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<document id="96ADB05FFECDA314475BA09A6D8E814A" ID-DOI="10.1649/0010-065x-63.1.86" ID-ISSN="1938-4394" ID-Zenodo-Dep="5496086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630073033676" checkinUser="valdenar" docAuthor="Douglas, Hume" docDate="2009" docId="7F0B879B8C77FF952118FE150A43BE14" docLanguage="en" docName="0010-065X-63.1.86.pdf.imf" docOrigin="The Coleopterists Bulletin 63 (1)" docSource="http://dx.doi.org/10.1649/0010-065x-63.1.86" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Blaiseus nothoafricanus Douglas, new species" docType="treatment" docVersion="4" lastPageNumber="97" masterDocId="8332FFE38C7CFF9E2072FF970865BB14" masterDocTitle="Revision of Blaiseus Fleutiaux, a Genus Now Known from Asia, Africa and North America (Coleoptera, Elateridae, Cardiophorinae)" masterLastPageNumber="100" masterPageNumber="86" pageNumber="97" updateTime="1699141175581" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="07BB2BA14A4BF10929B9609517469F23">Revision of Blaiseus Fleutiaux, a Genus Now Known from Asia, Africa and North America (Coleoptera, Elateridae, Cardiophorinae)</mods:title>
</mods:titleInfo>
<mods:name id="D1D415677FAC4A88D0BB7D071D418353" type="personal">
<mods:role id="B5ADE80CAC40F6682C9AD58DBCDE3CBC">
<mods:roleTerm id="A335C0EDB4BA6E79D2D12A432DB58E0C">Author</mods:roleTerm>
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<mods:namePart id="2D64F30AFF186F80EAE0CF655D671629">Douglas, Hume</mods:namePart>
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<mods:typeOfResource id="495DC3F925C72DB568951B04988498A4">text</mods:typeOfResource>
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<mods:titleInfo id="9DD30A89F2DDA3B1626A45755456471E">
<mods:title id="5C005DABB95752DA332ECBBB65C7D234">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="152B732027B5F77E76065B1FB4AA95F1">
<mods:date id="972BD59D5FDDD653C24F09A3DC013D5D">2009</mods:date>
<mods:detail id="0AC630FB123FABBB297B604E6C3BE514" type="pubDate">
<mods:number id="F95AA56741D96E293E638A4442037DD5">2009-03-31</mods:number>
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<mods:detail id="AFBB636991812256E0BD26DB39002F1F" type="volume">
<mods:number id="3CDA2DA524FAA0DD29C4259E14A9C0A0">63</mods:number>
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<mods:number id="798354597CDE251F4FF95174EFEA6FBA">1</mods:number>
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<mods:url id="143A368A34AD8913E38D7E6E3BD6D1C2">http://dx.doi.org/10.1649/0010-065x-63.1.86</mods:url>
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<mods:classification id="B961700A024BC5C4ED6371995ECEE153">journal article</mods:classification>
<mods:identifier id="613747B4574FD30AA00301B9449AF0CD" type="DOI">10.1649/0010-065x-63.1.86</mods:identifier>
<mods:identifier id="58A09F6A17A672DA894BBF57DDD0CB3F" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="4FEC65C035C664B04159D11723FCAC2C" type="Zenodo-Dep">5496086</mods:identifier>
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<treatment id="7F0B879B8C77FF952118FE150A43BE14" ID-DOI="http://doi.org/10.5281/zenodo.5300768" ID-Zenodo-Dep="5300768" LSID="urn:lsid:plazi:treatment:7F0B879B8C77FF952118FE150A43BE14" httpUri="http://treatment.plazi.org/id/7F0B879B8C77FF952118FE150A43BE14" lastPageNumber="97" pageId="11" pageNumber="97">
<subSubSection id="BFB865068C77FF952118FE150B54BA8C" box="[362,817,386,408]" pageId="11" pageNumber="97" type="nomenclature">
<paragraph id="F71D368D8C77FF952118FE150B54BA8C" blockId="11.[362,817,386,435]" box="[362,817,386,408]" pageId="11" pageNumber="97">
<heading id="AC5581E18C77FF952118FE150B54BA8C" box="[362,817,386,408]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="97" reason="2">
<emphasis id="C5D6EA9F8C77FF952118FE150B54BA8C" bold="true" box="[362,817,386,408]" pageId="11" pageNumber="97">
<taxonomicName id="30A24D0E8C77FF952118FE150AD7BA8C" authority="Douglas" authorityName="Douglas" box="[362,690,386,408]" class="Insecta" family="Elateridae" genus="Blaiseus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="species" species="nothoafricanus" status="sp. nov.">
<emphasis id="C5D6EA9F8C77FF952118FE150A32BA8C" bold="true" box="[362,599,386,408]" italics="true" pageId="11" pageNumber="97">Blaiseus nothoafricanus</emphasis>
Douglas
</taxonomicName>
,
<taxonomicNameLabel id="DEE557E48C77FF9522CDFE150B54BA8C" box="[703,817,386,408]" pageId="11" pageNumber="97" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BFB865068C77FF95218EFE0A0AFABAA7" box="[508,671,413,435]" pageId="11" pageNumber="97" type="description">
<paragraph id="F71D368D8C77FF95218EFE0A0AFABAA7" blockId="11.[362,817,386,435]" box="[508,671,413,435]" pageId="11" pageNumber="97">
(
<figureCitation id="6F992A088C77FF952276FE0A0A29BAA7" box="[516,588,413,435]" captionStart="Figs" captionStartId="2.[167,206,1083,1102]" captionTargetBox="[151,970,194,1061]" captionTargetId="figure-118@2.[151,970,194,1061]" captionTargetPageId="2" captionText="Figs. 1–2. Lateral view of pronotum of Blaiseus spp. 1) B. malayensis new species; 2) B. nothoafricanus new species. Arrows indicate concavity of hypomeron immediately mesoventrad of hind angles. Scale bar 5 1 mm. Figs. 3–4. Dorsal view of scutellum of Blaiseus spp. 3) B. mexicanus new species; 4) B. bedeli Fleutiaux. Scale bar 5 1 mm. Figs. 5– 6. Dorsal view of protibia of Blaiseus spp. 5) B. mexicanus new species; 6) B. guatemalensis new species. m 5 tooth at midlength; a 5 apical tooth. Scale bar 5 1 mm. Fig. 7. Wing of Blaiseus bedeli Fleutiaux. Arrows indicate forked vein CuA1. Scale bar 5 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5496088" httpUri="https://zenodo.org/record/5496088/files/figure.png" pageId="11" pageNumber="97">Figs. 2</figureCitation>
,
<figureCitation id="6F992A088C77FF95222BFE0A0A16BAA7" box="[601,627,413,435]" captionStart="Figs" captionStartId="4.[167,206,1430,1449]" captionTargetBox="[149,973,194,1408]" captionTargetId="figure-1@4.[149,973,194,1408]" captionTargetPageId="4" captionText="Figs. 8–16. Dorsal habitus of Blaiseus spp. 8) B. bedeli Fleutiaux; 9) B. chiapasensis new species; 10) B. daklakensis new species; 11) B. daklakensis new species, side view; 12) B. malayensis new species; 13) B. mexicanus new species; 14) B. nothoafricanus new species; 15) B. zamoranoensis new species female; 16) B. zamoranoensis new species female, side view. Scale bar 5 1.0 mm. All photos are dorsal view of male specimens unless specified otherwise." figureDoi="http://doi.org/10.5281/zenodo.5496090" httpUri="https://zenodo.org/record/5496090/files/figure.png" pageId="11" pageNumber="97">14</figureCitation>
,
<figureCitation id="6F992A088C77FF95220DFE0A0AFDBAA7" box="[639,664,413,435]" captionStart="Figs" captionStartId="5.[196,235,1249,1268]" captionTargetBox="[188,994,194,1226]" captionTargetId="figure-60@5.[186,994,194,1226]" captionTargetPageId="5" captionText="Figs. 17–26. Dorsal view of aedeagi of Blaiseus spp. 17) B. bedeli Fleutiaux; 18) B. chiapasensis new species; 19) B. daklakensis new species; 20) B. fujianus new species; 21) B. guatemalensis new species; 22) B. laoensis new species; 23) B. malayensis new species; 24) B. mexicanus new species; 25) B. nothoafricanus new species; 26) B. zamoranoensis new species. d 5 dorsal apex of parameres; i 5 inner apex; v 5 ventral apex. Scale bar 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5496093" httpUri="https://zenodo.org/record/5496093/files/figure.png" pageId="11" pageNumber="97">25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BFB865068C77FF9520B5FE5E0919B98E" pageId="11" pageNumber="97" type="materials_examined">
<paragraph id="F71D368D8C77FF9520B5FE5E0A69BACB" blockId="11.[175,1005,457,1280]" box="[199,524,457,479]" pageId="11" pageNumber="97">
<emphasis id="C5D6EA9F8C77FF9520B5FE5E09E8BACB" bold="true" box="[199,397,457,479]" pageId="11" pageNumber="97">Material examined.</emphasis>
Two males.
</paragraph>
<paragraph id="F71D368D8C77FF9520B5FE730BB2B970" blockId="11.[175,1005,457,1280]" pageId="11" pageNumber="97">
<materialsCitation id="47CA3CD08C77FF9520B5FE73095EB901" collectorName="Janson" location="Lydenburg dist." pageId="11" pageNumber="97" specimenCount="1" typeStatus="holotype">
<emphasis id="C5D6EA9F8C77FF9520B5FE73094FBAEE" bold="true" box="[199,298,484,506]" pageId="11" pageNumber="97">
<typeStatus id="2819882F8C77FF9520B5FE730943BAEE" box="[199,294,484,506]" pageId="11" pageNumber="97" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male labeled: ‘‘
<location id="F27D60568C77FF9521AAFE730AE7BAEE" LSID="urn:lsid:plazi:treatment:7F0B879B8C77FF952118FE150A43BE14:F27D60568C77FF9521AAFE730AE7BAEE" box="[472,642,484,506]" name="Lydenburg dist." pageId="11" pageNumber="97">Lydenburg dist.</location>
/ Transvaal’’; ‘‘
<collectorName id="5A57535B8C77FF952350FE730B08BAEE" box="[802,877,484,506]" pageId="11" pageNumber="97">Janson</collectorName>
coll./ 1903- 130.’’; [red] ‘‘
</materialsCitation>
<materialsCitation id="47CA3CD08C77FF952149FE680B8DB901" box="[315,1000,511,533]" collectionCode="NHM" pageId="11" pageNumber="97" specimenCount="1" typeStatus="holotype">
<typeStatus id="2819882F8C77FF952149FE6809A2B901" box="[315,455,511,533]" pageId="11" pageNumber="97" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="30A24D0E8C77FF9521A9FE680A48B901" authorityName="FLEUTIAUX, A" authorityYear="1931" box="[475,557,511,533]" class="Insecta" family="Elateridae" genus="Blaiseus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="genus">Blaiseus</taxonomicName>
/ nothoafricanus/ Douglas 2006’’ (
<collectionCode id="91B3AE488C77FF9523ECFE680B84B901" box="[926,993,511,533]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="11" pageNumber="97" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
.
<typeStatus id="2819882F8C77FF9520DDFD8E096BB93B" box="[175,270,537,559]" pageId="11" pageNumber="97" type="paratype">Paratype</typeStatus>
:
<specimenCount id="E1A4FD048C77FF952152FD8E0909B93B" box="[288,364,537,559]" pageId="11" pageNumber="97" type="male">1 male</specimenCount>
labeled: [handwritten] ‘‘Ayres’’; [handwritten] ‘‘P.B. Spei/ Transvaal’’; ‘‘Fry Coll./ 1905-100’’; [yellow] ‘‘
<materialsCitation id="47CA3CD08C77FF9522D2FDA30BB7B970" collectionCode="NHM" pageId="11" pageNumber="97" specimenCount="1" typeStatus="paratype">
<typeStatus id="2819882F8C77FF9522D2FDA30B4CB95E" box="[672,809,564,586]" pageId="11" pageNumber="97" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="30A24D0E8C77FF952332FDA30BF7B95E" authorityName="FLEUTIAUX, A" authorityYear="1931" box="[832,914,564,586]" class="Insecta" family="Elateridae" genus="Blaiseus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="genus">Blaiseus</taxonomicName>
/ nothoafricanus/ Douglas 2006,’’ with genitalia in microvial attached to pin (
<collectionCode id="91B3AE488C77FF9523FAFDD90BAEB970" box="[904,971,590,612]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="11" pageNumber="97" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F71D368D8C77FF9520B5FDFD0919B98E" blockId="11.[175,1005,457,1280]" pageId="11" pageNumber="97">
Type locality:
<collectingCountry id="8FB5761D8C77FF95211BFDFD0992B994" box="[361,503,618,640]" name="South Africa" pageId="11" pageNumber="97">South Africa</collectingCountry>
,
<collectingRegion id="3566F86F8C77FF952278FDFD0A9FB994" box="[522,762,618,640]" country="South Africa" name="Mpumalanga" pageId="11" pageNumber="97">Mpumalanga Province</collectingRegion>
(formerly Transvaal), Lydenburg District.
</paragraph>
</subSubSection>
<subSubSection id="BFB865068C77FF9520B5FD370B6FB9FF" pageId="11" pageNumber="97" type="diagnosis">
<paragraph id="F71D368D8C77FF9520B5FD370B6FB9FF" blockId="11.[175,1005,457,1280]" pageId="11" pageNumber="97">
<emphasis id="C5D6EA9F8C77FF9520B5FD3709DAB9A2" bold="true" box="[199,447,672,694]" pageId="11" pageNumber="97">Diagnosis. Pterothorax:</emphasis>
Elytra with apices not upturned (
<figureCitation id="6F992A088C77FF95233DFD370BFFB9A2" box="[847,922,672,694]" captionStart="Figs" captionStartId="4.[167,206,1430,1449]" captionTargetBox="[149,973,194,1408]" captionTargetId="figure-1@4.[149,973,194,1408]" captionTargetPageId="4" captionText="Figs. 8–16. Dorsal habitus of Blaiseus spp. 8) B. bedeli Fleutiaux; 9) B. chiapasensis new species; 10) B. daklakensis new species; 11) B. daklakensis new species, side view; 12) B. malayensis new species; 13) B. mexicanus new species; 14) B. nothoafricanus new species; 15) B. zamoranoensis new species female; 16) B. zamoranoensis new species female, side view. Scale bar 5 1.0 mm. All photos are dorsal view of male specimens unless specified otherwise." figureDoi="http://doi.org/10.5281/zenodo.5496090" httpUri="https://zenodo.org/record/5496090/files/figure.png" pageId="11" pageNumber="97">Fig. 11</figureCitation>
).
<emphasis id="C5D6EA9F8C77FF9523C4FD370B89B9A2" bold="true" box="[950,1004,672,694]" pageId="11" pageNumber="97">Legs:</emphasis>
Femora and tibiae expanded (fossorial); protibiae without posterior tooth at midlength or apex (Fig. 6, although strong spurs present).
</paragraph>
</subSubSection>
<subSubSection id="BFB865068C77FF9520B5FD670AF4BF6E" pageId="11" pageNumber="97" type="description">
<paragraph id="F71D368D8C77FF9520B5FD670AF4BF6E" blockId="11.[175,1005,457,1280]" pageId="11" pageNumber="97">
<emphasis id="C5D6EA9F8C77FF9520B5FD6709E4B812" bold="true" box="[199,385,752,774]" pageId="11" pageNumber="97">Description. Male:</emphasis>
Length 6.0 mm. Integument brown without lighter coloured patches on elytra.
<emphasis id="C5D6EA9F8C77FF95210EFC9C09DDB835" bold="true" box="[380,440,779,801]" pageId="11" pageNumber="97">Head:</emphasis>
Nasale with circular pits antero-mesad of antennal fossae; antennae reaching beyond mesocoxae; labrum convex.
<emphasis id="C5D6EA9F8C77FF95230CFCB20B89B82F" bold="true" box="[894,1004,805,827]" pageId="11" pageNumber="97">Prothorax:</emphasis>
Posterior edge of pronotum with 2 apices mesally; hypomeron with hind edge with semicircular emargination immediately meso-ventrad of hind angles (
<figureCitation id="6F992A088C77FF9523D2FCCC0B85B865" box="[928,992,859,881]" captionStart="Figs" captionStartId="2.[167,206,1083,1102]" captionTargetBox="[151,970,194,1061]" captionTargetId="figure-118@2.[151,970,194,1061]" captionTargetPageId="2" captionText="Figs. 1–2. Lateral view of pronotum of Blaiseus spp. 1) B. malayensis new species; 2) B. nothoafricanus new species. Arrows indicate concavity of hypomeron immediately mesoventrad of hind angles. Scale bar 5 1 mm. Figs. 3–4. Dorsal view of scutellum of Blaiseus spp. 3) B. mexicanus new species; 4) B. bedeli Fleutiaux. Scale bar 5 1 mm. Figs. 5– 6. Dorsal view of protibia of Blaiseus spp. 5) B. mexicanus new species; 6) B. guatemalensis new species. m 5 tooth at midlength; a 5 apical tooth. Scale bar 5 1 mm. Fig. 7. Wing of Blaiseus bedeli Fleutiaux. Arrows indicate forked vein CuA1. Scale bar 5 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5496088" httpUri="https://zenodo.org/record/5496088/files/figure.png" pageId="11" pageNumber="97">Fig. 2</figureCitation>
). Prosternum with ventral surface of prosternal process surface not carinate laterally.
<emphasis id="C5D6EA9F8C77FF952167FC0709F0B8B2" bold="true" box="[277,405,912,934]" pageId="11" pageNumber="97">Pterothorax:</emphasis>
Scutellum with sides straight before rounded posterior apex. Elytra not upturned at apex; interval 6 costate apically; epipleurae not serrate. Hind wings with CuA1 forked at junction with MP3+4, forming an additional closed cell (Fig. 7).
<emphasis id="C5D6EA9F8C77FF95218EFC480A54B8E1" bold="true" box="[508,561,991,1013]" pageId="11" pageNumber="97">Legs:</emphasis>
Femora and tibiae expanded (fossorial); protibiae without posterior tooth at midlength or apex (Fig. 6, although strong spurs present); tarsi simple.
<emphasis id="C5D6EA9F8C77FF9521A2FB820A0DBF3F" bold="true" box="[464,616,1045,1067]" pageId="11" pageNumber="97">Male genitalia:</emphasis>
Aedeagus with median lobe broadest near apex, not concave medially (
<figureCitation id="6F992A088C77FF95226AFBB80A00BF51" box="[536,613,1071,1093]" captionStart="Figs" captionStartId="5.[196,235,1249,1268]" captionTargetBox="[188,994,194,1226]" captionTargetId="figure-60@5.[186,994,194,1226]" captionTargetPageId="5" captionText="Figs. 17–26. Dorsal view of aedeagi of Blaiseus spp. 17) B. bedeli Fleutiaux; 18) B. chiapasensis new species; 19) B. daklakensis new species; 20) B. fujianus new species; 21) B. guatemalensis new species; 22) B. laoensis new species; 23) B. malayensis new species; 24) B. mexicanus new species; 25) B. nothoafricanus new species; 26) B. zamoranoensis new species. d 5 dorsal apex of parameres; i 5 inner apex; v 5 ventral apex. Scale bar 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5496093" httpUri="https://zenodo.org/record/5496093/files/figure.png" pageId="11" pageNumber="97">Fig. 25</figureCitation>
); parameres with dorsal (setose) and ventral apices both more than twice longer than broad (width at measured at midlength of free portion).
<emphasis id="C5D6EA9F8C77FF9521BEFBF30A7FBF6E" bold="true" box="[460,538,1124,1146]" pageId="11" pageNumber="97">Female:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="BFB865068C77FF9520B5FB170A43BE14" pageId="11" pageNumber="97" type="discussion">
<paragraph id="F71D368D8C77FF9520B5FB170A43BE14" blockId="11.[175,1005,457,1280]" pageId="11" pageNumber="97">
<emphasis id="C5D6EA9F8C77FF9520B5FB170952BF82" bold="true" box="[199,311,1152,1174]" pageId="11" pageNumber="97">Comments.</emphasis>
The hypothesis that this species is distinct from all other known
<taxonomicName id="30A24D0E8C77FF9520DDFB0C0964BFA4" authorityName="FLEUTIAUX, A" authorityYear="1931" box="[175,257,1179,1200]" class="Insecta" family="Elateridae" genus="Blaiseus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="C5D6EA9F8C77FF9520DDFB0C0964BFA4" box="[175,257,1179,1200]" italics="true" pageId="11" pageNumber="97">Blaiseus</emphasis>
</taxonomicName>
is supported by its unique combination of diagnostic characters. The known distribution of this species is limited to the vaguely defined
<typeStatus id="2819882F8C77FF952315FB220BF6BFDF" box="[871,915,1205,1227]" pageId="11" pageNumber="97">type</typeStatus>
locality, and may include higher elevation forests in Lydenburg District. This is the only
<taxonomicName id="30A24D0E8C77FF9520DDFB7D092BBE14" box="[175,334,1258,1280]" class="Insecta" family="Elateridae" genus="Blaiseus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C5D6EA9F8C77FF9520DDFB7D0964BFEB" box="[175,257,1258,1279]" italics="true" pageId="11" pageNumber="97">Blaiseus</emphasis>
species
</taxonomicName>
known from Africa.
</paragraph>
</subSubSection>
</treatment>
</document>
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