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<document id="B101759EFAF3367C60C6C0DEE05E309A" ID-DOI="10.5281/zenodo.185971" ID-GBIF-Dataset="1db39382-b4f0-46fa-ae08-dd5e57e0e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="185971" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460034098030" checkinUser="plazi" docAuthor="Pimvichai, Piyatida, Enghoff, Henrik &amp; Panha, Somsak" docDate="2009" docId="1644D538F448FFCEFF49FA239B93F8B3" docLanguage="en" docName="zt02016p050.pdf" docOrigin="Zootaxa 2016" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Thyropygus opinatus Karsch 1881, n. comb." docType="treatment" docVersion="10" lastPageNumber="29" masterDocId="EA7DAD40F442FFC2FFDEFFED9F66FFCD" masterDocTitle="A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)" masterLastPageNumber="50" masterPageNumber="17" pageNumber="27" updateTime="1698234296060" updateUser="plazi">
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<mods:title id="B39A71C24C6AAA5058A4D30C422B8E4B">A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)</mods:title>
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<mods:namePart id="66F643781697A2F2EB122F3EFBF53C5A">Pimvichai, Piyatida</mods:namePart>
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<mods:namePart id="4F5A31D967D176E3BC79788461CC962D">Enghoff, Henrik</mods:namePart>
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<mods:namePart id="8A3D2EEFCC2862B6F8F34A29CB9B661C">Panha, Somsak</mods:namePart>
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<mods:date id="3B92CE554985E517E8E2850347F4688D">2009</mods:date>
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<treatment id="1644D538F448FFCEFF49FA239B93F8B3" ID-DOI="http://doi.org/10.5281/zenodo.3509579" ID-GBIF-Taxon="119382579" ID-Zenodo-Dep="3509579" LSID="urn:lsid:plazi:treatment:1644D538F448FFCEFF49FA239B93F8B3" httpUri="http://treatment.plazi.org/id/1644D538F448FFCEFF49FA239B93F8B3" lastPageId="12" lastPageNumber="29" pageId="10" pageNumber="27">
<subSubSection id="D6F737A5F448FFC8FF49FA239E0BF9C6" pageId="10" pageNumber="27" type="nomenclature">
<paragraph id="9E52642EF448FFC8FF49FA239DA8FA25" blockId="10.[151,718,1486,1547]" box="[151,718,1486,1512]" pageId="10" pageNumber="27">
<heading id="C51AD342F448FFC8FF49FA239DA8FA25" bold="true" box="[151,718,1486,1512]" fontSize="11" level="1" pageId="10" pageNumber="27" reason="1">
<emphasis id="AC99B83CF448FFC8FF49FA239DA8FA25" bold="true" box="[151,718,1486,1512]" pageId="10" pageNumber="27">
<taxonomicName id="59ED1FADF448FFC8FF49FA239D33FA25" ID-CoL="56T3L" authority="Karsch, 1881" authorityName="Karsch" authorityYear="1881" box="[151,597,1486,1512]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="opinatus" status="comb. nov.">
<emphasis id="AC99B83CF448FFC8FF49FA239EF3FA25" bold="true" box="[151,405,1486,1512]" italics="true" pageId="10" pageNumber="27">Thyropygus opinatus</emphasis>
(
<bibRefCitation id="FA7C19DFF448FFC8FE7BFA239D2BFA25" author="Karsch" box="[421,589,1486,1512]" pageId="10" pageNumber="50" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die Gesamten Naturwissenschaften, Dritte Folge, 6, 3 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="B7AA0547F448FFC8FDBDFA239DA8FA25" box="[611,718,1486,1512]" pageId="10" pageNumber="27">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9E52642EF448FFC8FF49FA1C9E0BF9C6" blockId="10.[151,718,1486,1547]" box="[151,365,1521,1547]" pageId="10" pageNumber="27">
(
<figureCitation id="06D678ABF448FFC8FF41FA1C9F96F9C6" box="[159,240,1521,1547]" captionStart="FIGURE 6" captionStartId="11.[151,257,1658,1682]" captionTargetBox="[172,1391,359,1628]" captionTargetId="figure@11.[151,1436,349,1634]" captionTargetPageId="11" captionText="FIGURE 6. (A, D, E, F): Thyropygus opinatus, gonopods. A: anterior view, left telopodite removed (specimen from Malewoon). B: anterior view, anterior coxal fold (specimen from Nakwang cave). C: anterior view, anterior coxal fold (specimen from Ban Krude). D: right coxa, posterior view. E: right telopodite, anterior view. F: right telopodite, posterior view." httpUri="https://zenodo.org/record/185977/files/figure.png" pageId="10" pageNumber="27">Figs. 6</figureCitation>
AF, 18A)
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F448FFC8FF49F9D29D0CF918" pageId="10" pageNumber="27" type="reference_group">
<paragraph id="9E52642EF448FFC8FF49F9D29D25F998" blockId="10.[151,836,1598,1750]" box="[151,579,1598,1622]" pageId="10" pageNumber="27">
<treatmentCitationGroup id="BEFD4300F448FFC8FF49F9D29D25F998" box="[151,579,1598,1622]" pageId="10" pageNumber="27">
<taxonomicName id="59ED1FADF448FFC8FF49F9D29D25F998" authority="Karsch, 1881: 23" authorityName="Karsch" authorityPageNumber="23" authorityYear="1881" box="[151,579,1598,1622]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF448FFC8FF49F9D29EE0F99B" box="[151,390,1599,1622]" italics="true" pageId="10" pageNumber="27">Spirostreptus opinatus</emphasis>
<treatmentCitation id="1F4C423FF448FFC8FE52F9D39D25F998" author="Karsch" box="[396,579,1598,1621]" page="23" pageId="10" pageNumber="27" year="1881">
<bibRefCitation id="FA7C19DFF448FFC8FE52F9D39D78F998" author="Karsch" box="[396,542,1598,1621]" pageId="10" pageNumber="50" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die Gesamten Naturwissenschaften, Dritte Folge, 6, 3 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
: 23
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="9E52642EF448FFC8FF49F9B29D36F958" blockId="10.[151,836,1598,1750]" pageId="10" pageNumber="27">
<treatmentCitationGroup id="BEFD4300F448FFC8FF49F9B29D36F958" pageId="10" pageNumber="27">
<taxonomicName id="59ED1FADF448FFC8FF49F9B29DABF9B8" authority="Pocock 1889: 294" authorityName="Pocock" authorityPageNumber="294" authorityYear="1889" box="[151,717,1630,1654]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF448FFC8FF49F9B29EE0F9BB" box="[151,390,1631,1654]" italics="true" pageId="10" pageNumber="27">Spirostreptus opinatus</emphasis>
:
<treatmentCitation id="1F4C423FF448FFC8FE4AF9B39D30F9B8" author="Pocock" box="[404,598,1630,1653]" page="294" pageId="10" pageNumber="27" year="1889">
<bibRefCitation id="FA7C19DFF448FFC8FE4AF9B39D44F9B8" author="Pocock" box="[404,546,1630,1653]" pageId="10" pageNumber="50" refString="Pocock, R. I. (1889) Report on the Myriopoda of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta by Dr. John Anderson, F. R. S., Superintendent of the museum. Journal of the Linnean Society, 21, 287 - 302." type="journal article" year="1889">Pocock 1889</bibRefCitation>
: 294
</treatmentCitation>
, 1893: 401
</taxonomicName>
<taxonomicName id="59ED1FADF448FFC8FF49F9929D4AF958" authority="Pocock, 1889: 297" authorityName="Pocock" authorityPageNumber="297" authorityYear="1889" box="[151,556,1662,1686]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="regis">
<emphasis id="AC99B83CF448FFC8FF49F9929E38F95B" box="[151,350,1663,1686]" italics="true" pageId="10" pageNumber="27">Spirostreptus regis</emphasis>
<treatmentCitation id="1F4C423FF448FFC8FEBAF9939D4AF958" author="Pocock" box="[356,556,1662,1685]" page="297" pageId="10" pageNumber="27" year="1889">
<bibRefCitation id="FA7C19DFF448FFC8FEBAF9939E9FF958" author="Pocock" box="[356,505,1662,1685]" pageId="10" pageNumber="50" refString="Pocock, R. I. (1889) Report on the Myriopoda of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta by Dr. John Anderson, F. R. S., Superintendent of the museum. Journal of the Linnean Society, 21, 287 - 302." type="journal article" year="1889">Pocock, 1889</bibRefCitation>
: 297
</treatmentCitation>
</taxonomicName>
(?)
</treatmentCitationGroup>
</paragraph>
<paragraph id="9E52642EF448FFC8FF49F9729D0CF918" blockId="10.[151,836,1598,1750]" pageId="10" pageNumber="27">
<treatmentCitationGroup id="BEFD4300F448FFC8FF49F9729D0CF918" pageId="10" pageNumber="27">
<taxonomicName id="59ED1FADF448FFC8FF49F9729D04F978" authority="Pocock, 1893: 401" authorityName="Pocock" authorityPageNumber="401" authorityYear="1893" box="[151,610,1694,1718]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="AC99B83CF448FFC8FF49F9729EF3F97B" box="[151,405,1695,1718]" italics="true" pageId="10" pageNumber="27">Spirostreptus andersoni</emphasis>
<treatmentCitation id="1F4C423FF448FFC8FE45F9739D04F978" author="Pocock" box="[411,610,1694,1717]" page="401" pageId="10" pageNumber="27" year="1893">
<bibRefCitation id="FA7C19DFF448FFC8FE45F9739D56F978" author="Pocock" box="[411,560,1694,1717]" pageId="10" pageNumber="50" refString="Pocock, R. I. (1893) Viaggio di Leonardo Fea in Birmania e Regioni Vicine. LV. On the Myriopoda of Burma. Pt. 3. Report upon the Iulidae, Chordeumidae and Polyzonidae collected by Sig. L, Fea and Mr. E. W. Oates. Annali del Museo Civico di Storia Naturale di Genova, 13: 386 - 406." type="journal article" year="1893">Pocock, 1893</bibRefCitation>
: 401
</treatmentCitation>
</taxonomicName>
(nomen nudum?) (?)
<taxonomicName id="59ED1FADF448FFC8FF49F9529D0CF918" ID-CoL="YGDD" authority="Demange, 1961: 178" authorityName="Demange" authorityPageNumber="178" authorityYear="1961" box="[151,618,1726,1750]" class="Diplopoda" family="Harpagophoridae" genus="Cornugonus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF448FFC8FF49F9529E1BF91B" box="[151,381,1727,1750]" italics="true" pageId="10" pageNumber="27">Cornugonus opinatus</emphasis>
:
<treatmentCitation id="1F4C423FF448FFC8FE55F9539D0CF918" author="Demange" box="[395,618,1726,1749]" page="178" pageId="10" pageNumber="27" year="1961">
<bibRefCitation id="FA7C19DFF448FFC8FE55F9539D5EF918" author="Demange" box="[395,568,1726,1749]" pageId="10" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange, 1961</bibRefCitation>
: 178
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F448FFC9FF49F8E89A63FEEB" lastPageId="11" lastPageNumber="28" pageId="10" pageNumber="27" type="materials_examined">
<paragraph id="9E52642EF448FFC9FF49F8E89A63FEEB" blockId="10.[151,1436,1797,2016]" lastBlockId="11.[151,1436,152,294]" lastPageId="11" lastPageNumber="28" pageId="10" pageNumber="27">
<emphasis id="AC99B83CF448FFC8FF49F8E89E77F8D2" bold="true" box="[151,273,1797,1823]" pageId="10" pageNumber="27">Material:</emphasis>
<specimenCount id="88EBAFA7F448FFC8FEC7F8E89E0DF8D2" box="[281,363,1797,1823]" pageId="10" pageNumber="27" type="male">1 male</specimenCount>
<collectingCountry id="E6FA24BEF448FFC8FEAAF8E89D75F8D2" box="[372,531,1797,1823]" name="Myanmar" pageId="10" pageNumber="27">MYANMAR</collectingCountry>
, Malewoon, 10° 14ˏ 0˝ N, 98° 37ˏ 0˝ E. Fea leg., (
<collectionCode id="F8FCFCEBF448FFC8FBA4F8E89BBEF8D2" box="[1146,1240,1797,1823]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="10" pageNumber="27">ZMUC</collectionCode>
). This specimen comes from the same series as that illustrated by
<bibRefCitation id="FA7C19DFF448FFC8FD04F8C19CC5F88B" author="Demange" box="[730,931,1836,1862]" pageId="10" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange (1961)</bibRefCitation>
, studied earlier by
<bibRefCitation id="FA7C19DFF448FFC8FB5AF8C19A56F88B" author="Pocock" box="[1156,1328,1836,1862]" pageId="10" pageNumber="50" refString="Pocock, R. I. (1893) Viaggio di Leonardo Fea in Birmania e Regioni Vicine. LV. On the Myriopoda of Burma. Pt. 3. Report upon the Iulidae, Chordeumidae and Polyzonidae collected by Sig. L, Fea and Mr. E. W. Oates. Annali del Museo Civico di Storia Naturale di Genova, 13: 386 - 406." type="journal article" year="1893">Pocock (1893)</bibRefCitation>
and now belonging to
<collectionCode id="F8FCFCEBF448FFC8FEE9F8BF9EF3F8A1" LSID="urn:lsid:biocol.org:col:34988" box="[311,405,1874,1900]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="27">MNHN</collectionCode>
,
<specimenCount id="88EBAFA7F448FFC8FE7CF8BF9D64F8A1" box="[418,514,1874,1900]" pageId="10" pageNumber="27" type="male">3 males</specimenCount>
,
<specimenCount id="88EBAFA7F448FFC8FDD1F8BF9DE5F8A1" box="[527,643,1874,1900]" pageId="10" pageNumber="27" type="female">2 females</specimenCount>
<collectingCountry id="E6FA24BEF448FFC8FD52F8BF9C4CF8A1" box="[652,810,1874,1900]" name="Thailand" pageId="10" pageNumber="27">THAILAND</collectingCountry>
, Petchaburi Province, Cha-Um district, at Nakwang cave, 12° 51ˏ 26˝ N, 99° 56ˏ 29˝ E.
<date id="EA5342EEF448FFC8FD8CF8949C73F85E" box="[594,789,1913,1939]" pageId="10" pageNumber="27" value="2008-10-06">6 October 2008</date>
. H. Enghoff, S. Panha, P. Pimvichai and members of Animal Systematics Research Unit leg., (
<collectionCode id="F8FCFCEBF448FFC8FD5EF84D9DBDF877" box="[640,731,1952,1978]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="10" pageNumber="27">CUMZ</collectionCode>
),
<specimenCount id="88EBAFA7F448FFC8FD2EF84D9C26F877" box="[752,832,1952,1978]" pageId="10" pageNumber="27" type="male">1 male</specimenCount>
<collectingCountry id="E6FA24BEF448FFC8FC96F84D9C82F877" box="[840,996,1952,1978]" name="Thailand" pageId="10" pageNumber="27">THAILAND</collectingCountry>
, Prachuap Khirikhan Province, Bang Saphan district, Ban Grude, 11° 16ˏ 25˝ N, 99° 33ˏ 1˝ E.
<date id="EA5342EEF448FFC8FCE7F82B9B65F82D" box="[825,1027,1990,2016]" pageId="10" pageNumber="27" value="2008-10-12">12 October 2008</date>
. C. Sutcharit and P. Tongkerd leg., (
<collectionCode id="F8FCFCEBF449FFC9FF41FF759F9CFF7F" box="[159,250,152,178]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="11" pageNumber="28">CUMZ</collectionCode>
),
<specimenCount id="88EBAFA7F449FFC9FED1FF759E0DFF7F" box="[271,363,152,178]" pageId="11" pageNumber="28" type="male">3 males</specimenCount>
<collectingCountry id="E6FA24BEF449FFC9FEADFF759D69FF7F" box="[371,527,152,178]" name="Thailand" pageId="11" pageNumber="28">THAILAND</collectingCountry>
, Prachuap Khirikhan Province, Khao Sam Roi Yot National Park, ca.
<quantity id="5915C9CBF449FFC9FA8FFF759AFAFF7F" box="[1361,1436,152,178]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="28" unit="m" value="100.0">100 m</quantity>
before Kaeo cave, 12° 11ˏ 17˝ N, 99° 59ˏ 27˝ E.
<date id="EA5342EEF449FFC9FD0CFF539C0BFF15" box="[722,877,190,216]" pageId="11" pageNumber="28" value="2006-07-21">21 July 2006</date>
. G. Hantke &amp; F. Brand leg., (
<collectionCode id="F8FCFCEBF449FFC9FB1CFF539A66FF15" LSID="urn:lsid:biocol.org:col:34838" box="[1218,1280,190,216]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="28">SMF</collectionCode>
) and
<specimenCount id="88EBAFA7F449FFC9FA9FFF539AFAFF15" box="[1345,1436,190,216]" pageId="11" pageNumber="28" type="male">4 males</specimenCount>
<collectingCountry id="E6FA24BEF449FFC9FF49FF089E54FF32" box="[151,306,229,255]" name="Thailand" pageId="11" pageNumber="28">THAILAND</collectingCountry>
, Prachuap Khirikhan Province, Khao Sam Roi Yot National Park, ca. surrounding of
<quantity id="5915C9CBF449FFC9FAC1FF089A3DFF32" box="[1311,1371,229,255]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="28" unit="m" value="10.0">10 m</quantity>
from Kaeo cave, 12° 12ˏ 6˝ N, 99° 59ˏ 25˝ E.
<date id="EA5342EEF449FFC9FDAEFEE19C6AFEEB" box="[624,780,268,294]" pageId="11" pageNumber="28" value="2006-07-22">22 July 2006</date>
. G. Hantke &amp; F. Brand leg., (
<collectionCode id="F8FCFCEBF449FFC9FB80FEE19BFDFEEB" LSID="urn:lsid:biocol.org:col:34838" box="[1118,1179,268,294]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="28">SMF</collectionCode>
,
<collectionCode id="F8FCFCEBF449FFC9FB78FEE19B9BFEEB" box="[1190,1277,268,294]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="11" pageNumber="28">ZMUC</collectionCode>
)
</paragraph>
</subSubSection>
<caption id="CA9234A6F449FFC9FF49F9979FB7F934" httpUri="https://zenodo.org/record/185977/files/figure.png" pageId="11" pageNumber="28" targetBox="[172,1391,359,1628]" targetPageId="11">
<paragraph id="9E52642EF449FFC9FF49F9979FB7F934" blockId="11.[151,1436,1658,1785]" pageId="11" pageNumber="28">
<emphasis id="AC99B83CF449FFC9FF49F9979E7BF95F" bold="true" box="[151,285,1658,1682]" pageId="11" pageNumber="28">FIGURE 6.</emphasis>
(A, D, E, F):
<taxonomicName id="59ED1FADF449FFC9FE66F9979DF8F95C" box="[440,670,1658,1681]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF449FFC9FE66F9979DF8F95C" box="[440,670,1658,1681]" italics="true" pageId="11" pageNumber="28">Thyropygus opinatus</emphasis>
</taxonomicName>
, gonopods. A: anterior view, left telopodite removed (specimen from Malewoon). B: anterior view, anterior coxal fold (specimen from Nakwang cave). C: anterior view, anterior coxal fold (specimen from Ban Krude). D: right coxa, posterior view. E: right telopodite, anterior view. F: right telopodite, posterior view.
</paragraph>
</caption>
<subSubSection id="D6F737A5F449FFC9FF1BF8CA9D71F811" pageId="11" pageNumber="28" type="diagnosis">
<paragraph id="9E52642EF449FFC9FF1BF8CA9D71F811" blockId="11.[151,1436,1831,2012]" pageId="11" pageNumber="28">
<emphasis id="AC99B83CF449FFC9FF1BF8CA9E21F88C" bold="true" box="[197,327,1831,1857]" pageId="11" pageNumber="28">Diagnosis:</emphasis>
A species of the
<taxonomicName id="59ED1FADF449FFC9FDCDF8C59D1EF88C" box="[531,632,1832,1857]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF449FFC9FDCDF8C59D1EF88C" box="[531,632,1832,1857]" italics="true" pageId="11" pageNumber="28">opinatus</emphasis>
</taxonomicName>
subgroup. Spatulate lobe (
<emphasis id="AC99B83CF449FFC9FC6AF8C59CA1F88C" box="[948,967,1832,1857]" italics="true" pageId="11" pageNumber="28">sl</emphasis>
) at apical part of telopodite terminating in a sharp brown spine. Similar in this respect to
<taxonomicName id="59ED1FADF449FFC9FD04F8A39C2EF8AA" box="[730,840,1870,1895]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="AC99B83CF449FFC9FD04F8A39C2EF8AA" box="[730,840,1870,1895]" italics="true" pageId="11" pageNumber="28">T. floweri</emphasis>
</taxonomicName>
,
<taxonomicName id="59ED1FADF449FFC9FC87F8A39C96F8AA" box="[857,1008,1870,1895]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="implicatus">
<emphasis id="AC99B83CF449FFC9FC87F8A39C96F8AA" box="[857,1008,1870,1895]" italics="true" pageId="11" pageNumber="28">T. implicatus</emphasis>
</taxonomicName>
and
<taxonomicName id="59ED1FADF449FFC9FBF5F8A39BFAF8AA" box="[1067,1180,1870,1895]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="erectus">
<emphasis id="AC99B83CF449FFC9FBF5F8A39BFAF8AA" box="[1067,1180,1870,1895]" italics="true" pageId="11" pageNumber="28">T. erectus</emphasis>
</taxonomicName>
. Differing from these species by mesal margin of anterior coxal fold (
<emphasis id="AC99B83CF449FFC9FD12F8989D97F843" box="[716,753,1909,1934]" italics="true" pageId="11" pageNumber="28">alp</emphasis>
) serrated or with a single denticle, with a small spine (
<emphasis id="AC99B83CF449FFC9FAA5F8989AF4F843" box="[1403,1426,1909,1934]" italics="true" pageId="11" pageNumber="28">ss</emphasis>
) at base of apical part, opposite the origin of tibial spine, and the mesal process of anterior coxal fold (
<emphasis id="AC99B83CF449FFC9FA81F8719AF4F878" box="[1375,1426,1948,1973]" italics="true" pageId="11" pageNumber="28">amp</emphasis>
) almost as long as lateral process.
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F44EFFCEFF1BFF7A9CD6FCD1" pageId="12" pageNumber="29" type="description">
<paragraph id="9E52642EF44EFFCEFF1BFF7A9D62FEEB" blockId="12.[151,1437,151,1918]" pageId="12" pageNumber="29">
<emphasis id="AC99B83CF44EFFCEFF1BFF7A9E05FF7C" bold="true" box="[197,355,151,177]" pageId="12" pageNumber="29">Description:</emphasis>
Adult males with 6270 podous rings, no apodous rings. Length ca.
<quantity id="5915C9CBF44EFFCEFB78FF759A46FF7F" box="[1190,1312,152,178]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="12" pageNumber="29" unit="cm" value="20.0" valueMax="21.0" valueMin="19.0">1921 cm</quantity>
, width ca.
<quantity id="5915C9CBF44EFFCEFF49FF539E20FF15" box="[151,326,190,216]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.15" metricValueMin="1.05" pageId="12" pageNumber="29" unit="mm" value="11.0" valueMax="11.5" valueMin="10.5">10.511.5 mm</quantity>
. Adult females with 70 podous rings, no apodous rings. Length ca.
<quantity id="5915C9CBF44EFFCEFBB9FF539BD5FF15" box="[1127,1203,190,216]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="29" unit="cm" value="18.0">18 cm</quantity>
, width ca.
<quantity id="5915C9CBF44EFFCEFAEBFF539FAFFF32" metricMagnitude="-2" metricUnit="m" metricValue="1.03" metricValueMax="1.08" metricValueMin="0.98" pageId="12" pageNumber="29" unit="mm" value="10.3" valueMax="10.8" valueMin="9.8">9.810.8 mm</quantity>
. Overall color of living animal (
<figureCitation id="06D678ABF44EFFCEFD9EFF089DCCFF32" box="[576,682,229,255]" captionStart="FIGURE 18. A" captionStartId="29.[151,256,1564,1588]" captionTargetBox="[163,1421,241,1493]" captionTargetId="figure@29.[151,1436,229,1506]" captionTargetPageId="29" captionText="FIGURE 18. A: Thyropygus opinatus, living male. B: Thyropygus bearti, living male. C: Thyropygus bispinispatula, living male. D: Thyropygus brachyacanthus, living male." httpUri="https://zenodo.org/record/185989/files/figure.png" pageId="12" pageNumber="29">Fig. 18A</figureCitation>
) black. Legs and antennae reddish brown. Epiproct and margins of paraprocts yellowish brown.
</paragraph>
<paragraph id="9E52642EF44EFFCEFF1BFEDF9CD6FCD1" blockId="12.[151,1437,151,1918]" pageId="12" pageNumber="29">
Gonopods (
<figureCitation id="06D678ABF44EFFCEFE8CFEDF9EC1FE81" box="[338,423,306,332]" captionStart="FIGURE 6" captionStartId="11.[151,257,1658,1682]" captionTargetBox="[172,1391,359,1628]" captionTargetId="figure@11.[151,1436,349,1634]" captionTargetPageId="11" captionText="FIGURE 6. (A, D, E, F): Thyropygus opinatus, gonopods. A: anterior view, left telopodite removed (specimen from Malewoon). B: anterior view, anterior coxal fold (specimen from Nakwang cave). C: anterior view, anterior coxal fold (specimen from Ban Krude). D: right coxa, posterior view. E: right telopodite, anterior view. F: right telopodite, posterior view." httpUri="https://zenodo.org/record/185977/files/figure.png" pageId="12" pageNumber="29">Figs. 6</figureCitation>
AF): Anterior coxal fold (
<emphasis id="AC99B83CF44EFFCEFD2AFEDF9C76FE86" box="[756,784,306,331]" italics="true" pageId="12" pageNumber="29">ac</emphasis>
): lateral process (
<emphasis id="AC99B83CF44EFFCEFC30FEDF9B73FE86" box="[1006,1045,306,331]" italics="true" pageId="12" pageNumber="29">alp</emphasis>
) broadly expanded and apically gradually narrowed, mesal margin with fine serrations (
<bibRefCitation id="FA7C19DFF44EFFCEFCE0FEB49B0DFEBE" author="Demange" box="[830,1131,345,371]" pageId="12" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange 1961: fig. 244</bibRefCitation>
) or smooth with a single denticle (
<figureCitation id="06D678ABF44EFFCEFED8FE6D9E31FE57" box="[262,343,384,410]" captionStart="FIGURE 6" captionStartId="11.[151,257,1658,1682]" captionTargetBox="[172,1391,359,1628]" captionTargetId="figure@11.[151,1436,349,1634]" captionTargetPageId="11" captionText="FIGURE 6. (A, D, E, F): Thyropygus opinatus, gonopods. A: anterior view, left telopodite removed (specimen from Malewoon). B: anterior view, anterior coxal fold (specimen from Nakwang cave). C: anterior view, anterior coxal fold (specimen from Ban Krude). D: right coxa, posterior view. E: right telopodite, anterior view. F: right telopodite, posterior view." httpUri="https://zenodo.org/record/185977/files/figure.png" pageId="12" pageNumber="29">Figs. 6</figureCitation>
AC,
<bibRefCitation id="FA7C19DFF44EFFCEFE41FE6D9D59FE57" author="Pocock" box="[415,575,384,410]" pageId="12" pageNumber="50" refString="Pocock, R. I. (1889) Report on the Myriopoda of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta by Dr. John Anderson, F. R. S., Superintendent of the museum. Journal of the Linnean Society, 21, 287 - 302." type="journal article" year="1889">Pocock 1889</bibRefCitation>
: fig. 2b); mesal process (
<emphasis id="AC99B83CF44EFFCEFCB5FE6D9CF8FE54" box="[875,926,384,409]" italics="true" pageId="12" pageNumber="29">amp</emphasis>
) almost as long as lateral process, slender, straight, directed distad, terminally slightly curved, pointed, sometimes crossing over with opposite
<emphasis id="AC99B83CF44EFFCEFAB6FE4A9AFAFE0D" box="[1384,1436,423,448]" italics="true" pageId="12" pageNumber="29">amp</emphasis>
(
<figureCitation id="06D678ABF44EFFCEFF41FE209F97FE2A" box="[159,241,461,487]" captionStart="FIGURE 6" captionStartId="11.[151,257,1658,1682]" captionTargetBox="[172,1391,359,1628]" captionTargetId="figure@11.[151,1436,349,1634]" captionTargetPageId="11" captionText="FIGURE 6. (A, D, E, F): Thyropygus opinatus, gonopods. A: anterior view, left telopodite removed (specimen from Malewoon). B: anterior view, anterior coxal fold (specimen from Nakwang cave). C: anterior view, anterior coxal fold (specimen from Ban Krude). D: right coxa, posterior view. E: right telopodite, anterior view. F: right telopodite, posterior view." httpUri="https://zenodo.org/record/185977/files/figure.png" pageId="12" pageNumber="29">Figs. 6</figureCitation>
AC). Posterior coxal fold (
<emphasis id="AC99B83CF44EFFCEFDE5FE209D30FE2B" box="[571,598,461,486]" italics="true" pageId="12" pageNumber="29">pc</emphasis>
) (
<figureCitation id="06D678ABF44EFFCEFDAFFE209DDEFE2A" box="[625,696,461,487]" captionStart="FIGURE 6" captionStartId="11.[151,257,1658,1682]" captionTargetBox="[172,1391,359,1628]" captionTargetId="figure@11.[151,1436,349,1634]" captionTargetPageId="11" captionText="FIGURE 6. (A, D, E, F): Thyropygus opinatus, gonopods. A: anterior view, left telopodite removed (specimen from Malewoon). B: anterior view, anterior coxal fold (specimen from Nakwang cave). C: anterior view, anterior coxal fold (specimen from Ban Krude). D: right coxa, posterior view. E: right telopodite, anterior view. F: right telopodite, posterior view." httpUri="https://zenodo.org/record/185977/files/figure.png" pageId="12" pageNumber="29">Fig. 6</figureCitation>
D) distally with two processes: mesal process (
<emphasis id="AC99B83CF44EFFCEFB3DFE209A73FE2B" box="[1251,1301,461,486]" italics="true" pageId="12" pageNumber="29">pmp</emphasis>
) a rounded lobe, shorter than lateral process (
<emphasis id="AC99B83CF44EFFCEFDF7FE199D29FDC0" box="[553,591,500,525]" italics="true" pageId="12" pageNumber="29">plp</emphasis>
) and separated from
<emphasis id="AC99B83CF44EFFCEFC92FE199C14FDC0" box="[844,882,500,525]" italics="true" pageId="12" pageNumber="29">plp</emphasis>
by deep, rounded sinus; lateral process (
<emphasis id="AC99B83CF44EFFCEFA89FE199A1AFDC0" box="[1367,1404,500,525]" italics="true" pageId="12" pageNumber="29">plp</emphasis>
) a large, smooth, rounded node, projecting further caudad than mesal process. Telopodite (
<figureCitation id="06D678ABF44EFFCEFB41FDF79B96FDF9" box="[1183,1264,538,564]" captionStart="FIGURE 6" captionStartId="11.[151,257,1658,1682]" captionTargetBox="[172,1391,359,1628]" captionTargetId="figure@11.[151,1436,349,1634]" captionTargetPageId="11" captionText="FIGURE 6. (A, D, E, F): Thyropygus opinatus, gonopods. A: anterior view, left telopodite removed (specimen from Malewoon). B: anterior view, anterior coxal fold (specimen from Nakwang cave). C: anterior view, anterior coxal fold (specimen from Ban Krude). D: right coxa, posterior view. E: right telopodite, anterior view. F: right telopodite, posterior view." httpUri="https://zenodo.org/record/185977/files/figure.png" pageId="12" pageNumber="29">Figs. 6</figureCitation>
EF): Femoral spine (
<emphasis id="AC99B83CF44EFFCEFF34FDAC9E66FD97" box="[234,256,577,602]" italics="true" pageId="12" pageNumber="29">fe</emphasis>
) long, slender, very distinctly crenulated along outer curvature,
<emphasis id="AC99B83CF44EFFCEFBC3FDAC9B0DFD97" box="[1053,1131,577,602]" italics="true" pageId="12" pageNumber="29">in situ</emphasis>
resting against posterior surface of
<emphasis id="AC99B83CF44EFFCEFECBFD859E5DFD4C" box="[277,315,616,641]" italics="true" pageId="12" pageNumber="29">alp</emphasis>
; telopodite distally to
<emphasis id="AC99B83CF44EFFCEFD96FD859D3BFD4C" box="[584,605,616,641]" italics="true" pageId="12" pageNumber="29">fe</emphasis>
with a massive, broad, round lobe (
<emphasis id="AC99B83CF44EFFCEFBD5FD859B44FD4C" box="[1035,1058,616,641]" italics="true" pageId="12" pageNumber="29">lo</emphasis>
) projecting distolaterally; tibial spine (
<emphasis id="AC99B83CF44EFFCEFF39FD629F91FD65" box="[231,247,655,680]" italics="true" pageId="12" pageNumber="29">ti</emphasis>
) long, slender, recurved, in anterior view crossing over with
<emphasis id="AC99B83CF44EFFCEFC0EFD629C83FD65" box="[976,997,655,680]" italics="true" pageId="12" pageNumber="29">fe</emphasis>
and reaching mesal end of
<emphasis id="AC99B83CF44EFFCEFAF0FD629A23FD65" box="[1326,1349,655,680]" italics="true" pageId="12" pageNumber="29">lo</emphasis>
; apical part: spatulate lobe (
<emphasis id="AC99B83CF44EFFCEFE4FFD589EC3FD03" box="[401,421,693,718]" italics="true" pageId="12" pageNumber="29">sl</emphasis>
) terminating in a sharp brown spine (very slender in Nakwang specimens); a small spine (
<emphasis id="AC99B83CF44EFFCEFF3BFD319F9AFD38" box="[229,252,732,757]" italics="true" pageId="12" pageNumber="29">ss</emphasis>
) at base of apical part, opposite the origin of
<emphasis id="AC99B83CF44EFFCEFCCEFD319C46FD38" box="[784,800,732,757]" italics="true" pageId="12" pageNumber="29">ti</emphasis>
; palette (
<emphasis id="AC99B83CF44EFFCEFC50FD319CCDFD38" box="[910,939,732,757]" italics="true" pageId="12" pageNumber="29">pa</emphasis>
) simple, with a longitudinal rounded crest (
<emphasis id="AC99B83CF44EFFCEFF7FFCEF9FDFFCD6" box="[161,185,770,795]" italics="true" pageId="12" pageNumber="29">cr</emphasis>
) near tip, distally with about ten brownish blepharochaetae (
<emphasis id="AC99B83CF44EFFCEFC5CFCEF9CF9FCD6" box="[898,927,770,795]" italics="true" pageId="12" pageNumber="29">bp</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F44EFFCEFF1BFCC59B93F8B3" pageId="12" pageNumber="29" type="distribution">
<paragraph id="9E52642EF44EFFCEFF1BFCC59BD6FBC9" blockId="12.[151,1437,151,1918]" pageId="12" pageNumber="29">
<emphasis id="AC99B83CF44EFFCEFF1BFCC59E03FC8F" bold="true" box="[197,357,808,834]" pageId="12" pageNumber="29">Distribution</emphasis>
(
<figureCitation id="06D678ABF44EFFCEFEA8FCC49EB2FC8E" box="[374,468,809,835]" captionStart="FIGURE 20" captionStartId="32.[151,255,1208,1232]" captionText="FIGURE 20. Known distribution of the species of the T. opinatus subgroup." pageId="12" pageNumber="29">Fig. 20</figureCitation>
): Described from
<collectingCountry id="E6FA24BEF44EFFCEFD6CFCC49C34FC8E" box="[690,850,809,835]" name="Myanmar" pageId="12" pageNumber="29">MYANMAR</collectingCountry>
, Tenasserim, without an exact locality (
<bibRefCitation id="FA7C19DFF44EFFCEFAE2FCC49FBFFCA7" author="Karsch" pageId="12" pageNumber="50" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die Gesamten Naturwissenschaften, Dritte Folge, 6, 3 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
). Subsequently recorded from Sullivan Island (10° 50ˏ 0˝ N, 98° 15ˏ 0˝ E) by
<bibRefCitation id="FA7C19DFF44EFFCEFB79FCBD9A06FCA7" author="Pocock" box="[1191,1376,848,874]" pageId="12" pageNumber="50" refString="Pocock, R. I. (1889) Report on the Myriopoda of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta by Dr. John Anderson, F. R. S., Superintendent of the museum. Journal of the Linnean Society, 21, 287 - 302." type="journal article" year="1889">Pocock (1889)</bibRefCitation>
. The
<collectingCountry id="E6FA24BEF44EFFCEFF49FC9B9E6FFC5D" box="[151,265,886,912]" name="Myanmar" pageId="12" pageNumber="29">Myanmar</collectingCountry>
specimens studied by us,
<bibRefCitation id="FA7C19DFF44EFFCEFDE6FC9B9D82FC5D" author="Pocock" box="[568,740,886,912]" pageId="12" pageNumber="50" refString="Pocock, R. I. (1893) Viaggio di Leonardo Fea in Birmania e Regioni Vicine. LV. On the Myriopoda of Burma. Pt. 3. Report upon the Iulidae, Chordeumidae and Polyzonidae collected by Sig. L, Fea and Mr. E. W. Oates. Annali del Museo Civico di Storia Naturale di Genova, 13: 386 - 406." type="journal article" year="1893">Pocock (1893)</bibRefCitation>
and
<bibRefCitation id="FA7C19DFF44EFFCEFCC2FC9B9C87FC5D" author="Demange" box="[796,993,886,912]" pageId="12" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange (1961)</bibRefCitation>
are from Malewoon, Tenasserim (10° 14ˏ 0˝ N, 98° 37ˏ 0˝ E). We here record the species as new to
<collectingCountry id="E6FA24BEF44EFFCEFCA6FC709C87FC7A" box="[888,993,925,951]" name="Thailand" pageId="12" pageNumber="29">Thailand</collectingCountry>
based on specimens from Petchaburi and Prachuap Khirikhan Provinces. The records from King Island (
<bibRefCitation id="FA7C19DFF44EFFCEFC6BFC299B32FC13" author="Pocock" box="[949,1108,964,990]" pageId="12" pageNumber="50" refString="Pocock, R. I. (1889) Report on the Myriopoda of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta by Dr. John Anderson, F. R. S., Superintendent of the museum. Journal of the Linnean Society, 21, 287 - 302." type="journal article" year="1889">Pocock 1889</bibRefCitation>
), and from &quot;S. Tenasserim&quot; (
<bibRefCitation id="FA7C19DFF44EFFCEFF40FC079E27FBC9" author="Pocock" box="[158,321,1002,1028]" pageId="12" pageNumber="50" refString="Pocock, R. I. (1893) Viaggio di Leonardo Fea in Birmania e Regioni Vicine. LV. On the Myriopoda of Burma. Pt. 3. Report upon the Iulidae, Chordeumidae and Polyzonidae collected by Sig. L, Fea and Mr. E. W. Oates. Annali del Museo Civico di Storia Naturale di Genova, 13: 386 - 406." type="journal article" year="1893">Pocock,1893</bibRefCitation>
) are fraught with serious doubts (see notes) and are not shown on the map.
</paragraph>
<paragraph id="9E52642EF44EFFCEFF1BFBFD9BA6F985" blockId="12.[151,1437,151,1918]" pageId="12" pageNumber="29">
<emphasis id="AC99B83CF44EFFCEFF1BFBFD9E70FBE7" bold="true" box="[197,278,1040,1066]" pageId="12" pageNumber="29">Notes:</emphasis>
This species presents several unresolved problems. The original description (
<bibRefCitation id="FA7C19DFF44EFFCEFB64FBFC9A3BFBE6" author="Karsch" box="[1210,1373,1041,1067]" pageId="12" pageNumber="50" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die Gesamten Naturwissenschaften, Dritte Folge, 6, 3 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
) was said to be based on a male, but as usual, Karsch did not describe the gonopods. The whereabouts of the
<typeStatus id="4156DA8CF44EFFCEFAB7FBD59AFAFB9F" box="[1385,1436,1080,1106]" pageId="12" pageNumber="29">type</typeStatus>
specimen are unknown. The next mention of
<taxonomicName id="59ED1FADF44EFFCEFD68FBB29C7BFBB5" box="[694,797,1119,1144]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF44EFFCEFD68FBB29C7BFBB5" box="[694,797,1119,1144]" italics="true" pageId="12" pageNumber="29">opinatus</emphasis>
</taxonomicName>
was by
<bibRefCitation id="FA7C19DFF44EFFCEFC5FFBB39B56FBB5" author="Pocock" box="[897,1072,1118,1144]" pageId="12" pageNumber="50" refString="Pocock, R. I. (1889) Report on the Myriopoda of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta by Dr. John Anderson, F. R. S., Superintendent of the museum. Journal of the Linnean Society, 21, 287 - 302." type="journal article" year="1889">Pocock (1889)</bibRefCitation>
who reported it from Sullivan Island. Pocock gave a relatively good description, including a small gonopod drawing. In 1893, Pocock recorded
<taxonomicName id="59ED1FADF44EFFCEFED4FB419E15FB08" box="[266,371,1196,1221]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF44EFFCEFED4FB419E15FB08" box="[266,371,1196,1221]" italics="true" pageId="12" pageNumber="29">opinatus</emphasis>
</taxonomicName>
from Malewoon and further noted that “Two young examples collected by Oates in S. Tenasserim are doubtfully referred to this species” and further that “This species was originally described from Tenasserim and was subsequently procured by Dr. Anderson from the Mergui Archipelago”. The latter statement must refer to the name “?
<taxonomicName id="59ED1FADF44EFFCEFD87FACD9CA6FAF7" authority="Pocock, 1889" authorityName="Pocock" authorityYear="1889" box="[601,960,1312,1338]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="AC99B83CF44EFFCEFD87FACD9D11FAF4" box="[601,631,1312,1337]" italics="true" pageId="12" pageNumber="29">Sp</emphasis>
.
<emphasis id="AC99B83CF44EFFCEFD56FACD9C64FAF4" box="[648,770,1312,1337]" italics="true" pageId="12" pageNumber="29">andersoni</emphasis>
<bibRefCitation id="FA7C19DFF44EFFCEFCC6FACD9CA6FAF7" author="Pocock" box="[792,960,1312,1338]" pageId="12" pageNumber="50" refString="Pocock, R. I. (1889) Report on the Myriopoda of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta by Dr. John Anderson, F. R. S., Superintendent of the museum. Journal of the Linnean Society, 21, 287 - 302." type="journal article" year="1889">Pocock, 1889</bibRefCitation>
</taxonomicName>
listed in the synonymy of
<taxonomicName id="59ED1FADF44EFFCEFAD2FACD9A10FAF4" box="[1292,1398,1312,1337]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF44EFFCEFAD2FACD9A10FAF4" box="[1292,1398,1312,1337]" italics="true" pageId="12" pageNumber="29">opinatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="FA7C19DFF44EFFCEFF49FAAB9E20FAAD" author="Pocock" box="[151,326,1350,1376]" pageId="12" pageNumber="50" refString="Pocock, R. I. (1893) Viaggio di Leonardo Fea in Birmania e Regioni Vicine. LV. On the Myriopoda of Burma. Pt. 3. Report upon the Iulidae, Chordeumidae and Polyzonidae collected by Sig. L, Fea and Mr. E. W. Oates. Annali del Museo Civico di Storia Naturale di Genova, 13: 386 - 406." type="journal article" year="1893">Pocock (1893)</bibRefCitation>
with an exact reference to the page and figure number in his 1889 paper and a note that this name is based on a young specimen. The strange thing is that in
<bibRefCitation id="FA7C19DFF44EFFCEFC6BFA809B0FFA4A" author="Pocock" box="[949,1129,1389,1415]" pageId="12" pageNumber="50" refString="Pocock, R. I. (1889) Report on the Myriopoda of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta by Dr. John Anderson, F. R. S., Superintendent of the museum. Journal of the Linnean Society, 21, 287 - 302." type="journal article" year="1889">Pocock (1889)</bibRefCitation>
there is no
<taxonomicName id="59ED1FADF44EFFCEFB25FA809E6AFA60" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="AC99B83CF44EFFCEFB25FA809E6AFA60" italics="true" pageId="12" pageNumber="29">Spirostreptus andersoni</emphasis>
</taxonomicName>
; instead there is, on the cited page and under the cited figure number, a
<taxonomicName id="59ED1FADF44EFFCEFB82FA799FB2FA19" authority="Pocock, 1889" authorityName="Pocock" authorityYear="1889" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="regis">
<emphasis id="AC99B83CF44EFFCEFB82FA799A51FA60" box="[1116,1335,1428,1453]" italics="true" pageId="12" pageNumber="29">Spirostreptus regis</emphasis>
<bibRefCitation id="FA7C19DFF44EFFCEFAE0FA799FB2FA19" author="Pocock" pageId="12" pageNumber="50" refString="Pocock, R. I. (1889) Report on the Myriopoda of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta by Dr. John Anderson, F. R. S., Superintendent of the museum. Journal of the Linnean Society, 21, 287 - 302." type="journal article" year="1889">Pocock, 1889</bibRefCitation>
</taxonomicName>
, based on a single female from King Island. The length of this specimen was given as
<quantity id="5915C9CBF44EFFCEFB61FA579A74FA19" box="[1215,1298,1466,1492]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="29" unit="mm" value="55.0">55 mm</quantity>
which does suggest a juvenile. It seems most likely that
<taxonomicName id="59ED1FADF44EFFCEFD75FA0C9C44FA37" box="[683,802,1505,1530]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="AC99B83CF44EFFCEFD75FA0C9C44FA37" box="[683,802,1505,1530]" italics="true" pageId="12" pageNumber="29">andersoni</emphasis>
</taxonomicName>
is a
<emphasis id="AC99B83CF44EFFCEFC85FA0C9B66FA37" box="[859,1024,1505,1530]" italics="true" pageId="12" pageNumber="29">lapsus calami</emphasis>
for
<taxonomicName id="59ED1FADF44EFFCEFBEDFA0C9B0BFA37" box="[1075,1133,1505,1530]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="regis">
<emphasis id="AC99B83CF44EFFCEFBEDFA0C9B0BFA37" box="[1075,1133,1505,1530]" italics="true" pageId="12" pageNumber="29">regis</emphasis>
</taxonomicName>
, maybe due to confusion with
<taxonomicName id="59ED1FADF44EFFCEFF0DF9E59D1EF9EF" authority="Pocock, 1889" authorityName="Pocock" authorityYear="1889" box="[211,632,1544,1570]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="AC99B83CF44EFFCEFF0DF9E59EABF9EC" box="[211,461,1544,1569]" italics="true" pageId="12" pageNumber="29">Spirobolus andersoni</emphasis>
<bibRefCitation id="FA7C19DFF44EFFCEFE0BF9E59D1EF9EF" author="Pocock" box="[469,632,1544,1570]" pageId="12" pageNumber="50" refString="Pocock, R. I. (1889) Report on the Myriopoda of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta by Dr. John Anderson, F. R. S., Superintendent of the museum. Journal of the Linnean Society, 21, 287 - 302." type="journal article" year="1889">Pocock, 1889</bibRefCitation>
</taxonomicName>
, described just two pages after
<taxonomicName id="59ED1FADF44EFFCEFC35F9E59BA1F9EC" box="[1003,1223,1544,1569]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="regis">
<emphasis id="AC99B83CF44EFFCEFC35F9E59BA1F9EC" box="[1003,1223,1544,1569]" italics="true" pageId="12" pageNumber="29">Spirostreptus regis</emphasis>
</taxonomicName>
.
<bibRefCitation id="FA7C19DFF44EFFCEFB08F9E59AFDF9EF" author="Demange" box="[1238,1435,1544,1570]" pageId="12" pageNumber="50" refString="Demange, J. M. (1960) Les Types d'Harpagophoridae de R. I. Pocock conserves au British Museum (Natural History) (Myriapodes, Diplopodes). Bulletin of the British Museum (Natural History) Zoology, 7 (2), 141 - 179." type="journal article" year="1960">Demange (1960)</bibRefCitation>
found specimens of neither
<taxonomicName id="59ED1FADF44EFFCEFE02F9C29D37F985" box="[476,593,1583,1608]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="AC99B83CF44EFFCEFE02F9C29D37F985" box="[476,593,1583,1608]" italics="true" pageId="12" pageNumber="29">andersoni</emphasis>
</taxonomicName>
nor
<taxonomicName id="59ED1FADF44EFFCEFD59F9C29DA6F985" box="[647,704,1583,1608]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="regis">
<emphasis id="AC99B83CF44EFFCEFD59F9C29DA6F985" box="[647,704,1583,1608]" italics="true" pageId="12" pageNumber="29">regis</emphasis>
</taxonomicName>
in the Natural History Museum of London.
</paragraph>
<paragraph id="9E52642EF44EFFCEFF18F9B89B93F8B3" blockId="12.[151,1437,151,1918]" pageId="12" pageNumber="29">
<bibRefCitation id="FA7C19DFF44EFFCEFF18F9B89EE8F9A2" author="Demange" box="[198,398,1621,1647]" pageId="12" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange (1961)</bibRefCitation>
based his description of
<taxonomicName id="59ED1FADF44EFFCEFD64F9B89C79F9A3" box="[698,799,1621,1646]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF44EFFCEFD64F9B89C79F9A3" box="[698,799,1621,1646]" italics="true" pageId="12" pageNumber="29">opinatus</emphasis>
</taxonomicName>
on a specimen from Malewoon examined by
<bibRefCitation id="FA7C19DFF44EFFCEFA98F9B89F8EF95B" author="Pocock" pageId="12" pageNumber="50" refString="Pocock, R. I. (1893) Viaggio di Leonardo Fea in Birmania e Regioni Vicine. LV. On the Myriopoda of Burma. Pt. 3. Report upon the Iulidae, Chordeumidae and Polyzonidae collected by Sig. L, Fea and Mr. E. W. Oates. Annali del Museo Civico di Storia Naturale di Genova, 13: 386 - 406." type="journal article" year="1893">Pocock (1893)</bibRefCitation>
, and we have carried on this understanding of
<taxonomicName id="59ED1FADF44EFFCEFCD2F9919C16F958" box="[780,880,1660,1685]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF44EFFCEFCD2F9919C16F958" box="[780,880,1660,1685]" italics="true" pageId="12" pageNumber="29">opinatus</emphasis>
</taxonomicName>
, being aware that rediscovery of Karschs
<typeStatus id="4156DA8CF44EFFCEFAB7F9919AFAF95B" box="[1385,1436,1660,1686]" pageId="12" pageNumber="29">type</typeStatus>
specimen may require a revision. Pococks (1889) gonopod drawing based on the specimen he described from Sullivan Island bears a strong resemblance to our
<figureCitation id="06D678ABF44EFFCEFD2EF9249C51F92E" box="[752,823,1737,1763]" captionStart="FIGURE 6" captionStartId="11.[151,257,1658,1682]" captionTargetBox="[172,1391,359,1628]" captionTargetId="figure@11.[151,1436,349,1634]" captionTargetPageId="11" captionText="FIGURE 6. (A, D, E, F): Thyropygus opinatus, gonopods. A: anterior view, left telopodite removed (specimen from Malewoon). B: anterior view, anterior coxal fold (specimen from Nakwang cave). C: anterior view, anterior coxal fold (specimen from Ban Krude). D: right coxa, posterior view. E: right telopodite, anterior view. F: right telopodite, posterior view." httpUri="https://zenodo.org/record/185977/files/figure.png" pageId="12" pageNumber="29">Fig. 6</figureCitation>
A, although the lateral process (
<emphasis id="AC99B83CF44EFFCEFB67F9249BB9F92F" box="[1209,1247,1737,1762]" italics="true" pageId="12" pageNumber="29">alp</emphasis>
) of the anterior gonopod fold looks slightly narrower in the Sullivan Island specimen. Considering that the specimen illustrated by
<bibRefCitation id="FA7C19DFF44EFFCEFEE4F8FB9D64F8FD" author="Demange" box="[314,514,1814,1840]" pageId="12" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange (1961)</bibRefCitation>
is again slightly different from the one from the same series studied by us, in this case with an even broader
<emphasis id="AC99B83CF44EFFCEFDDFF8D09D40F89B" box="[513,550,1853,1878]" italics="true" pageId="12" pageNumber="29">alp</emphasis>
, and further considering the variability among the new
<collectingCountry id="E6FA24BEF44EFFCEFB6AF8D09B8CF89A" box="[1204,1258,1853,1879]" name="Thailand" pageId="12" pageNumber="29">Thai</collectingCountry>
specimens, we accept the record of
<taxonomicName id="59ED1FADF44EFFCEFE58F8899E8CF8B0" box="[390,490,1892,1917]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF44EFFCEFE58F8899E8CF8B0" box="[390,490,1892,1917]" italics="true" pageId="12" pageNumber="29">opinatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="FA7C19DFF44EFFCEFDC9F8899DA4F8B3" author="Pocock" box="[535,706,1892,1918]" pageId="12" pageNumber="50" refString="Pocock, R. I. (1889) Report on the Myriopoda of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta by Dr. John Anderson, F. R. S., Superintendent of the museum. Journal of the Linnean Society, 21, 287 - 302." type="journal article" year="1889">Pocock (1889)</bibRefCitation>
as belonging to this species as understood here.
</paragraph>
</subSubSection>
</treatment>
</document>