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<document id="3C4BF61D8E17F55DBBAC817A123CC5E1" ID-DOI="10.5281/zenodo.4509100" ID-ISSN="2345-7600" ID-Zenodo-Dep="4509100" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612548350057" checkinUser="carolina" docAuthor="Tong, Yanfeng &amp; Li, Shuqiang" docDate="2008" docId="03FA6634FC53FFF08370FAD93728DAA7" docLanguage="en" docName="RafflesBZool.56.1.45-53.pdf" docOrigin="Raffles Bulletin of Zoology 56 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Belisana huberi Tong &amp; Li 2008, new species" docType="treatment" docVersion="4" lastPageNumber="47" masterDocId="FFC31E4CFC51FFF28241FFD43260DE22" masterDocTitle="Four New Species Of Six-Eyed Pholcid Spiders (Araneae: Pholcidae) From Hainan Island, China" masterLastPageNumber="53" masterPageNumber="45" pageNumber="47" updateTime="1698867895616" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F6962C778E97D6E12D2166DFE99CCF8D">Four New Species Of Six-Eyed Pholcid Spiders (Araneae: Pholcidae) From Hainan Island, China</mods:title>
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<mods:name id="EBC2EE999625E284F2804AE5B81B6630" type="personal">
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<mods:namePart id="423637934A645DD7BE4531E52097EF7E">Tong, Yanfeng</mods:namePart>
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<mods:namePart id="2A08AEC68C38B2962B2B4B2E5CC8632E">Li, Shuqiang</mods:namePart>
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<mods:title id="FEA530DA7134FF0B60CF1058B86D122A">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="94523A8BBC195E76FCF695704461D720">
<mods:date id="706B31C2466CA93A8202376DD694C63E">2008</mods:date>
<mods:detail id="7AEEB0C026700D9707943E2C05FC4A8B" type="pubDate">
<mods:number id="D9C0B1121ADA65A967FB754C3E2C123F">2008-02-29</mods:number>
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<mods:number id="1FC52B4F035357A2700A677630395CEF">56</mods:number>
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<mods:identifier id="E59F47C6558333499E2A3D5755E90A5D" type="ISSN">2345-7600</mods:identifier>
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<treatment id="03FA6634FC53FFF08370FAD93728DAA7" LSID="urn:lsid:plazi:treatment:03FA6634FC53FFF08370FAD93728DAA7" httpUri="http://treatment.plazi.org/id/03FA6634FC53FFF08370FAD93728DAA7" lastPageNumber="47" pageId="2" pageNumber="47">
<subSubSection id="C34984A9FC53FFF08370FAD93017DB07" box="[305,631,1293,1317]" pageId="2" pageNumber="47" type="nomenclature">
<paragraph id="8BECD722FC53FFF08370FAD93017DB07" blockId="2.[305,631,1293,1349]" box="[305,631,1293,1317]" pageId="2" pageNumber="47">
<heading id="D0A4604EFC53FFF08370FAD93017DB07" bold="true" box="[305,631,1293,1317]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="47" reason="2">
<emphasis id="B9270B30FC53FFF08370FAD93017DB07" bold="true" box="[305,631,1293,1317]" pageId="2" pageNumber="47">
<taxonomicName id="4C53ACA1FC53FFF08370FAD93383DB07" ID-CoL="68H94" authority="Tong &amp; Li, 2008" authorityName="Tong &amp; Li" authorityYear="2008" box="[305,483,1293,1317]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="B9270B30FC53FFF08370FAD93383DB07" bold="true" box="[305,483,1293,1317]" italics="true" pageId="2" pageNumber="47">Belisana huberi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A214B64BFC53FFF083B3FAD93017DB07" box="[498,631,1293,1317]" pageId="2" pageNumber="47" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34984A9FC53FFF08327FAFA3022DB67" box="[358,578,1326,1349]" pageId="2" pageNumber="47" type="description">
<paragraph id="8BECD722FC53FFF08327FAFA3022DB67" blockId="2.[305,631,1293,1349]" box="[358,578,1326,1349]" pageId="2" pageNumber="47">
(
<figureCitation id="1368CBA7FC53FFF0832FFAFA33D5DB67" box="[366,437,1326,1349]" captionStart="Fig" captionStartId="1.[127,162,2048,2069]" captionTargetBox="[126,1464,914,2028]" captionTargetId="figure-102@1.[126,1464,914,2031]" captionTargetPageId="1" captionText="Fig. 1. AC, Belisana aliformis, new species; DF, Belisana huberi, new species; GI, Belisana lamellaris, new species; JL, Khorata diaoluoshanensis, new species. A, D, G, J, male habitus, dorsal view; B, E, H, K, male habitus, lateral view; C, F, I, L, female abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4509102" httpUri="https://zenodo.org/record/4509102/files/figure.png" pageId="2" pageNumber="47">Figs. 1</figureCitation>
DF, 3AG)
</paragraph>
</subSubSection>
<subSubSection id="C34984A9FC53FFF082D5FAB93174DBD7" pageId="2" pageNumber="47" type="materials_examined">
<paragraph id="8BECD722FC53FFF082D5FAB930E9DB9E" blockId="2.[148,788,1389,1468]" pageId="2" pageNumber="47">
<emphasis id="B9270B30FC53FFF082D5FAB9331ADBA0" bold="true" box="[148,378,1389,1410]" pageId="2" pageNumber="47">
<emphasis id="B9270B30FC53FFF082D5FAB93304DBA0" bold="true" box="[148,356,1389,1410]" italics="true" pageId="2" pageNumber="47">Material examined.</emphasis>
</emphasis>
<materialsCitation id="3B3BDD7FFC53FFF083C4FAB930E9DB9E" collectingDate="2005-03-22" collectionCode="IZCAS" collectorName="Island &amp; G. Deng" country="China" county="Changjiang County" latitude="19.066668" location="Bawangling National Natural Reserve" longLatPrecision="1273" longitude="109.13333" pageId="2" pageNumber="47" specimenCount="1" stateProvince="Hainan" typeStatus="holotype">
<typeStatus id="54E86980FC53FFF083C4FAB93387DBA0" box="[389,487,1389,1410]" pageId="2" pageNumber="47" type="holotype">Holotype</typeStatus>
: male (
<collectionCode id="ED424FE7FC53FFF08077FAB930E4DBA0" box="[566,644,1389,1410]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="2" pageNumber="47">IZCAS</collectionCode>
),
<location id="8E8C81F9FC53FFF080D8FAB933E6DBBD" LSID="urn:lsid:plazi:treatment:03FA6634FC53FFF08370FAD93728DAA7:8E8C81F9FC53FFF080D8FAB933E6DBBD" country="China" county="Changjiang County" latitude="19.066668" longLatPrecision="1273" longitude="109.13333" name="Bawangling National Natural Reserve" pageId="2" pageNumber="47" stateProvince="Hainan">Bawangling National Natural Reserve</location>
(
<geoCoordinate id="EE67B1E5FC53FFF083D2FA5E3382DBBD" box="[403,482,1417,1439]" degrees="19" direction="north" minutes="04" orientation="latitude" pageId="2" pageNumber="47" precision="925" value="19.066668">
19
<emphasis id="B9270B30FC53FFF083EAFA5D33D5DBBC" bold="true" box="[427,437,1417,1438]" pageId="2" pageNumber="47">°</emphasis>
04'N
</geoCoordinate>
<geoCoordinate id="EE67B1E5FC53FFF083A6FA5E3023DBBD" box="[487,579,1417,1439]" degrees="109" direction="east" minutes="08" orientation="longitude" pageId="2" pageNumber="47" precision="925" value="109.13333">
109
<emphasis id="B9270B30FC53FFF0804AFA5D3075DBBC" bold="true" box="[523,533,1417,1438]" pageId="2" pageNumber="47">°</emphasis>
08'E
</geoCoordinate>
),
<collectingCounty id="628DAFAEFC53FFF08012FA5E3170DBBD" box="[595,784,1418,1439]" pageId="2" pageNumber="47">Changjiang County</collectingCounty>
,
<collectingRegion id="499719C0FC53FFF082D5FA7332B9DB9E" box="[148,217,1447,1468]" country="China" name="Hainan" pageId="2" pageNumber="47">Hainan</collectingRegion>
<collectorName id="26A6B2F4FC53FFF082A3FA73337FDB9E" box="[226,287,1447,1468]" pageId="2" pageNumber="47">Island</collectorName>
,
<collectingCountry id="F34497B2FC53FFF0836BFA733306DB9E" box="[298,358,1447,1468]" name="China" pageId="2" pageNumber="47">China</collectingCountry>
, coll.
<collectorName id="26A6B2F4FC53FFF083E2FA733398DB9E" box="[419,504,1447,1468]" pageId="2" pageNumber="47">G. Deng</collectorName>
,
<date id="FFEDF1E2FC53FFF08045FA7330E5DB9E" box="[516,645,1447,1468]" pageId="2" pageNumber="47" value="2005-03-22">
<collectingDate id="EFA9080AFC53FFF08045FA7330E5DB9E" box="[516,645,1447,1468]" pageId="2" pageNumber="47" value="2005-03-22">22 Mar.2005</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BECD722FC53FFF082D5FA343174DBD7" blockId="2.[148,788,1504,1525]" box="[148,788,1504,1525]" pageId="2" pageNumber="47">
<materialsCitation id="3B3BDD7FFC53FFF082D5FA343170DBD7" box="[148,784,1504,1525]" collectingDate="2005-03-22" collectionCode="IZCAS" collectorName="Island &amp; G. Deng" country="China" county="Changjiang County" latitude="19.066668" location="Bawangling National Natural Reserve" longLatPrecision="1273" longitude="109.13333" pageId="2" pageNumber="47" specimenCount="9" specimenCount-female="7" specimenCount-male="2" stateProvince="Hainan" typeStatus="paratype">
<typeStatus id="54E86980FC53FFF082D5FA343295DBD7" box="[148,245,1504,1525]" pageId="2" pageNumber="47" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D551CABFC53FFF082BFFA343328DBD7" box="[254,328,1504,1525]" pageId="2" pageNumber="47" type="male">2 males</specimenCount>
and
<specimenCount id="9D551CABFC53FFF08336FA3433B4DBD7" box="[375,468,1504,1525]" pageId="2" pageNumber="47" type="female">7 females</specimenCount>
(
<collectionCode id="ED424FE7FC53FFF083A1FA34304ADBD7" box="[480,554,1504,1525]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="2" pageNumber="47">IZCAS</collectionCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34984A9FC53FFF082D5F9C930A1D8B7" pageId="2" pageNumber="47" type="diagnosis">
<paragraph id="8BECD722FC53FFF082D5F9C930A1D8B7" blockId="2.[148,787,1565,1685]" pageId="2" pageNumber="47">
<emphasis id="B9270B30FC53FFF082D5F9C93340D817" bold="true" box="[148,288,1565,1589]" pageId="2" pageNumber="47">
<emphasis id="B9270B30FC53FFF082D5F9C93369D817" bold="true" box="[148,265,1565,1589]" italics="true" pageId="2" pageNumber="47">Diagnosis.</emphasis>
</emphasis>
The new species can be easily distinguished from known congeners by the modifications of chelicerae (pair of upward-directed proximal apophyses), the shape of procursus, and the unique female vulval structures.
</paragraph>
</subSubSection>
<subSubSection id="C34984A9FC53FFF082D5F96937AFDD87" pageId="2" pageNumber="47" type="description">
<paragraph id="8BECD722FC53FFF082D5F9693174D617" blockId="2.[148,788,1725,2101]" pageId="2" pageNumber="47">
<emphasis id="B9270B30FC53FFF082D5F969330DD8F7" bold="true" box="[148,365,1725,1749]" pageId="2" pageNumber="47">
<emphasis id="B9270B30FC53FFF082D5F969337AD8F7" bold="true" box="[148,282,1725,1749]" italics="true" pageId="2" pageNumber="47">Description.</emphasis>
Male
</emphasis>
(
<typeStatus id="54E86980FC53FFF08338F96A33BDD8F7" box="[377,477,1726,1749]" pageId="2" pageNumber="47" type="holotype">holotype</typeStatus>
). Total length 1.75 (1.86 with clypeus), carapace width 0.75. Leg 1: 11.87 (3.05+0.27+2.95+4.5+1.1), tibia 2: 1.92, tibia 3: 1.3, tibia 4: 1.87; tibia 1 L/d: 39. Habitus as in
<figureCitation id="1368CBA7FC53FFF08042F8CA30C7D917" box="[515,679,1822,1845]" captionStart="Fig" captionStartId="3.[126,161,2020,2041]" captionTargetBox="[230,1352,224,1970]" captionTargetId="figure-12@3.[214,1379,209,1992]" captionTargetPageId="3" captionText="Fig. 2. A, Belisana aliformis, new species, anterior part of male prosoma (arrow shows the duck-beak shaped apophysis), dorsal view; B C, male left palp, prolateral (B) and retrolateral (C) views; D, male left procursus, prolateral view; E, apical part of cheliceral frontal apophysis, internal view; F, male chelicerae, frontal view; GH, cleared epigynum, ventral (G) and dorsal (H) views. Scale bars: AD, FH = 0.2 mm; E = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4509104" httpUri="https://zenodo.org/record/4509104/files/figure.png" pageId="2" pageNumber="47">Fig. 2D and 2E</figureCitation>
. Carapace yellowish, brown margins laterally, sternum whitish. Legs yellow except brown on patella and joints of tibia to metatarsus, without darker rings. Opisthosoma whitish-grey, with some indistinct darker spots dorsally. Ocular area not elevated, thoracic furrow absent; distance PME-PME 0.12; diameter PME 0.08; distance PME-ALE 0.03; AME absent. Clypeus unmodified. Sternum slightly wider than long (0.51/ 0.47). Chelicerae as in
<figureCitation id="1368CBA7FC53FFF083CEF7CA3057D617" box="[399,567,2078,2101]" captionStart="Fig" captionStartId="4.[148,183,2048,2069]" captionTargetBox="[244,1404,200,2003]" captionTargetId="figure-9@4.[220,1424,182,2023]" captionTargetPageId="4" captionText="Fig. 3. A, Belisana huberi, new species, male left procursus, prolateral view; B, male chelicerae, lateral view; C, male chelicerae, frontal view; DE, male left palp, prolateral (D) and retrolateral (E) views; FG, cleared epigynum, ventral (F) and dorsal (G) views. Scale bars: AE = 0.2 mm; F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4509108" httpUri="https://zenodo.org/record/4509108/files/figure.png" pageId="2" pageNumber="47">Fig. 3B and 3C</figureCitation>
, with pair of curved
</paragraph>
<paragraph id="8BECD722FC53FFF08111FF1A363DDC27" blockId="2.[848,1487,206,517]" pageId="2" pageNumber="47">
frontal apophyses, tips with cuticular ridges laterally, 0.14 apart; with pair of upward-directed proximal apophyses and pair of proximolateral apophyses. Palps as in
<figureCitation id="1368CBA7FC53FFF0870BFEDA31FFDF67" captionStart="Fig" captionStartId="4.[148,183,2048,2069]" captionTargetBox="[244,1404,200,2003]" captionTargetId="figure-9@4.[220,1424,182,2023]" captionTargetPageId="4" captionText="Fig. 3. A, Belisana huberi, new species, male left procursus, prolateral view; B, male chelicerae, lateral view; C, male chelicerae, frontal view; DE, male left palp, prolateral (D) and retrolateral (E) views; FG, cleared epigynum, ventral (F) and dorsal (G) views. Scale bars: AE = 0.2 mm; F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4509108" httpUri="https://zenodo.org/record/4509108/files/figure.png" pageId="2" pageNumber="47">Fig. 3A, 3D and 3E</figureCitation>
; trochanter with short retrolateral apophysis, femur with distinctive hump proximo-dorsally, procursus complicated distally, with ventral membranous flap, bulb with hooked apophysis and complicated embolus. Retrolateral trichobothrium of tibia 1 at 16%; legs without spines, vertical hairs, and curved hairs; tarsus 1 with about 15 quite distinct pseudosegments distally.
</paragraph>
<paragraph id="8BECD722FC53FFF08111FDFA31D3DCE7" blockId="2.[848,1487,558,709]" pageId="2" pageNumber="47">
Variation. Tibia
<quantity id="4CAB7AC7FC53FFF081BFFDFA3645DC67" box="[1022,1061,558,581]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="47" unit="in" value="1.0">1 in</quantity>
other males (n = 2): 3.05, 3.07. Distance between tips of distal cheliceral apophyses in other males (n = 2): 0.09, 0.13. One male with obscure moustache-like marks on median of carapace; dorsal spots on opisthosoma nearly invisible.
</paragraph>
<paragraph id="8BECD722FC53FFF08111FD3937AFDD87" blockId="2.[848,1488,749,933]" pageId="2" pageNumber="47">
<emphasis id="B9270B30FC53FFF08111FD3931D4DD27" bold="true" box="[848,948,749,773]" pageId="2" pageNumber="47">Females.</emphasis>
In general, similar to male. Some specimens with distinctive moustache-like marks on median of carapace. Tibia
<quantity id="4CAB7AC7FC53FFF081CCFCFA31D4DD67" box="[909,948,814,837]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="47" unit="in" value="1.0">1 in</quantity>
females (n = 7): 2.272.87 (mean: 2.58). Epigynum simple and flat externally, with distinctive internal dark structures visible through cuticle (
<figureCitation id="1368CBA7FC53FFF086F7FCBA376DDDA7" box="[1206,1293,878,901]" captionStart="Fig" captionStartId="1.[127,162,2048,2069]" captionTargetBox="[126,1464,914,2028]" captionTargetId="figure-102@1.[126,1464,914,2031]" captionTargetPageId="1" captionText="Fig. 1. AC, Belisana aliformis, new species; DF, Belisana huberi, new species; GI, Belisana lamellaris, new species; JL, Khorata diaoluoshanensis, new species. A, D, G, J, male habitus, dorsal view; B, E, H, K, male habitus, lateral view; C, F, I, L, female abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4509102" httpUri="https://zenodo.org/record/4509102/files/figure.png" pageId="2" pageNumber="47">Figs. 1F</figureCitation>
and
<figureCitation id="1368CBA7FC53FFF0877EFCBA373EDDA7" box="[1343,1374,878,901]" captionStart="Fig" captionStartId="4.[148,183,2048,2069]" captionTargetBox="[244,1404,200,2003]" captionTargetId="figure-9@4.[220,1424,182,2023]" captionTargetPageId="4" captionText="Fig. 3. A, Belisana huberi, new species, male left procursus, prolateral view; B, male chelicerae, lateral view; C, male chelicerae, frontal view; DE, male left palp, prolateral (D) and retrolateral (E) views; FG, cleared epigynum, ventral (F) and dorsal (G) views. Scale bars: AE = 0.2 mm; F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4509108" httpUri="https://zenodo.org/record/4509108/files/figure.png" pageId="2" pageNumber="47">3F</figureCitation>
); with pair of posterior pockets, 0.09 apart. Dorsal view as in
<figureCitation id="1368CBA7FC53FFF08734FC5A37ABDD87" box="[1397,1483,910,933]" captionStart="Fig" captionStartId="4.[148,183,2048,2069]" captionTargetBox="[244,1404,200,2003]" captionTargetId="figure-9@4.[220,1424,182,2023]" captionTargetPageId="4" captionText="Fig. 3. A, Belisana huberi, new species, male left procursus, prolateral view; B, male chelicerae, lateral view; C, male chelicerae, frontal view; DE, male left palp, prolateral (D) and retrolateral (E) views; FG, cleared epigynum, ventral (F) and dorsal (G) views. Scale bars: AE = 0.2 mm; F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4509108" httpUri="https://zenodo.org/record/4509108/files/figure.png" pageId="2" pageNumber="47">Fig. 3G</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34984A9FC53FFF08111FC1931F5DA67" pageId="2" pageNumber="47" type="etymology">
<paragraph id="8BECD722FC53FFF08111FC1931F5DA67" blockId="2.[848,1487,973,1093]" pageId="2" pageNumber="47">
<emphasis id="B9270B30FC53FFF08111FC193182DDC7" bold="true" box="[848,994,973,997]" pageId="2" pageNumber="47">
<emphasis id="B9270B30FC53FFF08111FC1931ABDDC7" bold="true" box="[848,971,973,997]" italics="true" pageId="2" pageNumber="47">Etymology.</emphasis>
</emphasis>
Named for Dr. Bernhard A. Huber from the Alexander Koenig Zoological Research Museum who contributed to our understanding of the taxonomy of this genus.
</paragraph>
</subSubSection>
<subSubSection id="C34984A9FC53FFF08111FBB93728DAA7" box="[848,1352,1133,1157]" pageId="2" pageNumber="47" type="distribution">
<paragraph id="8BECD722FC53FFF08111FBB93728DAA7" blockId="2.[848,1352,1133,1157]" box="[848,1352,1133,1157]" pageId="2" pageNumber="47">
<emphasis id="B9270B30FC53FFF08111FBB93191DAA7" bold="true" box="[848,1009,1133,1157]" pageId="2" pageNumber="47">
<emphasis id="B9270B30FC53FFF08111FBB931BCDAA7" bold="true" box="[848,988,1133,1157]" italics="true" pageId="2" pageNumber="47">Distribution.</emphasis>
</emphasis>
<collectingRegion id="499719C0FC53FFF081BBFBBA362ADAA7" box="[1018,1098,1134,1157]" country="China" name="Hainan" pageId="2" pageNumber="47">Hainan</collectingRegion>
,
<collectingCountry id="F34497B2FC53FFF08616FBBA36F7DAA7" box="[1111,1175,1134,1157]" name="China" pageId="2" pageNumber="47">China</collectingCountry>
(present study).
</paragraph>
</subSubSection>
</treatment>
</document>