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<mods:title id="09235DC9A58C21EA36E62D4F31E61273">Contribution to the knowledge of the Colombian Batodesmini (Polydesmida: Chelodesmidae) with the description of a new genus and species, and taxonomic insights on the genus Cormodesmus Chamberlin, 1923</mods:title>
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<mods:namePart id="2014F0CB3752F7A6CC3D2B1977B6AC99">Alvear, Santiago</mods:namePart>
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<subSubSection id="D8946513D42FCA6FD9861F172619FAD5" box="[151,523,1303,1330]" pageId="1" pageNumber="554" type="nomenclature">
<paragraph id="90313698D42FCA6FD9861F172619FAD5" blockId="1.[151,523,1303,1330]" box="[151,523,1303,1330]" pageId="1" pageNumber="554">
<heading id="CB7981F4D42FCA6FD9861F172619FAD5" bold="true" box="[151,523,1303,1330]" fontSize="11" level="1" pageId="1" pageNumber="554" reason="1">
<emphasis id="A2FAEA8AD42FCA6FD9861F172619FAD5" bold="true" box="[151,523,1303,1330]" pageId="1" pageNumber="554">
Genus
<taxonomicName id="578E4D1BD42FCA6FD9FF1F172584FAD6" authorityName="Alvear &amp; Romero-Rincon" authorityYear="2024" box="[238,406,1303,1330]" class="Diplopoda" family="Chelodesmidae" genus="Caucadesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A2FAEA8AD42FCA6FD9FF1F172584FAD6" bold="true" box="[238,406,1303,1330]" italics="true" pageId="1" pageNumber="554">Caucadesmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B9C957F1D42FCA6FD88F1F172619FAD5" box="[414,523,1303,1329]" pageId="1" pageNumber="554" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="D8946513D42FCA6FD9861F5F27A7FA9D" box="[151,949,1375,1402]" pageId="1" pageNumber="554" type="type_taxon">
<paragraph id="90313698D42FCA6FD9861F5F27A7FA9D" blockId="1.[151,1437,1375,1942]" box="[151,949,1375,1402]" pageId="1" pageNumber="554">
<emphasis id="A2FAEA8AD42FCA6FD9861F5F2520FA9D" bold="true" box="[151,306,1375,1401]" pageId="1" pageNumber="554">
<typeStatus id="4F35883AD42FCA6FD9861F5F24C0FA9D" box="[151,210,1375,1401]" pageId="1" pageNumber="554">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="578E4D1BD42FCA6FD8281F5F2651FA9D" authorityName="Alvear &amp; Romero-Rincon" authorityYear="2024" box="[313,579,1375,1401]" class="Diplopoda" family="Chelodesmidae" genus="Caucadesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="lucreciae" status="sp. nov.">
<emphasis id="A2FAEA8AD42FCA6FD8281F5F2651FA9D" box="[313,579,1375,1401]" italics="true" pageId="1" pageNumber="554">Caucadesmus lucreciae</emphasis>
</taxonomicName>
<emphasis id="A2FAEA8AD42FCA6FDB5B1F5F26B1FA9D" bold="true" box="[586,675,1375,1401]" pageId="1" pageNumber="554">
<taxonomicNameLabel id="B9C957F1D42FCA6FDB5B1F5F26B1FA9D" box="[586,675,1375,1401]" pageId="1" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by present designation.
</paragraph>
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<subSubSection id="D8946513D42FCA6FD9D61F8325D2FA02" pageId="1" pageNumber="554" type="etymology">
<paragraph id="90313698D42FCA6FD9D61F8325D2FA02" blockId="1.[151,1437,1375,1942]" pageId="1" pageNumber="554">
<emphasis id="A2FAEA8AD42FCA6FD9D61F83255CFA79" bold="true" box="[199,334,1411,1437]" pageId="1" pageNumber="554">Etymology.</emphasis>
The generic name is formed by a combination of the word Cauca in reference to the
<collectingRegion id="524AF87AD42FCA6FDC001F832145FA79" box="[1297,1367,1411,1437]" country="Colombia" name="Cauca" pageId="1" pageNumber="554">Cauca</collectingRegion>
River in
<collectingCountry id="E8997608D42FCA6FD9A71FA72535FA25" box="[182,295,1447,1473]" name="Colombia" pageId="1" pageNumber="554">Colombia</collectingCountry>
, and the suffix -
<emphasis id="A2FAEA8AD42FCA6FD8FA1FA7262FFA25" box="[491,573,1447,1473]" italics="true" pageId="1" pageNumber="554">desmus</emphasis>
, which is commonly used in generic names in the order
<taxonomicName id="578E4D1BD42FCA6FDDC01FA72177FA25" authorityName="Pocock" authorityYear="1887" box="[1233,1381,1447,1473]" class="Diplopoda" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
. The name is a masculine noun.
</paragraph>
</subSubSection>
<subSubSection id="D8946513D42FCA6FD9D61FEF274CF871" pageId="1" pageNumber="554" type="diagnosis">
<paragraph id="90313698D42FCA6FD9D61FEF2081F992" blockId="1.[151,1437,1375,1942]" pageId="1" pageNumber="554">
<emphasis id="A2FAEA8AD42FCA6FD9D61FEF2550F9ED" bold="true" box="[199,322,1519,1545]" pageId="1" pageNumber="554">Diagnosis.</emphasis>
A batodesminine genus belonging to the
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<emphasis id="A2FAEA8AD42FCA6FDA251FEF27DAF9ED" box="[820,968,1519,1545]" italics="true" pageId="1" pageNumber="554">Alocodesmus</emphasis>
</taxonomicName>
group, characterized by the absence of significant body modifications. No evident epicranial processes nor modified collum. Metazonites smooth, without ornamentation or tubercles; ozopores opening laterally on the paranota without developed peritremata. The sternite of segment 5 features a large protruding tubercle densely surrounded by tufts of long setae.
</paragraph>
<paragraph id="90313698D42FCA6FD9D61C7F26E8F8CE" blockId="1.[151,1437,1375,1942]" pageId="1" pageNumber="554">
The gonopod of
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<emphasis id="A2FAEA8AD42FCA6FD89D1C7F263AF97D" box="[396,552,1663,1689]" italics="true" pageId="1" pageNumber="554">Caucadesmus</emphasis>
</taxonomicName>
<emphasis id="A2FAEA8AD42FCA6FDB231C7F268DF97D" bold="true" box="[562,671,1663,1689]" pageId="1" pageNumber="554">
<taxonomicNameLabel id="B9C957F1D42FCA6FDB231C7F268DF97D" box="[562,671,1663,1689]" pageId="1" pageNumber="554" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
differs from other batodesminine genera, except
<taxonomicName id="578E4D1BD42FCA6FDDCE1C7F2522F959" authority="Attems, 1931" authorityName="Attems" authorityYear="1931" class="Diplopoda" family="Chelodesmidae" genus="Cordilleronomus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FDDCE1C7F218EF97D" box="[1247,1436,1663,1689]" italics="true" pageId="1" pageNumber="554">Cordilleronomus</emphasis>
<bibRefCitation id="F41F4B69D42FCA6FD9861CA32522F959" author="Attems, C." box="[151,304,1699,1725]" pageId="1" pageNumber="554" pagination="1 - 150" refId="ref3730" refString="Attems, C. (1931) Die Familie Leptodesmidae und andere Polydesmiden. Zoologica, 79, 1 - 150." type="journal article" year="1931">Attems, 1931</bibRefCitation>
</taxonomicName>
, by the absence of numerous long setae on the dorsal surface of
<emphasis id="A2FAEA8AD42FCA6FDD161CA3203BF959" bold="true" box="[1031,1065,1699,1725]" pageId="1" pageNumber="554">Cx</emphasis>
(vs the presence of such setae in
<taxonomicName id="578E4D1BD42FCA6FD9861CC725C5F905" authority="Silvestri, 1896" authorityName="Silvestri" authorityYear="1896" box="[151,471,1735,1761]" class="Diplopoda" family="Chelodesmidae" genus="Alocodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FD9861CC72539F905" box="[151,299,1735,1761]" italics="true" pageId="1" pageNumber="554">Alocodesmus</emphasis>
<bibRefCitation id="F41F4B69D42FCA6FD8201CC725C5F905" author="Silvestri, F." box="[305,471,1735,1761]" pageId="1" pageNumber="554" pagination="1 - 6" refId="ref5180" refString="Silvestri, F. (1896) Chilopodi e Diplopodi raccolti dal Dott. E. Festa a La Guayra, nel Darien e a Cuenca. Bollettino del musei di zoologia e di anatomia comparata della Reale Universita di Torino, 11 (254), 1 - 6." type="journal article" year="1896">Silvestri, 1896</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="578E4D1BD42FCA6FD8F01CC7271DF905" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" box="[481,783,1735,1761]" class="Diplopoda" family="Chelodesmidae" genus="Carlopeltis" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FD8F01CC7264DF905" box="[481,607,1735,1761]" italics="true" pageId="1" pageNumber="554">Carlopeltis</emphasis>
<bibRefCitation id="F41F4B69D42FCA6FDB741CC7271DF905" author="Verhoeff, K. W." box="[613,783,1735,1761]" pageId="1" pageNumber="554" pagination="1 - 54" refId="ref5292" refString="Verhoeff, K. W. (1938) Uber Diplopoden des Zoologischen Museums in Munchen. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 71 (1 - 2), 1 - 54." type="journal article" year="1938">Verhoeff, 1938</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="578E4D1BD42FCA6FDA081CC72030F905" authority="Carl, 1914" authorityName="Carl" authorityYear="1914" box="[793,1058,1735,1761]" class="Diplopoda" family="Chelodesmidae" genus="Heteropeltis" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FDA081CC727B0F905" box="[793,930,1735,1761]" italics="true" pageId="1" pageNumber="554">Heteropeltis</emphasis>
<bibRefCitation id="F41F4B69D42FCA6FDAB81CC72030F905" author="Carl, J." box="[937,1058,1735,1761]" pageId="1" pageNumber="554" pagination="821 - 993" refId="ref4105" refString="Carl, J. (1914) Die Diplopoden von Columbien nebst Beitragen zur Morphologie der Stemmatoiuliden. Memoires de la Societe neuchateloise des Sciences Naturelles, 5, 821 - 993." type="journal article" year="1914">Carl, 1914</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="578E4D1BD42FCA6FDD3D1CC7217BF905" authority="Hoffman, 1969" authorityName="Hoffman" authorityYear="1969" box="[1068,1385,1735,1761]" class="Diplopoda" family="Chelodesmidae" genus="Dyoparyphe" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FDD3D1CC720A4F905" box="[1068,1206,1735,1761]" italics="true" pageId="1" pageNumber="554">Dyoparyphe</emphasis>
<bibRefCitation id="F41F4B69D42FCA6FDDAC1CC7217BF905" author="Hoffman, R. L." box="[1213,1385,1735,1761]" pageId="1" pageNumber="554" pagination="263 - 283" refId="ref4492" refString="Hoffman, R. L. (1969) Chelodesmid Studies IV. A summary of the tribe Batodesmini, with the description of a new species of Biporodesmus from Northwestern Brasil. Papeis Avulsos de Zoologia, 22, 263 - 283. https: // doi. org / 10.11606 / 0031 - 1049.1969.22 p 263 - 283" type="journal article" year="1969">Hoffman, 1969</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="578E4D1BD42FCA6FD9861CEB25F2F8E1" authority="Hoffman, 1982" authorityName="Hoffman" authorityYear="1982" box="[151,480,1771,1797]" class="Diplopoda" family="Chelodesmidae" genus="Cheirogonus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FD9861CEB2535F8E1" box="[151,295,1771,1797]" italics="true" pageId="1" pageNumber="554">Cheirogonus</emphasis>
<bibRefCitation id="F41F4B69D42FCA6FD8211CEB25F2F8E1" author="Hoffman, R. L." box="[304,480,1771,1797]" pageId="1" pageNumber="554" pagination="633 - 654" refId="ref4575" refString="Hoffman, R. L. (1982) Chelodesmid studies. XVIII. On some new or poorly-known taxa in the tribe Batodesmini (Polydesmida: Chelodesmidae). Journal of Natural History, 16, 633 - 654. https: // doi. org / 10.1080 / 00222938200770481" type="journal article" year="1982">Hoffman, 1982</bibRefCitation>
</taxonomicName>
). The
<emphasis id="A2FAEA8AD42FCA6FDB381CEB2647F8E1" bold="true" box="[553,597,1771,1797]" pageId="1" pageNumber="554">PfP</emphasis>
is bifid, contrasting with the three-branched
<emphasis id="A2FAEA8AD42FCA6FDD481CEB2097F8E1" bold="true" box="[1113,1157,1771,1797]" pageId="1" pageNumber="554">PfP</emphasis>
in
<taxonomicName id="578E4D1BD42FCA6FDDBD1CEB2152F8E1" authorityName="Silvestri" authorityYear="1896" box="[1196,1344,1771,1797]" class="Diplopoda" family="Chelodesmidae" genus="Alocodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FDDBD1CEB2152F8E1" box="[1196,1344,1771,1797]" italics="true" pageId="1" pageNumber="554">Alocodesmus</emphasis>
</taxonomicName>
and the unbranched
<emphasis id="A2FAEA8AD42FCA6FD8301D0F255FF8CD" bold="true" box="[289,333,1807,1833]" pageId="1" pageNumber="554">PfP</emphasis>
in
<taxonomicName id="578E4D1BD42FCA6FD8611D0F263FF8CD" authorityName="Attems" authorityYear="1931" box="[368,557,1807,1833]" class="Diplopoda" family="Chelodesmidae" genus="Cordilleronomus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FD8611D0F263FF8CD" box="[368,557,1807,1833]" italics="true" pageId="1" pageNumber="554">Cordilleronomus</emphasis>
</taxonomicName>
and
<taxonomicName id="578E4D1BD42FCA6FDB751D0F26E6F8CD" authorityName="Hoffman" authorityYear="1982" box="[612,756,1807,1833]" class="Diplopoda" family="Chelodesmidae" genus="Cheirogonus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FDB751D0F26E6F8CD" box="[612,756,1807,1833]" italics="true" pageId="1" pageNumber="554">Cheirogonus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="90313698D42FCA6FD9D61D33274CF871" blockId="1.[151,1437,1375,1942]" pageId="1" pageNumber="554">
<taxonomicName id="578E4D1BD42FCA6FD9D61D332571F8A9" authorityName="Alvear &amp; Romero-Rincon" authorityYear="2024" box="[199,355,1843,1869]" class="Diplopoda" family="Chelodesmidae" genus="Caucadesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FD9D61D332571F8A9" box="[199,355,1843,1869]" italics="true" pageId="1" pageNumber="554">Caucadesmus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="578E4D1BD42FCA6FD8EB1D332703F8A9" authority="Strand, 1928" authorityName="Strand" authorityYear="1928" box="[506,785,1843,1869]" class="Diplopoda" family="Chelodesmidae" genus="Attemsiella" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FD8EB1D33266AF8A9" box="[506,632,1843,1869]" italics="true" pageId="1" pageNumber="554">Attemsiella</emphasis>
<bibRefCitation id="F41F4B69D42FCA6FDB901D332703F8A9" author="Strand, E." box="[641,785,1843,1869]" pageId="1" pageNumber="554" pagination="30 - 75" refId="ref5261" refString="Strand, E. (1928) Miscellanea nomenclatoria zoologica et palaeontologica. I - II. Archiv fur Naturgeschichte, 92 A (8), 30 - 75." type="journal article" year="1928">Strand, 1928</bibRefCitation>
</taxonomicName>
by lacking small tubercles and transverse grooves on the dorsal surface of most body rings, and from
<taxonomicName id="578E4D1BD42FCA6FDB6F1D57201BF895" authority="Chamberlin, 1923" authorityName="Chamberlin" authorityYear="1923" box="[638,1033,1879,1905]" class="Diplopoda" family="Chelodesmidae" genus="Colombodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">
<emphasis id="A2FAEA8AD42FCA6FDB6F1D572725F895" box="[638,823,1879,1905]" italics="true" pageId="1" pageNumber="554">Colombodesmus</emphasis>
<bibRefCitation id="F41F4B69D42FCA6FDA2C1D57201BF895" author="Chamberlin, R. V." box="[829,1033,1879,1905]" pageId="1" pageNumber="554" pagination="1 - 83" refId="ref4308" refString="Chamberlin, R. V. (1923) Results of the Bryant Walker expeditions of the University of Michigan to Colombia, 1913, and British Guiana, 1914. Occasional Papers of the Museum of Zoology, University of Michigan, 133, 1 - 83." type="journal article" year="1923">Chamberlin, 1923</bibRefCitation>
</taxonomicName>
, by the absence of sternal spines and transverse rows of granules on the dorsal surface of body rings.
</paragraph>
</subSubSection>
</treatment>
</document>