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<document id="C8455ECD2AED8561FF6837B74B8CAE52" ID-CLB-Dataset="51561" ID-DOI="10.11646/zootaxa.4092.4.6" ID-GBIF-Dataset="9336f407-217f-436a-aaf5-211b9419f93a" ID-ISSN="1175-5326" ID-Zenodo-Dep="260976" ID-ZooBank="AB185179-B61C-4C30-8D67-579330563CD4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458297171994" checkinUser="plazi" docAuthor="Grazia, Jocelia, Barros, Lurdiana Dayse De &amp; Barão, Kim Ribeiro" docDate="2016" docId="0630AD03C10AFFE150FFFE64FA2F4E51" docLanguage="en" docName="zootaxa.4092.4.6.pdf.imf" docOrigin="Zootaxa 4092 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Elanela Rolston revisited (Heteroptera: Pentatomidae): new distributional records and description of new species" docType="treatment" docVersion="7" lastPageNumber="563" masterDocId="FA09D57BC10BFFE35068FF99FFB34C5C" masterDocTitle="Elanela Rolston revisited (Heteroptera: Pentatomidae): new distributional records and description of new species" masterLastPageNumber="571" masterPageNumber="561" pageNumber="562" updateTime="1698188163191" updateUser="ExternalLinkService">
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<mods:title id="40B656DACC63C7BFB09AC32581FF7352">Elanela Rolston revisited (Heteroptera: Pentatomidae): new distributional records and description of new species</mods:title>
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<mods:namePart id="553AEA18375BBE2C55145F0D7EA82C2F">Grazia, Jocelia</mods:namePart>
<mods:affiliation id="AD2A370EC5D6FE271B67EA8B7C08AC42">Depto. de Zoologia &amp; PPG Biologia Animal, Instituto de Biociências, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, 91501 - 970, Porto Alegre, RS, Brazil</mods:affiliation>
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<heading id="D56EAB79C10AFFE250FFFE64FE154E4B" bold="true" box="[151,422,509,535]" fontSize="11" level="1" pageId="1" pageNumber="562" reason="1">
<emphasis id="BCEDC007C10AFFE250FFFE64FE154E4B" bold="true" box="[151,422,509,535]" pageId="1" pageNumber="562">
<emphasis id="BCEDC007C10AFFE250FFFE64FF454E4B" bold="true" box="[151,246,509,535]" italics="true" pageId="1" pageNumber="562">Elanela</emphasis>
<bibRefCitation id="EA0861E4C10AFFE25094FE64FE154E4B" author="Rolston, L. H. &amp; McDonald, F. J. D. &amp; Thomas, D. B." box="[252,422,509,535]" journalOrPublisher="Journal of the New York Entomological Society" pageId="1" pageNumber="562" pagination="120 - 132" part="88" refId="ref7617" refString="Rolston, L. H., McDonald, F. J. D. &amp; Thomas, D. B. (1980) A conspectus of Pentatomini genera of the Western hemisphere. Part I. (Hemiptrea: Pentatomidae). Journal of the New York Entomological Society, 88, 120 - 132." title="A conspectus of Pentatomini genera of the Western hemisphere. Part I. (Hemiptrea: Pentatomidae)" type="journal article" year="1980">Rolston, 1980</bibRefCitation>
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<paragraph id="8E261C15C10AFFE250FFFDDDFB774E06" blockId="1.[151,1220,580,602]" box="[151,1220,580,602]" pageId="1" pageNumber="562">
<treatmentCitationGroup id="AE893B3BC10AFFE250FFFDDDFB734E06" box="[151,1216,580,602]" pageId="1" pageNumber="562">
<emphasis id="BCEDC007C10AFFE250FFFDDDFF554E06" box="[151,230,580,602]" italics="true" pageId="1" pageNumber="562">Elanela</emphasis>
<bibRefCitation id="EA0861E4C10AFFE25084FDDDFEC94E06" author="Rolston, L. H. &amp; McDonald, F. J. D. &amp; Thomas, D. B." box="[236,378,580,602]" journalOrPublisher="Journal of the New York Entomological Society" pageId="1" pageNumber="562" pagination="120 - 132" part="88" refId="ref7617" refString="Rolston, L. H., McDonald, F. J. D. &amp; Thomas, D. B. (1980) A conspectus of Pentatomini genera of the Western hemisphere. Part I. (Hemiptrea: Pentatomidae). Journal of the New York Entomological Society, 88, 120 - 132." title="A conspectus of Pentatomini genera of the Western hemisphere. Part I. (Hemiptrea: Pentatomidae)" type="journal article" year="1980">
Rolston,
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<emphasis id="BCEDC007C10AFFE251E8FDDCFE274E06" box="[384,404,581,602]" italics="true" pageId="1" pageNumber="562">in</emphasis>
<treatmentCitation id="0F383A04C10AFFE251F2FDDDFD674E06" box="[410,724,580,602]" pageId="1" pageNumber="562">
<bibRefCitation id="EA0861E4C10AFFE251F2FDDDFDC34E06" author="Rolston, L. H. &amp; McDonald, F. J. D. &amp; Thomas, D. B." box="[410,624,580,602]" journalOrPublisher="Journal of the New York Entomological Society" pageId="1" pageNumber="562" pagination="120 - 132" part="88" refId="ref7617" refString="Rolston, L. H., McDonald, F. J. D. &amp; Thomas, D. B. (1980) A conspectus of Pentatomini genera of the Western hemisphere. Part I. (Hemiptrea: Pentatomidae). Journal of the New York Entomological Society, 88, 120 - 132." title="A conspectus of Pentatomini genera of the Western hemisphere. Part I. (Hemiptrea: Pentatomidae)" type="journal article" year="1980">
Rolston
<emphasis id="BCEDC007C10AFFE25187FDDCFDAF4E06" box="[495,540,580,602]" italics="true" pageId="1" pageNumber="562">et al</emphasis>
. (1980)
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: 125–126
</treatmentCitation>
, figs. 8, 12–13;
<treatmentCitation id="0F383A04C10AFFE2531FFDDDFB734E06" box="[887,1216,580,602]" pageId="1" pageNumber="562">
<bibRefCitation id="EA0861E4C10AFFE2531FFDDDFB734E06" author="Grazia, J. &amp; Greve, C." box="[887,1216,580,602]" journalOrPublisher="Heteropterus Revista de Entomologia" pageId="1" pageNumber="562" pagination="261 - 266" part="11" refId="ref7532" refString="Grazia, J. &amp; Greve, C. (2011) Contributions to the knowledge of Elanela: Elanela jordi sp. nov., from Amazonas, Brazil (Hemiptera: Heteroptera: Pentatomidae). Heteropterus Revista de Entomologia, 11, 261 - 266." title="Contributions to the knowledge of Elanela: Elanela jordi sp. nov., from Amazonas, Brazil (Hemiptera: Heteroptera: Pentatomidae)" type="journal article" year="2011">Grazia &amp; Greve, 2011: 261–262</bibRefCitation>
</treatmentCitation>
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.
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<paragraph id="8E261C15C10AFFE250FFFD1FFC2B4EC3" blockId="1.[151,1437,646,1427]" box="[151,920,646,671]" pageId="1" pageNumber="562">
<emphasis id="BCEDC007C10AFFE250FFFD1FFE814EC3" bold="true" box="[151,306,646,671]" inLineHeading="true" pageId="1" pageNumber="562" reason="1">
<typeStatus id="5122A2B7C10AFFE250FFFD1FFF614EC3" box="[151,210,646,671]" pageId="1" pageNumber="562">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="49996796C10AFFE25151FD1FFD344EC3" authority="Rolston, 1980" authorityName="Rolston" authorityYear="1980" box="[313,647,646,671]" class="Insecta" family="Pentatomidae" genus="Elanela" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="hevera">
<emphasis id="BCEDC007C10AFFE25151FD1FFE504EC2" box="[313,483,646,670]" italics="true" pageId="1" pageNumber="562">Elanela hevera</emphasis>
<bibRefCitation id="EA0861E4C10AFFE25182FD1FFD344EC3" author="Rolston, L. H. &amp; McDonald, F. J. D. &amp; Thomas, D. B." box="[490,647,646,671]" journalOrPublisher="Journal of the New York Entomological Society" pageId="1" pageNumber="562" pagination="120 - 132" part="88" refId="ref7617" refString="Rolston, L. H., McDonald, F. J. D. &amp; Thomas, D. B. (1980) A conspectus of Pentatomini genera of the Western hemisphere. Part I. (Hemiptrea: Pentatomidae). Journal of the New York Entomological Society, 88, 120 - 132." title="A conspectus of Pentatomini genera of the Western hemisphere. Part I. (Hemiptrea: Pentatomidae)" type="journal article" year="1980">Rolston, 1980</bibRefCitation>
</taxonomicName>
by original designation.
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<paragraph id="8E261C15C10AFFE250AFFD33FDA24F2B" blockId="1.[151,1437,646,1427]" pageId="1" pageNumber="562">
<emphasis id="BCEDC007C10AFFE250AFFD33FEEA4E9F" bold="true" box="[199,345,682,707]" inLineHeading="true" pageId="1" pageNumber="562" reason="1">Description.</emphasis>
Body castaneous with brown punctuations. Lateral margin of each mandibular plate outlined in black; each buccula with black, longitudinal stripe with a row of deep punctures basally; distal half of antennal articles IV and V black. Legs yellow, area surrounding each setae brown; scutellum with small yellow calluses on each basal angle and on basal margin medially; disc of scutellum black, reddish, or ivory, calloused; corium of hemelytra black medially, radial vein with a subapical yellow spot; membrane of hemelytra infuscate. Outline of each abdominal urosternite black.
</paragraph>
<paragraph id="8E261C15C10AFFE250FFFC1BFA3048CA" blockId="1.[151,1437,646,1427]" pageId="1" pageNumber="562">Head wider than long, strongly sloping ventrally; apex of each mandibular plate not surpassing clypeus; margins of mandibular plates strongly concave before eyes; posterior margin of each buccula rounded; first labial segment surpassing bucculae; first antennal segment stout; proportion of antennal segments variable. Pronotum trapezoidal; anterolateral margins straight; humeral angles rounded; posterior margin slightly convex, nearly straight; apex of scutellum obtuse; scutellum shorter than corium; apex of corium rounded, reaching posterior margin of abdominal segment VI; femora unarmed; mesosternum carinate; metasternum produced, humped towards the midline, carinate anteriorly and bifurcate posteriorly; ostiole of metasternal gland elliptical, peritreme spout-shaped; evaporatorium covering most of mesopleura and metepisternum. Posterolateral angle of each connexivum acute.</paragraph>
<paragraph id="8E261C15C10AFFE250AFFB3BFE29495F" blockId="1.[151,1437,646,1427]" pageId="1" pageNumber="562">
<emphasis id="BCEDC007C10AFFE250AFFB3BFECF48E7" bold="true" box="[199,380,1186,1211]" inLineHeading="true" pageId="1" pageNumber="562" reason="1">Male genitalia.</emphasis>
Ventral wall of pygophore impressed. Parameres covering segment X basally. Segment X elongate or quadrate; if quadrate, each posterolateral angle of segment X produced posteriorly. Phallus: conjunctiva absent, vesica reduced.
</paragraph>
<paragraph id="8E261C15C10AFFE250AFFA97FBF94917" blockId="1.[151,1437,646,1427]" pageId="1" pageNumber="562">
<emphasis id="BCEDC007C10AFFE250AFFA97FE2A497B" bold="true" box="[199,409,1294,1319]" inLineHeading="true" pageId="1" pageNumber="562" reason="1">Female genitalia.</emphasis>
Gonocoxites 8 covering most of gonocoxites 9. Each laterotergite 8 with a spiracle; posterior margin of laterotergite 8 with small process. Gonocoxites 9 flat or swollen.
</paragraph>
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<subSubSection id="C6834F9EC10AFFE250AFFACFFC3C49CF" pageId="1" pageNumber="562" type="distribution">
<paragraph id="8E261C15C10AFFE250AFFACFFC3C49CF" blockId="1.[151,1437,646,1427]" pageId="1" pageNumber="562">
<emphasis id="BCEDC007C10AFFE250AFFACFFED34933" bold="true" box="[199,352,1366,1391]" inLineHeading="true" pageId="1" pageNumber="562" reason="1">Distribution.</emphasis>
South America: Suriname (Paramaribo), French Guiana, Colombia (Amazonas), Ecuador (Napo), Peru (Loreto), Brazil (Amapá, Amazonas, Pará, Rondônia).
</paragraph>
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<subSubSection id="C6834F9EC10AFFE250FFFA7FFDCE4A5C" box="[151,637,1510,1536]" pageId="1" pageNumber="562" type="discussion">
<paragraph id="8E261C15C10AFFE250FFFA7FFDCE4A5C" blockId="1.[151,637,1510,1536]" box="[151,637,1510,1536]" pageId="1" pageNumber="562">
<heading id="D56EAB79C10AFFE250FFFA7FFDCE4A5C" bold="true" box="[151,637,1510,1536]" fontSize="11" level="1" pageId="1" pageNumber="562" reason="1">
<emphasis id="BCEDC007C10AFFE250FFFA7FFDCE4A5C" bold="true" box="[151,637,1510,1536]" pageId="1" pageNumber="562">
Key to species of
<emphasis id="BCEDC007C10AFFE25105FA7FFE784A5C" bold="true" box="[365,459,1510,1536]" italics="true" pageId="1" pageNumber="562">Elanela</emphasis>
<bibRefCitation id="EA0861E4C10AFFE251BBFA7FFDCE4A5C" author="Rolston, L. H. &amp; McDonald, F. J. D. &amp; Thomas, D. B." box="[467,637,1510,1536]" journalOrPublisher="Journal of the New York Entomological Society" pageId="1" pageNumber="562" pagination="120 - 132" part="88" refId="ref7617" refString="Rolston, L. H., McDonald, F. J. D. &amp; Thomas, D. B. (1980) A conspectus of Pentatomini genera of the Western hemisphere. Part I. (Hemiptrea: Pentatomidae). Journal of the New York Entomological Society, 88, 120 - 132." title="A conspectus of Pentatomini genera of the Western hemisphere. Part I. (Hemiptrea: Pentatomidae)" type="journal article" year="1980">Rolston, 1980</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C6834F9EC10AFFE150FFF9B5FA2F4E51" lastPageId="2" lastPageNumber="563" pageId="1" pageNumber="562" type="key">
<key id="FA084B24C10AFFE150FFF9B5FA2F4E51" lastPageId="2" lastPageNumber="563" pageId="1" pageNumber="562">
<keyStep id="356D04B0C10AFFE250FFF9B5FA284AC6" pageId="1" pageNumber="562">
<paragraph id="8E261C15C10AFFE250FFF9B5FA2F4A3C" blockId="1.[151,1436,1580,1690]" pageId="1" pageNumber="562">
<keyLead id="3568BF20C10AFFE250FFF9B5FA2F4A3C" pageId="1" pageNumber="562">
1. Legs yellow; few, scattered brown spots on each leg; urosternite VII black medially. Gonocoxites 8 as long as wide (
<figureCitation id="16A20090C10AFFE25556F9B5FA374A1E" box="[1342,1412,1580,1602]" captionStart="FIGURES 17–21" captionStartId="7.[151,264,1796,1819]" captionTargetBox="[396,1192,449,1774]" captionTargetId="figure@7.[394,1192,445,1776]" captionTargetPageId="7" captionText="FIGURES 17–21. Genitalia of species of Elanela Rolston, 1980. 17–18, pygophore in dorsal and ventral views, respectively, of E. rafaeli sp. nov.; 19–20, pygophore in dorsal and ventral views, respectively, of E. fernandesi sp. nov.; 21, female genital plates of E. scutomaculata sp. nov. in posteroventral view. Abbreviations: gc8, gonocoxites 8, gc9, gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; mdr, marginal process of dorsal rim; pa, paramere; X, abdominal segment X; vr, ventral rim. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/260980/files/figure.png" pageId="1" pageNumber="562">Fig. 21</figureCitation>
).................................................................................
<emphasis id="BCEDC007C10AFFE254F5F9D3FA2F4A3C" bold="true" box="[1181,1436,1610,1632]" pageId="1" pageNumber="562">
<taxonomicName id="49996796C10AFFE254F5F9D3FAFA4A3C" box="[1181,1353,1610,1632]" class="Insecta" family="Pentatomidae" genus="Elanela" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="scutomaculata" status="sp. nov.">
<emphasis id="BCEDC007C10AFFE254F5F9D3FAFA4A3C" bold="true" box="[1181,1353,1610,1632]" italics="true" pageId="1" pageNumber="562">E. scutomaculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7DE7D7CC10AFFE25527F9D3FA2F4A3C" box="[1359,1436,1610,1632]" pageId="1" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8E261C15C10AFFE250FFF9FEFA284AC6" blockId="1.[151,1436,1580,1690]" pageId="1" pageNumber="562">
<keyLead id="3568BF20C10AFFE250FFF9FEFA284AC6" pageId="1" pageNumber="562">- Legs yellow; several brown spots on each leg; urosternite VII concolour with remaining urosternites. Gonocoxites 8 longer than wide or wider than long............................................................................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="356D04B0C10AFFE250FFF938FA284B16" pageId="1" pageNumber="562">
<paragraph id="8E261C15C10AFFE250FFF938FA2F4AAE" blockId="1.[151,1436,1697,1866]" pageId="1" pageNumber="562">
<keyLead id="3568BF20C10AFFE250FFF938FA2F4AAE" pageId="1" pageNumber="562">
2(1). Pygophore with marginal processes on each side of dorsal rim medially, each process projected posteriorly and developed inwards genital cup; segment X elongate, carinate dorsally, sinuous posteriorly, and blunt apically (
<figureCitation id="16A20090C10AFFE254DAF926FB494A89" box="[1202,1274,1727,1749]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1640,1663]" captionTargetBox="[153,1435,703,1617]" captionTargetId="figure@6.[151,1435,700,1619]" captionTargetPageId="6" captionText="FIGURES 13–16. Genitalia of Elanela diamphidiosa sp. nov. 13–14, pygophore in dorsal and ventral views, respectively; 15, female genital plates in posteroventral view; 16, female receptaculum seminis and ausenwand. Abbreviations: cs, capsula seminalis; dr, ductus receptaculli; g9, gonapophyses 9; gc8, gonocoxites 8, gc9, gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; mdr, marginal process of dorsal rim; pa, paramere; X, abdominal segment X; tvi, thickening of vaginal intima; va, vesicular area; vr, ventral rim. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/260979/files/figure.png" pageId="1" pageNumber="562">Fig. 13</figureCitation>
). Gonocoxites 9 flat; distal margin of gonocoxites 9 deeply notched medially (
<figureCitation id="16A20090C10AFFE25347F945FCC64AAE" box="[815,885,1756,1778]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1640,1663]" captionTargetBox="[153,1435,703,1617]" captionTargetId="figure@6.[151,1435,700,1619]" captionTargetPageId="6" captionText="FIGURES 13–16. Genitalia of Elanela diamphidiosa sp. nov. 13–14, pygophore in dorsal and ventral views, respectively; 15, female genital plates in posteroventral view; 16, female receptaculum seminis and ausenwand. Abbreviations: cs, capsula seminalis; dr, ductus receptaculli; g9, gonapophyses 9; gc8, gonocoxites 8, gc9, gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; mdr, marginal process of dorsal rim; pa, paramere; X, abdominal segment X; tvi, thickening of vaginal intima; va, vesicular area; vr, ventral rim. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/260979/files/figure.png" pageId="1" pageNumber="562">Fig. 15</figureCitation>
)........................
<emphasis id="BCEDC007C10AFFE254C0F945FA2F4AAE" bold="true" box="[1192,1436,1756,1778]" pageId="1" pageNumber="562">
<taxonomicName id="49996796C10AFFE254C0F945FAFA4AAE" box="[1192,1353,1756,1778]" class="Insecta" family="Pentatomidae" genus="Elanela" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="diamphidiosa" status="sp. nov.">
<emphasis id="BCEDC007C10AFFE254C0F945FAFA4AAE" bold="true" box="[1192,1353,1756,1778]" italics="true" pageId="1" pageNumber="562">E. diamphidiosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7DE7D7CC10AFFE25527F945FA2F4AAE" box="[1359,1436,1756,1778]" pageId="1" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8E261C15C10AFFE250FFF963FA284B16" blockId="1.[151,1436,1697,1866]" pageId="1" pageNumber="562">
<keyLead id="3568BF20C10AFFE250FFF963FA284B16" pageId="1" pageNumber="562">- Pygophore without marginal processes on each side of dorsal rim; segment X quadrate, convex or concave dorsally, each posterolateral angle of segment X produced posteriorly. Gonocoxites 9 flat or strongly convex; distal margin of gonocoxites 9 shallowly notched medially or straight.................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="356D04B0C10AFFE250FFF8C8FA284B9C" pageId="1" pageNumber="562">
<paragraph id="8E261C15C10AFFE250FFF8C8FA2F4BD9" blockId="1.[151,1436,1873,1984]" pageId="1" pageNumber="562">
<keyLead id="3568BF20C10AFFE250FFF8C8FA2F4BD9" pageId="1" pageNumber="562">
3(2). Distal half of femur of metathoracic legs black. Ventral rim of pygophore with acute process medially (
<figureCitation id="16A20090C10AFFE254D0F8C8FAB34B3B" box="[1208,1280,1873,1895]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1865,1888]" captionTargetBox="[151,1436,987,1842]" captionTargetId="figure@4.[151,1436,983,1843]" captionTargetPageId="4" captionText="FIGURES 9–12. Genitalia of Elanela nigrofemorata sp. nov. 9–10, pygophore in dorsal and ventral views, respectively; 11, female genital plates in posteroventral view; 12, female receptaculum seminis and ausenwand. Abbreviations: cs, capsula seminalis; dr, ductus receptaculli; g9, gonapophyses 9; gc8, gonocoxites 8, gc9, gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; pa, paramere; X, abdominal segment X; tvi, thickening of vaginal intima; va, vesicular area; vr, ventral rim. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/260978/files/figure.png" pageId="1" pageNumber="562">Fig. 10</figureCitation>
). Gonocoxites 9 swollen, strongly convex, higher in profile than gonocoxites 8 (
<figureCitation id="16A20090C10AFFE25329F8F6FC364BD9" box="[833,901,1903,1925]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1865,1888]" captionTargetBox="[151,1436,987,1842]" captionTargetId="figure@4.[151,1436,983,1843]" captionTargetPageId="4" captionText="FIGURES 9–12. Genitalia of Elanela nigrofemorata sp. nov. 9–10, pygophore in dorsal and ventral views, respectively; 11, female genital plates in posteroventral view; 12, female receptaculum seminis and ausenwand. Abbreviations: cs, capsula seminalis; dr, ductus receptaculli; g9, gonapophyses 9; gc8, gonocoxites 8, gc9, gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; pa, paramere; X, abdominal segment X; tvi, thickening of vaginal intima; va, vesicular area; vr, ventral rim. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/260978/files/figure.png" pageId="1" pageNumber="562">Fig. 11</figureCitation>
)......................
<emphasis id="BCEDC007C10AFFE254F7F8F6FA2F4BD9" bold="true" box="[1183,1436,1903,1925]" pageId="1" pageNumber="562">
<taxonomicName id="49996796C10AFFE254F7F8F6FAFA4BD9" box="[1183,1353,1903,1925]" class="Insecta" family="Pentatomidae" genus="Elanela" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="nigrofemorata" status="sp. nov.">
<emphasis id="BCEDC007C10AFFE254F7F8F6FAFA4BD9" bold="true" box="[1183,1353,1903,1925]" italics="true" pageId="1" pageNumber="562">E. nigrofemorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7DE7D7CC10AFFE25527F8F6FA2F4BD9" box="[1359,1436,1903,1925]" pageId="1" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8E261C15C10AFFE250FFF814FA284B9C" blockId="1.[151,1436,1873,1984]" pageId="1" pageNumber="562">
<keyLead id="3568BF20C10AFFE250FFF814FA284B9C" pageId="1" pageNumber="562">- Distal half of femur of each metathoracic leg castaneous with brown spots. Ventral rim of pygophore lacking acute process medially. Gonocoxites 9 flat, if swollen, lower than gonocoxites 8.............................................. 4</keyLead>
</paragraph>
</keyStep>
<paragraph id="8E261C15C10AFFE150FFF85EFA284CF0" blockId="1.[151,1435,1991,2013]" lastBlockId="2.[151,1435,151,173]" lastPageId="2" lastPageNumber="563" pageId="1" pageNumber="562">
<keyStep id="356D04B0C10AFFE150FFF85EFA284CF0" lastPageId="2" lastPageNumber="563" pageId="1" pageNumber="562">
<keyLead id="3568BF20C10AFFE150FFF85EFA284CF0" lastPageId="2" lastPageNumber="563" pageId="1" pageNumber="562">4(3). Segment X surpassing posterior margin of pygophore........................................................ 5 - Segment X not surpassing posterior margin of pygophore.................................................... 7</keyLead>
</keyStep>
</paragraph>
<keyStep id="356D04B0C109FFE150FFFF2DFA284D7E" pageId="2" pageNumber="563">
<paragraph id="8E261C15C109FFE150FFFF2DFA284CB4" blockId="2.[151,1436,180,290]" pageId="2" pageNumber="563">
<keyLead id="3568BF20C109FFE150FFFF2DFA284CB4" pageId="2" pageNumber="563">
5(4). Parameres acute apically; distal margin of segment X with deep U-shaped notch. Posterior margins of gonocoxites 8 sinuate, concave medially; apex of each laterotergite 9 acute......................................
<taxonomicName id="49996796C109FFE154CEFF4BFA284CB4" authority="Grazia &amp; Greve" authorityName="Grazia &amp; Greve" box="[1190,1435,210,232]" class="Insecta" family="Pentatomidae" genus="Elanela" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="species" species="jordi">
<emphasis id="BCEDC007C109FFE154CEFF4BFA284CB4" bold="true" box="[1190,1435,210,232]" pageId="2" pageNumber="563">
<emphasis id="BCEDC007C109FFE154CEFF4BFB424CB4" bold="true" box="[1190,1265,210,232]" italics="true" pageId="2" pageNumber="563">E. jordi</emphasis>
Grazia &amp; Greve
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8E261C15C109FFE150FFFF76FA284D7E" blockId="2.[151,1436,180,290]" pageId="2" pageNumber="563">
<keyLead id="3568BF20C109FFE150FFFF76FA284D7E" pageId="2" pageNumber="563">- Parameres quadrate apically; distal margin of segment X with deep V-shaped notch. Posterior margins of gonocoxites 8 straight; posterior margin of gonocoxites 9 sinuate; apex of each laterotergite 9 obtuse.............................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="356D04B0C109FFE150FFFEB0FA2F4DC4" pageId="2" pageNumber="563">
<paragraph id="8E261C15C109FFE150FFFEB0FA284D01" blockId="2.[151,1436,297,408]" pageId="2" pageNumber="563">
<keyLead id="3568BF20C109FFE150FFFEB0FA284D01" pageId="2" pageNumber="563">
6(5). Pygophore globose; ventral rim rounded laterally; parameres with apices not juxtaposed. Posterior margin of gonocoxites 9 sinuate; laterotergites 9 surpassing tergite VIII.................................................
<taxonomicName id="49996796C109FFE15481FEDEFA284D01" authority="Rolston" authorityName="Rolston" box="[1257,1435,327,349]" class="Insecta" family="Pentatomidae" genus="Elanela" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="species" species="hevera">
<emphasis id="BCEDC007C109FFE15481FEDEFA284D01" bold="true" box="[1257,1435,327,349]" pageId="2" pageNumber="563">
<emphasis id="BCEDC007C109FFE15481FEDEFAF44D01" bold="true" box="[1257,1351,327,349]" italics="true" pageId="2" pageNumber="563">E. hevera</emphasis>
Rolston
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8E261C15C109FFE150FFFEFCFA2F4DC4" blockId="2.[151,1436,297,408]" pageId="2" pageNumber="563">
<keyLead id="3568BF20C109FFE150FFFEFCFA2F4DC4" pageId="2" pageNumber="563">
- Pygophore quadrate; ventral rim triangular laterally; parameres with apices juxtaposed. Posterior margin of gonocoxites 9 convex; laterotergites 9 not surpassing tergite VIII.......................................
<taxonomicName id="49996796C109FFE154EAFE1BFA2F4DC4" authority="Grazia &amp; Silva" authorityName="Grazia &amp; Silva" box="[1154,1436,386,408]" class="Insecta" family="Pentatomidae" genus="Elanela" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="species" species="kerzhneri">
<emphasis id="BCEDC007C109FFE154EAFE1BFA2F4DC4" bold="true" box="[1154,1436,386,408]" pageId="2" pageNumber="563">
<emphasis id="BCEDC007C109FFE154EAFE1BFB4F4DC4" bold="true" box="[1154,1276,386,408]" italics="true" pageId="2" pageNumber="563">E. kerzhneri</emphasis>
Grazia &amp; Silva
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="356D04B0C109FFE150FFFE06FA2F4E51" pageId="2" pageNumber="563">
<keyLead id="3568BF20C109FFE150FFFE06FA2F4D8E" pageId="2" pageNumber="563">
<paragraph id="8E261C15C109FFE150FFFE06FA244DE8" blockId="2.[151,1436,415,525]" box="[151,1431,415,437]" pageId="2" pageNumber="563">
7(4). Parameres reduced, scarcely visible dorsally; projections of ventral rim of pygophore stout, quadrate at apex (
<figureCitation id="16A20090C109FFE15493FE06FADB4DE9" box="[1275,1384,415,437]" captionStart="FIGURES 17–21" captionStartId="7.[151,264,1796,1819]" captionTargetBox="[396,1192,449,1774]" captionTargetId="figure@7.[394,1192,445,1776]" captionTargetPageId="7" captionText="FIGURES 17–21. Genitalia of species of Elanela Rolston, 1980. 17–18, pygophore in dorsal and ventral views, respectively, of E. rafaeli sp. nov.; 19–20, pygophore in dorsal and ventral views, respectively, of E. fernandesi sp. nov.; 21, female genital plates of E. scutomaculata sp. nov. in posteroventral view. Abbreviations: gc8, gonocoxites 8, gc9, gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; mdr, marginal process of dorsal rim; pa, paramere; X, abdominal segment X; vr, ventral rim. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/260980/files/figure.png" pageId="2" pageNumber="563">Figs 17–18</figureCitation>
)...
</paragraph>
<paragraph id="8E261C15C109FFE1508BFE24FA2F4D8E" blockId="2.[151,1436,415,525]" box="[227,1436,444,466]" pageId="2" pageNumber="563">
......................................................................................
<emphasis id="BCEDC007C109FFE15484FE25FA2F4D8E" bold="true" box="[1260,1436,444,466]" pageId="2" pageNumber="563">
<taxonomicName id="49996796C109FFE15484FE25FAF94D8E" box="[1260,1354,444,466]" class="Insecta" family="Pentatomidae" genus="Elanela" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="species" species="rafaeli" status="sp. nov.">
<emphasis id="BCEDC007C109FFE15484FE25FAF94D8E" bold="true" box="[1260,1354,444,466]" italics="true" pageId="2" pageNumber="563">E. rafaeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7DE7D7CC109FFE15527FE25FA2F4D8E" box="[1359,1436,444,466]" pageId="2" pageNumber="563" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</keyLead>
<paragraph id="8E261C15C109FFE150FFFE43FA2F4E51" blockId="2.[151,1436,415,525]" pageId="2" pageNumber="563">
<keyLead id="3568BF20C109FFE150FFFE43FA2F4E51" pageId="2" pageNumber="563">
- Parameres scythe-like, projected mesially, not reaching posterior margin of pygophore; projections of ventral rim of pygophore rounded at apex (
<figureCitation id="16A20090C109FFE151EFFE6EFE474E51" box="[391,500,503,525]" captionStart="FIGURES 17–21" captionStartId="7.[151,264,1796,1819]" captionTargetBox="[396,1192,449,1774]" captionTargetId="figure@7.[394,1192,445,1776]" captionTargetPageId="7" captionText="FIGURES 17–21. Genitalia of species of Elanela Rolston, 1980. 17–18, pygophore in dorsal and ventral views, respectively, of E. rafaeli sp. nov.; 19–20, pygophore in dorsal and ventral views, respectively, of E. fernandesi sp. nov.; 21, female genital plates of E. scutomaculata sp. nov. in posteroventral view. Abbreviations: gc8, gonocoxites 8, gc9, gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; mdr, marginal process of dorsal rim; pa, paramere; X, abdominal segment X; vr, ventral rim. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/260980/files/figure.png" pageId="2" pageNumber="563">Figs 19–20</figureCitation>
)..........................................................
<emphasis id="BCEDC007C109FFE154ACFE6EFA2F4E51" bold="true" box="[1220,1436,503,525]" pageId="2" pageNumber="563">
<taxonomicName id="49996796C109FFE154ACFE6EFAFA4E51" box="[1220,1353,503,525]" class="Insecta" family="Pentatomidae" genus="Elanela" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="species" species="fernandesi" status="sp. nov.">
<emphasis id="BCEDC007C109FFE154ACFE6EFAFA4E51" bold="true" box="[1220,1353,503,525]" italics="true" pageId="2" pageNumber="563">E. fernandesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7DE7D7CC109FFE15527FE6EFA2F4E51" box="[1359,1436,503,525]" pageId="2" pageNumber="563" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</subSection>
</treatment>
</document>
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