503 lines
61 KiB
XML
503 lines
61 KiB
XML
<document id="D937CCCCDD41D51D2288ADCDED12345A" ID-DOI="10.3897/zookeys.1223.139346" ID-ZooBank="451749E9-009E-43E9-A6F1-892035BDF1B0" ID-publisher-id="139346" URI-arpha="6EA2BAFE-82AB-5570-95B8-EE1F5FA60D46" URI-zoobank="http://zoobank.org/451749E9-009E-43E9-A6F1-892035BDF1B0" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1736349736103" checkinUser="pensoft" docAuthor="Wesener, Thomas, Akkari, Nesrine & Golovatch, Sergei I." docDate="2025" docId="51691BCD50735AB5A5F1FFC3AC39BCB4" docLanguage="en" docName="ZooKeys 1223: 185-220" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/139346/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Phymatodesmus sakalava" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="220" masterDocId="6EA2BAFE82AB557095B8EE1F5FA60D46" masterDocTitle="Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida)" masterLastPageNumber="220" masterPageNumber="185" pageNumber="185" updateTime="1736355520980" updateUser="admin">
|
||
<mods:mods id="3428C035282E13D5A82C222839A81CFF" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="8F62DAD679F5F2EF6611ADB374CA9BAA">
|
||
<mods:title id="9A56661422E3983054812C13E3F52E79">Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="624029389D99C9C9771E5D7E81D7C556" type="personal">
|
||
<mods:role id="1B3F91F0026BB3EC72F7FE76AF2485BE">
|
||
<mods:roleTerm id="E2609DECDB099D9D14065BC726835F02">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="79081E58069D96076EE1D442040951EC">Wesener, Thomas</mods:namePart>
|
||
<mods:nameIdentifier id="EA91E6D17FCD7CC0F43859412E5EBD69" type="ORCID">0000-0002-2028-3541</mods:nameIdentifier>
|
||
<mods:affiliation id="807ABBD5980E2B3F86FA69DDACF4E650">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, D- 53113 Bonn, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="F95957A57423EFF5BAB088D6249E1AD5" type="personal">
|
||
<mods:role id="DE7B3976BC34A94B35B5B75E1C29EAC3">
|
||
<mods:roleTerm id="19C096796887497E42C17CD578EE3CC4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="99C9D1F9F29311054E5F45B6C0B69C37">Akkari, Nesrine</mods:namePart>
|
||
<mods:nameIdentifier id="3694AE3DFAFF969E6E5F76D8C7E3F42D" type="ORCID">0000-0001-5019-4833</mods:nameIdentifier>
|
||
<mods:affiliation id="84CD8B9A3AD93A5C0B848EDA54BB813F">Third Zoological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="EAE0B2D2559A9B84C15CCA6F4083D29A" type="personal">
|
||
<mods:role id="49644E2DFE0CFE82CBA2BA725ECDE95F">
|
||
<mods:roleTerm id="91A17F1D3312417CB3B6D36F56BEEAA9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="067E74DED08887CA92AE25D6F321C1A2">Golovatch, Sergei I.</mods:namePart>
|
||
<mods:affiliation id="2C5186A7AC4CAF8959C0B007A9D50448">Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="D55BB74EF1D5E97E15A6E3847880FE3B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="71B057AD639364F8E6FB862855EF1E0F" type="host">
|
||
<mods:titleInfo id="D43E44B66FE33D67ED715EBC7BF5CBCC">
|
||
<mods:title id="0253D039EC48C75029CC8FD50F8C8FD8">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="955A648BAB55ED308B0CA452851AE66B">
|
||
<mods:date id="3E74FB1888B14F2D7F2C9C1E018465E2">2025</mods:date>
|
||
<mods:detail id="90F10578D0839B04148B1A4C91E12FEA" type="pubDate">
|
||
<mods:number id="0911C4DCC534B1C044F8395AFE960423">2025-01-08</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="1510B7787C978FB8D4E143752913A1C5" type="volume">
|
||
<mods:number id="ABA1A4FE22A3B5A3F9715C17A68D78AB">1223</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="806DB4B469A06E5454C472EF5913A292" unit="page">
|
||
<mods:start id="3539548B9863D6A34598CF8AE4C0B729">185</mods:start>
|
||
<mods:end id="9F3BB6F3B96CB58D9FEE2B0E43627DF7">220</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="0D7C819EB73461565DCB8B550D825EF6">journal article</mods:classification>
|
||
<mods:identifier id="26F0FE02F30D26D23C3C4F3208DD4E9F" type="DOI">10.3897/zookeys.1223.139346</mods:identifier>
|
||
<mods:identifier id="DB8F2360A3F65D01C8CD470BFCF8B4A4" type="ZooBank">451749E9-009E-43E9-A6F1-892035BDF1B0</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="51691BCD50735AB5A5F1FFC3AC39BCB4" ID-arpha="51691BCD-5073-5AB5-A5F1-FFC3AC39BCB4" LSID="urn:lsid:plazi:treatment:51691BCD50735AB5A5F1FFC3AC39BCB4" httpUri="http://treatment.plazi.org/id/51691BCD50735AB5A5F1FFC3AC39BCB4" scope_class="Diplopoda" scope_family="Dalodesmidae" scope_order="Polydesmida">
|
||
<subSubSection id="C669D06AA7492B3E186FF17CD7049C8B" type="nomenclature">
|
||
<paragraph id="764FB1B6B3CFD998063E4997BB44BFBE">
|
||
<taxonomicName id="E1F7F8D396AA5A23A8CE30C6A6D7A418" ID-CoL="77GLZ" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="sakalava">
|
||
<emphasis id="3106C85FD391F2DC80A83B62AB0812EB" italics="true">Phymatodesmus sakalava</emphasis>
|
||
(
|
||
<bibRefCitation id="D46F5D27C81B9F9AE82E84B747018172" author="de Saussure H & Zehntner L" refId="B7" refString="de Saussure H, Zehntner L (1897) Atlas de l'histoire naturelle des Myriapodes. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Paris: Imprimerie nationale, Pl 1 - 15." year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="665E96BC91D1245EABC4B164BAE69D6E" type="description">
|
||
<paragraph id="6C7BD90F2213BA004A53E6A25335C54A">
|
||
<figureCitation id="D6C5FCE1DF24CC83CE8BEF4131091DE1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♀ holotype MHNG (G, H), ♂ specimen (ZFMK MYR 12217). Multi-layer photographs and drawings A habitus, dorsal view B habitus, lateral view C habitus, ventral view D detail of midbody rings, dorsal view E detail of anterior body rings and gonopod, lateral view F detail of anterior rings and gonopods, ventral view G drawing from the original description in de Saussure and Zehntner (1897) H detail of body ring, ♀ holotype I gonopod still attached, ventral view. Abbreviations: lb = lateral branch; mb = mesal branch; sl = solenomere branch. Scale bar: 0.3 mm (H)." figureDoi="10.3897/zookeys.1223.139346.figure1" httpUri="https://binary.pensoft.net/fig/1215825">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="9A1FF9240435F76AD3B7A8A203B2C552" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♂ specimen (ZFMK MYR 12217). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, lateral view C midbody ring, paranota with ozopore D details of surface structures, dorsal view E detail of surface structures of prozonite and metazonite F endotergum. Scale bars: 100 µm (A – C); 10 µm (D); 20 µm (E); 10 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure2" httpUri="https://binary.pensoft.net/fig/1215826">2</figureCitation>
|
||
,
|
||
<figureCitation id="4C77E4E1596954EB134853F072D9BE8A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♂ specimen (ZFMK MYR 12217). Multi-layered photographs and drawings, gonopod A posterior view B lateral and mesal view C lateral view D posterior view E anterior view F lateral view. Abbreviations: fe = femorite; lb = lateral branch; mb = mesal branch; sl = solenomere branch." figureDoi="10.3897/zookeys.1223.139346.figure3" httpUri="https://binary.pensoft.net/fig/1215827">3</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BA8CD652C4287C709495675E9B6AD87" type="reference_group">
|
||
<paragraph id="3432CC052A1B81F84A27DCA1A5FBB72A">
|
||
<treatmentCitationGroup id="7601407ABD0A6CD6272AE1CD396C60B0">
|
||
<treatmentCitation id="043EF87BFCFD64DF9E042D099E0BD00F">
|
||
<taxonomicName id="9794C14F3C47074BAE48BDC4B92CA678" authority="de Saussure & Zehntner, 1897" authorityName="de Saussure & Zehntner" authorityYear="1897" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="sakalava">
|
||
<emphasis id="B2D9737BBF4FF66136B4C770F2724984" italics="true">Polydesmus sakalava</emphasis>
|
||
<bibRefCitation id="1C31F8D2A93B32519BA17C0F41589653" author="de Saussure H & Zehntner L" refId="B7" refString="de Saussure H, Zehntner L (1897) Atlas de l'histoire naturelle des Myriapodes. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Paris: Imprimerie nationale, Pl 1 - 15." year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
: plate 5, fig. 22 (figure and caption only).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="F493C35427BD108B42455209A8291DCE">
|
||
<treatmentCitationGroup id="4E5FCBAC32800ECB53E0809BD069502F">
|
||
<taxonomicName id="FFF317D26FA118B50FDCF9511C52E48D" authorityName="de Saussure & Zehntner" authorityYear="1897" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="sakalava" subGenus="Phymatodesmus">
|
||
<emphasis id="56CB04610A3B07C442925F4EFF708ABA" italics="true">Polydesmus</emphasis>
|
||
(
|
||
<emphasis id="12103A5892042FD42B022A3CF99B3B47" italics="true">Phymatodesmus</emphasis>
|
||
)
|
||
<emphasis id="84541B338123FCE88C93797E6E503420" italics="true">sakalava</emphasis>
|
||
</taxonomicName>
|
||
<comment id="8D299D45FA4B10A05AFB565398DF502A">
|
||
–
|
||
<treatmentCitation id="87440DBBAA63AFB7592472CE9FF8AB27" author="de Saussure H & Zehntner L" page="95" year="1902">
|
||
<bibRefCitation id="68AA649F85DD91A7A968C549B3348722" author="de Saussure H & Zehntner L" editor="Grandidier A" firstAuthor="de Saussure" journalOrPublisher="Imprimérie nationale, Paris" page="95" pagination="1-356" refId="B9" refString="de Saussure H, Zehntner L (1902) Myriapodes de Madagascar. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Imprimérie nationale, Paris, 1–356." title="Myriapodes de Madagascar." volumeTitle="Histoire physique, naturelle et politique de Madagascar." year="1902">de Saussure and Zehntner 1902: 95</bibRefCitation>
|
||
</treatmentCitation>
|
||
(D).
|
||
</comment>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7ED88C0C9BE5515606745D22C17E628B">
|
||
<treatmentCitationGroup id="9DBAB07006FC38D043BE1988649AED9F">
|
||
<taxonomicName id="250E21226BC94BB3CB57402CE1B5A29F" ID-CoL="8N7ND" baseAuthorityName="De Saussure & Zehntner" baseAuthorityYear="1901" class="Diplopoda" family="Dalodesmidae" genus="Eutubercularium" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="sakalava">
|
||
<emphasis id="C75C940862563276E55D20AF12C55B2F" italics="true">Eutubercularium sakalava</emphasis>
|
||
</taxonomicName>
|
||
<comment id="2DAB351AE7E568D8BE108E1427407152">
|
||
–
|
||
<treatmentCitation id="FBFB435E4738430EF9F83D4BB82237F3" author="Brölemann HW" page="605" year="1916">
|
||
<bibRefCitation id="A612C883CA7EE261765AFA170C845F00" DOI="10.1080/21686351.1915.12279415" author="Brölemann HW" firstAuthor="Brolemann" issue="4" journalOrPublisher="Annales de la Société Entomologique de France" page="605" pagination="523-617" refId="B5" refString="Brölemann HW (1916) Essai de classification des polydesmiens (Myriapodes). Annales de la Société Entomologique de France 84 (4): 523–617. https://doi.org/10.1080/21686351.1915.12279415" title="Essai de classification des polydesmiens (Myriapodes)." volume="84" year="1916">Brölemann 1916: 605</bibRefCitation>
|
||
</treatmentCitation>
|
||
(D);
|
||
<treatmentCitation id="D18170FFA2382160BBAAF805CDB610D5" author="Hollier J & Wesener T" page="62" year="2017">
|
||
<bibRefCitation id="1DAEA85C65C56E1C1ACEBA0C9539218B" DOI="10.5281/zenodo.322665" author="Hollier J & Wesener T" firstAuthor="Hollier" issue="1" journalOrPublisher="Revue suisse de Zoologie" page="62" pagination="53-65" refId="B16" refString="Hollier J, Wesener T (2017) The Diplopoda (Myriapoda) of Madagascar described by Henri de Saussure and Leo Zehntner. Revue suisse de Zoologie 124 (1): 53–65. https://doi.org/10.5281/zenodo.322665" title="The Diplopoda (Myriapoda) of Madagascar described by Henri de Saussure and Leo Zehntner." volume="124" year="2017">Hollier and Wesener 2017: 62</bibRefCitation>
|
||
</treatmentCitation>
|
||
(L).
|
||
</comment>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="4971A6BC72019A4189F5A500DBD7E46D">
|
||
<treatmentCitationGroup id="C12361AA19F0F4C62B44C3B20CC6F84C">
|
||
<taxonomicName id="98641A3115A8BE6ED6F7E8028C2324A1" ID-CoL="8N7ND" authorityName="de Saussure & Zehntner" authorityYear="1897" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="sakalava">
|
||
<emphasis id="2293E67F741D5E91E8BB95FE54056BC0" italics="true">Dalodesmus sakalava</emphasis>
|
||
</taxonomicName>
|
||
<comment id="B2961F5EF5A994B93766631F49EBDA42">
|
||
–
|
||
<treatmentCitation id="F6A55EFBC8350FA43DEADF5BA553C425" author="Jeekel CAW" page="238" year="1965">
|
||
<bibRefCitation id="1F3BE58D8B6F12554C9C5C7AEB3F8722" author="Jeekel CAW" firstAuthor="Jeekel" journalOrPublisher="Entomologische Berichten" page="238" pagination="236-239" refId="B17" refString="Jeekel CAW (1965) The identity of Dalodesmus tectus Cook, 1896, and the status of the family names Dalodesmidae Cook, 1896, Vanhoeffeniidae Attems, 1914 and Sphaeriotrichopodidae Attems, 1914 (Diplopoda, Polydesmida). Entomologische Berichten 25: 236–239." title="The identity of Dalodesmus tectus Cook, 1896, and the status of the family names Dalodesmidae Cook, 1896, Vanhoeffeniidae Attems, 1914 and Sphaeriotrichopodidae Attems, 1914 (Diplopoda, Polydesmida)." volume="25" year="1965">Jeekel 1965: 238</bibRefCitation>
|
||
</treatmentCitation>
|
||
(L);
|
||
<treatmentCitation id="F126EC2C8E99D0CA5BC8A4AD100534C0" author="Hoffman RL" page="230" year="1974">
|
||
<bibRefCitation id="07850F9873FEDD6B78A8B9D53F3B3BC0" author="Hoffman RL" firstAuthor="Hoffman" issue="2" journalOrPublisher="Wasmann Journal of Biology" page="230" pagination="221-246" refId="B14" refString="Hoffman RL (1974) Short studies on dalodesmid millipeds from South Africa and Madagascar. Wasmann Journal of Biology 32 (2): 221–246." title="Short studies on dalodesmid millipeds from South Africa and Madagascar." volume="32" year="1974">Hoffman 1974: 230</bibRefCitation>
|
||
</treatmentCitation>
|
||
(L);
|
||
<treatmentCitation id="FD78F950A84D372D58BD2204C17C2B18" author="Golovatch SI & Hoffman RL" page="162" year="1989">
|
||
<bibRefCitation id="D72095E7E2831BA7AD871D3AE731D3B5" DOI="10.1080/03946975.1989.10539436" author="Golovatch SI & Hoffman RL" firstAuthor="Golovatch" journalOrPublisher="Tropical Zoology" page="162" pagination="159-164" refId="B12" refString="Golovatch SI, Hoffman RL (1989) Identity of Polydesmus hamatus Brandt 1841, a Malagasy milliped (Diplopoda Polydesmida Dalodesmidae). Tropical Zoology 2: 159–164. https://doi.org/10.1080/03946975.1989.10539436" title="Identity of Polydesmus hamatus Brandt 1841, a Malagasy milliped (Diplopoda Polydesmida Dalodesmidae)." volume="2" year="1989">Golovatch and Hoffman 1989: 162</bibRefCitation>
|
||
</treatmentCitation>
|
||
(L).
|
||
</comment>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="CDF3E27435C21ECFE0C4E5F22F97F17C">
|
||
<treatmentCitationGroup id="87158C0E6B76148F8E03585B99AACA20">
|
||
<taxonomicName id="4EC04B641ED26684EE1923E1390ECECA" ID-CoL="77GLZ" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="sakalava">
|
||
<emphasis id="10959CA6A6431FC4D792AF6309209A04" italics="true">Phymatodesmus sakalava</emphasis>
|
||
</taxonomicName>
|
||
<comment id="A6B717777D0E595523679D6F7D1EE897">
|
||
–
|
||
<treatmentCitation id="AE490D1BE34CB1E45F178030094632F7" author="Attems C" page="490" year="1940">
|
||
<bibRefCitation id="C38306FD9A7ED2FAA626C106EAC768D2" DOI="10.1515/9783111609645.1" author="Attems C" firstAuthor="Attems" page="490" refId="B2" refString="Attems C (1940) Myriapoda 3. Polydesmoidea III. Fam. Polydesmidae, Vanhoeffeniidae, Cryptodesmidae, Oniscodesmidae, Sphaerotrichopidae, Peridontodesmidae, Rhachidesmidae, Macellolophidae, Pandirodesmidae. Das Tierreich 70, i – xvii + 1–577. https://doi.org/10.1515/9783111609645.1" year="1940">Attems 1940: 490</bibRefCitation>
|
||
</treatmentCitation>
|
||
(L);
|
||
<treatmentCitation id="41C002E08134C13E513EF1744D4C77BE" author="Enghoff H" page="623" year="2003">
|
||
<bibRefCitation id="B8D3DF3AA3C974DC52F8A171A1944812" author="Enghoff H" editor="Goodman SM & Benstead JP" firstAuthor="Enghoff" journalOrPublisher="University of Chicago Press, Chicago and London" page="623" pagination="617-627" refId="B10" refString="Enghoff H (2003) Diplopoda, millipedes. In: Goodman SM, Benstead JP (Eds) The Natural History of Madagascar. University of Chicago Press, Chicago and London, 617–627." title="Diplopoda, millipedes." volumeTitle="The Natural History of Madagascar." year="2003">Enghoff 2003: 623</bibRefCitation>
|
||
</treatmentCitation>
|
||
(L);
|
||
<treatmentCitation id="28FEC93742B0E47475E519369F2647FF" ID-DOI="http://doi.org/10.5281/zenodo.5964246" ID-Zenodo-Dep="5964246" author="Mesibov R & Wesener T & Hollier J" httpUri="http://treatment.plazi.org/id/0382CE12F230FFFCF18BF962FC98FEA3" page="389" year="2018">
|
||
<bibRefCitation id="BC5BC3891912A8FE68E62C7658B48585" DOI="10.11646/zootaxa.4413.2.11" author="Mesibov R & Wesener T & Hollier J" etAl="et al." firstAuthor="Mesibov" issue="2" journalOrPublisher="Zootaxa" page="389" pagination="389-391" refId="B20" refString="Mesibov R, Wesener T, Hollier J (2018) A replacement name for Dalodesmus sakalava (de Saussure & Zehntner, 1901) (Diplopoda: Polydesmida: Dalodesmidae). Zootaxa 4413 (2): 389–391. https://doi.org/10.11646/zootaxa.4413.2.11" title="A replacement name for Dalodesmus sakalava (de Saussure & Zehntner, 1901) (Diplopoda: Polydesmida: Dalodesmidae)." volume="4413" year="2018">Mesibov et al. 2018: 389</bibRefCitation>
|
||
</treatmentCitation>
|
||
(N);
|
||
<treatmentCitation id="194A81099A0BCD6DD2C8FFB826B19C06" author="Wesener T & Enghoff H" page="926" year="2022">
|
||
<bibRefCitation id="4C0DB8FFC47C5359270A23DBC4FE1246" DOI="10.2307/j.ctv2ks6tbb.118" author="Wesener T & Enghoff H" editor="Fisher BL & Goodman SM" firstAuthor="Wesener" journalOrPublisher="Princeton University Press USA" page="926" pagination="918-933" refId="B22" refString="Wesener T, Enghoff H (2022) Diplopoda, millipedes, Menavetraka, Ankodabitra, Ankodiavitry, Marotanana, Sakolavitsy. In: Fisher BL, Goodman SM (Eds) The New Natural History of Madagascar. Princeton University Press USA, 918–933. https://doi.org/10.2307/j.ctv2ks6tbb.118" title="Diplopoda, millipedes, Menavetraka, Ankodabitra, Ankodiavitry, Marotanana, Sakolavitsy." volumeTitle="The New Natural History of Madagascar." year="2022">Wesener and Enghoff 2022: 926</bibRefCitation>
|
||
</treatmentCitation>
|
||
(L).
|
||
</comment>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="01CF1E5A501CFB82F3E4E02152FBE2A6">
|
||
<treatmentCitationGroup id="435E1F154D3B26F6288161236BCA3531">
|
||
<taxonomicName id="0059E534DFA5ABBF4E4ACE7B41C943F1" ID-CoL="7NJ72" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Dalodesmidae" genus="Undetermined" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="undetermined">
|
||
Dalodesmidae
|
||
<comment id="B98CD76950884573363C8E4EC6E1CA4C">
|
||
sp. –
|
||
<treatmentCitation id="0B8D3D199F8118723438480DBF400C3C" author="Spelzhausen L & Wesener T & Schütte K" page="4" year="2020">
|
||
<bibRefCitation id="3D70DF74990A648FF4F2B432E226F151" DOI="10.4314/mcd.v15i1.3" author="Spelzhausen L & Wesener T & Schütte K" etAl="et al." firstAuthor="Spelzhausen" issue="1" journalOrPublisher="Madagascar Conservation and Development" page="4" pagination="1-8" refId="B21" refString="Spelzhausen L, Wesener T, Schütte K (2020) Vegetation thresholds for the occurrence of millipedes (Diplopoda) in different tropical forest types in Andasibe, Madagascar. Madagascar Conservation and Development 15 (1): 1–8. http://doi.org/10.4314/mcd.v15i1.3" title="Vegetation thresholds for the occurrence of millipedes (Diplopoda) in different tropical forest types in Andasibe, Madagascar." volume="15" year="2020">Spelzhausen et al., 2020: 4</bibRefCitation>
|
||
</treatmentCitation>
|
||
(L).
|
||
</comment>
|
||
</taxonomicName>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ENKAE" type="notes">
|
||
<paragraph id="169A1202EB1E03699350B0FE8A22020B">
|
||
<heading id="7B50F3579423583F69132E3B6B857EA9" reason="title">Note.</heading>
|
||
</paragraph>
|
||
<paragraph id="C8C50D6A0DE0B3086B996720854DCE2F">
|
||
As iterated above, the confusion
|
||
<bibRefCitation id="DDDA34182C40E394D8587D7BCB0DACB7" author="de Saussure and Zehntner" firstAuthor="de Saussure" refId="B7" refString="de Saussure H, Zehntner L (1897) Atlas de l’histoire naturelle des Myriapodes. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Paris: Imprimérie nationale, Pl 1–15." year="1897">de Saussure and Zehntner (1897</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="707610046709EC4C10DBD46F52659CA6" author="de Saussure and Zehntner" firstAuthor="de Saussure" issue="4" journalOrPublisher="Abhandlungen der Senckenbergischen naturforschenden Gesellschaft" pagination="429-460" refId="B8" refString="de Saussure H, Zehntner L (1901) Myriopoden aus Madagaskar und Zanzibar, gesammelt von Dr. A. Voeltzkow. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 26 (4): 429–460." title="Myriopoden aus Madagaskar und Zanzibar, gesammelt von Dr. A. Voeltzkow." volume="26" year="1901">1901</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="ACE0F0CF0A3F4070BCB51F5B83231AE9" author="de Saussure and Zehntner" editor="Grandidier A" firstAuthor="de Saussure" journalOrPublisher="Imprimérie nationale, Paris" pagination="1-356" refId="B9" refString="de Saussure H, Zehntner L (1902) Myriapodes de Madagascar. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Imprimérie nationale, Paris, 1–356." title="Myriapodes de Madagascar." volumeTitle="Histoire physique, naturelle et politique de Madagascar." year="1902">1902</bibRefCitation>
|
||
) created by the initial usage of the specific epithet
|
||
<taxonomicName id="952EE57C9C68D173712CB237C1BBF769" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="sakalava">
|
||
<emphasis id="4727FF1F4644F954DBDE8361065804E1" italics="true">sakalava</emphasis>
|
||
</taxonomicName>
|
||
for two species of
|
||
<taxonomicName id="674CEB224A1115EF0A148804AC1E8F89" authority="Latreille, 1761" authorityName="Latreille" authorityYear="1761" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E59B9666CF1D956310C4DBDF37AA148D" italics="true">Polydesmus</emphasis>
|
||
Latreille, 1761
|
||
</taxonomicName>
|
||
has been resolved only recently (
|
||
<bibRefCitation id="727B5DF402A847354FE7E748314EE06D" DOI="10.11646/zootaxa.4413.2.11" author="Mesibov" etAl="et al." firstAuthor="Mesibov" issue="2" journalOrPublisher="Zootaxa" pagination="389-391" refId="B20" refString="Mesibov R, Wesener T, Hollier J (2018) A replacement name for Dalodesmus sakalava (de Saussure & Zehntner, 1901) (Diplopoda: Polydesmida: Dalodesmidae). Zootaxa 4413 (2): 389–391. https://doi.org/10.11646/zootaxa.4413.2.11" title="A replacement name for Dalodesmus sakalava (de Saussure & Zehntner, 1901) (Diplopoda: Polydesmida: Dalodesmidae)." volume="4413" year="2018">Mesibov et al. 2018</bibRefCitation>
|
||
). While the original type locality of the species is unknown, Franz Sikora is known to have collected mainly around the capital,
|
||
<collectingRegion id="06D6F27721EA38924389D9370CD47638" country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
|
||
. The freshly collected male specimen comes from Andasibe (
|
||
<bibRefCitation id="F3AA011916AF8D4CD0B268D56C32D21D" DOI="10.4314/mcd.v15i1.3" author="Spelzhausen" etAl="et al." firstAuthor="Spelzhausen" issue="1" journalOrPublisher="Madagascar Conservation and Development" pagination="1-8" refId="B21" refString="Spelzhausen L, Wesener T, Schütte K (2020) Vegetation thresholds for the occurrence of millipedes (Diplopoda) in different tropical forest types in Andasibe, Madagascar. Madagascar Conservation and Development 15 (1): 1–8. http://doi.org/10.4314/mcd.v15i1.3" title="Vegetation thresholds for the occurrence of millipedes (Diplopoda) in different tropical forest types in Andasibe, Madagascar." volume="15" year="2020">Spelzhausen et al. 2020</bibRefCitation>
|
||
), one of the largest remaining blocks of natural vegetation, on the old way from the coast to the capital.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EYLAE" type="material">
|
||
<paragraph id="A008980FE7A47883019BDBC82FFA1569">
|
||
<heading id="EFF15624B1CE8612AB61B3224F1015BC" reason="title">Material examined.</heading>
|
||
</paragraph>
|
||
<paragraph id="483A4531EF0BB391AF98BD86EF1B5FBC">
|
||
•
|
||
<materialsCitation id="7CEB67C9B751BE21CB9133633301D171" collectionCode="MHNG" collectorName="Madagascar' & Sikora" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<specimenCount id="59020AD341DAE2ABAA8CACE09E8590F1" count="1" type="female">♀</specimenCount>
|
||
<emphasis id="1948529E68543BE101170A0C3D5E5916" bold="true" italics="true">
|
||
<typeStatus id="651443F9A4D3ACACD2D36662B89F2481" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
, fragmented (
|
||
<named-content id="958ACC63865B31687D48AAC6371173E0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-dhistoire-naturelle-6" xlink_title="Muséum d’histoire naturelle de la Ville de Genève, Geneva, Switzerland">
|
||
<collectionCode id="6F4F3AD1607B8FE8203E46F5D36F9D1D" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" type="Museum">MHNG</collectionCode>
|
||
</named-content>
|
||
), ‘
|
||
<collectorName id="A4C9F3238825CE39D32DCC41418F4F21">Madagascar’</collectorName>
|
||
, coll.
|
||
<collectorName id="311A1237EF17466088BE6FBE2409ADFA">Sikora</collectorName>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="BE72F458286FFDB34853D580328FE4F1" collectingDate="2017-04" collectionCode="ZFMK" collectorName="L. Spelzhausen & G. Rakotonirina" country="Madagascar" county="Analamazaotra Forest Station" location="Moramanga District" municipality="Andasibe National Park" specimenCode="MYR 12217" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<specimenCount id="46664B883200B2FFA0C1D3D3D4C94C01" count="1" type="male">1 ♂</specimenCount>
|
||
,
|
||
<named-content id="025DCF1E1B882F3C1AA267D4AB43DEB4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoologisches Forschungsinstitut und Museum Alexander Koenig, Germany">
|
||
<collectionCode id="DA9864250BFF2D4C2DB1ADBAC1A95C9D" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||
</named-content>
|
||
<specimenCode id="C360DEA580F6F7927E49D8B8D1C24B9D" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">MYR 12217</specimenCode>
|
||
;
|
||
<collectingCountry id="7A95516C939ABEE5BE57DD61A688E5D3" name="Madagascar">Madagascar</collectingCountry>
|
||
,
|
||
<location id="F08ADB46DBBB850AB393115CF1774A45" LSID="urn:lsid:plazi:treatment:51691BCD50735AB5A5F1FFC3AC39BCB4:F08ADB46DBBB850AB393115CF1774A45" country="Madagascar" county="Analamazaotra Forest Station" municipality="Andasibe National Park" name="Moramanga District">Moramanga District</location>
|
||
,
|
||
<collectingMunicipality id="4D8151A21855A99EE07AAF2721933271">Andasibe National Park</collectingMunicipality>
|
||
(= Périnet),
|
||
<collectingCounty id="CA4F7C7FC11CD20E4943D057B0886CDC">Analamazaotra Forest Station</collectingCounty>
|
||
, secondary forest,
|
||
<taxonomicName id="BBFB2E4B5ED044F69E40ED1C4C6CF774" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="11881DBF8C080ED7B174C327561510A3" italics="true">Eucalyptus</emphasis>
|
||
</taxonomicName>
|
||
1909 plantation, sifted leaf litter,
|
||
<date id="69AF92F0EF411C530F60904FD11EF124" value="2017-04">
|
||
<collectingDate id="72053861127B42FA1A41B52F1647B265" value="2017-04">IV. 2017</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="E39801884FB1F0039DDBA223D1ACF6E4">L. Spelzhausen</collectorName>
|
||
and
|
||
<collectorName id="48C220C81D370EEAB57793CC27AB7DB2">G. Rakotonirina</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ESMAE" type="description">
|
||
<paragraph id="F17E116DB5C60FDA7C53CB98AB708D5A">
|
||
<heading id="22EA2B0B3B061327DD6E59B60A5FDEAE" reason="title">Brief description.</heading>
|
||
</paragraph>
|
||
<paragraph id="80354ADFB46760F74E8A3E11BD19541C">
|
||
(After
|
||
<bibRefCitation id="C1ABC0B5E20C1F2A4006169AD6FFE2A3" author="de Saussure and Zehntner" firstAuthor="de Saussure" refId="B7" refString="de Saussure H, Zehntner L (1897) Atlas de l’histoire naturelle des Myriapodes. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Paris: Imprimérie nationale, Pl 1–15." year="1897">de Saussure and Zehntner 1897</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="8140D7410C9B046D3A88811E1D2FC9FE" author="de Saussure and Zehntner" editor="Grandidier A" firstAuthor="de Saussure" journalOrPublisher="Imprimérie nationale, Paris" pagination="1-356" refId="B9" refString="de Saussure H, Zehntner L (1902) Myriapodes de Madagascar. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Imprimérie nationale, Paris, 1–356." title="Myriapodes de Madagascar." volumeTitle="Histoire physique, naturelle et politique de Madagascar." year="1902">1902</bibRefCitation>
|
||
.) A single line drawing (Fig.
|
||
<figureCitation id="FB478DD1B8EBA022F8007C69DD0C369C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♀ holotype MHNG (G, H), ♂ specimen (ZFMK MYR 12217). Multi-layer photographs and drawings A habitus, dorsal view B habitus, lateral view C habitus, ventral view D detail of midbody rings, dorsal view E detail of anterior body rings and gonopod, lateral view F detail of anterior rings and gonopods, ventral view G drawing from the original description in de Saussure and Zehntner (1897) H detail of body ring, ♀ holotype I gonopod still attached, ventral view. Abbreviations: lb = lateral branch; mb = mesal branch; sl = solenomere branch. Scale bar: 0.3 mm (H)." figureDoi="10.3897/zookeys.1223.139346.figure1" httpUri="https://binary.pensoft.net/fig/1215825">1 G</figureCitation>
|
||
) and its caption reading “
|
||
<taxonomicName id="8F6E0601BBDA0979BB1B3DD7814150E8" authority="" in de Saussure and Zehntner (1897)" authorityName="" in de Saussure and Zehntner" authorityYear="1897" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="sakalava">
|
||
<emphasis id="A12EE301D37BB5E8AA415C6565500AA4" italics="true">Polydesmus sakalava</emphasis>
|
||
” in
|
||
<bibRefCitation id="326699CDE9EDE0D24AF7C929771DAF9B" author="de Saussure H & Zehntner L" firstAuthor="de Saussure" refId="B7" refString="de Saussure H, Zehntner L (1897) Atlas de l’histoire naturelle des Myriapodes. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Paris: Imprimérie nationale, Pl 1–15." year="1897">de Saussure and Zehntner (1897)</bibRefCitation>
|
||
</taxonomicName>
|
||
serve as the original description. Combined with a later verbal description of the adult
|
||
<specimenCount id="6287D20BE33A7AF330F1FBFC2FE1B4EF" count="1" type="female">♀</specimenCount>
|
||
<typeStatus id="ACA31AEBFB4F0655068272594FE3B58E" type="holotype">holotype</typeStatus>
|
||
, by
|
||
<bibRefCitation id="CA0BE03B8C12E0DEAC6D7122417CE194" author="de Saussure H & Zehntner L" editor="Grandidier A" firstAuthor="de Saussure" journalOrPublisher="Imprimérie nationale, Paris" pagination="1-356" refId="B9" refString="de Saussure H, Zehntner L (1902) Myriapodes de Madagascar. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Imprimérie nationale, Paris, 1–356." title="Myriapodes de Madagascar." volumeTitle="Histoire physique, naturelle et politique de Madagascar." year="1902">de Saussure and Zehntner (1902)</bibRefCitation>
|
||
, the following relevant information can be obtained: body pale reddish, subcylindrical,
|
||
<quantity id="6DCC532A858D465051FE1A007F3F64B6" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
|
||
long,
|
||
<quantity id="8AA40EE2180B0CAA8440BB51655BD222" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
wide, with a strongly convex dorsum and very narrow, declivous, and subrectangular paraterga.
|
||
</paragraph>
|
||
<caption id="EEAC7374801FCCCFB5CA4996DD2A5CE5" ID-DOI="10.3897/zookeys.1223.139346.figure1" ID-arpha="20ADB95F-127B-57C7-AA30-1C985F8C8C2D" httpUri="https://binary.pensoft.net/fig/1215825" startId="F1">
|
||
<paragraph id="17B48536476F048D4E77FA6A7B8DDC4A">
|
||
<label id="750915E3DBD08B12D8556AD4B156F6A1">Figure 1.</label>
|
||
</paragraph>
|
||
<paragraph id="90060F362D8FF00BDAB92A787AED0273">
|
||
<taxonomicName id="56B72D06FCA746F58C5541EB3CDDB2A5" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="sakalava">
|
||
<emphasis id="28B0DEAA0DBC7693FAC3F7A173D6481B" italics="true">Phymatodesmus sakalava</emphasis>
|
||
(
|
||
<bibRefCitation id="4CDD1C861B5CBADA98FD11AFFA581B1E" author="de Saussure H & Zehntner L" refId="B7" refString="de Saussure H, Zehntner L (1897) Atlas de l'histoire naturelle des Myriapodes. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Paris: Imprimerie nationale, Pl 1 - 15." year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ♀ holotype
|
||
<named-content id="8B9EE8537904DD2023F4E04BC203BC87" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-dhistoire-naturelle-6" xlink_title="Muséum d’histoire naturelle de la Ville de Genève, Geneva, Switzerland">MHNG</named-content>
|
||
(G, H), ♂ specimen (
|
||
<named-content id="4ECDA029D96FA2A74380B89F9143273F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoologisches Forschungsinstitut und Museum Alexander Koenig, Germany">ZFMK</named-content>
|
||
MYR 12217). Multi-layer photographs and drawings
|
||
<emphasis id="CBA9D7D097ADCFF8F956876C0CAC7ABA" bold="true">A</emphasis>
|
||
habitus, dorsal view
|
||
<emphasis id="37F97ABE20D5D5971BB03BC679C3DFE2" bold="true">B</emphasis>
|
||
habitus, lateral view
|
||
<emphasis id="2A011AE019BD13281D8DC37FAE8E8AD2" bold="true">C</emphasis>
|
||
habitus, ventral view
|
||
<emphasis id="1B1ED0F62C42808214CDB28BD614B3B2" bold="true">D</emphasis>
|
||
detail of midbody rings, dorsal view
|
||
<emphasis id="B502B1BA2CAEB1C9B2CC381ED7567A69" bold="true">E</emphasis>
|
||
detail of anterior body rings and gonopod, lateral view
|
||
<emphasis id="177298E763350D142305B4E62FF19BB7" bold="true">F</emphasis>
|
||
detail of anterior rings and gonopods, ventral view
|
||
<emphasis id="45AE21929EDE216C653ACDF98EE45D8F" bold="true">G</emphasis>
|
||
drawing from the original description in
|
||
<bibRefCitation id="CE31A5B06A4FB66BBFFEE461FBEDBC83" author="de Saussure H & Zehntner L" firstAuthor="de Saussure" refId="B7" refString="de Saussure H, Zehntner L (1897) Atlas de l’histoire naturelle des Myriapodes. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Paris: Imprimérie nationale, Pl 1–15." year="1897">de Saussure and Zehntner (1897)</bibRefCitation>
|
||
<emphasis id="660AAF2DA0BE7C5BD2DB39074586E9CC" bold="true">H</emphasis>
|
||
detail of body ring, ♀ holotype
|
||
<emphasis id="4C5B057B0A337505C10E2A3EA592D8A1" bold="true">I</emphasis>
|
||
gonopod still attached, ventral view. Abbreviations: lb = lateral branch; mb = mesal branch; sl = solenomere branch. Scale bar: 0.3 mm (
|
||
<emphasis id="3E4C14F0612A77388680CA681C9F054B" bold="true">H</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E7C274A58D02A011421CDA8BB4C8070">
|
||
<taxonomicName id="3BDF6FE6A8F65300C0F4ABC7B9C475B5" authorityName="de Saussure & Zehntner" authorityYear="1902" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="15F7095661BC07DD28F2499850148CBD" italics="true">Phymatodesmus</emphasis>
|
||
</taxonomicName>
|
||
s
|
||
<emphasis id="4FCA02DAB56858AE8F78979AA5E6E4C8" italics="true">akalava</emphasis>
|
||
can be readily distinguished from other known Malagasy
|
||
<taxonomicName id="5F324EFF7B2A84E13EBA1E1091F65D61" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Dalodesmidae" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="family">Dalodesmidae</taxonomicName>
|
||
: the small size, a subcylindrical body, the mostly small, conical, and sharp tuberculations on midbody metaterga arranged in four unusually regular transverse rows, and strikingly small, narrow, and rectangular paraterga.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EBQAE" type="description">
|
||
<paragraph id="2BC1153304C7893878FDB48CCBA121D0">
|
||
<heading id="3FE29887B2BD6C0FC18E1545127620E3" reason="title">First description of the male.</heading>
|
||
</paragraph>
|
||
<paragraph id="F33501303799B0B6867C8EE641B9E55F">
|
||
Length ca
|
||
<quantity id="9E619CF29DBAA387C0AD7F20FE416AC1" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
|
||
, width of midbody pro- and metazona 0.75 and
|
||
<quantity id="F3A1BAA66267EF76020DE7DF36244B3E" metricMagnitude="-3" metricUnit="m" metricValue="1.05" unit="mm" value="1.05">1.05 mm</quantity>
|
||
, respectively.
|
||
</paragraph>
|
||
<paragraph id="4F1D61A3DED0CD9519F2B44F7DFB2141">Colouration (freshly preserved in ethanol) brown; prozona, basal parts of legs, mandibles, and paraprocts paler. Epicranium grey, antennae and apical parts of legs faded grey.</paragraph>
|
||
<paragraph id="402B95697C6D1620625B6E34D3D69714">
|
||
Body with 20 rings. Tegument mainly dull (Fig.
|
||
<figureCitation id="BDEB5AA1152C98EA97DD587CA16CFB12" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♀ holotype MHNG (G, H), ♂ specimen (ZFMK MYR 12217). Multi-layer photographs and drawings A habitus, dorsal view B habitus, lateral view C habitus, ventral view D detail of midbody rings, dorsal view E detail of anterior body rings and gonopod, lateral view F detail of anterior rings and gonopods, ventral view G drawing from the original description in de Saussure and Zehntner (1897) H detail of body ring, ♀ holotype I gonopod still attached, ventral view. Abbreviations: lb = lateral branch; mb = mesal branch; sl = solenomere branch. Scale bar: 0.3 mm (H)." figureDoi="10.3897/zookeys.1223.139346.figure1" httpUri="https://binary.pensoft.net/fig/1215825">1 A – F</figureCitation>
|
||
), microgranulate to microtuberculate throughout (Fig.
|
||
<figureCitation id="E5C3636DDDE34AACC7B4FE0A04F98B1B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♀ holotype MHNG (G, H), ♂ specimen (ZFMK MYR 12217). Multi-layer photographs and drawings A habitus, dorsal view B habitus, lateral view C habitus, ventral view D detail of midbody rings, dorsal view E detail of anterior body rings and gonopod, lateral view F detail of anterior rings and gonopods, ventral view G drawing from the original description in de Saussure and Zehntner (1897) H detail of body ring, ♀ holotype I gonopod still attached, ventral view. Abbreviations: lb = lateral branch; mb = mesal branch; sl = solenomere branch. Scale bar: 0.3 mm (H)." figureDoi="10.3897/zookeys.1223.139346.figure1" httpUri="https://binary.pensoft.net/fig/1215825">1 G, H</figureCitation>
|
||
), even surfaces of prozona and of metazona below paraterga finely microgranulate, sterna granulate.
|
||
</paragraph>
|
||
<paragraph id="7BFDFA6F116D5BD8BB2FD0738F58FD6F">
|
||
Head also densely microtuberculate or granulate throughout, micropilose; epicranial suture thin, but distinct; genae squarish, set off ventrally from gnathochilarial stipes by a small, but evident ridge. Interantennal isthmus ~ 2 × diameter of antennal socket (Fig.
|
||
<figureCitation id="7A8E9B2AFC904CA0EB6CE1CE89DC3914" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♀ holotype MHNG (G, H), ♂ specimen (ZFMK MYR 12217). Multi-layer photographs and drawings A habitus, dorsal view B habitus, lateral view C habitus, ventral view D detail of midbody rings, dorsal view E detail of anterior body rings and gonopod, lateral view F detail of anterior rings and gonopods, ventral view G drawing from the original description in de Saussure and Zehntner (1897) H detail of body ring, ♀ holotype I gonopod still attached, ventral view. Abbreviations: lb = lateral branch; mb = mesal branch; sl = solenomere branch. Scale bar: 0.3 mm (H)." figureDoi="10.3897/zookeys.1223.139346.figure1" httpUri="https://binary.pensoft.net/fig/1215825">1 C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D6E2D5ACBEB146DAB62DE80B63D6AF57">Antennae very short and rather clavate, in situ reaching back past ring 2 when stretched dorsally, very densely setose and microgranulate. In length, antennomere 6> 5 = 3> 4> 2> 1 = 7; antennomere 6 the largest and the highest, antennomeres 5 and 6 each with a small, round, distodorsal knob, most likely beset with sensory cones.</paragraph>
|
||
<paragraph id="B0F04E6DA11E4C0E595DF30E1B15933A">
|
||
In width, collum = head <ring 2> 3 = 4–16; thereafter body gradually tapering towards telson (Fig.
|
||
<figureCitation id="9D92F134B95689C88D6335DD18ACB308" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♀ holotype MHNG (G, H), ♂ specimen (ZFMK MYR 12217). Multi-layer photographs and drawings A habitus, dorsal view B habitus, lateral view C habitus, ventral view D detail of midbody rings, dorsal view E detail of anterior body rings and gonopod, lateral view F detail of anterior rings and gonopods, ventral view G drawing from the original description in de Saussure and Zehntner (1897) H detail of body ring, ♀ holotype I gonopod still attached, ventral view. Abbreviations: lb = lateral branch; mb = mesal branch; sl = solenomere branch. Scale bar: 0.3 mm (H)." figureDoi="10.3897/zookeys.1223.139346.figure1" httpUri="https://binary.pensoft.net/fig/1215825">1 A – C</figureCitation>
|
||
). Collum transversely suboval, regularly and broadly rounded laterally, densely tuberculate, most tuberculations being circular, evident, equipped with very short, mostly subclavate setae and arranged in 15–17 lateral, 8–9 transverse, rather irregular, arcuated rows. Metaterga 2–4 narrow, each with four, similar, transverse, arcuated, circular rows of setigerous tubercles, following metaterga each largely with five such rows (Fig.
|
||
<figureCitation id="F412FB83202F8EF19BD03C58554CAE3D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♂ specimen (ZFMK MYR 12217). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, lateral view C midbody ring, paranota with ozopore D details of surface structures, dorsal view E detail of surface structures of prozonite and metazonite F endotergum. Scale bars: 100 µm (A – C); 10 µm (D); 20 µm (E); 10 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure2" httpUri="https://binary.pensoft.net/fig/1215826">2 A, B</figureCitation>
|
||
). Paraterga short, rectangular, strongly declivous, posterior margin straight (Fig.
|
||
<figureCitation id="BE0733E1D7E6C221E33322812A8009AF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♂ specimen (ZFMK MYR 12217). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, lateral view C midbody ring, paranota with ozopore D details of surface structures, dorsal view E detail of surface structures of prozonite and metazonite F endotergum. Scale bars: 100 µm (A – C); 10 µm (D); 20 µm (E); 10 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure2" httpUri="https://binary.pensoft.net/fig/1215826">2 B, C</figureCitation>
|
||
). Lateral margin of paraterga beset with 5–6 similarly circular, setigerous (Fig.
|
||
<figureCitation id="D92C313EE30DF1734D34C8D47D85DC00" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♂ specimen (ZFMK MYR 12217). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, lateral view C midbody ring, paranota with ozopore D details of surface structures, dorsal view E detail of surface structures of prozonite and metazonite F endotergum. Scale bars: 100 µm (A – C); 10 µm (D); 20 µm (E); 10 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure2" httpUri="https://binary.pensoft.net/fig/1215826">2 D</figureCitation>
|
||
) tubercles / lobulations. Ozopores inconspicuous, opening laterally near penultimate lateral lobulation on pore-bearing rings 5, 7, 9, 10, 12, 13, 15–19. Strictures between pro- and metazona narrow and rather deep, nearly smooth (Fig.
|
||
<figureCitation id="C4D72DAB2C28120C5DF5F68053136F25" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♂ specimen (ZFMK MYR 12217). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, lateral view C midbody ring, paranota with ozopore D details of surface structures, dorsal view E detail of surface structures of prozonite and metazonite F endotergum. Scale bars: 100 µm (A – C); 10 µm (D); 20 µm (E); 10 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure2" httpUri="https://binary.pensoft.net/fig/1215826">2 E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="2C53E717EDD437F51D7E324034937291" ID-DOI="10.3897/zookeys.1223.139346.figure2" ID-arpha="282CE3CB-CECC-5784-8949-74E7C0A1BD8B" httpUri="https://binary.pensoft.net/fig/1215826" startId="F2">
|
||
<paragraph id="B9C844FFDA5E9CB2DEB046FD7DCF42AF">
|
||
<label id="A9FBE8E403C5185D4BB1752600FDDE14">Figure 2.</label>
|
||
</paragraph>
|
||
<paragraph id="5C97FF19978F4F54F408773312487EFF">
|
||
<taxonomicName id="DCABBC27FAF27DA220A27D78A0DE0BAE" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="sakalava">
|
||
<emphasis id="A683D81B0F4ECF8EADDCD74E829A6CE9" italics="true">Phymatodesmus sakalava</emphasis>
|
||
(
|
||
<bibRefCitation id="0EB2D3876CEF8CFA3D02648393FFE03B" author="de Saussure H & Zehntner L" refId="B7" refString="de Saussure H, Zehntner L (1897) Atlas de l'histoire naturelle des Myriapodes. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Paris: Imprimerie nationale, Pl 1 - 15." year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ♂ specimen (
|
||
<named-content id="FB37EC1A8DE59AC4E61AA3DCBDAC2019" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoologisches Forschungsinstitut und Museum Alexander Koenig, Germany">ZFMK</named-content>
|
||
MYR 12217).
|
||
<abbrev id="ABBRID0EGSAE" xlink_title="scanning electron microscopy">SEM</abbrev>
|
||
micrographs
|
||
<emphasis id="FCCA3A52546783FFD40B63609774DE51" bold="true">A</emphasis>
|
||
midbody ring with pro- and metazonite, dorsal view
|
||
<emphasis id="B14751930D2C29C548CD499FF546E962" bold="true">B</emphasis>
|
||
midbody ring, lateral view
|
||
<emphasis id="42BF68D6A069A32C285F36BC0897401D" bold="true">C</emphasis>
|
||
midbody ring, paranota with ozopore
|
||
<emphasis id="A7DDA879F3ADB1AB7318C4EFC592E299" bold="true">D</emphasis>
|
||
details of surface structures, dorsal view
|
||
<emphasis id="4A510467B243FFD283ED2F6EC96B9D46" bold="true">E</emphasis>
|
||
detail of surface structures of prozonite and metazonite
|
||
<emphasis id="F1A6166F09162C6BB7061B9834EEFA49" bold="true">F</emphasis>
|
||
endotergum. Scale bars: 100 µm (
|
||
<emphasis id="DEFE8386DFFCAD6D88367F563C422D24" bold="true">A – C</emphasis>
|
||
); 10 µm (
|
||
<emphasis id="46E53F71EAC1D06335033248031C8042" bold="true">D</emphasis>
|
||
); 20 µm (
|
||
<emphasis id="590579AAA35A036299FE9A68B488D8CD" bold="true">E</emphasis>
|
||
); 10 µm (
|
||
<emphasis id="0DCA67989702A16CF3221529A161BCCC" bold="true">F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="429CAFCABA2437C282C2FE2616F33D96">
|
||
Telson: Epiproct small, conical, and subtruncate at tip. Hypoproct trapeziform, with 1 + 1 setae borne on distinct oblong knobs at caudal margin. Paraprocts with 2 + 2 setae on triangular and projecting knobs (Fig.
|
||
<figureCitation id="B9126ED145B4F33D5A3E8F1267DF9AE4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♂ specimen (ZFMK MYR 12217). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, lateral view C midbody ring, paranota with ozopore D details of surface structures, dorsal view E detail of surface structures of prozonite and metazonite F endotergum. Scale bars: 100 µm (A – C); 10 µm (D); 20 µm (E); 10 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure2" httpUri="https://binary.pensoft.net/fig/1215826">2 C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E9DA74C2F1FE28B75DD2B8C485D4BD16">
|
||
Limbus very thin, small, and entire. Neither an axial line nor pleurosternal carinae (Fig.
|
||
<figureCitation id="308F994C75D8AF9F13F8AE6DCE7E25D5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♀ holotype MHNG (G, H), ♂ specimen (ZFMK MYR 12217). Multi-layer photographs and drawings A habitus, dorsal view B habitus, lateral view C habitus, ventral view D detail of midbody rings, dorsal view E detail of anterior body rings and gonopod, lateral view F detail of anterior rings and gonopods, ventral view G drawing from the original description in de Saussure and Zehntner (1897) H detail of body ring, ♀ holotype I gonopod still attached, ventral view. Abbreviations: lb = lateral branch; mb = mesal branch; sl = solenomere branch. Scale bar: 0.3 mm (H)." figureDoi="10.3897/zookeys.1223.139346.figure1" httpUri="https://binary.pensoft.net/fig/1215825">1 A – C</figureCitation>
|
||
). Posterior margin of metazona a row of dense, elongate, apically microdenticulate (with 6–9 indentations (Fig.
|
||
<figureCitation id="945ECE7D2322407416E014AA1C7F37F6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♂ specimen (ZFMK MYR 12217). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, lateral view C midbody ring, paranota with ozopore D details of surface structures, dorsal view E detail of surface structures of prozonite and metazonite F endotergum. Scale bars: 100 µm (A – C); 10 µm (D); 20 µm (E); 10 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure2" httpUri="https://binary.pensoft.net/fig/1215826">2 F</figureCitation>
|
||
), subrectangular projections (Fig.
|
||
<figureCitation id="70EA1429363BC13476F50F85FD236A45" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♂ specimen (ZFMK MYR 12217). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, lateral view C midbody ring, paranota with ozopore D details of surface structures, dorsal view E detail of surface structures of prozonite and metazonite F endotergum. Scale bars: 100 µm (A – C); 10 µm (D); 20 µm (E); 10 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure2" httpUri="https://binary.pensoft.net/fig/1215826">2 F</figureCitation>
|
||
)).
|
||
</paragraph>
|
||
<paragraph id="0C0CF4B56DC649C9C34F7E81023CC6F3">
|
||
Gonopodal aperture roundly pentagonal, large, taking up ~ 2 / 3 width of metazonum 7, clearly open and drawn into metazonum 6 (Fig.
|
||
<figureCitation id="8A25FA223963FDA935757E32B310E607" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♀ holotype MHNG (G, H), ♂ specimen (ZFMK MYR 12217). Multi-layer photographs and drawings A habitus, dorsal view B habitus, lateral view C habitus, ventral view D detail of midbody rings, dorsal view E detail of anterior body rings and gonopod, lateral view F detail of anterior rings and gonopods, ventral view G drawing from the original description in de Saussure and Zehntner (1897) H detail of body ring, ♀ holotype I gonopod still attached, ventral view. Abbreviations: lb = lateral branch; mb = mesal branch; sl = solenomere branch. Scale bar: 0.3 mm (H)." figureDoi="10.3897/zookeys.1223.139346.figure1" httpUri="https://binary.pensoft.net/fig/1215825">1 I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="744C3B46CC2CE37F0768E9F54A4BBE96">First two leg pairs shorter and thicker than other legs. Midbody legs incrassate, medium in length, as long as body height, with small, stout, abundant, and usually curved setae with admixture of sphaerotrichomes ventrally on all podomeres; gonopores on coxae 2 inconspicuous, prefemora not bulged laterally; claws simple, very small; in length, tarsus> femur> prefemur> coxa> tibia = postfemur.</paragraph>
|
||
<paragraph id="305F3DDEC95FFFC160077E92EBCAD9B5">
|
||
Gonopods (Figs
|
||
<figureCitation id="D1DBA5A448662F4CBCFDEE21B8A71ACE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♀ holotype MHNG (G, H), ♂ specimen (ZFMK MYR 12217). Multi-layer photographs and drawings A habitus, dorsal view B habitus, lateral view C habitus, ventral view D detail of midbody rings, dorsal view E detail of anterior body rings and gonopod, lateral view F detail of anterior rings and gonopods, ventral view G drawing from the original description in de Saussure and Zehntner (1897) H detail of body ring, ♀ holotype I gonopod still attached, ventral view. Abbreviations: lb = lateral branch; mb = mesal branch; sl = solenomere branch. Scale bar: 0.3 mm (H)." figureDoi="10.3897/zookeys.1223.139346.figure1" httpUri="https://binary.pensoft.net/fig/1215825">1 I</figureCitation>
|
||
,
|
||
<figureCitation id="E1FD3F88AF072E46455EA03ED8503F55" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phymatodesmus sakalava (de Saussure & Zehntner, 1897), ♂ specimen (ZFMK MYR 12217). Multi-layered photographs and drawings, gonopod A posterior view B lateral and mesal view C lateral view D posterior view E anterior view F lateral view. Abbreviations: fe = femorite; lb = lateral branch; mb = mesal branch; sl = solenomere branch." figureDoi="10.3897/zookeys.1223.139346.figure3" httpUri="https://binary.pensoft.net/fig/1215827">3 A – F</figureCitation>
|
||
) relatively simple. Both coxite and prefemorite very short, fused medially, prefemorite setose. Femorites (fe) contiguous medially, densely setose both ventrally and laterally, rather stout (~ 2 × as long as acropodites), suberect and clearly flattened dorsoventrally. Acropodite tripartite, divided into a dorsomedial, long, simple and subsecuriform solenomere (sl), a simple and subspiniform lateral branch (lb), both sl and lb being subequal in length, but clearly shorter than the longest, simple, rather finger-shaped, apically roundly and irregularly trifid, mesal branch (mb).
|
||
</paragraph>
|
||
<caption id="1ED81C530346DFFCA74A11A0B91AB895" ID-DOI="10.3897/zookeys.1223.139346.figure3" ID-arpha="F7FF77AA-17CC-5BC1-86CD-35C4782B0D2C" httpUri="https://binary.pensoft.net/fig/1215827" startId="F3">
|
||
<paragraph id="ADE38D6F04B5955BD92B6A4C6E130CED">
|
||
<label id="3284136540C7512980C9789A4110481E">Figure 3.</label>
|
||
</paragraph>
|
||
<paragraph id="6068028D2900B3CE6C24DD2A5BF1DCD5">
|
||
<taxonomicName id="CB2ACDD272DAC4B725EBD6BE9F3964E3" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="sakalava">
|
||
<emphasis id="2E3FD1CF9E711351B3A45BF845750B28" italics="true">Phymatodesmus sakalava</emphasis>
|
||
(
|
||
<bibRefCitation id="32F6C56F1B257EEABE0A511C26FBC238" author="de Saussure H & Zehntner L" refId="B7" refString="de Saussure H, Zehntner L (1897) Atlas de l'histoire naturelle des Myriapodes. In: Grandidier A (Ed.) Histoire physique, naturelle et politique de Madagascar. Paris: Imprimerie nationale, Pl 1 - 15." year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ♂ specimen (
|
||
<named-content id="C179E78C47126DFF83C98C88D0795DBE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoologisches Forschungsinstitut und Museum Alexander Koenig, Germany">ZFMK</named-content>
|
||
MYR 12217). Multi-layered photographs and drawings, gonopod
|
||
<emphasis id="CE7A88748618E9A7F722066A4EFE1EDE" bold="true">A</emphasis>
|
||
posterior view
|
||
<emphasis id="807C24CF842F798AEEA4A2B13052976C" bold="true">B</emphasis>
|
||
lateral and mesal view
|
||
<emphasis id="36A4AD284D2A8F3A54E1CB2BE28832BC" bold="true">C</emphasis>
|
||
lateral view
|
||
<emphasis id="B8FCFDBE89F13A12F572484C034740EF" bold="true">D</emphasis>
|
||
posterior view
|
||
<emphasis id="5D9F2DBB9F22DD0C36B05007688DC5C0" bold="true">E</emphasis>
|
||
anterior view
|
||
<emphasis id="0A310B4EAE23D2311F869EC214171D5A" bold="true">F</emphasis>
|
||
lateral view. Abbreviations: fe = femorite; lb = lateral branch; mb = mesal branch; sl = solenomere branch.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ERVAE" type="remarks">
|
||
<paragraph id="9948F62FC44F3E75A67BFB1F2413F411">
|
||
<heading id="C1C8F839FA2B6F6FAF47B0C75C20E52A" reason="title">Remarks.</heading>
|
||
</paragraph>
|
||
<paragraph id="826CA7AE21249295292B6974582976C8">
|
||
Aside from the very obvious differences between
|
||
<taxonomicName id="786367DED23E58DCA1B069DE39A7441F" authorityName="de Saussure & Zehntner" authorityYear="1902" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F1026BDAAD84F18C3D9AC69DE41A9383" italics="true">Phymatodesmus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3E2B05EB0286A210895CD3DB586B5BA1" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AED1FAA38E2203559DE3030EA85890F3" italics="true">Dalodesmus</emphasis>
|
||
</taxonomicName>
|
||
in the development and shape of paraterga, clear-cut discrepancies also concern leg lengths (in
|
||
<taxonomicName id="94343F48DA19295AE4CCDE6F1F45B537" authorityName="de Saussure & Zehntner" authorityYear="1902" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BCA34E11CA0F56760A8DE942806C6A51" italics="true">Phymatodesmus</emphasis>
|
||
</taxonomicName>
|
||
, the whole legs, especially both postfemora and tibiae are shorter), the antennae are also shorter, the collum is as wide as the head, the head is completely micropilose, including the epicranium (vs glabrous in
|
||
<taxonomicName id="BF2E3528440C15A74D510CE3A460126D" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3CA55DF2F2147A7531B5B8D9DCB0719A" italics="true">Dalodesmus</emphasis>
|
||
</taxonomicName>
|
||
), the paraprocts of the telson lack triangular setiferous knobs / projections (vs 2 + 2 setae borne on knobs on the paraprocts), and the 1 + 1 setae on the hypoproct are borne on prominent knobs and better separated (vs placed closer to one another in
|
||
<taxonomicName id="2C1D7F108179E00DECDF252D38351632" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6E2EF12AB817B53A4509330A0D4E24F0" italics="true">Dalodesmus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |