1438,193,1479]" targetPageId="12">
<paragraph id="B608822E5B5BEC1FFF60FA74FE32D824" blockId="12.[132,1457,1514,1555]" pageId="12" pageNumber="67">
FIG. 8. — Reconstructed growth series of
<taxonomicName id="71B7F9AD5B5BEC1FFE3FFA74FD0DDBCB" box="[475,714,1514,1532]" class="Insecta" family="Tettigoniidae" genus="Gryllobencain" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="67" phylum="Arthropoda" rank="species" species="patrickmuelleri" status="n. gen., n. sp.">
<emphasis id="84C35E3C5B5BEC1FFE3FFA74FD0DDBCB" box="[475,714,1514,1532]" italics="true" pageId="12" pageNumber="67">Gryllobencain patrickmuelleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FF0E3475B5BEC1FFD2BFA74FC87DBCB" box="[719,832,1514,1532]" pageId="12" pageNumber="67" rank="species">n. gen., n. sp.</taxonomicNameLabel>
based on specimens investigated in this study:
<emphasis id="84C35E3C5B5BEC1FFB2FFA74FB1FDBCB" bold="true" box="[1227,1240,1514,1532]" pageId="12" pageNumber="67">A</emphasis>
, BUB 3073;
<emphasis id="84C35E3C5B5BEC1FFAA4FA74FA8ADBCB" bold="true" box="[1344,1357,1514,1532]" pageId="12" pageNumber="67">B</emphasis>
, PED 0147;
<emphasis id="84C35E3C5B5BEC1FFF60F99FFF55D824" bold="true" box="[132,146,1537,1555]" pageId="12" pageNumber="67">C</emphasis>
, PED 0178 + BUB 3072. Scale bar: 1 mm.
</paragraph>
</caption>
<subSection id="DF389BC55B5BEC1FFF60F9EDFE6CD97B" pageId="12" pageNumber="67" type="multiple">
<paragraph id="B608822E5B5BEC1FFF60F9EDFEF6D8BB" blockId="12.[132,305,1650,1677]" box="[132,305,1650,1677]" pageId="12" pageNumber="67">
<heading id="ED4035425B5BEC1FFF60F9EDFEF6D8BB" box="[132,305,1650,1677]" fontSize="11" level="2" pageId="12" pageNumber="67" reason="8">
<emphasis id="84C35E3C5B5BEC1FFF60F9EDFEF6D8BB" box="[132,305,1650,1677]" italics="true" pageId="12" pageNumber="67">Size of specimens</emphasis>
</heading>
</paragraph>
<paragraph id="B608822E5B5BEC1FFF60F90CFE6CD97B" blockId="12.[132,777,1682,1868]" pageId="12" pageNumber="67">
While the specimens are very similar in overall appearance, they differ in size. Though it remains difficult to measure, it appears that the
<specimenCount id="A0B149A75B5BEC1FFEB6F94CFE32D8DB" box="[338,501,1746,1772]" pageId="12" pageNumber="67" type="generic">four specimens</specimenCount>
fall into three size classes (
<figureCitation id="2E8C9EAB5B5BEC1FFF69F96CFF0BD93B" box="[141,204,1778,1804]" captionStart="FIG" captionStartId="12.[132,143,1514,1532]" captionTargetBox="[152,1438,193,1479]" captionTargetId="figure-190@12.[703,980,744,1020]" captionTargetPageId="12" captionText="FIG. 8. — Reconstructed growth series of Gryllobencain patrickmuelleri n. gen., n. sp. based on specimens investigated in this study: A, BUB 3073; B, PED 0147; C, PED 0178 + BUB 3072. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6000927" httpUri="https://zenodo.org/record/6000927/files/figure.png" pageId="12" pageNumber="67">Fig. 8</figureCitation>
). It seems likely that these represent three successive stages. It remains unclear whether the largest size class corresponds to the adult stage.
</paragraph>
</subSection>
</subSubSection>
</treatment>
</document>