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<document id="EA726AB785C1374E565867FE1D5BD67B" ID-CLB-Dataset="60704" ID-DOI="10.5852/ejt.2023.885.2199" ID-GBIF-Dataset="d205aec5-8b36-4f96-b447-afe09885bdf0" ID-ISSN="2118-9773" ID-Zenodo-Dep="8222417" ID-ZooBank="8432EEFF-391F-4778-81F6-14F3F6ECAA5F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691437419139" checkinUser="felipe" docAuthor="Korn, Dieter &amp; Klug, Christian" docDate="2023" docId="AE7E8780077DFF85FDB3104BFA8FF9CD" docLanguage="en" docName="ejt-885_156-194.pdf" docOrigin="European Journal of Taxonomy 885" docSource="http://dx.doi.org/10.5852/ejt.2023.885.2199" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Solenochilus Meek &amp; Worthen 1870" docType="treatment" docVersion="7" lastPageNumber="184" masterDocId="5247FFF80761FF99FFA71124FF8FFFA1" masterDocTitle="Early Carboniferous coiled nautiloids from the Anti-Atlas (Morocco)" masterLastPageNumber="194" masterPageNumber="156" pageNumber="184" updateTime="1699551382698" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="6EFFC84A3AC246B141F5E0A22A4EC4C6">
<mods:title id="6794D11CED5F3DF4A8DA935C70460BF6">Early Carboniferous coiled nautiloids from the Anti-Atlas (Morocco)</mods:title>
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<mods:name id="98F0C6B9828710D68FE21D70492B598C" type="personal">
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<mods:namePart id="B42387A57E5D43478C17C218C859A6E5">Korn, Dieter</mods:namePart>
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<mods:affiliation id="B248E6CAA67C62ECE0C961990DD26210">Museum für Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
<mods:nameIdentifier id="49FCCECDE208622ACF8626A5BFFEA738" type="email">dieter.korn@mfn.berlin</mods:nameIdentifier>
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<mods:namePart id="FF2C84C3CB61116E165EE475E5BCEF9C">Klug, Christian</mods:namePart>
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<mods:affiliation id="725CC9D608340B35ED08B12F5D89A8AA">Paläontologisches Institut und Museum, Universität Zürich, Karl-Schmid-Strasse 4, 8006 Zürich, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="D36C936532290F560873A5563D7E6BD2" type="email">chklug@pim.uzh.ch</mods:nameIdentifier>
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<mods:title id="55A899781721670A34C7C38EC53BF490">European Journal of Taxonomy</mods:title>
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<mods:part id="4852628C28478C8391C1AB15BDAD54E3">
<mods:date id="9E38C44887CFB34B663A602F889499F6">2023</mods:date>
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<mods:number id="6ECF430AF5D1CB8E3D29ED3D3441AAF6">2023-08-02</mods:number>
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<mods:number id="40265B759A18BABA7D207E756A9AEFC7">885</mods:number>
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<treatment id="AE7E8780077DFF85FDB3104BFA8FF9CD" ID-DOI="http://doi.org/10.5281/zenodo.8222515" ID-GBIF-Taxon="212425332" ID-Zenodo-Dep="8222515" LSID="urn:lsid:plazi:treatment:AE7E8780077DFF85FDB3104BFA8FF9CD" httpUri="http://treatment.plazi.org/id/AE7E8780077DFF85FDB3104BFA8FF9CD" lastPageNumber="184" pageId="28" pageNumber="184">
<subSubSection id="6ECD651D077DFF85FDB3104BFBAFFE28" box="[532,1056,366,393]" pageId="28" pageNumber="184" type="nomenclature">
<paragraph id="26683696077DFF85FDB3104BFBAFFE28" blockId="28.[532,1056,366,393]" box="[532,1056,366,393]" pageId="28" pageNumber="184">
<heading id="7D2081FA077DFF85FDB3104BFBAFFE28" box="[532,1056,366,393]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="184" reason="2">
Genus
<taxonomicName id="E1D74D15077DFF85FDC1104AFBAFFE28" authority="Meek &amp; Worthen, 1870" authorityName="Meek &amp; Worthen" authorityYear="1870" box="[614,1056,366,393]" class="Cephalopoda" family="Solenochilidae" genus="Solenochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="28" pageNumber="184" phylum="Mollusca" rank="genus">
<emphasis id="14A3EA84077DFF85FDC1104AFC8DFE28" bold="true" box="[614,770,366,393]" italics="true" pageId="28" pageNumber="184">Solenochilus</emphasis>
Meek &amp; Worthen, 1870
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6ECD651D077DFF85FF1A1095FBEFFE5B" pageId="28" pageNumber="184" type="type_taxon">
<paragraph id="26683696077DFF85FF1A1095FED7FE6B" blockId="28.[189,344,432,459]" box="[189,344,432,459]" pageId="28" pageNumber="184">
<heading id="7D2081FA077DFF85FF1A1095FED7FE6B" bold="true" box="[189,344,432,459]" fontSize="11" level="3" pageId="28" pageNumber="184" reason="3">
<emphasis id="14A3EA84077DFF85FF1A1095FED7FE6B" bold="true" box="[189,344,432,459]" pageId="28" pageNumber="184">
<typeStatus id="F96C8834077DFF85FF1A1095FF75FE6A" box="[189,250,433,459]" pageId="28" pageNumber="184">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="26683696077DFF85FF1A10C4FBEFFE5B" blockId="28.[189,1120,479,506]" box="[189,1120,479,506]" pageId="28" pageNumber="184">
<taxonomicName id="E1D74D15077DFF85FF1A10C4FCEDFE58" authority="Springeri White &amp; St. John, 1868" authorityName="Springeri White &amp; St. John" authorityYear="1868" box="[189,866,479,506]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" kingdom="Animalia" order="Nautilida" pageId="28" pageNumber="184" phylum="Mollusca" rank="subGenus" subGenus="Cryptoceras">
<emphasis id="14A3EA84077DFF85FF1A10C4FEAFFE5B" box="[189,288,480,506]" italics="true" pageId="28" pageNumber="184">Nautilus</emphasis>
(
<emphasis id="14A3EA84077DFF85FE9510C4FE4CFE5B" box="[306,451,480,506]" italics="true" pageId="28" pageNumber="184">Cryptoceras</emphasis>
)
<emphasis id="14A3EA84077DFF85FE7310C4FDCCFE5B" box="[468,579,480,506]" italics="true" pageId="28" pageNumber="184">Springeri</emphasis>
White &amp; St. John, 1868
</taxonomicName>
; original designation.
</paragraph>
</subSubSection>
<subSubSection id="6ECD651D077DFF85FF1A1305FAF9FD5B" pageId="28" pageNumber="184" type="diagnosis">
<paragraph id="26683696077DFF85FF1A1305FEB9FD9A" blockId="28.[189,310,545,571]" box="[189,310,545,571]" pageId="28" pageNumber="184">
<heading id="7D2081FA077DFF85FF1A1305FEB9FD9A" bold="true" box="[189,310,545,571]" fontSize="11" level="3" pageId="28" pageNumber="184" reason="3">
<emphasis id="14A3EA84077DFF85FF1A1305FEB9FD9A" bold="true" box="[189,310,545,571]" pageId="28" pageNumber="184">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="26683696077DFF85FF1A1375FAF9FD5B" blockId="28.[188,1399,593,762]" pageId="28" pageNumber="184">
Genus of the family
<taxonomicName id="E1D74D15077DFF85FE1E1375FDE8FDCA" authorityName="Hyatt" authorityYear="1893" box="[441,615,593,619]" class="Cephalopoda" family="Solenochilidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="28" pageNumber="184" phylum="Mollusca" rank="family">Solenochilidae</taxonomicName>
with pachyconic to globular, involute or subinvolute conch. The conch is very rapidly increasing in height with an extremely high coiling rate (WER usually higher than 4.00). Adult body chamber with long lateral outgrowths. Whorls weakly without contact or embracing, their profile ranges from reniform to nearly circular. Suture line simple, nearly straight with a small annular lobe. The siphuncle has a position adjacent to the venter (after
<bibRefCitation id="42464B67077DFF85FC5013FBFB1AFD5B" author="Gordon M. Jr." box="[1015,1173,735,762]" pageId="28" pageNumber="184" pagination="1 - 322" refId="ref15357" refString="Gordon M. Jr. 1965. Carboniferous cephalopods of Arkansas. Professional Papers, U. S. Geological Survey 460: 1 - 322. https: // doi. org / 10.3133 / pp 460" type="journal article" year="1965">Gordon 1965</bibRefCitation>
;
<bibRefCitation id="42464B67077DFF85FB0613FBFAE9FD58" author="Shimansky V. N." box="[1185,1382,735,761]" pageId="28" pageNumber="184" pagination="1 - 244" refId="ref16954" refString="Shimansky V. N. 1967. Kamennougol'nye Nautilida. Trudy paleontologicheskogo Instituta Akademiya Nauk SSSR 115: 1 - 244." type="journal article" year="1967">Shimansky 1967</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6ECD651D077DFF85FF1A1206FE8CFB1E" pageId="28" pageNumber="184" type="distribution">
<paragraph id="26683696077DFF85FF1A1206FD1BFC9A" blockId="28.[189,660,801,828]" box="[189,660,801,828]" pageId="28" pageNumber="184">
<heading id="7D2081FA077DFF85FF1A1206FD1BFC9A" bold="true" box="[189,660,801,828]" fontSize="11" level="3" pageId="28" pageNumber="184" reason="3">
<emphasis id="14A3EA84077DFF85FF1A1206FD1BFC9A" bold="true" box="[189,660,801,828]" pageId="28" pageNumber="184">Included Early Carboniferous species</emphasis>
</heading>
</paragraph>
<paragraph id="26683696077DFF85FF1A1274FEA5FC13" blockId="28.[189,1398,848,946]" pageId="28" pageNumber="184">
<bibRefCitation id="42464B67077DFF85FF1A1274FE28FCCA" author="Gordon M. Jr." box="[189,423,848,875]" pageId="28" pageNumber="184" pagination="1 - 322" refId="ref15357" refString="Gordon M. Jr. 1965. Carboniferous cephalopods of Arkansas. Professional Papers, U. S. Geological Survey 460: 1 - 322. https: // doi. org / 10.3133 / pp 460" type="journal article" year="1965">Gordon (1965: 152)</bibRefCitation>
gave an overview of the previously described species of
<taxonomicName id="E1D74D15077DFF85FBE31274FB55FCCB" authorityName="Meek &amp; Worthen" authorityYear="1870" box="[1092,1242,848,874]" class="Cephalopoda" family="Solenochilidae" genus="Solenochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="28" pageNumber="184" phylum="Mollusca" rank="genus">
<emphasis id="14A3EA84077DFF85FBE31274FB55FCCB" box="[1092,1242,848,874]" italics="true" pageId="28" pageNumber="184">Solenochilus</emphasis>
</taxonomicName>
. He listed 24 species (some with question marks), most of which are known from Late Carboniferous strata of North America.
</paragraph>
<paragraph id="26683696077DFF85FF1A12FEFBB3FC55" blockId="28.[189,1084,986,1012]" box="[189,1084,986,1012]" pageId="28" pageNumber="184">So far, only a few Early Carboniferous species are known from two regions:</paragraph>
<paragraph id="26683696077DFF85FF1A1538FBC5FBF8" blockId="28.[189,1396,1052,1114]" pageId="28" pageNumber="184">
British Isles(
<bibRefCitation id="42464B67077DFF85FEF61538FE60FB97" author="Phillips J." box="[337,495,1052,1078]" pageId="28" pageNumber="184" refId="ref16710" refString="Phillips J. 1836. Illustrations of the Geology of Yorkshire, Part II. The Mountain Limestone District. John Murray, London." type="book" year="1836">Phillips 1836</bibRefCitation>
;
<bibRefCitation id="42464B67077DFF85FE5E1538FD0FFB97" author="Foord A. H." box="[505,640,1052,1078]" pageId="28" pageNumber="184" refId="ref15174" refString="Foord A. H. 1891. Catalogue of the Fossil Cephalopoda in the British Museum, Part II. Containing the Remainder of the Suborder Nautiloidea, Consisting of the Families Lituitidae, Trochoceratidae, and Nautilidae, with a Supplement. Order of the Trustees, London." type="book" year="1891">Foord 1891</bibRefCitation>
,
<bibRefCitation id="42464B67077DFF85FD2D1538FD49FB97" author="Foord A. H." box="[650,710,1052,1078]" pageId="28" pageNumber="184" pagination="127 - 146" refId="ref15283" refString="Foord A. H. 1901. Monograph of the Carboniferous Cephalopoda of Ireland, Part IV. Containing the families Solenocheilidae (concluded) and Glyphioceratidae. Monographs of the Palaeontographical Society 55: 127 - 146. https: // doi. org / 10.1080 / 02693445.1901.12035497" type="journal article" year="1901">1901</bibRefCitation>
):
<taxonomicName id="E1D74D15077DFF85FD791538FBC3FB97" authority="Phillips, 1836" authorityName="Phillips" authorityYear="1836" box="[734,1100,1052,1078]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" kingdom="Animalia" order="Nautilida" pageId="28" pageNumber="184" phylum="Mollusca" rank="species" species="dorsalis">
<emphasis id="14A3EA84077DFF85FD791538FC2AFB97" box="[734,933,1052,1078]" italics="true" pageId="28" pageNumber="184">Nautilus dorsalis</emphasis>
<bibRefCitation id="42464B67077DFF85FC0E1538FBC3FB97" author="Phillips J." box="[937,1100,1052,1078]" pageId="28" pageNumber="184" refId="ref16710" refString="Phillips J. 1836. Illustrations of the Geology of Yorkshire, Part II. The Mountain Limestone District. John Murray, London." type="book" year="1836">Phillips, 1836</bibRefCitation>
</taxonomicName>
,Yorkshire;
<taxonomicName id="E1D74D15077DFF85FB761538FE57FBFB" authority="Hibernicus Foord, 1891" authorityName="Hibernicus Foord" authorityYear="1891" class="Cephalopoda" family="Solenochilidae" genus="Solenocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="28" pageNumber="184" phylum="Mollusca" rank="genus">
<emphasis id="14A3EA84077DFF85FB761538FEB0FBF8" italics="true" pageId="28" pageNumber="184">Solenocheilus Hibernicus</emphasis>
<bibRefCitation id="42464B67077DFF85FEE0151BFE57FBFB" author="Foord A. H." box="[327,472,1087,1114]" pageId="28" pageNumber="184" refId="ref15174" refString="Foord A. H. 1891. Catalogue of the Fossil Cephalopoda in the British Museum, Part II. Containing the Remainder of the Suborder Nautiloidea, Consisting of the Families Lituitidae, Trochoceratidae, and Nautilidae, with a Supplement. Order of the Trustees, London." type="book" year="1891">Foord, 1891</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="5EC07606077DFF85FE42151BFDB3FBF8" box="[485,572,1087,1113]" name="Ireland" pageId="28" pageNumber="184">Ireland</collectingCountry>
;
<taxonomicName id="E1D74D15077DFF85FDE0151BFC6CFBFB" authority="Foord, 1901" authorityName="Foord" authorityYear="1901" box="[583,995,1087,1114]" class="Cephalopoda" family="Solenochilidae" genus="Solenocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="28" pageNumber="184" phylum="Mollusca" rank="species" species="clausus">
<emphasis id="14A3EA84077DFF85FDE0151BFCC5FBF8" box="[583,842,1087,1113]" italics="true" pageId="28" pageNumber="184">Solenocheilus clausus</emphasis>
<bibRefCitation id="42464B67077DFF85FCF6151BFC6CFBFB" author="Foord A. H." box="[849,995,1087,1114]" pageId="28" pageNumber="184" pagination="127 - 146" refId="ref15283" refString="Foord A. H. 1901. Monograph of the Carboniferous Cephalopoda of Ireland, Part IV. Containing the families Solenocheilidae (concluded) and Glyphioceratidae. Monographs of the Palaeontographical Society 55: 127 - 146. https: // doi. org / 10.1080 / 02693445.1901.12035497" type="journal article" year="1901">Foord, 1901</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="5EC07606077DFF85FC48151BFBC9FBF8" box="[1007,1094,1087,1113]" name="Ireland" pageId="28" pageNumber="184">Ireland</collectingCountry>
.
</paragraph>
<paragraph id="26683696077DFF85FF1A15A5FE8CFB1E" blockId="28.[189,1399,1153,1215]" pageId="28" pageNumber="184">
North Africa (this paper):
<taxonomicName id="E1D74D15077DFF85FE5715A5FD67FB3A" authority="Korn &amp; Klug, 2023" authorityName="Korn &amp; Klug" authorityYear="2023" box="[496,744,1153,1179]" class="Cephalopoda" family="Solenochilidae" genus="Solenochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="28" pageNumber="184" phylum="Mollusca" rank="species" species="lucynae" status="sp. nov.">
<emphasis id="14A3EA84077DFF85FE5715A5FD67FB3A" box="[496,744,1153,1179]" italics="true" pageId="28" pageNumber="184">Solenochilus lucynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0F9057FF077DFF85FD5715A6FCC2FB3D" box="[752,845,1154,1180]" pageId="28" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="E413F874077DFF85FCF015A5FC58FB3A" box="[855,983,1153,1179]" pageId="28" pageNumber="184">Anti-Atlas</collectingRegion>
;
<taxonomicName id="E1D74D15077DFF85FC4415A5FB46FB3A" authority="Korn &amp; Klug, 2023" authorityName="Korn &amp; Klug" authorityYear="2023" box="[995,1225,1153,1179]" class="Cephalopoda" family="Solenochilidae" genus="Solenochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="28" pageNumber="184" phylum="Mollusca" rank="species" species="pohlei" status="sp. nov.">
<emphasis id="14A3EA84077DFF85FC4415A5FB46FB3A" box="[995,1225,1153,1179]" italics="true" pageId="28" pageNumber="184">Solenochilus pohlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0F9057FF077DFF85FB7615A6FAA1FB3D" box="[1233,1326,1154,1180]" pageId="28" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="E413F874077DFF85FA9E15A5FF70FB1E" pageId="28" pageNumber="184">Anti-Atlas</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="6ECD651D077DFF85FF1A15C3FA8FF9CD" pageId="28" pageNumber="184" type="discussion">
<paragraph id="26683696077DFF85FF1A15C3FEA0FAA0" blockId="28.[189,303,1255,1281]" box="[189,303,1255,1281]" pageId="28" pageNumber="184">
<heading id="7D2081FA077DFF85FF1A15C3FEA0FAA0" bold="true" box="[189,303,1255,1281]" fontSize="11" level="3" pageId="28" pageNumber="184" reason="3">
<emphasis id="14A3EA84077DFF85FF1A15C3FEA0FAA0" bold="true" box="[189,303,1255,1281]" pageId="28" pageNumber="184">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="26683696077DFF85FF1A1432FC0AFA3D" blockId="28.[186,1399,1302,1436]" pageId="28" pageNumber="184">
<taxonomicName id="E1D74D15077DFF85FF1A1432FEDCFA91" authorityName="Meek &amp; Worthen" authorityYear="1870" box="[189,339,1302,1328]" class="Cephalopoda" family="Solenochilidae" genus="Solenochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="28" pageNumber="184" phylum="Mollusca" rank="genus">
<emphasis id="14A3EA84077DFF85FF1A1432FEDCFA91" box="[189,339,1302,1328]" italics="true" pageId="28" pageNumber="184">Solenochilus</emphasis>
</taxonomicName>
is a genus whose species are mainly known from late Carboniferous strata of North America.
<bibRefCitation id="42464B67077DFF85FF1D141EFEE3FAF4" author="Gordon M. Jr." box="[186,364,1338,1365]" pageId="28" pageNumber="184" pagination="1 - 322" refId="ref15357" refString="Gordon M. Jr. 1965. Carboniferous cephalopods of Arkansas. Professional Papers, U. S. Geological Survey 460: 1 - 322. https: // doi. org / 10.3133 / pp 460" type="journal article" year="1965">Gordon (1965)</bibRefCitation>
published an identification key for these species, in which the species described from Europe can also be included. However, it is clear that this key does not represent a phylogenetic scheme and that it contains species with different ontogenetic stages.
</paragraph>
<paragraph id="26683696077DFF85FF1A14E0FA8FF9CD" blockId="28.[188,1399,1475,1645]" pageId="28" pageNumber="184">
Most occurrences of specimens of the genus are in sediments of the shallow shelf; therefore,
<taxonomicName id="E1D74D15077DFF85FB7914E7FAFBFA7C" authorityName="Meek &amp; Worthen" authorityYear="1870" box="[1246,1396,1475,1501]" class="Cephalopoda" family="Solenochilidae" genus="Solenochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="28" pageNumber="184" phylum="Mollusca" rank="genus">
<emphasis id="14A3EA84077DFF85FB7914E7FAFBFA7C" box="[1246,1396,1475,1501]" italics="true" pageId="28" pageNumber="184">Solenochilus</emphasis>
</taxonomicName>
is usually only rarely found together with ammonoids. For example, the genus is apparently absent from the assemblages of the Dalle à
<taxonomicName id="E1D74D15077DFF85FD8A172FFD30F985" authorityName="Schindewolf" authorityYear="1922" box="[557,703,1547,1572]" class="Cephalopoda" family="Prolecanitidae" genus="Merocanites" kingdom="Animalia" pageId="28" pageNumber="184" phylum="Mollusca" rank="genus">
<emphasis id="14A3EA84077DFF85FD8A172FFD30F985" box="[557,703,1547,1572]" italics="true" pageId="28" pageNumber="184">Merocanites</emphasis>
</taxonomicName>
of Timimoun (Korn
<emphasis id="14A3EA84077DFF85FC10172FFC7DF984" box="[951,1010,1547,1573]" italics="true" pageId="28" pageNumber="184">et al.</emphasis>
2022) and Gara el Itima N; both are occurrences characterised by a high number of ammonoid specimens. The comparatively frequent occurrence of the genus in layers with many ammonoids at Rissani is therefore an exception.
</paragraph>
</subSubSection>
</treatment>
</document>
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