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<document id="927885042FBDA710436A68568F8548FF" ID-DOI="10.5281/zenodo.202371" ID-GBIF-Dataset="f74f4953-be75-44cb-89e1-9e5c822d6579" ID-ISSN="1175-5326" ID-Zenodo-Dep="202371" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460316144790" checkinUser="plazi" docAuthor="Komatsu, Hironori &amp; Ng, Peter K. L." docDate="2011" docId="88260608E62B986FFF7096164400FDAF" docLanguage="en" docName="zt02881p038.pdf" docOrigin="Zootaxa 2881" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Kabutos Komatsu &amp; Ng, 2011, n. gen." docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="741F7E70E62A986DFFE797014636FFF0" masterDocTitle="Kabutos, a new genus for the western Pacific leucosiid crab, Merocryptus durandi Serène, 1955 (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="38" masterPageNumber="31" pageNumber="32" updateTime="1698255129783" updateUser="plazi">
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<mods:title id="95B97E2998C7966182A9AED241706B70">Kabutos, a new genus for the western Pacific leucosiid crab, Merocryptus durandi Serène, 1955 (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:namePart id="E4B5B02FDDA5EB1D009B27E8685D197C">Komatsu, Hironori</mods:namePart>
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<mods:namePart id="AD4752B1E60B6C49F8EF97E7B088EBCD">Ng, Peter K. L.</mods:namePart>
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<mods:date id="DEA900A2D00ABF695853174CD6B48F47">2011</mods:date>
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<treatment id="88260608E62B986FFF7096164400FDAF" ID-DOI="http://doi.org/10.5281/zenodo.6184775" ID-GBIF-Taxon="119401718" ID-Zenodo-Dep="6184775" LSID="urn:lsid:plazi:treatment:88260608E62B986FFF7096164400FDAF" httpUri="http://treatment.plazi.org/id/88260608E62B986FFF7096164400FDAF" lastPageId="2" lastPageNumber="33" pageId="1" pageNumber="32">
<subSubSection id="4895E495E62B986CFF7096164765FEC1" box="[151,339,279,305]" pageId="1" pageNumber="32" type="nomenclature">
<paragraph id="0030B71EE62B986CFF7096164765FEC1" blockId="1.[151,339,279,305]" box="[151,339,279,305]" pageId="1" pageNumber="32">
<heading id="5B780072E62B986CFF7096164765FEC1" bold="true" box="[151,339,279,305]" fontSize="11" level="1" pageId="1" pageNumber="32" reason="1">
<emphasis id="32FB6B0CE62B986CFF7096164765FEC1" bold="true" box="[151,339,279,305]" pageId="1" pageNumber="32">
<taxonomicName id="C78FCC9DE62B986CFF70961646CDFEC1" ID-CoL="7PLYN" box="[151,251,279,305]" class="Malacostraca" family="Leucosiidae" genus="Kabutos" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="32FB6B0CE62B986CFF70961646CDFEC1" bold="true" box="[151,251,279,305]" italics="true" pageId="1" pageNumber="32">Kabutos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29C8D677E62B986CFEE696164765FEC1" box="[257,339,279,305]" pageId="1" pageNumber="32" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="4895E495E62B986CFF7096564418FD27" pageId="1" pageNumber="32" type="diagnosis">
<paragraph id="0030B71EE62B986CFF7096564418FD27" blockId="1.[151,1436,343,1628]" pageId="1" pageNumber="32">
<emphasis id="32FB6B0CE62B986CFF7096564727FE80" bold="true" box="[151,273,343,368]" pageId="1" pageNumber="32">Diagnosis.</emphasis>
Carapace distinctly broader than long, dorsal surface with distinct median, epibranchial ridges; front produced; mesogastric region longitudinally positioned, broadly convex; gastro-cardiac region raised, with 2 gastric tubercles, single cardiac tubercle; intestinal region produced; branchial region expanded laterally, gently sloping anterolaterally from ridge. Basal segment of antennule not occupying entire fossa when closed. Basal segment of antenna lodged in orbital hiatus. Maxillule with tongue-shaped endopod. First, second maxillipeds with exopodal flagellum. Third maxilliped exopod with triangular tooth at proximal end of lateral margin. Chelipeds relatively slender, not elongate. Ambulatory legs relatively short. Thoracic sternum covered with numerous rounded granules; sutures between s3/4, s4/5, s5/6, s6/7, s7/8 interrupted medially; medial suture absent. Male abdomen with first, second somites free, third to fifth somites completely fused, sixth somite, telson free. Female abdomen with second, third somites free, fourth to sixth somites fused, telson free. G1 relatively slender, straight, compressed. G2 short, about 0.3 times as long as G1.
</paragraph>
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<subSubSection id="4895E495E62B986CFF2095E545AEFD0D" box="[199,920,739,765]" pageId="1" pageNumber="32" type="materials_examined">
<paragraph id="0030B71EE62B986CFF2095E545AEFD0D" blockId="1.[151,1436,343,1628]" box="[199,920,739,765]" pageId="1" pageNumber="32">
<emphasis id="32FB6B0CE62B986CFF2095E54754FD0C" bold="true" box="[199,354,739,764]" pageId="1" pageNumber="32">
<typeStatus id="DF3409BCE62B986CFF2095E54734FD0C" box="[199,258,740,764]" pageId="1" pageNumber="32">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="C78FCC9DE62B986CFE8E95E444C5FD0C" authority="Serene, 1955" authorityName="Serene" authorityYear="1955" box="[361,755,740,764]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="durandi">
<emphasis id="32FB6B0CE62B986CFE8E95E44461FD0C" box="[361,599,740,764]" italics="true" pageId="1" pageNumber="32">Merocryptus durandi</emphasis>
<bibRefCitation id="641ECAEFE62B986CFDBA95E544C5FD0C" author="Serene" box="[605,755,740,764]" pageId="1" pageNumber="32" refString="Serene, R. (1955) Sur quelques especes rares de Brachyures (Leucosidae) de l'Indo-Pacifique (2 e partie). Treubia, 23 (1), 137 - 218, pls. 6 - 11." type="journal article" year="1955">Serène, 1955</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="4895E495E62B986CFF2094094492FCB4" pageId="1" pageNumber="32" type="etymology">
<paragraph id="0030B71EE62B986CFF2094094492FCB4" blockId="1.[151,1436,343,1628]" pageId="1" pageNumber="32">
<emphasis id="32FB6B0CE62B986CFF2094094778FCD0" bold="true" box="[199,334,776,800]" pageId="1" pageNumber="32">Etymology.</emphasis>
The name of the genus is derived from
<emphasis id="32FB6B0CE62B986CFCF494094569FCD0" box="[787,863,776,800]" italics="true" pageId="1" pageNumber="32">kabuto</emphasis>
, the Japanese word for a Samurai helmet, alluding to the form of the carapace. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="4895E495E62B986CFF20945145C9F81A" pageId="1" pageNumber="32" type="discussion">
<paragraph id="0030B71EE62B986CFF2094514767FB5C" blockId="1.[151,1436,343,1628]" pageId="1" pageNumber="32">
<emphasis id="32FB6B0CE62B986CFF209451470DFC98" bold="true" box="[199,315,848,872]" pageId="1" pageNumber="32">Remarks.</emphasis>
<bibRefCitation id="641ECAEFE62B986CFEA4945147D4FC98" author="Serene" box="[323,482,848,872]" pageId="1" pageNumber="32" refString="Serene, R. (1955) Sur quelques especes rares de Brachyures (Leucosidae) de l'Indo-Pacifique (2 e partie). Treubia, 23 (1), 137 - 218, pls. 6 - 11." type="journal article" year="1955">Serène (1955)</bibRefCitation>
discussed the characters and composition of
<taxonomicName id="C78FCC9DE62B986CFC0094504243FC98" box="[999,1141,849,872]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="32FB6B0CE62B986CFC0094504243FC98" box="[999,1141,849,872]" italics="true" pageId="1" pageNumber="32">Merocryptus</emphasis>
</taxonomicName>
at length. He commented that while there existed substantial variation in the form of the antero- and posterolateral margins and the structure of the carapace regions in the four species (including
<taxonomicName id="C78FCC9DE62B986CFD0A9498455AFC40" box="[749,876,920,944]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="durandi">
<emphasis id="32FB6B0CE62B986CFD0A9498455AFC40" box="[749,876,920,944]" italics="true" pageId="1" pageNumber="32">M. durandi</emphasis>
</taxonomicName>
), they are nevertheless related and the differences are only in degree. The main generic characters according to
<bibRefCitation id="641ECAEFE62B986CFCA494BD45D7FC24" author="Serene" box="[835,993,956,980]" pageId="1" pageNumber="32" refString="Serene, R. (1955) Sur quelques especes rares de Brachyures (Leucosidae) de l'Indo-Pacifique (2 e partie). Treubia, 23 (1), 137 - 218, pls. 6 - 11." type="journal article" year="1955">Serène (1955)</bibRefCitation>
are the adjoined base of the orbit to the elongated antennular peduncle; the 5-lobed lateral margins of carapace; the well developed pterygostomial region forming a distinct tooth or lobe; and the swollen or raised branchial regions forming a prominent lateral lobe or projection on the carapace. He noted that the diagnosis for the genus by
<bibRefCitation id="641ECAEFE62B986CFC2D932942C6FBB0" author="Milne-Edwards" box="[970,1264,1064,1088]" pageId="1" pageNumber="32" refString="Milne-Edwards, A. (1873) Description de quelques Crustaces nouveaux ou peu connus provenant du Musee de M. C. Godeffroy. Journal des Museum Godeffroy, 1 (4), 77 - 88, pls. 1, 2 [pages 1 - 12, plates 12, 13 on separate]." type="journal article" year="1873">A. Milne-Edwards (1873)</bibRefCitation>
, later modified by
<bibRefCitation id="641ECAEFE62B986CFF5B934D4762FB94" author="Miers" box="[188,340,1100,1124]" pageId="1" pageNumber="32" refString="Miers, E. J. (1886) Report on the Brachyura collected by H. M. S. Challenger during the years 1873 - 76. In: Murray, J. (Ed.), Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876 under the command of Captain George S. Nares, N. R., F. R. S. and the late Captain Frank Tourle Thomson, R. N. prepared under the Superintendence of the late Sir C. Wyville Thomson, Knt., F. R. S. &amp; c. Regius Professor of Natural history in the University of Edinburgh of the civilian scientific staff on board and now of John Murray one of the naturalists of the Expedition. Zoology - Vol. XVII, Published by Order of Her Majesty's Government. HMSO, London, Edinburgh and Dublin, pp. i - l, 1 - 362, pls. 1 - 29." type="book chapter" year="1886">Miers (1886)</bibRefCitation>
, was generally effective, but with the description of
<taxonomicName id="C78FCC9DE62B986CFC50934C420EFB94" box="[951,1080,1100,1124]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="durandi">
<emphasis id="32FB6B0CE62B986CFC50934C420EFB94" box="[951,1080,1100,1124]" italics="true" pageId="1" pageNumber="32">M. durandi</emphasis>
</taxonomicName>
, the posterior carapace margin may be entire (not with two teeth or projections) and the anterior distal margin of the endostome may be emarginated (not entire).
</paragraph>
<paragraph id="0030B71EE62B986CFF2293B94786F9AC" blockId="1.[151,1436,343,1628]" pageId="1" pageNumber="32">
Comparison of the present material with the
<typeStatus id="DF3409BCE62B986CFD2B93B844CAFB21" box="[716,764,1209,1233]" pageId="1" pageNumber="32">type</typeStatus>
species,
<taxonomicName id="C78FCC9DE62B986CFC8093B84217FB20" box="[871,1057,1208,1232]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="lambriformis">
<emphasis id="32FB6B0CE62B986CFC8093B84217FB20" box="[871,1057,1208,1232]" italics="true" pageId="1" pageNumber="32">M. lambriformis</emphasis>
</taxonomicName>
, however, revealed major differences between the two species, indicating they are not congeneric. In
<taxonomicName id="C78FCC9DE62B986CFC4493DC4217FB04" box="[931,1057,1244,1268]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="durandi">
<emphasis id="32FB6B0CE62B986CFC4493DC4217FB04" box="[931,1057,1244,1268]" italics="true" pageId="1" pageNumber="32">M. durandi</emphasis>
</taxonomicName>
, the basal segment of the antenna fills the orbital hiatus (leaves a gape in the orbital hiatus in
<taxonomicName id="C78FCC9DE62B986CFCCF920045E8FAE8" box="[808,990,1280,1304]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="lambriformis">
<emphasis id="32FB6B0CE62B986CFCCF920045E8FAE8" box="[808,990,1280,1304]" italics="true" pageId="1" pageNumber="32">M. lambriformis</emphasis>
</taxonomicName>
), the exopod of the third maxilliped has a well developed tooth on the lateral margin (absent in
<taxonomicName id="C78FCC9DE62B986CFCF5922445FCFACC" box="[786,970,1316,1340]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="lambriformis">
<emphasis id="32FB6B0CE62B986CFCF5922445FCFACC" box="[786,970,1316,1340]" italics="true" pageId="1" pageNumber="32">M. lambriformis</emphasis>
</taxonomicName>
), and the arrangement of the fused male and female abdominal somites are quite different (Table 1). These characters, together with the different carapace features, argue for the establishment of a separate genus for
<taxonomicName id="C78FCC9DE62B986CFCB3926C4241FA74" authority="Serene, 1955" authorityName="Serene" authorityYear="1955" box="[852,1143,1388,1412]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="durandi">
<emphasis id="32FB6B0CE62B986CFCB3926C45E3FA74" box="[852,981,1388,1412]" italics="true" pageId="1" pageNumber="32">M. durandi</emphasis>
<bibRefCitation id="641ECAEFE62B986CFC38926D4241FA74" author="Serene" box="[991,1143,1388,1412]" pageId="1" pageNumber="32" refString="Serene, R. (1955) Sur quelques especes rares de Brachyures (Leucosidae) de l'Indo-Pacifique (2 e partie). Treubia, 23 (1), 137 - 218, pls. 6 - 11." type="journal article" year="1955">Serène, 1955</bibRefCitation>
</taxonomicName>
. On the basis of the
<typeStatus id="DF3409BCE62B986CFA8B926C43AAFA75" box="[1388,1436,1389,1413]" pageId="1" pageNumber="32">type</typeStatus>
descriptions of the Atlantic
<taxonomicName id="C78FCC9DE62B986CFE349290446FFA58" box="[467,601,1424,1448]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="boletifer">
<emphasis id="32FB6B0CE62B986CFE349290446FFA58" box="[467,601,1424,1448]" italics="true" pageId="1" pageNumber="32">M. boletifer</emphasis>
</taxonomicName>
and
<taxonomicName id="C78FCC9DE62B986CFD7792904514FA58" box="[656,802,1424,1448]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="obsoletus">
<emphasis id="32FB6B0CE62B986CFD7792904514FA58" box="[656,802,1424,1448]" italics="true" pageId="1" pageNumber="32">M. obsoletus</emphasis>
</taxonomicName>
(see also
<bibRefCitation id="641ECAEFE62B986CFC75929142ABFA58" author="Guinot" box="[914,1181,1424,1448]" pageId="1" pageNumber="32" refString="Guinot, D. &amp; Ribeiro, A. (1962) Sur une collection de Crustaces Brachyoures des Iles du Cap-Vert et de 1 ' Angola. Memorias da Junta de Investigac o es do Ultramar, (2) 40, 9 - 89, pls. 1 - 4." type="journal article" year="1962">Guinot &amp; Ribeiro 1962</bibRefCitation>
;
<bibRefCitation id="641ECAEFE62B986CFB4E929146E4FA3C" author="Manning" pageId="1" pageNumber="32" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contribution of Zoology, 306, 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis 1981</bibRefCitation>
), their key generic diagnostic characters are the same as those of
<taxonomicName id="C78FCC9DE62B986CFC2392B4424BFA3C" box="[964,1149,1460,1484]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="lambriformis">
<emphasis id="32FB6B0CE62B986CFC2392B4424BFA3C" box="[964,1149,1460,1484]" italics="true" pageId="1" pageNumber="32">M. lambriformis</emphasis>
</taxonomicName>
.
<bibRefCitation id="641ECAEFE62B986CFB6A92B443AAFA3C" author="Forest" box="[1165,1436,1460,1484]" pageId="1" pageNumber="32" refString="Forest, J. &amp; Guinot, D. (1966) Crustaces Decapodes: Brachyoures. In: Campagne de la Calypso dans le golfe de Guinee et aux iles Principe, Sao Tome et Annobon (1956), 16. Annales de I'Institut Oceanographique, Monaco, 44, 23 - 124." type="journal article" year="1966">Forest &amp; Guinot (1966)</bibRefCitation>
had suggested that the Atlantic genus
<taxonomicName id="C78FCC9DE62B986CFDD992D9456DFA00" authority="Doflein, 1904" authorityName="Doflein" authorityYear="1904" box="[574,859,1496,1520]" class="Malacostraca" family="Leucosiidae" genus="Atlantotlos" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="32FB6B0CE62B986CFDD992D9448FFA00" box="[574,697,1496,1520]" italics="true" pageId="1" pageNumber="32">Atlantotlos</emphasis>
<bibRefCitation id="641ECAEFE62B986CFD2792D9456DFA00" author="Doflein" box="[704,859,1496,1520]" pageId="1" pageNumber="32" refString="Doflein, F. (1904) Brachyura. In: Chun, C. (Ed.) Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899, Band VI. Verlag von Gustav Fischer, Jena, pp. i - xiv, 314, pls. 1 - 57." type="journal article" year="1904">Doflein, 1904</bibRefCitation>
</taxonomicName>
(
<typeStatus id="DF3409BCE62B986CFC8D92D945AAFA00" box="[874,924,1496,1520]" pageId="1" pageNumber="32">type</typeStatus>
and only species
<taxonomicName id="C78FCC9DE62B986CFB8592D943A2FA00" authority="Doflein, 1904" authorityName="Doflein" authorityYear="1904" box="[1122,1428,1496,1520]" class="Malacostraca" family="Leucosiidae" genus="Atlantotlos" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="rhombifer">
<emphasis id="32FB6B0CE62B986CFB8592D942C7FA00" box="[1122,1265,1496,1520]" italics="true" pageId="1" pageNumber="32">A. rhombifer</emphasis>
<bibRefCitation id="641ECAEFE62B986CFB1092D943A2FA00" author="Doflein" box="[1271,1428,1496,1520]" pageId="1" pageNumber="32" refString="Doflein, F. (1904) Brachyura. In: Chun, C. (Ed.) Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899, Band VI. Verlag von Gustav Fischer, Jena, pp. i - xiv, 314, pls. 1 - 57." type="journal article" year="1904">Doflein, 1904</bibRefCitation>
</taxonomicName>
) was synonymous with
<taxonomicName id="C78FCC9DE62B986CFE4D92FC440EF9E4" box="[426,568,1533,1556]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="32FB6B0CE62B986CFE4D92FC440EF9E4" box="[426,568,1533,1556]" italics="true" pageId="1" pageNumber="32">Merocryptus</emphasis>
</taxonomicName>
but
<bibRefCitation id="641ECAEFE62B986CFD9092FD45F3F9E4" author="Manning" box="[631,965,1532,1556]" pageId="1" pageNumber="32" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contribution of Zoology, 306, 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis (1981)</bibRefCitation>
argued that there were several notable differences, especially in the form of the G1, that indicated both were distinct genera. These differences are also valid for
<taxonomicName id="C78FCC9DE62B986CFF1A9145476DF9AC" box="[253,347,1604,1628]" class="Malacostraca" family="Leucosiidae" genus="Kabutos" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="32FB6B0CE62B986CFF1A9145476DF9AC" box="[253,347,1604,1628]" italics="true" pageId="1" pageNumber="32">Kabutos</emphasis>
</taxonomicName>
<emphasis id="32FB6B0CE62B986CFE8591454786F9AC" bold="true" box="[354,432,1604,1628]" pageId="1" pageNumber="32">
<taxonomicNameLabel id="29C8D677E62B986CFE8591454786F9AC" box="[354,432,1604,1628]" pageId="1" pageNumber="32" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="0030B71EE62B986CFF4A918F42CCF953" blockId="1.[173,1418,1677,2027]" box="[173,1274,1677,1700]" pageId="1" pageNumber="32">
<emphasis id="32FB6B0CE62B986CFF4A918F4736F953" bold="true" box="[173,256,1678,1699]" pageId="1" pageNumber="32">Table 1.</emphasis>
Comparison of
<taxonomicName id="C78FCC9DE62B986CFE41918F47CCF954" box="[422,506,1678,1700]" class="Malacostraca" family="Leucosiidae" genus="Kabutos" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="32FB6B0CE62B986CFE41918F47CCF954" box="[422,506,1678,1700]" italics="true" pageId="1" pageNumber="32">Kabutos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29C8D677E62B986CFDE6918F4470F953" box="[513,582,1678,1699]" pageId="1" pageNumber="32" rank="genus">n. gen.</taxonomicNameLabel>
with its allied genera. Abbreviations: R = fused somites; T = telson.
</paragraph>
<paragraph id="0030B71EE62B986CFE3B91AF432AF934" blockId="1.[173,1418,1677,2027]" box="[476,1308,1710,1732]" pageId="1" pageNumber="32">
<taxonomicName id="C78FCC9DE62B986CFE3B91AF4407F934" box="[476,561,1710,1732]" class="Malacostraca" family="Leucosiidae" genus="Kabutos" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="32FB6B0CE62B986CFE3B91AF4407F934" box="[476,561,1710,1732]" italics="true" pageId="1" pageNumber="32">Kabutos</emphasis>
</taxonomicName>
<emphasis id="32FB6B0CE62B986CFDD191AE4448F934" bold="true" box="[566,638,1711,1732]" pageId="1" pageNumber="32">
<taxonomicNameLabel id="29C8D677E62B986CFDD191AE4448F934" box="[566,638,1711,1732]" pageId="1" pageNumber="32" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
<emphasis id="32FB6B0CE62B986CFCFD91AE432AF934" box="[794,1308,1710,1732]" italics="true" pageId="1" pageNumber="32">
<taxonomicName id="C78FCC9DE62B986CFCFD91AE45ACF934" box="[794,922,1711,1732]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">Merocryptus</taxonomicName>
<taxonomicName id="C78FCC9DE62B986CFC2B91AF425DF934" box="[972,1131,1710,1732]" class="Malacostraca" family="Leucosiidae" genus="Merocryptoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">Merocryptoides</taxonomicName>
<taxonomicName id="C78FCC9DE62B986CFB2391AF432AF934" box="[1220,1308,1710,1732]" class="Malacostraca" family="Leucosiidae" genus="Oreotlos" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">Oreotlos</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="0030B71EE62B986CFF4A91CE435EF915" blockId="1.[173,1418,1677,2027]" box="[173,1384,1743,1765]" pageId="1" pageNumber="32">Orbital hiatus filled with basal segment of leaving a gape filled with basal filled with basal</paragraph>
<paragraph id="0030B71EE62B986CFE3B91F0435FF839" blockId="1.[173,1418,1677,2027]" pageId="1" pageNumber="32">
antenna segment of antenna segment of antenna Second maxilliped exopod with flagellum with flagellum without flagellum with flagellum Third maxilliped exopod with triangular tooth without tooth with triangular tooth without tooth Chela normal normal normal blade-like Male abdomen 1+2+R+6+T 2+R+T 1+2+R+T 1+2+R+6+T Female abdomen 2+3+R+T 1+2+R+T 2+R+T 2+3+R+T References
<bibRefCitation id="641ECAEFE62B986CFE3C90B24455F839" author="Serene" box="[475,611,1971,1993]" pageId="1" pageNumber="32" refString="Serene, R. (1955) Sur quelques especes rares de Brachyures (Leucosidae) de l'Indo-Pacifique (2 e partie). Treubia, 23 (1), 137 - 218, pls. 6 - 11." type="journal article" year="1955">Serène, 1955</bibRefCitation>
; present study
<bibRefCitation id="641ECAEFE62B986CFCFD90B24597F839" author="Serene" box="[794,929,1971,1993]" pageId="1" pageNumber="32" refString="Serene, R. (1955) Sur quelques especes rares de Brachyures (Leucosidae) de l'Indo-Pacifique (2 e partie). Treubia, 23 (1), 137 - 218, pls. 6 - 11." type="journal article" year="1955">Serène, 1955</bibRefCitation>
; Komatsu &amp; Takeda,
<bibRefCitation id="641ECAEFE62B986CFB2390B5435FF839" author="Tan" box="[1220,1385,1971,1993]" pageId="1" pageNumber="32" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1995) A revision of the Indo-Pacific genus Oreophorus Ruppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae). In: Richer de Forges B. (Ed.), Les fonds meubles des lagons de Nouvelle-Caledonie (Sedimentologie, benthos). ORSTOM, Paris, pp. 101 - 189." type="book chapter" year="1995">Tan &amp; Ng, 1995</bibRefCitation>
</paragraph>
<paragraph id="0030B71EE62B986CFCFD90D745C9F81A" blockId="1.[173,1418,1677,2027]" box="[794,1023,2005,2027]" pageId="1" pageNumber="32">present study 2001</paragraph>
</subSubSection>
<subSubSection id="4895E495E628986FFF22979947CCFEEC" pageId="2" pageNumber="33" type="reference_group">
<paragraph id="0030B71EE628986FFF22979947CCFEEC" blockId="2.[151,1436,151,608]" pageId="2" pageNumber="33">
<taxonomicName id="C78FCC9DE628986FFF2297994715FF40" ID-CoL="7PLYN" box="[197,291,152,176]" class="Malacostraca" family="Leucosiidae" genus="Kabutos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="32FB6B0CE628986FFF2297994715FF40" box="[197,291,152,176]" italics="true" pageId="2" pageNumber="33">Kabutos</emphasis>
</taxonomicName>
<emphasis id="32FB6B0CE628986FFECD9799474CFF40" bold="true" box="[298,378,152,176]" pageId="2" pageNumber="33">
<taxonomicNameLabel id="29C8D677E628986FFECD9799474CFF40" box="[298,378,152,176]" pageId="2" pageNumber="33" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
shows intermediate morphological features between
<taxonomicName id="C78FCC9DE628986FFC3297994323FF5F" ID-CoL="7PNSF" authority="Sakai, 1963" authorityName="Sakai" authorityYear="1963" box="[981,1301,151,176]" class="Malacostraca" family="Leucosiidae" genus="Merocryptoides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="32FB6B0CE628986FFC32979942B3FF40" box="[981,1157,152,176]" italics="true" pageId="2" pageNumber="33">Merocryptoides</emphasis>
<bibRefCitation id="641ECAEFE628986FFB6A97964323FF5F" author="Sakai" box="[1165,1301,151,175]" pageId="2" pageNumber="33" refString="Sakai, T. (1963) Description of two new genera and fourteen new species of Japanese crabs from the collection of his Majesty the Emperor of Japan. Crustaceana, 5, 213 - 233." type="journal article" year="1963">Sakai, 1963</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="C78FCC9DE628986FFAB597994704FF24" ID-CoL="7PNMR" authority="Ihle, 1918" authorityName="Ihle" authorityYear="1918" class="Malacostraca" family="Leucosiidae" genus="Oreotlos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="32FB6B0CE628986FFAB59799468EFF24" italics="true" pageId="2" pageNumber="33">Oreotlos</emphasis>
<bibRefCitation id="641ECAEFE628986FFF5997BD4704FF24" author="Ihle" box="[190,306,188,212]" pageId="2" pageNumber="33" refString="Ihle, J. E. W. (1918) Die Decapoda Brachyura der Siboga-Expedition. III. Oxystomata: Calappidae, Leucosiidae, Raninidae. Siboga Expeditie, 39 b 2, 1 - 322." type="journal article" year="1918">Ihle, 1918</bibRefCitation>
</taxonomicName>
, but can be differentiated from them by the structure of their second and third maxillipeds and chelae as well as the degree of fusion of male and female abdominal somites (see
<bibRefCitation id="641ECAEFE628986FFC1E97E14287FF07" author="Tan" box="[1017,1201,223,248]" pageId="2" pageNumber="33" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1995) A revision of the Indo-Pacific genus Oreophorus Ruppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae). In: Richer de Forges B. (Ed.), Les fonds meubles des lagons de Nouvelle-Caledonie (Sedimentologie, benthos). ORSTOM, Paris, pp. 101 - 189." type="book chapter" year="1995">Tan &amp; Ng 1995</bibRefCitation>
;
<bibRefCitation id="641ECAEFE628986FFB5997E146E5FEEC" author="Komatsu" pageId="2" pageNumber="33" refString="Komatsu, H. &amp; Takeda, M. (2001) On Merocryptoides, a leucosiid crab genus (Crustacea: Decapoda: Brachyura) endemic to Japan, with description of two new species. Zoological Science, 18, 993 - 1002." type="journal article" year="2001">Komatsu &amp; Takeda 2001</bibRefCitation>
; Ng
<emphasis id="32FB6B0CE628986FFEE096044709FEEC" box="[263,319,260,284]" italics="true" pageId="2" pageNumber="33">et al.</emphasis>
2009) (Table 1).
</paragraph>
</subSubSection>
<subSubSection id="4895E495E628986FFF2096264400FDAF" pageId="2" pageNumber="33" type="materials_examined">
<paragraph id="0030B71EE628986FFF2096264400FDAF" blockId="2.[151,1436,151,608]" pageId="2" pageNumber="33">
<emphasis id="32FB6B0CE628986FFF20962647E1FEB0" bold="true" box="[199,471,295,320]" pageId="2" pageNumber="33">Comparative material.</emphasis>
<taxonomicName id="C78FCC9DE628986FFE3996284277FECF" authority="A. Milne-Edwards, 1873: 1" authorityName="A. Milne-Edwards" authorityPageNumber="1" authorityYear="1873" box="[478,1089,295,320]" class="Malacostraca" family="Leucosiidae" genus="Merocryptus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="lambriformis">
<emphasis id="32FB6B0CE628986FFE3996284533FEB0" box="[478,773,296,320]" italics="true" pageId="2" pageNumber="33">Merocryptus lambriformis</emphasis>
<bibRefCitation id="641ECAEFE628986FFCEA9629421EFECF" author="Milne-Edwards" box="[781,1064,295,320]" pageId="2" pageNumber="33" refString="Milne-Edwards, A. (1873) Description de quelques Crustaces nouveaux ou peu connus provenant du Musee de M. C. Godeffroy. Journal des Museum Godeffroy, 1 (4), 77 - 88, pls. 1, 2 [pages 1 - 12, plates 12, 13 on separate]." type="journal article" year="1873">A. Milne-Edwards, 1873</bibRefCitation>
:
<specimenCount id="16897C97E628986FFBD496264277FECF" box="[1075,1089,295,319]" pageId="2" pageNumber="33" type="male">1</specimenCount>
</taxonomicName>
male (10.0 × 12.0 mm), 1 ovigerous female (12.3 × 15.0 mm), NSMT-Cr 13196, Tosa Bay,
<collectingCountry id="7898F78EE628986FFCB2964C45AEFE94" box="[853,920,333,356]" name="Japan" pageId="2" pageNumber="33">Japan</collectingCountry>
,
<quantity id="C7771AFBE628986FFC43964D4213FE94" box="[932,1061,332,356]" metricMagnitude="2" metricUnit="m" metricValue="1.54" metricValueMax="1.55" metricValueMin="1.53" pageId="2" pageNumber="33" unit="m" value="154.0" valueMax="155.0" valueMin="153.0">153155 m</quantity>
, coll. RV
<emphasis id="32FB6B0CE628986FFB71964D431AFE94" box="[1174,1324,332,356]" italics="true" pageId="2" pageNumber="33">Kotaka Maru</emphasis>
,
<date id="743191DEE628986FFADC964D46CBFE77" pageId="2" pageNumber="33" value="1998-01-21">21 January 1998</date>
.
<taxonomicName id="C78FCC9DE628986FFEEE967144FEFE77" authority="Sakai, 1963: 2" authorityName="Sakai" authorityPageNumber="2" authorityYear="1963" box="[265,712,367,392]" class="Malacostraca" family="Leucosiidae" genus="Merocryptoides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="32FB6B0CE628986FFEEE96714429FE78" box="[265,543,368,392]" italics="true" pageId="2" pageNumber="33">Merocryptoides frontalis</emphasis>
<bibRefCitation id="641ECAEFE628986FFDC1966E4499FE77" author="Sakai" box="[550,687,367,391]" pageId="2" pageNumber="33" refString="Sakai, T. (1963) Description of two new genera and fourteen new species of Japanese crabs from the collection of his Majesty the Emperor of Japan. Crustaceana, 5, 213 - 233." type="journal article" year="1963">Sakai, 1963</bibRefCitation>
:
<specimenCount id="16897C97E628986FFD5D966E44FEFE77" box="[698,712,367,391]" pageId="2" pageNumber="33" type="male">2</specimenCount>
</taxonomicName>
males (3.5 ×
<quantity id="C7771AFBE628986FFC83966E4588FE77" box="[868,958,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="33" unit="mm" value="3.7">3.7 mm</quantity>
, 4.8 ×
<quantity id="C7771AFBE628986FFBEC966E4253FE77" box="[1035,1125,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="33" unit="mm" value="4.4">4.4 mm</quantity>
), 1 ovigerous female (5.3 ×
<quantity id="C7771AFBE628986FFF70969546C4FE5C" box="[151,242,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="33" unit="mm" value="6.6">6.6 mm</quantity>
), NSMT-Cr 10406, off Kisami, Shimoda,
<collectingCountry id="7898F78EE628986FFD3696944522FE5C" box="[721,788,405,428]" name="Japan" pageId="2" pageNumber="33">Japan</collectingCountry>
,
<quantity id="C7771AFBE628986FFCC6969545B1FE5C" box="[801,903,404,428]" metricMagnitude="1" metricUnit="m" metricValue="3.15" metricValueMax="4.5" metricValueMin="1.8" pageId="2" pageNumber="33" unit="m" value="31.5" valueMax="45.0" valueMin="18.0">1845 m</quantity>
, coll. K. Nakamura,
<date id="743191DEE628986FFB9296954318FE5C" box="[1141,1326,404,428]" pageId="2" pageNumber="33" value="1982-08-23">23 August 1982</date>
.
<taxonomicName id="C78FCC9DE628986FFADC969547FEFE3F" authority="Rathbun, 1906" authorityName="Rathbun" authorityYear="1906" class="Malacostraca" family="Leucosiidae" genus="Oreotlos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="32FB6B0CE628986FFADC96954730FE20" italics="true" pageId="2" pageNumber="33">Oreotlos angulatus</emphasis>
(
<bibRefCitation id="641ECAEFE628986FFEF396B647F6FE3F" author="Rathbun" box="[276,448,439,463]" pageId="2" pageNumber="33" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3) [1903], 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun, 1906</bibRefCitation>
)
</taxonomicName>
: 1 ovigerous female (10.7 ×
<quantity id="C7771AFBE628986FFCEE96B64547FE3F" box="[777,881,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="2" pageNumber="33" unit="mm" value="16.5">16.5 mm</quantity>
), NSMT-Cr S 928, Chichi-jima I., Ogasawara Is.,
<collectingCountry id="7898F78EE628986FFF7096DC46ECFE04" box="[151,218,477,500]" name="Japan" pageId="2" pageNumber="33">Japan</collectingCountry>
, 27°04.55ʹN 142°09.16ʹE 27°04.73ʹN 142°09.31ʹE, 83
<quantity id="C7771AFBE628986FFC8A96DD459FFE04" box="[877,937,476,500]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="2" pageNumber="33" unit="m" value="81.0">81 m</quantity>
, coll. RV
<emphasis id="32FB6B0CE628986FFBF896DC426CFE04" box="[1055,1114,477,500]" italics="true" pageId="2" pageNumber="33">Koyo</emphasis>
,
<date id="743191DEE628986FFB8D96DD4318FE04" box="[1130,1326,476,500]" pageId="2" pageNumber="33" value="2008-10-28">28 October 2008</date>
.
<taxonomicName id="C78FCC9DE628986FFADC96DD47F8FDE7" authority="Tan &amp; Ng, 1995: 1" authorityName="Tan &amp; Ng" authorityPageNumber="1" authorityYear="1995" class="Malacostraca" family="Leucosiidae" genus="Oreotlos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="heuretos">
<emphasis id="32FB6B0CE628986FFADC96DD46C0FDE8" italics="true" pageId="2" pageNumber="33">Oreotlos heuretos</emphasis>
<bibRefCitation id="641ECAEFE628986FFF1A95014780FDE7" author="Tan" box="[253,438,511,536]" pageId="2" pageNumber="33" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1995) A revision of the Indo-Pacific genus Oreophorus Ruppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae). In: Richer de Forges B. (Ed.), Les fonds meubles des lagons de Nouvelle-Caledonie (Sedimentologie, benthos). ORSTOM, Paris, pp. 101 - 189." type="book chapter" year="1995">Tan &amp; Ng, 1995</bibRefCitation>
:
<specimenCount id="16897C97E628986FFE2796FE47F8FDE7" box="[448,462,511,535]" pageId="2" pageNumber="33" type="male">1</specimenCount>
</taxonomicName>
male (4.1 × 6.0 mm), 1 ovigerous female (7.5 ×
<quantity id="C7771AFBE628986FFC0996FE4260FDE7" box="[1006,1110,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="2" pageNumber="33" unit="mm" value="12.3">12.3 mm</quantity>
), NSMT-Cr S 931, northwest of Ototo-jima I., Ogasawara Is.,
<collectingCountry id="7898F78EE628986FFDC595244453FDCC" box="[546,613,549,572]" name="Japan" pageId="2" pageNumber="33">Japan</collectingCountry>
, 27°13.09ʹN 142°09.19ʹE 27°13.19ʹN 142°09.23ʹE, 135.8
<quantity id="C7771AFBE628986FFAD4952543A1FDCC" box="[1331,1431,548,572]" metricMagnitude="2" metricUnit="m" metricValue="1.355" pageId="2" pageNumber="33" unit="m" value="135.5">135.5 m</quantity>
, dredge, coll. RV
<emphasis id="32FB6B0CE628986FFEB3954847B9FD90" box="[340,399,585,608]" italics="true" pageId="2" pageNumber="33">Koyo</emphasis>
,
<date id="743191DEE628986FFE7995464404FDAF" box="[414,562,583,607]" pageId="2" pageNumber="33" value="2009-07-15">15 July 2009</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>