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<mods:title id="D2F7FBE79B515310B640245A83C97DBD">On the identity of some poorly known lithobiid centipedes described by Karl Verhoeff (Chilopoda: Lithobiomorpha)</mods:title>
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<heading id="FA2A81FAE107FFDCFEA6FBC6CD38FB79" bold="true" box="[264,869,1148,1174]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<taxonomicName id="66DD4D15E107FFDCFEA6FBC6CCAFFB79" ID-CoL="6LG8K" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" box="[264,754,1148,1174]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dollfusi" status="comb. nov.">
<emphasis id="93A9EA84E107FFDCFEA6FBC6CC7FFB79" bold="true" box="[264,546,1148,1174]" italics="true" pageId="1" pageNumber="2">Harpolithobius dollfusi</emphasis>
(
<bibRefCitation id="C54C4B67E107FFDCFD9CFBC6CCB7FB79" author="Verhoeff" box="[562,746,1148,1174]" firstAuthor="Verhoeff" pageId="1" pageNumber="12" refString="Verhoeff, K. (1901) Chilopoden und Diplopoden aus Rumanien. In: Faune de la Roumanie par M. Jaquet. Bulletin de la Societe des Sciences, Bucarest, 10, 168 - 177." type="journal article" year="1901">Verhoeff, 1901</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="889A57FFE107FFDCFD54FBC6CD38FB79" box="[762,869,1148,1174]" pageId="1" pageNumber="2">comb. n.</taxonomicNameLabel>
</emphasis>
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<paragraph id="A1623696E107FFDCFEA6FB25CFC4FB56" blockId="1.[264,869,1148,1209]" box="[264,409,1183,1209]" pageId="1" pageNumber="2">
<figureCitation id="39E62A13E107FFDCFEA6FB25CF2DFB56" box="[264,368,1183,1209]" captionStart="FIGURES 1 4" captionStartId="3.[264,382,1133,1157]" captionTargetBox="[351,1217,299,1101]" captionTargetId="figure@3.[351,1233,291,1102]" captionTargetPageId="3" captionText="FIGURES 1 4. Harpolithobius dollfusi (Verhoeff, 1901): 1 head, dorsal view; 2 maxillipede, ventral view; 3 coxa, ventro­lateral view; 4 female gonopods, ventral view. Scale bars: 1 mm (Figs 1 2), 0.5 mm (Fig. 4)." httpUri="https://zenodo.org/record/170549/files/figure.png" pageId="1" pageNumber="2">Figs 14</figureCitation>
,
<figureCitation id="39E62A13E107FFDCFED2FB25CFC4FB56" box="[380,409,1183,1209]" captionStart="FIGURES 14 16" captionStartId="9.[264,382,1822,1846]" captionTargetBox="[286,1300,1209,1800]" captionTargetId="figure@9.[625,963,1248,1805]" captionTargetPageId="9" captionText="FIGURES 14 16: 14 H. dollfusi, lectotype; 15 L. lapadensis, lectotype; 16 L. karamani, paralectotype." httpUri="https://zenodo.org/record/170551/files/figure.png" pageId="1" pageNumber="2">14</figureCitation>
</paragraph>
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<paragraph id="A1623696E107FFDCFEA6FB58CCF1FB17" blockId="1.[264,1322,1249,1593]" box="[264,684,1249,1273]" pageId="1" pageNumber="2">
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<taxonomicName id="66DD4D15E107FFDCFEA6FB58CCF5FB17" ID-CoL="62XZ5" authority="Dollfusi Verhoeff, 1901: 173" authorityName="Dollfusi Verhoeff" authorityPageNumber="173" authorityYear="1901" box="[264,680,1249,1273]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="93A9EA84E107FFDCFEA6FB58CF94FB16" box="[264,457,1250,1273]" italics="true" pageId="1" pageNumber="2">Lithobius Dollfusi</emphasis>
<treatmentCitation id="207C1087E107FFDCFE7EFB5BCCF5FB17" author="Verhoeff" box="[464,680,1249,1272]" page="173" pageId="1" pageNumber="2" year="1901">
<bibRefCitation id="C54C4B67E107FFDCFE7EFB5BCCF5FB17" author="Verhoeff" box="[464,680,1249,1272]" firstAuthor="Verhoeff" pageId="1" pageNumber="2" refString="Verhoeff, K. (1901) Chilopoden und Diplopoden aus Rumanien. In: Faune de la Roumanie par M. Jaquet. Bulletin de la Societe des Sciences, Bucarest, 10, 168 - 177." type="journal article" year="1901">Verhoeff, 1901: 173</bibRefCitation>
</treatmentCitation>
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.
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</paragraph>
<paragraph id="A1623696E107FFDCFEA6FAB8CD1CFAF7" blockId="1.[264,1322,1249,1593]" box="[264,833,1281,1305]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="81CD11B8E107FFDCFEA6FAB8CD1CFAF7" box="[264,833,1281,1305]" pageId="1" pageNumber="2">
<taxonomicName id="66DD4D15E107FFDCFEA6FAB8CFE1FAF6" box="[264,444,1282,1305]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dolfusi">
<emphasis id="93A9EA84E107FFDCFEA6FAB8CFE1FAF6" box="[264,444,1282,1305]" italics="true" pageId="1" pageNumber="2">Lithobius dolfusi</emphasis>
</taxonomicName>
[sic!]:
<treatmentCitation id="207C1087E107FFDCFDA7FABBCC9EFAF7" author="Matic" box="[521,707,1281,1304]" page="188" pageId="1" pageNumber="2" year="1964">
<bibRefCitation id="C54C4B67E107FFDCFDA7FABBCC9EFAF7" author="Matic" box="[521,707,1281,1304]" firstAuthor="Matic" pageId="1" pageNumber="2" refString="Matic, Z. (1964) Nota critica asupra unor specii de Lithobiidae (Chilopoda) din fauna Republicii Populare Romine. Studii S i cercet a ri de Biologie, Seria Zoologie, Cluj, 16 (3), 187 - 191." type="journal article" year="1964">Matic, 1964: 188</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="C54C4B67E107FFDCFD60FABBCD61FAF7" author="Matic, Z." box="[718,828,1281,1304]" pageId="1" pageNumber="2" pagination="1 - 272" refId="ref5658" refString="Matic, Z. (1966) Classa Chilopoda. Subclassa Anamorpha. In: Fauna Republicii Socialiste Romania. Editura Academiei Republicii Socialiste Romania, Bucuresti, 6 (1), 1 - 272." type="journal article" year="1966">1966: 252</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A1623696E107FFDCFEA6FA98CD57FAD7" blockId="1.[264,1322,1249,1593]" box="[264,778,1313,1337]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="81CD11B8E107FFDCFEA6FA98CD57FAD7" box="[264,778,1313,1337]" pageId="1" pageNumber="2">
<taxonomicName id="66DD4D15E107FFDCFEA6FA98CD58FAD7" authority="Moritz &amp; Fischer, 1979: 318" authorityName="Moritz &amp; Fischer" authorityPageNumber="318" authorityYear="1979" box="[264,773,1313,1337]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dollfusi">
<emphasis id="93A9EA84E107FFDCFEA6FA98CF9EFAD6" box="[264,451,1314,1337]" italics="true" pageId="1" pageNumber="2">Lithobius dollfusi</emphasis>
:
<treatmentCitation id="207C1087E107FFDCFE7CFA9BCD58FAD7" author="Moritz" box="[466,773,1313,1336]" page="318" pageId="1" pageNumber="2" year="1979">
<bibRefCitation id="C54C4B67E107FFDCFE7CFA9BCD58FAD7" author="Moritz" box="[466,773,1313,1336]" firstAuthor="Moritz" pageId="1" pageNumber="2" refString="Moritz, M., &amp; Fischer, S. - Ch. (1979) Die Typen der Myriapoden-Sammlung des zoologischen Museums Berlin. II. Chilopoda. Mitteilungen aus dem Zoologischen Museum in Berlin, 55 (2), 297 - 352." type="journal article" year="1979">Moritz &amp; Fischer, 1979: 318</bibRefCitation>
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.
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</paragraph>
<paragraph id="A1623696E107FFDCFEA6FAF8CC1EFA97" blockId="1.[264,1322,1249,1593]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="81CD11B8E107FFDCFEA6FAF8CC1EFA97" pageId="1" pageNumber="2">
<taxonomicName id="66DD4D15E107FFDCFEA6FAF8CCBEFAB7" ID-CoL="6LG8N" authority="Matic, 1958: 91" authorityName="Matic" authorityPageNumber="91" authorityYear="1958" box="[264,739,1345,1369]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E107FFDCFEA6FAF8CC6DFAB6" box="[264,560,1346,1369]" italics="true" pageId="1" pageNumber="2">Harpolithobius intermedius</emphasis>
<treatmentCitation id="207C1087E107FFDCFD99FAFBCCBEFAB7" author="Matic" box="[567,739,1345,1368]" page="91" pageId="1" pageNumber="2" year="1958">
<bibRefCitation id="C54C4B67E107FFDCFD99FAFBCCBEFAB7" author="Matic" box="[567,739,1345,1368]" firstAuthor="Matic" pageId="1" pageNumber="2" refString="Matic, Z. (1958) Zur Kenntnis der Gattung Harpolithobius in Siebenburgen (Chilopoda, Lithobiidae). Senckenbergiana biologica, 39 (1 - 2), 91 - 95." type="journal article" year="1958">Matic, 1958: 91</bibRefCitation>
</treatmentCitation>
</taxonomicName>
,
<figureCitation id="39E62A13E107FFDCFD41FAFBCD16FAB7" box="[751,843,1345,1368]" captionStart="FIGURES 1 4" captionStartId="3.[264,382,1133,1157]" captionTargetBox="[351,1217,299,1101]" captionTargetId="figure@3.[351,1233,291,1102]" captionTargetPageId="3" captionText="FIGURES 1 4. Harpolithobius dollfusi (Verhoeff, 1901): 1 head, dorsal view; 2 maxillipede, ventral view; 3 coxa, ventro­lateral view; 4 female gonopods, ventral view. Scale bars: 1 mm (Figs 1 2), 0.5 mm (Fig. 4)." httpUri="https://zenodo.org/record/170549/files/figure.png" pageId="1" pageNumber="2">Figs 13</figureCitation>
syn. n.; 1961a: 166; 1961b: 8184, Figs 20 22; 1966: 90, Fig. 35ac.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A1623696E107FFDCFEA6FA38CD84FA77" blockId="1.[264,1322,1249,1593]" box="[264,985,1409,1433]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="81CD11B8E107FFDCFEA6FA38CD84FA77" box="[264,985,1409,1433]" pageId="1" pageNumber="2">
<taxonomicName id="66DD4D15E107FFDCFEA6FA38CDD1FA77" ID-CoL="5HLLF" authority="Matic, 1958: 93" authorityName="Matic" authorityPageNumber="93" authorityYear="1958" box="[264,908,1409,1433]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="subSpecies" species="intermedius" subSpecies="transsylvanicus">
<emphasis id="93A9EA84E107FFDCFEA6FA38CC81FA76" box="[264,732,1410,1433]" italics="true" pageId="1" pageNumber="2">Harpolithobius intermedius transsylvanicus</emphasis>
<treatmentCitation id="207C1087E107FFDCFD4DFA3BCDD1FA77" author="Matic" box="[739,908,1409,1432]" page="93" pageId="1" pageNumber="2" year="1958">
<bibRefCitation id="C54C4B67E107FFDCFD4DFA3BCDD1FA77" author="Matic" box="[739,908,1409,1432]" firstAuthor="Matic" pageId="1" pageNumber="2" refString="Matic, Z. (1958) Zur Kenntnis der Gattung Harpolithobius in Siebenburgen (Chilopoda, Lithobiidae). Senckenbergiana biologica, 39 (1 - 2), 91 - 95." type="journal article" year="1958">Matic, 1958: 93</bibRefCitation>
</treatmentCitation>
</taxonomicName>
<taxonomicNameLabel id="889A57FFE107FFDCFC3DFA3BCD84FA77" box="[915,985,1409,1432]" pageId="1" pageNumber="2">syn. n.</taxonomicNameLabel>
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</paragraph>
<paragraph id="A1623696E107FFDCFEA6FA18CDD2FA57" blockId="1.[264,1322,1249,1593]" box="[264,911,1441,1465]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="81CD11B8E107FFDCFEA6FA18CDD2FA57" box="[264,911,1441,1465]" pageId="1" pageNumber="2">
<taxonomicName id="66DD4D15E107FFDCFEA6FA18CDD6FA57" ID-CoL="6LG8N" authority="Dobroruka, 1960: 200" authorityName="Dobroruka" authorityPageNumber="200" authorityYear="1960" box="[264,907,1441,1465]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="subSpecies" species="spinipes" subSpecies="intermedius">
<emphasis id="93A9EA84E107FFDCFEA6FA18CCD0FA56" box="[264,653,1442,1465]" italics="true" pageId="1" pageNumber="2">Harpolithobius spinipes intermedius</emphasis>
:
<treatmentCitation id="207C1087E107FFDCFD32FA1BCDD6FA57" author="Dobroruka" box="[668,907,1441,1464]" page="200" pageId="1" pageNumber="2" year="1960">
<bibRefCitation id="C54C4B67E107FFDCFD32FA1BCDD6FA57" author="Dobroruka" box="[668,907,1441,1464]" firstAuthor="Dobroruka" pageId="1" pageNumber="2" refString="Dobroruka, L. (1960) Bemerkungen zur Gattung Harpolithobius (Chilopoda). Zoologischer Anzeiger, 164 (5 - 6), 198 - 201." type="journal article" year="1960">Dobroruka, 1960: 200</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A1623696E107FFDCFEA6FA78CD24FA37" blockId="1.[264,1322,1249,1593]" box="[264,889,1473,1497]" pageId="1" pageNumber="2">
<taxonomicName id="66DD4D15E107FFDCFEA6FA78CD28FA37" ID-CoL="6LG8N" authority="Deltshev et al., 2000: 535" authorityName="Deltshev et al." authorityPageNumber="535" authorityYear="2000" box="[264,885,1473,1497]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E107FFDCFEA6FA78CFF6FA36" box="[264,427,1474,1497]" italics="true" pageId="1" pageNumber="2">Harpolithobius</emphasis>
cf.
<emphasis id="93A9EA84E107FFDCFE7AFA78CC0FFA36" box="[468,594,1474,1497]" italics="true" pageId="1" pageNumber="2">intermedius</emphasis>
: Deltshev et al., 2000: 535
</taxonomicName>
.
</paragraph>
<paragraph id="A1623696E107FFDCFEA6FA58CD51FA17" blockId="1.[264,1322,1249,1593]" box="[264,780,1505,1529]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="81CD11B8E107FFDCFEA6FA58CD51FA17" box="[264,780,1505,1529]" pageId="1" pageNumber="2">
<taxonomicName id="66DD4D15E107FFDCFEA6FA58CD55FA17" ID-CoL="6LG8N" authority="Stoev, 2002: 55" authorityName="Stoev" authorityPageNumber="55" authorityYear="2002" box="[264,776,1505,1529]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E107FFDCFEA6FA58CFF6FA16" box="[264,427,1506,1529]" italics="true" pageId="1" pageNumber="2">Harpolithobius</emphasis>
cf.
<emphasis id="93A9EA84E107FFDCFE7AFA58CC0FFA16" box="[468,594,1506,1529]" italics="true" pageId="1" pageNumber="2">intermedius</emphasis>
:
<treatmentCitation id="207C1087E107FFDCFDCEFA5BCD55FA17" author="Stoev" box="[608,776,1505,1528]" page="55" pageId="1" pageNumber="2" year="2002">
<bibRefCitation id="C54C4B67E107FFDCFDCEFA5BCD55FA17" author="Stoev" box="[608,776,1505,1528]" firstAuthor="Stoev" pageId="1" pageNumber="2" refString="Stoev, P. (2002) A Catalogue and Key to the centipedes (Chilopoda) of Bulgaria. Pensoft Publishers, Sofia-Moscow, 103 pp." type="book" year="2002">Stoev, 2002: 55</bibRefCitation>
</treatmentCitation>
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.
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</paragraph>
<paragraph id="A1623696E107FFDCFEA6F9B8CD3BF9F7" blockId="1.[264,1322,1249,1593]" box="[264,870,1537,1561]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="81CD11B8E107FFDCFEA6F9B8CD3BF9F7" box="[264,870,1537,1561]" pageId="1" pageNumber="2">
<taxonomicName id="66DD4D15E107FFDCFEA6F9B8CC86F9F7" ID-CoL="6LG8N" authority="Ilie, 2003a: 89" authorityName="Ilie" authorityPageNumber="89" authorityYear="2003" box="[264,731,1537,1561]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E107FFDCFEA6F9B8CC72F9F6" box="[264,559,1538,1561]" italics="true" pageId="1" pageNumber="2">Harpolithobius intermedius</emphasis>
:
<treatmentCitation id="207C1087E107FFDCFD90F9BBCC86F9F7" author="Ilie" box="[574,731,1537,1560]" page="89" pageId="1" pageNumber="2" year="2003">
<bibRefCitation id="C54C4B67E107FFDCFD90F9BBCC86F9F7" author="Ilie" box="[574,731,1537,1560]" firstAuthor="Ilie" pageId="1" pageNumber="2" refString="Ilie, V. (2003 a) Chilopoda from the edaphic and subterranean environments in the Resita-Carasova area (Banat, Romania). A preliminary note. Archive Biological Sciences, Belgrade, 55 (3 - 4), 87 - 92." type="journal article" year="2003" yearSuffix="a">Ilie, 2003a: 89</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<bibRefCitation id="C54C4B67E107FFDCFD49F9BBCD3FF9F7" author="Ilie, V." box="[743,866,1537,1560]" pageId="1" pageNumber="2" pagination="129 - 137" refId="ref5245" refString="Ilie, V. (2003 b) Chilopoda species from the edaphic and subterranean environments of the Closani karstic area (Romania). Travaux du Museum National d'Histoire Naturelle &quot; Grigore Antipa &quot;, 45, 129 - 137." type="journal article" year="2003">2003b: 132</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A1623696E107FFDCFEA6F998CD6DF9D7" blockId="1.[264,1322,1249,1593]" box="[264,816,1569,1593]" pageId="1" pageNumber="2">
<taxonomicName id="66DD4D15E107FFDCFEA6F998CD71F9D7" ID-CoL="6LG8N" authority="Ilie et al., 2003: 93" authorityName="Ilie et al." authorityPageNumber="93" authorityYear="2003" box="[264,812,1569,1593]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E107FFDCFEA6F998CFF6F9D6" box="[264,427,1570,1593]" italics="true" pageId="1" pageNumber="2">Harpolithobius</emphasis>
cf.
<emphasis id="93A9EA84E107FFDCFE7AF998CC0FF9D6" box="[468,594,1570,1593]" italics="true" pageId="1" pageNumber="2">intermedius</emphasis>
:
<bibRefCitation id="C54C4B67E107FFDCFDCEF99BCD71F9D7" author="Ilie, V. &amp; Negrea, S. &amp; Mitic, B." box="[608,812,1569,1592]" pageId="1" pageNumber="2" pagination="93 - 99" refId="ref5287" refString="Ilie, V., Negrea, S. &amp; Mitic, B. (2003) The diversity of the Chilopoda species from the karstic area of the Anina Mountains (Banat, Romania). Archive Biological Sciences, Belgrade, 55 (3 - 4), 93 - 99." type="journal article" year="2003">Ilie et al., 2003: 93</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E9C7651DE107FFDEFEA6F9D3CB45F88A" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="A1623696E107FFDCFEA6F9D3CC69F914" blockId="1.[264,1323,1641,1867]" pageId="1" pageNumber="2">
Material examined: female
<typeStatus id="7E668834E107FFDCFDFFF9D3CCF1F96C" box="[593,684,1641,1667]" pageId="1" pageNumber="2" type="syntype">syntype</typeStatus>
mounted on a slide No. 249, Coll. Verhoeff, labelled “
<collectingCountry id="D9CA7606E107FFDCFEBCF92BCFCDF944" box="[274,400,1681,1707]" name="Romania" pageId="1" pageNumber="2">Rumänien</collectingCountry>
,
<collectionCode id="C7CCAE53E107FFDCFE32F92BCF81F944" box="[412,476,1681,1707]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="1" pageNumber="2">ZMB</collectionCode>
No 13 529”;
<typeStatus id="7E668834E107FFDCFDD4F92BCCB8F944" box="[634,741,1681,1707]" pageId="1" pageNumber="2" type="lectotype">lectotype</typeStatus>
by present designation. The fate of the other
<typeStatus id="7E668834E107FFDCFB56F92BCF1BF93C" pageId="1" pageNumber="2" type="syntype">syntypes</typeStatus>
is unknown, so herein I designate as
<typeStatus id="7E668834E107FFDCFCADF903CD33F93C" box="[771,878,1721,1747]" pageId="1" pageNumber="2" type="lectotype">lectotype</typeStatus>
the only available and comparatively well­preserved specimen.
</paragraph>
<paragraph id="A1623696E107FFDFFE96F8B3CAABFB46" blockId="1.[264,1323,1641,1867]" lastBlockId="2.[264,1325,284,1833]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<bibRefCitation id="C54C4B67E107FFDCFE96F8B3CFA9F8CC" author="Verhoeff" box="[312,500,1801,1827]" firstAuthor="Verhoeff" pageId="1" pageNumber="12" refString="Verhoeff, K. (1901) Chilopoden und Diplopoden aus Rumanien. In: Faune de la Roumanie par M. Jaquet. Bulletin de la Societe des Sciences, Bucarest, 10, 168 - 177." type="journal article" year="1901">Verhoeff (1901)</bibRefCitation>
described
<taxonomicName id="66DD4D15E107FFDCFDDDF8B3CD1FF8CD" box="[627,834,1801,1826]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="dollfusi">
<emphasis id="93A9EA84E107FFDCFDDDF8B3CD1FF8CD" box="[627,834,1801,1826]" italics="true" pageId="1" pageNumber="2">Lithobius dollfusi</emphasis>
</taxonomicName>
from Laculeţe and Azuga,
<collectingCountry id="D9CA7606E107FFDCFB2AF8B3CAACF8CC" box="[1156,1265,1801,1827]" name="Romania" pageId="1" pageNumber="2">Romania</collectingCountry>
. The taxonomic status of the species has never been re­considered, and subsequent researchers never found it.
<taxonomicName id="66DD4D15E104FFDFFE6FFEA6CC67FEDA" box="[449,570,284,309]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dollfusi">
<emphasis id="93A9EA84E104FFDFFE6FFEA6CC67FEDA" box="[449,570,284,309]" italics="true" pageId="2" pageNumber="3">L. dollfusi</emphasis>
</taxonomicName>
was even missed in
<bibRefCitation id="C54C4B67E104FFDFFC9BFEA6CA55FED9" author="Verhoeff, K." box="[821,1032,284,310]" pageId="2" pageNumber="3" pagination="171 - 257" refId="ref5936" refString="Verhoeff, K. (1937) Chilopoden-Studien. Zur Kenntnis der Lithobiiden. Archiv fur Naturgeschichte (Neue Folge), Berlin, 6, 171 - 257." type="journal article" year="1937">Verhoeffs (1937)</bibRefCitation>
sophisticated key to the species of the genus
<taxonomicName id="66DD4D15E104FFDFFE55FEFECC37FEB2" box="[507,618,324,349]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="93A9EA84E104FFDFFE55FEFECC37FEB2" box="[507,618,324,349]" italics="true" pageId="2" pageNumber="3">Lithobius</emphasis>
</taxonomicName>
, an argument used by
<bibRefCitation id="C54C4B67E104FFDFFCDDFEFECA5BFEB1" author="Matic" box="[883,1030,324,350]" firstAuthor="Matic" pageId="2" pageNumber="12" refString="Matic, Z. (1964) Nota critica asupra unor specii de Lithobiidae (Chilopoda) din fauna Republicii Populare Romine. Studii S i cercet a ri de Biologie, Seria Zoologie, Cluj, 16 (3), 187 - 191." type="journal article" year="1964">Matic (1964</bibRefCitation>
,
<bibRefCitation id="C54C4B67E104FFDFFBBCFEFECA13FEB1" author="Matic" box="[1042,1102,324,350]" firstAuthor="Matic" pageId="2" pageNumber="12" refString="Matic, Z. (1966) Classa Chilopoda. Subclassa Anamorpha. In: Fauna Republicii Socialiste Romania. Editura Academiei Republicii Socialiste Romania, Bucuresti, 6 (1), 1 - 272." type="journal article" year="1966">1966</bibRefCitation>
) to declare it, mistakenly, as a “nomen nudum”! The examined material, though broken into pieces and mounted on a permanent microscope slide, allowed observation of some important characters revealing its identity. Immediately evident is the fact that
<taxonomicName id="66DD4D15E104FFDFFC52FE06CA2BFE3A" box="[1020,1142,444,469]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dollfusi">
<emphasis id="93A9EA84E104FFDFFC52FE06CA2BFE3A" box="[1020,1142,444,469]" italics="true" pageId="2" pageNumber="3">L. dollfusi</emphasis>
</taxonomicName>
belongs to the genus
<taxonomicName id="66DD4D15E104FFDFFEFAFE5ECC9DFE11" authority="Verhoeff, 1904" authorityName="Verhoeff" authorityYear="1904" box="[340,704,484,510]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="93A9EA84E104FFDFFEFAFE5ECC5AFE12" box="[340,519,484,509]" italics="true" pageId="2" pageNumber="3">Harpolithobius</emphasis>
Verhoeff, 1904
</taxonomicName>
, which at the time of the original species description had not yet been established. The genus
<taxonomicName id="66DD4D15E104FFDFFD5CFDB6CDF8FDCA" box="[754,933,524,549]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="93A9EA84E104FFDFFD5CFDB6CDF8FDCA" box="[754,933,524,549]" italics="true" pageId="2" pageNumber="3">Harpolithobius</emphasis>
</taxonomicName>
is characterized by a number of synapomorphies, e.g. prosternal edge not incised medially, porodonts always thickened; male posterior legs with modifications, etc., and is hitherto known to comprise more than 30 species and subspecies. The genus range includes
<collectingCountry id="D9CA7606E104FFDFFC29FD3ECA4CFD71" box="[903,1041,644,670]" name="Turkey" pageId="2" pageNumber="3">Asia Minor</collectingCountry>
, the Caucasus, the Balkans, the Carpathians, and the Alps, with one species,
<taxonomicName id="66DD4D15E104FFDFFC23FD16CAE4FD29" authority="Latzel, 1880" authorityName="Latzel" authorityYear="1880" box="[909,1209,684,710]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="anodus">
<emphasis id="93A9EA84E104FFDFFC23FD16CA5AFD2A" box="[909,1031,684,709]" italics="true" pageId="2" pageNumber="3">H. anodus</emphasis>
(Latzel, 1880)
</taxonomicName>
, reaching the Ligurian Apennines to the west (
<bibRefCitation id="C54C4B67E104FFDFFD1BFD6ECD2AFD01" author="Zapparoli" box="[693,887,724,750]" firstAuthor="Zapparoli" pageId="2" pageNumber="12" refString="Zapparoli, M. (2003) The present knowledge on the European fauna of Lithobiomorpha (Chilopoda). Bulletin of the British Myriapod and Isopod Group, 19, 20 - 41." type="journal article" year="2003">Zapparoli, 2003</bibRefCitation>
). It has never been an object of comprehensive revision, and many taxa, especially from the Balkans and
<collectingCountry id="D9CA7606E104FFDFFBCCFD46CAB2FCF9" box="[1122,1263,764,790]" name="Turkey" pageId="2" pageNumber="3">Asia Minor</collectingCountry>
, still await a proper re­description. So far, eight species are known from
<collectingCountry id="D9CA7606E104FFDFFBF9FC9ECA99FCD1" box="[1111,1220,804,830]" name="Romania" pageId="2" pageNumber="3">Romania</collectingCountry>
, four of which occur exclusively within the country boundaries (cf.
<bibRefCitation id="C54C4B67E104FFDFFC60FCF6CA3CFC89" author="Matic" box="[974,1121,844,870]" firstAuthor="Matic" pageId="2" pageNumber="12" refString="Matic, Z. (1966) Classa Chilopoda. Subclassa Anamorpha. In: Fauna Republicii Socialiste Romania. Editura Academiei Republicii Socialiste Romania, Bucuresti, 6 (1), 1 - 272." type="journal article" year="1966">Matic, 1966</bibRefCitation>
). One of these is
<taxonomicName id="66DD4D15E104FFDFFEA6FCCECC11FC62" box="[264,588,884,909]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E104FFDFFEA6FCCECC11FC62" box="[264,588,884,909]" italics="true" pageId="2" pageNumber="3">Harpolithobius intermedius</emphasis>
</taxonomicName>
(including its junior synonym
<taxonomicName id="66DD4D15E104FFDFFC1BFCCECB7CFC62" box="[949,1313,884,909]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="intermedius" subSpecies="transsylvanicus">
<emphasis id="93A9EA84E104FFDFFC1BFCCECB7CFC62" box="[949,1313,884,909]" italics="true" pageId="2" pageNumber="3">H. intermedius transsylvanicus</emphasis>
</taxonomicName>
) described from Valea Ord ă ncuŞei (Ord ă ncuŞei Valley), the Apuseni Mts. and Torda (= Turzii) Gorge,
<collectingCountry id="D9CA7606E104FFDFFE14FC7DCC79FC0E" box="[442,548,967,993]" name="Romania" pageId="2" pageNumber="3">Romania</collectingCountry>
(
<bibRefCitation id="C54C4B67E104FFDFFD9BFC7DCC97FC0E" author="Matic" box="[565,714,967,993]" firstAuthor="Matic" pageId="2" pageNumber="12" refString="Matic, Z. (1958) Zur Kenntnis der Gattung Harpolithobius in Siebenburgen (Chilopoda, Lithobiidae). Senckenbergiana biologica, 39 (1 - 2), 91 - 95." type="journal article" year="1958">Matic, 1958</bibRefCitation>
). It was downgraded to a subspecies of
<taxonomicName id="66DD4D15E104FFDFFB0BFC7DCFB5FBE6" authority="Folkmanova, 1958" authorityName="Folkmanova" authorityYear="1958" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="93A9EA84E104FFDFFB0BFC7DCB77FC0F" box="[1189,1322,967,992]" italics="true" pageId="2" pageNumber="3">H. spinipes</emphasis>
Folkmanova, 1958
</taxonomicName>
by
<bibRefCitation id="C54C4B67E104FFDFFDB2FC55CCAAFBE6" author="Dobroruka" box="[540,759,1007,1033]" firstAuthor="Dobroruka" pageId="2" pageNumber="11" refString="Dobroruka, L. (1960) Bemerkungen zur Gattung Harpolithobius (Chilopoda). Zoologischer Anzeiger, 164 (5 - 6), 198 - 201." type="journal article" year="1960">Dobroruka (1960)</bibRefCitation>
, but
<bibRefCitation id="C54C4B67E104FFDFFC96FC55CDBEFBE6" author="Matic" box="[824,995,1007,1033]" firstAuthor="Matic" pageId="2" pageNumber="12" refString="Matic, Z. (1961 a) Nota critica asupra speciilor Harpolithobius spinipes Folkm. 1958 si Harpolithobius intermedius Matic 1958 (Chilopoda-Lithobiidae). Studia Universitatis Babe S - Bolyai, Cluj, Series II, fasc. 2, 166 - 169." type="book chapter" year="1961" yearSuffix="a">Matic (1961a)</bibRefCitation>
resurrected its full species rank and added new data on its morphology (
<bibRefCitation id="C54C4B67E104FFDFFC88FBADCD97FBDE" author="Matic" box="[806,970,1047,1073]" firstAuthor="Matic" pageId="2" pageNumber="12" refString="Matic, Z. (1961 b) Genul Harpolithobius (Chilopoda, Lithobiidae) in fauna Republicii Populare Romine. Studii S i cercet a ri de Biologie, Cluj, 12 (1), 73 - 89." type="journal article" year="1961" yearSuffix="b">Matic, 1961b</bibRefCitation>
). According to
<bibRefCitation id="C54C4B67E104FFDFFB2AFBADCB70FBDE" author="Matic" box="[1156,1325,1047,1073]" firstAuthor="Matic" pageId="2" pageNumber="12" refString="Matic, Z. (1961 a) Nota critica asupra speciilor Harpolithobius spinipes Folkm. 1958 si Harpolithobius intermedius Matic 1958 (Chilopoda-Lithobiidae). Studia Universitatis Babe S - Bolyai, Cluj, Series II, fasc. 2, 166 - 169." type="book chapter" year="1961" yearSuffix="a">Matic (1961a)</bibRefCitation>
<taxonomicName id="66DD4D15E104FFDFFEA6FB85CFD0FBB7" box="[264,397,1087,1112]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="93A9EA84E104FFDFFEA6FB85CFD0FBB7" box="[264,397,1087,1112]" italics="true" pageId="2" pageNumber="3">H. spinipes</emphasis>
</taxonomicName>
is well distinguished from
<taxonomicName id="66DD4D15E104FFDFFD7BFB85CDD9FBB7" box="[725,900,1087,1112]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E104FFDFFD7BFB85CDD9FBB7" box="[725,900,1087,1112]" italics="true" pageId="2" pageNumber="3">H. intermedius</emphasis>
</taxonomicName>
by the tripartite female gonopodial claw (vs. single), shorter and thicker gonopodial spurs, first leg­pair richer in ventral spines and having unmodified tibiae (vs. single ventral spine and enlarged tibiae), etc.
</paragraph>
<paragraph id="A1623696E104FFDFFE96FB0DCC88F936" blockId="2.[264,1325,284,1833]" pageId="2" pageNumber="3">
Recently,
<taxonomicName id="66DD4D15E104FFDFFE00FB0DCC03FB3F" box="[430,606,1207,1232]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E104FFDFFE00FB0DCC03FB3F" box="[430,606,1207,1232]" italics="true" pageId="2" pageNumber="3">H. intermedius</emphasis>
</taxonomicName>
was recorded also from the Anina Mts., Banat (
<bibRefCitation id="C54C4B67E104FFDFFB0DFB0DCB7BFB3E" author="Ilie" box="[1187,1318,1207,1233]" firstAuthor="Ilie" pageId="2" pageNumber="12" refString="Ilie, V. (2003 a) Chilopoda from the edaphic and subterranean environments in the Resita-Carasova area (Banat, Romania). A preliminary note. Archive Biological Sciences, Belgrade, 55 (3 - 4), 87 - 92." type="journal article" year="2003" yearSuffix="a">Ilie, 2003a</bibRefCitation>
,
<bibRefCitation id="C54C4B67E104FFDFFEA6FB65CFE1FB16" author="Ilie, V. &amp; Negrea, S. &amp; Mitic, B." box="[264,444,1247,1273]" pageId="2" pageNumber="3" pagination="93 - 99" refId="ref5287" refString="Ilie, V., Negrea, S. &amp; Mitic, B. (2003) The diversity of the Chilopoda species from the karstic area of the Anina Mountains (Banat, Romania). Archive Biological Sciences, Belgrade, 55 (3 - 4), 93 - 99." type="journal article" year="2003">Ilie et al., 2003</bibRefCitation>
) and the region of CloŞani, Oltenia (
<bibRefCitation id="C54C4B67E104FFDFFCC3FB65CDAFFB16" author="Ilie" box="[877,1010,1247,1273]" firstAuthor="Ilie" pageId="2" pageNumber="12" refString="Ilie, V. (2003 b) Chilopoda species from the edaphic and subterranean environments of the Closani karstic area (Romania). Travaux du Museum National d'Histoire Naturelle &quot; Grigore Antipa &quot;, 45, 129 - 137." type="journal article" year="2003" yearSuffix="b">Ilie, 2003b</bibRefCitation>
). Stoev (in Deltshev et al., 2000) mentioned it for the Central Stara Planina Mts.,
<collectingCountry id="D9CA7606E104FFDFFC23FABDCDAFFACE" box="[909,1010,1287,1313]" name="Bulgaria" pageId="2" pageNumber="3">Bulgaria</collectingCountry>
(record repeated in
<bibRefCitation id="C54C4B67E104FFDFFB4EFABDCF19FAA6" author="Stoev" firstAuthor="Stoev" pageId="2" pageNumber="12" refString="Stoev, P. (2002) A Catalogue and Key to the centipedes (Chilopoda) of Bulgaria. Pensoft Publishers, Sofia-Moscow, 103 pp." type="book" year="2002">Stoev, 2002</bibRefCitation>
) without confidence about its identification.
<taxonomicName id="66DD4D15E104FFDFFCF3FA95CA50FAA7" box="[861,1037,1327,1352]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E104FFDFFCF3FA95CA50FAA7" box="[861,1037,1327,1352]" italics="true" pageId="2" pageNumber="3">H. intermedius</emphasis>
</taxonomicName>
was distinguished from the other congeners by a number of characters, the most striking being the presence of a single female gonopodial claw. Though apparent, Matic failed to mention its close resemblance to
<taxonomicName id="66DD4D15E104FFDFFED4FA1DCFADFA2F" box="[378,496,1447,1472]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dollfusi">
<emphasis id="93A9EA84E104FFDFFED4FA1DCFADFA2F" box="[378,496,1447,1472]" italics="true" pageId="2" pageNumber="3">L. dollfusi</emphasis>
</taxonomicName>
. A direct comparison between the two species, which is based on both the literature (cf.
<bibRefCitation id="C54C4B67E104FFDFFE76FA75CCD1FA06" author="Verhoeff" box="[472,652,1487,1513]" firstAuthor="Verhoeff" pageId="2" pageNumber="12" refString="Verhoeff, K. (1901) Chilopoden und Diplopoden aus Rumanien. In: Faune de la Roumanie par M. Jaquet. Bulletin de la Societe des Sciences, Bucarest, 10, 168 - 177." type="journal article" year="1901">Verhoeff, 1901</bibRefCitation>
;
<bibRefCitation id="C54C4B67E104FFDFFD34FA75CD76FA06" author="Matic" box="[666,811,1487,1513]" firstAuthor="Matic" pageId="2" pageNumber="12" refString="Matic, Z. (1958) Zur Kenntnis der Gattung Harpolithobius in Siebenburgen (Chilopoda, Lithobiidae). Senckenbergiana biologica, 39 (1 - 2), 91 - 95." type="journal article" year="1958">Matic, 1958</bibRefCitation>
,
<bibRefCitation id="C54C4B67E104FFDFFC97FA75CDD8FA06" author="Matic" box="[825,901,1487,1513]" firstAuthor="Matic" pageId="2" pageNumber="12" refString="Matic, Z. (1961 a) Nota critica asupra speciilor Harpolithobius spinipes Folkm. 1958 si Harpolithobius intermedius Matic 1958 (Chilopoda-Lithobiidae). Studia Universitatis Babe S - Bolyai, Cluj, Series II, fasc. 2, 166 - 169." type="book chapter" year="1961" yearSuffix="a">1961a</bibRefCitation>
,
<bibRefCitation id="C54C4B67E104FFDFFC24FA75CDC7FA06" author="Matic" box="[906,922,1487,1513]" pageId="2" pageNumber="12" refString="Matic, Z. (1961 b) Genul Harpolithobius (Chilopoda, Lithobiidae) in fauna Republicii Populare Romine. Studii S i cercet a ri de Biologie, Cluj, 12 (1), 73 - 89." type="journal article" year="1961" yearSuffix="b">b</bibRefCitation>
,
<bibRefCitation id="C54C4B67E104FFDFFC09FA75CDBEFA06" author="Matic" box="[935,995,1487,1513]" firstAuthor="Matic" pageId="2" pageNumber="12" refString="Matic, Z. (1966) Classa Chilopoda. Subclassa Anamorpha. In: Fauna Republicii Socialiste Romania. Editura Academiei Republicii Socialiste Romania, Bucuresti, 6 (1), 1 - 272." type="journal article" year="1966">1966</bibRefCitation>
) and original data from the
<typeStatus id="7E668834E104FFDFFEA6FA4DCF2BF9FE" box="[264,374,1527,1553]" pageId="2" pageNumber="3" type="lectotype">lectotype</typeStatus>
, is given below in
<tableCitation id="EC5F032DE104FFDFFDF8FA4DCCF2F9FE" box="[598,687,1527,1553]" captionStart="TABLE 1. A" captionStartId="3.[264,352,1319,1343]" captionTargetBox="[277,1249,1381,1753]" captionTargetPageId="3" captionText="TABLE 1. A morphological comparison between H. dollfusi and H. intermedius." httpUri="http://table.plazi.org/id/F5A2661EE105FFDEFEA6FA9DCA37FAAF" pageId="2" pageNumber="3" tableUuid="F5A2661EE105FFDEFEA6FA9DCA37FAAF">Table 1</tableCitation>
. Data deriving from the personal examination of Victoria Ilie (in litt.) of both the
<typeStatus id="7E668834E104FFDFFDCEF9A5CCC3F9D6" box="[608,670,1567,1593]" pageId="2" pageNumber="3">types</typeStatus>
of
<taxonomicName id="66DD4D15E104FFDFFD69F9A5CD28F9D7" box="[711,885,1567,1592]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E104FFDFFD69F9A5CD28F9D7" box="[711,885,1567,1592]" italics="true" pageId="2" pageNumber="3">H. intermedius</emphasis>
</taxonomicName>
in the Zoological Museum, Cluj and his own material were also taken into account. The table shows that the main taxonomic characters of the two species overlap considerably, which is a reason to propose the following new synonymy and combination:
<taxonomicName id="66DD4D15E104FFDFFD59F92DCA87F95E" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" box="[759,1242,1687,1713]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dollfusi" status="comb. nov.">
<emphasis id="93A9EA84E104FFDFFD59F92DCA56F95F" box="[759,1035,1687,1712]" italics="true" pageId="2" pageNumber="3">Harpolithobius dollfusi</emphasis>
(
<bibRefCitation id="C54C4B67E104FFDFFBB2F92DCA8CF95E" author="Verhoeff" box="[1052,1233,1687,1713]" firstAuthor="Verhoeff" pageId="2" pageNumber="12" refString="Verhoeff, K. (1901) Chilopoden und Diplopoden aus Rumanien. In: Faune de la Roumanie par M. Jaquet. Bulletin de la Societe des Sciences, Bucarest, 10, 168 - 177." type="journal article" year="1901">Verhoeff, 1901</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="889A57FFE104FFDFFB4CF92DCF43F936" pageId="2" pageNumber="3">comb. n.</taxonomicNameLabel>
=
<taxonomicName id="66DD4D15E104FFDFFE93F905CCDCF936" authority="Matic, 1958" authorityName="Matic" authorityYear="1958" box="[317,641,1727,1753]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E104FFDFFE93F905CFB6F937" box="[317,491,1727,1752]" italics="true" pageId="2" pageNumber="3">H. intermedius</emphasis>
<bibRefCitation id="C54C4B67E104FFDFFE5DF905CCDCF936" author="Matic" box="[499,641,1727,1753]" firstAuthor="Matic" pageId="2" pageNumber="12" refString="Matic, Z. (1958) Zur Kenntnis der Gattung Harpolithobius in Siebenburgen (Chilopoda, Lithobiidae). Senckenbergiana biologica, 39 (1 - 2), 91 - 95." type="journal article" year="1958">Matic, 1958</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="889A57FFE104FFDFFD26F905CC88F936" box="[648,725,1727,1753]" pageId="2" pageNumber="3">syn. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="A1623696E104FFDFFE96F95DCCC9F8C6" blockId="2.[264,1325,284,1833]" pageId="2" pageNumber="3">
General distribution.
<collectingCountry id="D9CA7606E104FFDFFD9CF95DCCFCF8EE" box="[562,673,1767,1793]" name="Romania" pageId="2" pageNumber="3">Romania</collectingCountry>
: Apuseni Mts., Torda Gorge, Anina Mts., CloŞani;
<collectingCountry id="D9CA7606E104FFDFFB58F95DCF14F8C6" name="Bulgaria" pageId="2" pageNumber="3">Bulgaria</collectingCountry>
?: Central Stara Planina Mts.
</paragraph>
<caption id="F5A2661EE105FFDEFEA6FBD7CC6FFB25" httpUri="https://zenodo.org/record/170549/files/figure.png" pageId="3" pageNumber="4" targetBox="[351,1217,299,1101]" targetPageId="3">
<paragraph id="A1623696E105FFDEFEA6FBD7CC6FFB25" blockId="3.[264,1323,1133,1226]" pageId="3" pageNumber="4">
<emphasis id="93A9EA84E105FFDEFEA6FBD7CFE8FB6A" bold="true" box="[264,437,1133,1157]" pageId="3" pageNumber="4">FIGURES 14.</emphasis>
<taxonomicName id="66DD4D15E105FFDEFE13FBD4CD2AFB6B" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" box="[445,887,1133,1157]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dollfusi">
<emphasis id="93A9EA84E105FFDEFE13FBD4CCE4FB6A" box="[445,697,1134,1157]" italics="true" pageId="3" pageNumber="4">Harpolithobius dollfusi</emphasis>
(
<bibRefCitation id="C54C4B67E105FFDEFD69FBD7CD32FB6B" author="Verhoeff, K." box="[711,879,1133,1156]" pageId="3" pageNumber="4" pagination="168 - 177" refId="ref5897" refString="Verhoeff, K. (1901) Chilopoden und Diplopoden aus Rumanien. In: Faune de la Roumanie par M. Jaquet. Bulletin de la Societe des Sciences, Bucarest, 10, 168 - 177." type="journal article" year="1901">Verhoeff, 1901</bibRefCitation>
)
</taxonomicName>
: 1 head, dorsal view; 2 maxillipede, ventral view; 3 coxa, ventro­lateral view; 4 female gonopods, ventral view. Scale bars: 1 mm (Figs 12), 0.5 mm (Fig. 4).
</paragraph>
</caption>
<caption id="F5A2661EE105FFDEFEA6FA9DCA37FAAF" ID-Table-UUID="F5A2661EE105FFDEFEA6FA9DCA37FAAF" box="[264,1130,1318,1344]" httpUri="http://table.plazi.org/id/F5A2661EE105FFDEFEA6FA9DCA37FAAF" pageId="3" pageNumber="4" targetBox="[277,1249,1381,1753]" targetIsTable="true" targetPageId="3">
<paragraph id="A1623696E105FFDEFEA6FA9DCA37FAAF" blockId="3.[264,1130,1318,1344]" box="[264,1130,1318,1344]" pageId="3" pageNumber="4">
<emphasis id="93A9EA84E105FFDEFEA6FA9DCF26FAD0" bold="true" box="[264,379,1319,1343]" pageId="3" pageNumber="4">TABLE 1.</emphasis>
A morphological comparison between
<taxonomicName id="66DD4D15E105FFDEFC8EFA92CDCCFAD0" box="[800,913,1320,1343]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dollfusi">
<emphasis id="93A9EA84E105FFDEFC8EFA92CDCCFAD0" box="[800,913,1320,1343]" italics="true" pageId="3" pageNumber="4">H. dollfusi</emphasis>
</taxonomicName>
and
<taxonomicName id="66DD4D15E105FFDEFC6BFA92CA3EFAD0" box="[965,1123,1320,1343]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E105FFDEFC6BFA92CA3EFAD0" box="[965,1123,1320,1343]" italics="true" pageId="3" pageNumber="4">H. intermedius</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="A1623696E105FFDEFE0CFADFCAFDF936" pageId="3" pageNumber="4">
<table id="D3DDC436E1050022FEBBFADFCABCF936" box="[277,1249,1381,1753]" gridcols="3" gridrows="9" pageId="3" pageNumber="4">
<tr id="1FED34D4E1050022FEBBFADFCABCFA95" box="[277,1249,1381,1402]" gridrow="0" pageId="3" pageNumber="4">
<th id="5C3C5DA8E1050022FEBBFADFCC3FFA95" box="[277,610,1381,1402]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="93A9EA84E105FFDEFE0CFADFCC48FA95" bold="true" box="[418,533,1381,1402]" pageId="3" pageNumber="4">Characters</emphasis>
</th>
<th id="5C3C5DA8E1050022FD75FADFCDFEFA95" box="[731,931,1381,1402]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="66DD4D15E105FFDEFCA5FADFCD29FA95" box="[779,884,1381,1402]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dollfusi">
<emphasis id="93A9EA84E105FFDEFCA5FADFCD29FA95" bold="true" box="[779,884,1381,1402]" italics="true" pageId="3" pageNumber="4">H. dollfusi</emphasis>
</taxonomicName>
</th>
<th id="5C3C5DA8E1050022FBB4FADFCABCFA95" box="[1050,1249,1381,1402]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="66DD4D15E105FFDEFB9AFADFCA9AFA95" box="[1076,1223,1381,1402]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="93A9EA84E105FFDEFB9AFADFCA9AFA95" bold="true" box="[1076,1223,1381,1402]" italics="true" pageId="3" pageNumber="4">H. intermedius</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="1FED34D4E1050022FEBBFA2ACABCFA4A" box="[277,1249,1424,1445]" gridrow="1" pageId="3" pageNumber="4">
<th id="5C3C5DA8E1050022FEBBFA2ACC3FFA4A" box="[277,610,1424,1445]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">Body size</th>
<td id="5C3C5DA8E1050022FD75FA2ACDFEFA4A" box="[731,931,1424,1445]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">16 mm</td>
<td id="5C3C5DA8E1050022FBB4FA2ACABCFA4A" box="[1050,1249,1424,1445]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">1821 mm</td>
</tr>
<tr id="1FED34D4E1050022FEBBFA01CABCFA3F" box="[277,1249,1467,1488]" gridrow="2" pageId="3" pageNumber="4">
<th id="5C3C5DA8E1050022FEBBFA01CC3FFA3F" box="[277,610,1467,1488]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">Number of antennal articles</th>
<td id="5C3C5DA8E1050022FD75FA01CDFEFA3F" box="[731,931,1467,1488]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">4548</td>
<td id="5C3C5DA8E1050022FBB4FA01CABCFA3F" box="[1050,1249,1467,1488]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">(33) 4449 (57)</td>
</tr>
<tr id="1FED34D4E1050022FEBBFA5FCABCFA15" box="[277,1249,1509,1530]" gridrow="3" pageId="3" pageNumber="4">
<th id="5C3C5DA8E1050022FEBBFA5FCC3FFA15" box="[277,610,1509,1530]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">Number of ocelli</th>
<td id="5C3C5DA8E1050022FD75FA5FCDFEFA15" box="[731,931,1509,1530]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">1517</td>
<td id="5C3C5DA8E1050022FBB4FA5FCABCFA15" box="[1050,1249,1509,1530]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">(14) 1721</td>
</tr>
<tr id="1FED34D4E1050022FEBBF9B4CABCF9C5" box="[277,1249,1550,1578]" gridrow="4" pageId="3" pageNumber="4">
<th id="5C3C5DA8E1050022FEBBF9B4CC3FF9C5" box="[277,610,1550,1578]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">Teeth on maxillipede coxosternum</th>
<td id="5C3C5DA8E1050022FD75F9B4CDFEF9C5" box="[731,931,1550,1578]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">2+2a</td>
<td id="5C3C5DA8E1050022FBB4F9B4CABCF9C5" box="[1050,1249,1550,1578]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">2+2</td>
</tr>
<tr id="1FED34D4E1050022FEBBF985CABCF9BB" box="[277,1249,1599,1620]" gridrow="5" pageId="3" pageNumber="4">
<th id="5C3C5DA8E1050022FEBBF985CC3FF9BB" box="[277,610,1599,1620]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">Coxal pores</th>
<td id="5C3C5DA8E1050022FD75F985CDFEF9BB" box="[731,931,1599,1620]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">6, 6, 7, 6</td>
<td id="5C3C5DA8E1050022FBB4F985CABCF9BB" box="[1050,1249,1599,1620]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">6, 6, 7, 5 (6)</td>
</tr>
<tr id="1FED34D4E1050022FEBBF9D3CABCF96B" box="[277,1249,1641,1668]" gridrow="6" pageId="3" pageNumber="4">
<th id="5C3C5DA8E1050022FEBBF9D3CC3FF96B" box="[277,610,1641,1668]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">Ventral spinulation of 15th leg­pair</th>
<td id="5C3C5DA8E1050022FD75F9D3CDFEF96B" box="[731,931,1641,1668]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">0, 1, 3, 3, 1</td>
<td id="5C3C5DA8E1050022FBB4F9D3CABCF96B" box="[1050,1249,1641,1668]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">0, 1, 3, 3, 1</td>
</tr>
<tr id="1FED34D4E1050022FEBBF920CABCF940" box="[277,1249,1690,1711]" gridrow="7" pageId="3" pageNumber="4">
<th id="5C3C5DA8E1050022FEBBF920CC3FF940" box="[277,610,1690,1711]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">Female gonopodial spurs</th>
<td id="5C3C5DA8E1050022FD75F920CDFEF940" box="[731,931,1690,1711]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">2+2 moderately long</td>
<td id="5C3C5DA8E1050022FBB4F920CABCF940" box="[1050,1249,1690,1711]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">2+2 moderately long</td>
</tr>
<tr id="1FED34D4E1050022FEBBF97ECABCF936" box="[277,1249,1732,1753]" gridrow="8" pageId="3" pageNumber="4">
<th id="5C3C5DA8E1050022FEBBF97ECC3FF936" box="[277,610,1732,1753]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">Female gonopodial claw</th>
<td id="5C3C5DA8E1050022FD75F97ECDFEF936" box="[731,931,1732,1753]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">Simple</td>
<td id="5C3C5DA8E1050022FBB4F97ECABCF936" box="[1050,1249,1732,1753]" gridcol="2" gridrow="8" pageId="3" pageNumber="4">Simple</td>
</tr>
</table>
</paragraph>
<paragraph id="A1623696E105FFDEFEA6F8BBCB45F88A" blockId="3.[264,1324,1793,1893]" pageId="3" pageNumber="4">
a There are not prosternal teeth in the examined
<typeStatus id="7E668834E105FFDEFCA5F8B2CD03F8F0" box="[779,862,1800,1823]" pageId="3" pageNumber="4" type="syntype">syntype</typeStatus>
although in the original description
<bibRefCitation id="C54C4B67E105FFDEFB4DF8B2CF29F8AD" author="Verhoeff" firstAuthor="Verhoeff" pageId="3" pageNumber="12" refString="Verhoeff, K. (1901) Chilopoden und Diplopoden aus Rumanien. In: Faune de la Roumanie par M. Jaquet. Bulletin de la Societe des Sciences, Bucarest, 10, 168 - 177." type="journal article" year="1901">Verhoeff (1901)</bibRefCitation>
reported 2+2 teeth as characterizing the species. It could be due to an individual variation, which is known also in other congeners, namely
<taxonomicName id="66DD4D15E105FFDEFCE9F8F4CA0FF88A" authority="Latzel, 1882" authorityName="Latzel" authorityYear="1882" box="[839,1106,1870,1893]" class="Chilopoda" family="Lithobiidae" genus="Harpolithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="anodus">
<emphasis id="93A9EA84E105FFDEFCE9F8F4CDE8F88A" box="[839,949,1870,1893]" italics="true" pageId="3" pageNumber="4">H. anodus</emphasis>
(Latzel, 1882)
</taxonomicName>
(cf.
<bibRefCitation id="C54C4B67E105FFDEFB2AF8F4CB50F88A" author="Eason" box="[1156,1293,1870,1893]" firstAuthor="Eason" pageId="3" pageNumber="11" refString="Eason, E. H. (1982) A review of the north-west European species of Lithobiomorpha with a revised key to their identification. Zoological Journal of the Linnean Society, 74, 9 - 33." type="journal article" year="1982">Eason, 1982</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>