191 lines
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191 lines
25 KiB
XML
<document id="687809139CA39C78DB366762CB4B73C5" ID-DOI="10.11646/zootaxa.5523.5.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="13949910" ID-ZooBank=":CD0617A6-5CF4-438E-9778-7E3291BE1708" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729236646768" checkinUser="plazi" docAuthor="Alvear, Santiago & Romero-Rincon, Juan" docDate="2024" docId="1827878ED42BCA68D9861C6F24F7FE6D" docLanguage="en" docName="Zootaxa.5523.5.5.pdf" docOrigin="Zootaxa 5523 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5523.5.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cormodesmus Chamberlin 1923" docType="treatment" docVersion="2" lastPageNumber="559" masterDocId="E41EFFF6D42ECA6ED9111A002412FFE4" masterDocTitle="Contribution to the knowledge of the Colombian Batodesmini (Polydesmida: Chelodesmidae) with the description of a new genus and species, and taxonomic insights on the genus Cormodesmus Chamberlin, 1923" masterLastPageNumber="563" masterPageNumber="553" pageNumber="558" updateTime="1729237254395" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FE0883ACD0F8F38E44F77583B61A32F9">Contribution to the knowledge of the Colombian Batodesmini (Polydesmida: Chelodesmidae) with the description of a new genus and species, and taxonomic insights on the genus Cormodesmus Chamberlin, 1923</mods:title>
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<mods:namePart id="0D49BB09F14F9589F6271E2069D1946B">Alvear, Santiago</mods:namePart>
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<mods:affiliation id="C7C0CAC3111A1265C4E28C1B49BB3E42">Semillero de Investigación en Diversidad Funcional y Servicios Ecosistémicos-Grupo de Estudios Ambientales, Universidad del Cauca, Popayán, Colombia.</mods:affiliation>
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<mods:nameIdentifier id="828E44000B7969EC57E53261EFC6AAF3" type="email">alexs.alvear@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8711BA031AA4E64A3257E3CF628CF719">Romero-Rincon, Juan</mods:namePart>
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<mods:affiliation id="388F7E00500C323D3AA8FAAD384C1B3E">Museo de Historia Natural, Universidad Pedagógica Nacional, Bogotá, Colombia.</mods:affiliation>
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<mods:date id="E8A1F1B77A9E55EF6938E9A6DA2AE53B">2024</mods:date>
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<subSubSection id="D8946513D42BCA6BD9861C6F2696F96D" box="[151,644,1647,1674]" pageId="5" pageNumber="558" type="nomenclature">
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<paragraph id="90313698D42BCA6BD9861C6F2696F96D" blockId="5.[151,644,1647,1674]" box="[151,644,1647,1674]" pageId="5" pageNumber="558">
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<heading id="CB7981F4D42BCA6BD9861C6F2696F96D" bold="true" box="[151,644,1647,1674]" fontSize="11" level="1" pageId="5" pageNumber="558" reason="1">
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<emphasis id="A2FAEA8AD42BCA6BD9861C6F2696F96D" bold="true" box="[151,644,1647,1674]" pageId="5" pageNumber="558">
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Genus
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<taxonomicName id="578E4D1BD42BCA6BD9FF1C6F2696F96D" ID-CoL="7P4SL" authority="Chamberlin, 1923" authorityName="Chamberlin" authorityYear="1923" box="[238,644,1647,1674]" class="Diplopoda" family="Chelodesmidae" genus="Cormodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="558" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD42BCA6BD9FF1C6F2589F96E" bold="true" box="[238,411,1647,1674]" italics="true" pageId="5" pageNumber="558">Cormodesmus</emphasis>
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<bibRefCitation id="F41F4B69D42BCA6BD8B21C6F2696F96D" author="Chamberlin, R. V." box="[419,644,1647,1673]" pageId="5" pageNumber="558" pagination="1 - 83" refId="ref4308" refString="Chamberlin, R. V. (1923) Results of the Bryant Walker expeditions of the University of Michigan to Colombia, 1913, and British Guiana, 1914. Occasional Papers of the Museum of Zoology, University of Michigan, 133, 1 - 83." type="journal article" year="1923">Chamberlin, 1923</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="D8946513D42BCA6BD9861CB42047F928" box="[151,1109,1716,1740]" pageId="5" pageNumber="558" type="reference_group">
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<paragraph id="90313698D42BCA6BD9861CB42047F928" blockId="5.[151,1109,1716,1740]" box="[151,1109,1716,1740]" pageId="5" pageNumber="558">
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<treatmentCitation id="112F1089D42BCA6BD9861CB4261DF928" author="Chamberlin, R. V." box="[151,527,1716,1740]" page="56" pageId="5" pageNumber="558" year="1923">
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<taxonomicName id="578E4D1BD42BCA6BD9861CB4261DF928" ID-CoL="7P4SL" authority="Chamberlin, 1923: 56" authorityName="Chamberlin" authorityPageNumber="56" authorityYear="1923" box="[151,527,1716,1740]" class="Diplopoda" family="Chelodesmidae" genus="Cormodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="558" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD42BCA6BD9861CB4253AF928" box="[151,296,1716,1740]" italics="true" pageId="5" pageNumber="558">Cormodesmus</emphasis>
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<bibRefCitation id="F41F4B69D42BCA6BD83F1CB4261DF928" author="Chamberlin, R. V." box="[302,527,1716,1740]" pageId="5" pageNumber="558" pagination="1 - 83" refId="ref4308" refString="Chamberlin, R. V. (1923) Results of the Bryant Walker expeditions of the University of Michigan to Colombia, 1913, and British Guiana, 1914. Occasional Papers of the Museum of Zoology, University of Michigan, 133, 1 - 83." type="journal article" year="1923">Chamberlin, 1923: 56</bibRefCitation>
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</taxonomicName>
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.
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<treatmentCitation id="112F1089D42BCA6BDB061CB426D1F928" author="Attems, C." box="[535,707,1716,1740]" page="69" pageId="5" pageNumber="558" year="1938">
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<bibRefCitation id="F41F4B69D42BCA6BDB061CB426D1F928" author="Attems, C." box="[535,707,1716,1740]" pageId="5" pageNumber="558" pagination="1 - 147" refId="ref3752" refString="Attems, C. (1938) Myriapoda 3. Polydesmoidea II. Fam. Leptodesmidae, Platyrhachidae, Oxydesmidae, Gomphodesmidae. Das Tierreich, 69, 1 - 147. https: // doi. org / 10.1515 / 9783111430645" type="journal article" year="1938">Attems 1938: 69</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="112F1089D42BCA6BDBDF1CB4278AF928" author="Hoffman, R. L." box="[718,920,1716,1740]" page="280" pageId="5" pageNumber="558" year="1969">
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<bibRefCitation id="F41F4B69D42BCA6BDBDF1CB4278AF928" author="Hoffman, R. L." box="[718,920,1716,1740]" pageId="5" pageNumber="558" pagination="263 - 283" refId="ref4492" refString="Hoffman, R. L. (1969) Chelodesmid Studies IV. A summary of the tribe Batodesmini, with the description of a new species of Biporodesmus from Northwestern Brasil. Papeis Avulsos de Zoologia, 22, 263 - 283. https: // doi. org / 10.11606 / 0031 - 1049.1969.22 p 263 - 283" type="journal article" year="1969">Hoffman 1969: 280</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="112F1089D42BCA6BDAB31CB42043F928" author="Jeekel, C. A. W." box="[930,1105,1716,1740]" page="255" pageId="5" pageNumber="558" year="1971">
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<bibRefCitation id="F41F4B69D42BCA6BDAB31CB42043F928" author="Jeekel, C. A. W." box="[930,1105,1716,1740]" pageId="5" pageNumber="558" pagination="1 - 412" refId="ref4662" refString="Jeekel, C. A. W. (1971) Nomenclator generum et familiarum Diplopodorum: a list of the genus and family-group names in the Class Diplopoda from the 10 th edition of Linnaeus, 1758, to the end of 1957. Monografieen van de Nederlandse Entomologische Vereniging, 5, 1 - 412." type="journal article" year="1971">Jeekel 1971: 255</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="D8946513D42BCA6BD9861CF527A7F8EB" box="[151,949,1781,1807]" pageId="5" pageNumber="558" type="materials_examined">
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<paragraph id="90313698D42BCA6BD9861CF527A7F8EB" blockId="5.[151,1437,1781,2024]" box="[151,949,1781,1807]" pageId="5" pageNumber="558">
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<typeStatus id="4F35883AD42BCA6BD9861CF524DDF8EB" box="[151,207,1781,1807]" pageId="5" pageNumber="558">Type</typeStatus>
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species:
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<taxonomicName id="578E4D1BD42BCA6BD8241CF526B3F8EB" authority="Chamberlin, 1923" authorityName="Chamberlin" authorityYear="1923" box="[309,673,1781,1807]" class="Diplopoda" family="Chelodesmidae" genus="Cormodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="hirrutellus">
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<emphasis id="A2FAEA8AD42BCA6BD8241CF525DEF8EB" box="[309,460,1781,1807]" italics="true" pageId="5" pageNumber="558">C. hirrutellus</emphasis>
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<bibRefCitation id="F41F4B69D42BCA6BD8C21CF526B3F8EB" author="Chamberlin, R. V." box="[467,673,1781,1807]" pageId="5" pageNumber="558" pagination="1 - 83" refId="ref4308" refString="Chamberlin, R. V. (1923) Results of the Bryant Walker expeditions of the University of Michigan to Colombia, 1913, and British Guiana, 1914. Occasional Papers of the Museum of Zoology, University of Michigan, 133, 1 - 83." type="journal article" year="1923">Chamberlin, 1923</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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<subSubSection id="D8946513D42BCA68D9D61D1924F7FE6D" lastPageId="6" lastPageNumber="559" pageId="5" pageNumber="558" type="diagnosis">
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<paragraph id="90313698D42BCA6BD9D61D192030F89F" blockId="5.[151,1437,1781,2024]" pageId="5" pageNumber="558">
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<emphasis id="A2FAEA8AD42BCA6BD9D61D192528F8D7" bold="true" box="[199,314,1817,1843]" pageId="5" pageNumber="558">Diagnosis</emphasis>
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. A batodesminine genus belonging to the
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<taxonomicName id="578E4D1BD42BCA6BDA331D1927D3F8D7" authorityName="Attems" authorityYear="1898" box="[802,961,1817,1843]" class="Diplopoda" family="Chelodesmidae" genus="Biporodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="558" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD42BCA6BDA331D1927D3F8D7" box="[802,961,1817,1843]" italics="true" pageId="5" pageNumber="558">Biporodesmus</emphasis>
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</taxonomicName>
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group, characterized by having modified metazonites, with large, irregularly shaped tubercles (
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<figureCitation id="08B52A1DD42BCA6BDBFC1D3D2794F8B3" box="[749,902,1853,1879]" captionStart="FIGURE 6" captionStartId="7.[151,250,1892,1916]" captionTargetBox="[164,1422,453,1869]" captionTargetId="figure-138@7.[164,1422,453,1869]" captionTargetPageId="7" captionText="FIGURE 6. Cormodesmus hirrutellus Chamberlin, 1923 (MHNUC-MD-0302). A, C, E. Anterior, middle and posterior parts of body respectively, dorsal views. B, D, F. Anterior, middle and posterior parts of body respectively, ventral views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13949930" httpUri="https://zenodo.org/record/13949930/files/figure.png" pageId="5" pageNumber="558">Fig. 6A, C, E</figureCitation>
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). Paranota are elevated above the dorsum, welldeveloped, thick, and beset with large, round tubercles. Ozopores inconspicuous.
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</paragraph>
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<paragraph id="90313698D42BCA68D9D61D852763FF1D" blockId="5.[151,1437,1781,2024]" lastBlockId="6.[151,1437,151,393]" lastPageId="6" lastPageNumber="559" pageId="5" pageNumber="558">
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The gonopod of
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<taxonomicName id="578E4D1BD42BCA6BD86E1D85260DF87B" authorityName="Chamberlin" authorityYear="1923" box="[383,543,1925,1951]" class="Diplopoda" family="Chelodesmidae" genus="Cormodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="558" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD42BCA6BD86E1D85260DF87B" box="[383,543,1925,1951]" italics="true" pageId="5" pageNumber="558">Cormodesmus</emphasis>
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</taxonomicName>
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differs from other genera in the
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<taxonomicName id="578E4D1BD42BCA6BDA941D852036F87B" authorityName="Attems" authorityYear="1898" box="[901,1060,1925,1951]" class="Diplopoda" family="Chelodesmidae" genus="Biporodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="558" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD42BCA6BDA941D852036F87B" box="[901,1060,1925,1951]" italics="true" pageId="5" pageNumber="558">Biporodesmus</emphasis>
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</taxonomicName>
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group by the absence of a field of numerous long setae on the dorsal side of the
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<emphasis id="A2FAEA8AD42BCA6BDB811DA926A0F827" bold="true" box="[656,690,1961,1987]" pageId="5" pageNumber="558">Cx</emphasis>
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(
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<figureCitation id="08B52A1DD42BCA6BDBD11DA92716F827" box="[704,772,1961,1987]" captionStart="FIGURE 7" captionStartId="8.[151,250,1008,1032]" captionTargetBox="[151,1435,181,984]" captionTargetId="figure-19@8.[151,1435,181,984]" captionTargetPageId="8" captionText="FIGURE 7. Cormodesmus hirrutellus Chamberlin, 1923 (MHNUC-MD-0302). A, B. Left gonopod, mesal view. C, D. Lateral view. Abbreviations: ca: cannula, Cx: gonocoxa, Pfe: prefemoral region, PfP: prefemoral process, S: solenomere, t: tibiotarsal branch. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13949938" httpUri="https://zenodo.org/record/13949938/files/figure.png" pageId="5" pageNumber="558">Fig. 7</figureCitation>
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), contrasting with their presence in
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<taxonomicName id="578E4D1BD42BCA6BDD811DA924DDF80C" authority="Chamberlin, 1950" authorityName="Chamberlin" authorityYear="1950" class="Diplopoda" family="Chelodesmidae" genus="Tunochilus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="558" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD42BCA6BDD811DA92119F827" box="[1168,1291,1961,1987]" italics="true" pageId="5" pageNumber="558">Tunochilus</emphasis>
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<bibRefCitation id="F41F4B69D42BCA6BDC001DA924DDF80C" author="Chamberlin, R. V." pageId="5" pageNumber="558" pagination="133 - 144" refId="ref4357" refString="Chamberlin, R. V. (1950) Neotropical Chilopods and Diplopods in the collection of the department of tropical research, New York. Zoologica, 35 (2), 133 - 144. https: // doi. org / 10.5962 / p. 184615" type="journal article" year="1950">Chamberlin, 1950</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="578E4D1BD42BCA6BD8141DCD260BF803" authority="Cook, 1896" authorityName="Cook" authorityYear="1896" box="[261,537,1997,2023]" class="Diplopoda" family="Chelodesmidae" genus="Batodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="558" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD42BCA6BD8141DCD259EF803" box="[261,396,1997,2023]" italics="true" pageId="5" pageNumber="558">Batodesmus</emphasis>
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<bibRefCitation id="F41F4B69D42BCA6BD8851DCD260BF803" author="Cook, O. F." box="[404,537,1997,2023]" pageId="5" pageNumber="558" pagination="15 - 17" refId="ref4138" refString="Cook, O. F. (1896) On the Xystodesmidae, a new family. Brandtia (A series of occasional papers on Diplopoda and other Arthropoda), 4, 15 - 17. https: // doi. org / 10.5962 / bhl. title. 125177" type="journal article" year="1896">Cook, 1896</bibRefCitation>
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</taxonomicName>
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. The telopodite has a large, expanded
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<emphasis id="A2FAEA8AD42BCA6BDAD51DCD27E2F803" bold="true" box="[964,1008,1997,2023]" pageId="5" pageNumber="558">PfP</emphasis>
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covering the acropodite dorsally (
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<figureCitation id="08B52A1DD42BCA68DC7E1DCD24CDFF55" captionStart="FIGURE 7" captionStartId="8.[151,250,1008,1032]" captionTargetBox="[151,1435,181,984]" captionTargetId="figure-19@8.[151,1435,181,984]" captionTargetPageId="8" captionText="FIGURE 7. Cormodesmus hirrutellus Chamberlin, 1923 (MHNUC-MD-0302). A, B. Left gonopod, mesal view. C, D. Lateral view. Abbreviations: ca: cannula, Cx: gonocoxa, Pfe: prefemoral region, PfP: prefemoral process, S: solenomere, t: tibiotarsal branch. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13949938" httpUri="https://zenodo.org/record/13949938/files/figure.png" lastPageId="6" lastPageNumber="559" pageId="5" pageNumber="558">Fig. 7C, D</figureCitation>
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).
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<emphasis id="A2FAEA8AD428CA68D9E51A9724EFFF55" bold="true" box="[244,253,151,177]" pageId="6" pageNumber="559">t</emphasis>
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is small and simple (
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<figureCitation id="08B52A1DD428CA68D8E21A97267DFF55" box="[499,623,151,177]" captionStart="FIGURE 7" captionStartId="8.[151,250,1008,1032]" captionTargetBox="[151,1435,181,984]" captionTargetId="figure-19@8.[151,1435,181,984]" captionTargetPageId="8" captionText="FIGURE 7. Cormodesmus hirrutellus Chamberlin, 1923 (MHNUC-MD-0302). A, B. Left gonopod, mesal view. C, D. Lateral view. Abbreviations: ca: cannula, Cx: gonocoxa, Pfe: prefemoral region, PfP: prefemoral process, S: solenomere, t: tibiotarsal branch. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13949938" httpUri="https://zenodo.org/record/13949938/files/figure.png" pageId="6" pageNumber="559">Fig. 7A, B</figureCitation>
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), differing from the multi-branched
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<emphasis id="A2FAEA8AD428CA68DD1D1A972007FF55" bold="true" box="[1036,1045,151,177]" pageId="6" pageNumber="559">t</emphasis>
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in
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<taxonomicName id="578E4D1BD428CA68DD2D1A97216EFF55" authority="Attems, 1898" authorityName="Attems" authorityYear="1898" box="[1084,1404,151,177]" class="Diplopoda" family="Chelodesmidae" genus="Biporodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="559" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD428CA68DD2D1A9720C9FF55" box="[1084,1243,151,177]" italics="true" pageId="6" pageNumber="559">Biporodesmus</emphasis>
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<bibRefCitation id="F41F4B69D428CA68DDF51A97216EFF55" author="Attems, C." box="[1252,1404,151,177]" pageId="6" pageNumber="559" pagination="221 - 482" refId="ref3699" refString="Attems, C. (1898) System der Polydesmiden I. Theil. Denkschriften der Akademie der Wissenschaften Wien, Mathematisch- Naturwissenschaftliche Klassen, 67, 221 - 482." type="journal article" year="1898">Attems, 1898</bibRefCitation>
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</taxonomicName>
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or absent in
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<taxonomicName id="578E4D1BD428CA68D81B1ABB2597FF31" authorityName="Chamberlin" authorityYear="1950" box="[266,389,187,213]" class="Diplopoda" family="Chelodesmidae" genus="Tunochilus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="559" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD428CA68D81B1ABB2597FF31" box="[266,389,187,213]" italics="true" pageId="6" pageNumber="559">Tunochilus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="578E4D1BD428CA68D8D21ABB2709FF31" authority="Hoffman, 2007" authorityName="Hoffman" authorityYear="2007" box="[451,795,187,213]" class="Diplopoda" family="Chelodesmidae" genus="Curimaguana" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="559" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD428CA68D8D21ABB264DFF31" box="[451,607,187,213]" italics="true" pageId="6" pageNumber="559">Curimaguana</emphasis>
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<bibRefCitation id="F41F4B69D428CA68DB781ABB2709FF31" author="Hoffman, R. L." box="[617,795,187,213]" pageId="6" pageNumber="559" pagination="1 - 1" refId="ref4626" refString="Hoffman, R. L. (2007) Replacement of the preoccupied generic name of a Neotropical milliped taxon (Polydesmida: Chelodesmidae: Batodesmini). Myriapodologica, 9, 1 - 1." type="journal article" year="2007">Hoffman, 2007</bibRefCitation>
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</taxonomicName>
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.
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<emphasis id="A2FAEA8AD428CA68DA381ABB2720FF31" bold="true" box="[809,818,187,213]" pageId="6" pageNumber="559">t</emphasis>
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is directed ventrad vs. anteriad in
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<taxonomicName id="578E4D1BD428CA68DDDF1ABB2529FF1D" authority="Silvestri, 1898" authorityName="Silvestri" authorityYear="1898" class="Diplopoda" family="Chelodesmidae" genus="Plusioporodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="559" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD428CA68DDDF1ABB218EFF31" box="[1230,1436,187,213]" italics="true" pageId="6" pageNumber="559">Plusioporodesmus</emphasis>
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<bibRefCitation id="F41F4B69D428CA68D9861ADF2529FF1D" author="Silvestri, F." box="[151,315,223,249]" pageId="6" pageNumber="559" pagination="53 - 79" refId="ref5231" refString="Silvestri, F. (1898) Diagnosticos de nuevos Diplopodos Sudamericanos. Anales del Museo Nacional de historia natural de Buenos Aires, 6, 53 - 79." type="journal article" year="1898">Silvestri, 1898</bibRefCitation>
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</taxonomicName>
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and lateroventrad in
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<taxonomicName id="578E4D1BD428CA68DB391ADF277EFF1D" authority="Hoffman, 1982" authorityName="Hoffman" authorityYear="1982" box="[552,876,223,249]" class="Diplopoda" family="Chelodesmidae" genus="Igaraparana" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="559" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD428CA68DB391ADF26A5FF1C" box="[552,695,223,248]" italics="true" pageId="6" pageNumber="559">Igaraparana</emphasis>
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<bibRefCitation id="F41F4B69D428CA68DBAE1ADF277EFF1D" author="Hoffman, R. L." box="[703,876,223,249]" pageId="6" pageNumber="559" pagination="633 - 654" refId="ref4575" refString="Hoffman, R. L. (1982) Chelodesmid studies. XVIII. On some new or poorly-known taxa in the tribe Batodesmini (Polydesmida: Chelodesmidae). Journal of Natural History, 16, 633 - 654. https: // doi. org / 10.1080 / 00222938200770481" type="journal article" year="1982">Hoffman, 1982</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="90313698D428CA68D9D61B0324F7FE6D" blockId="6.[151,1437,151,393]" pageId="6" pageNumber="559">
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<taxonomicName id="578E4D1BD428CA68D9D61B032575FEF9" authorityName="Chamberlin" authorityYear="1923" box="[199,359,259,285]" class="Diplopoda" family="Chelodesmidae" genus="Cormodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="559" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD428CA68D9D61B032575FEF9" box="[199,359,259,285]" italics="true" pageId="6" pageNumber="559">Cormodesmus</emphasis>
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</taxonomicName>
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differs from
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|
<taxonomicName id="578E4D1BD428CA68DB101B03266DFEF9" authorityName="Strand" authorityYear="1928" box="[513,639,259,285]" class="Diplopoda" family="Chelodesmidae" genus="Attemsiella" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="559" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD428CA68DB101B03266DFEF9" box="[513,639,259,285]" italics="true" pageId="6" pageNumber="559">Attemsiella</emphasis>
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</taxonomicName>
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|
by lacking transverse grooves on the tergites and having large, round tubercles instead of small, irregular ones, and from
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<taxonomicName id="578E4D1BD428CA68DBCA1B272786FEA5" authorityName="Chamberlin" authorityYear="1923" box="[731,916,295,321]" class="Diplopoda" family="Chelodesmidae" genus="Colombodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="559" phylum="Arthropoda" rank="genus">
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<emphasis id="A2FAEA8AD428CA68DBCA1B272786FEA5" box="[731,916,295,321]" italics="true" pageId="6" pageNumber="559">Colombodesmus</emphasis>
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</taxonomicName>
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by the absence of transverse rows of granules on the dorsal surface of the body rings and by having small pointed sternal projections instead of large sternal spines.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |