198 lines
19 KiB
XML
198 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1150.93760" ID-Pensoft-Pub="1313-2970-1150-1" ID-Pensoft-UUID="2D0F861C78665B9BABB241437CA5ED53" ID-ZooBank="A38C5FB69F664F858788AAA53D21704D" ModsDocID="1313-2970-1150-1" checkinTime="1677664009133" checkinUser="pensoft" docAuthor="Crews, Sarah C." docDate="2023" docId="C6E89A76CA27507196EFD86EE9149EF6" docLanguage="en" docName="ZooKeys 1150: 1-189" docOrigin="ZooKeys 1150" docPubDate="2023-02-27" docSource="http://dx.doi.org/10.3897/zookeys.1150.93760" docTitle="Karaops kariyarra Crews 2013" docType="treatment" docVersion="1" id="2D0F861C78665B9BABB241437CA5ED53" lastPageNumber="1" masterDocId="2D0F861C78665B9BABB241437CA5ED53" masterDocTitle="But wait, there's more! Descriptions of new species and undescribed sexes of flattie spiders (Araneae, Selenopidae, Karaops) from Australia" masterLastPageNumber="189" masterPageNumber="1" pageNumber="1" updateTime="1677664009133" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>But wait, there's more! Descriptions of new species and undescribed sexes of flattie spiders (Araneae, Selenopidae, Karaops) from Australia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Crews, Sarah C.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9360-6236</mods:nameIdentifier>
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<mods:affiliation>California Academy of Sciences, Department of Entomology, 55 Music Concourse Drive, San Francisco, CA, 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="email">screwsemail@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-27</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1150</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>189</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1150.93760</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1150.93760</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1150-1</mods:identifier>
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<mods:identifier type="ZooBank">A38C5FB69F664F858788AAA53D21704D</mods:identifier>
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<mods:identifier type="Pensoft-UUID">2D0F861C78665B9BABB241437CA5ED53</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:C6E89A76CA27507196EFD86EE9149EF6" httpUri="http://treatment.plazi.org/id/C6E89A76CA27507196EFD86EE9149EF6" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="3R2MC" LSID="C6E89A76-CA27-5071-96EF-D86EE9149EF6" authority="Crews, 2013" authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops kariyarra" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kariyarra">Karaops kariyarra Crews, 2013</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 73" captionStartId="F82" captionText="Figure 73. Members of the Pilbara-Gascoyne species group A Karaops nyiyaparli, BHP Rail Line (sel _ 1230, WAM T 155606) (Photo: J. DeJong) B habitat of Karaops nyiyaparli, Millstream-Chichester National Park C habitat of Karaops nyiyaparli along BHP Rail Line D Karaops nyiyaparli, holotype female (WAM T 111455) E Karaops kariyarra, holotype female, Wodgina Mine (WAM T 105208) F Karaops kariyarra, adult female, 55 km S of Port Hedland (WAM T 106657) G Karaops feedtime, holotype female, NNW Newman (WAM T 111456) H Karaops forteyi, holotype female, Cowra Line Camp (WAM T 79407) I same, paratype male, Cowra Line Camp (WAM T 79408)." figureDoi="10.3897/zookeys.1150.93760.figure73" httpUri="https://binary.pensoft.net/fig/816462" pageId="0" pageNumber="1">Fig. 73E</figureCitation>
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<figureCitation captionStart="Map 1" captionStartId="F3" captionText="Map 1. Map of Australia showing areas that are expanded in subsequent figures. CD = Central Desert group, D = Karaops dawara group, F = Karaops francesae group, K = Kimberley group, UA = species that are not assigned to a group, PG = Pilbara-Gascoyne group." figureDoi="10.3897/zookeys.1150.93760.map1" httpUri="https://binary.pensoft.net/fig/816383" pageId="0" pageNumber="1">, Maps 1</figureCitation>
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<figureCitation captionStart="Map 9" captionStartId="F57" captionText="Map 9. Species of the Pilbara-Gascoyne species group A diamonds (juveniles): blue = Erawandoo Hill, orange = Mt. Augustus, red = Nammuldi, yellow = Hillside-Corunna-Glenn Herring, black = Marble Bar-Glacier Valley, pink = Orebody 19, 31, teal = Wheelara North, orange with thick outline = Harding Dam (maybe Karaops ngarluma, see text); Circles: dark blue = Karaops julianneae, dark green = Karaops badgeradda, red = Karaops joehaeneri sp. nov., teal = Karaops karrawarla, blue = Karaops durrantorum sp. nov., brown = Karaops burbidgei, green = Karaops martamarta, purple = Karaops nyangumarta, black = Karaops banyjima, yellow = Karaops morganoconnelli sp. nov., gray = Karaops nyiyaparli, red-brown = Karaops feedtime, bright pink = Karaops nyamal (probably, see text), white = Karaops kariyarra, bright green = Karaops yurlburr, orange = Karaops ngarluma, tan = Karaops jaburrara, peach = Karaops forteyi, teal = new species yet undescribed; small white diamonds are unidentifiable immatures for which no molecular data are available B boxed area expanded for clarity." figureDoi="10.3897/zookeys.1150.93760.map9" httpUri="https://binary.pensoft.net/fig/816437" pageId="0" pageNumber="1">, 9A, B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops kariyarra" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kariyarra">Karaops kariyarra</taxonomicName>
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Crews, 2013: 451, figs 7, 8 (♀, examined).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The female is most similar to
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<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops nyiyaparli" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nyiyaparli">
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<emphasis italics="true" pageId="0" pageNumber="1">Karaops nyiyaparli</emphasis>
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</taxonomicName>
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in that the copulatory ducts are located in a depression of the median field of the epigyne, and the lateral lobes are easy to distinguish (
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3647.3.3" author="Crews, SC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="443 - 469" refId="B11" refString="Crews, SC, 2013. Thirteen new species of the spider genus Karaops (Araneae: Selenopidae) from Western Australia. Zootaxa 3647 (3): 443 - 469, DOI: https://doi.org/10.11646/zootaxa.3647.3.3" title="Thirteen new species of the spider genus Karaops (Araneae: Selenopidae) from Western Australia." url="https://doi.org/10.11646/zootaxa.3647.3.3" volume="3647" year="2013">Crews 2013</bibRefCitation>
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: figs 5, 7). The endogynes are also similar in that the spermathecae are dumbbell shaped, and the accessory bulbs (erroneously labeled as spermathecae in
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3647.3.3" author="Crews, SC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="443 - 469" refId="B11" refString="Crews, SC, 2013. Thirteen new species of the spider genus Karaops (Araneae: Selenopidae) from Western Australia. Zootaxa 3647 (3): 443 - 469, DOI: https://doi.org/10.11646/zootaxa.3647.3.3" title="Thirteen new species of the spider genus Karaops (Araneae: Selenopidae) from Western Australia." url="https://doi.org/10.11646/zootaxa.3647.3.3" volume="3647" year="2013">Crews (2013)</bibRefCitation>
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) are long and thin. They can be differentiated by the shape of the depression on the endogyne. In
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<taxonomicName lsidName="K. nyiyaparli" pageId="0" pageNumber="1" rank="species" species="nyiyaparli">
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<emphasis italics="true" pageId="0" pageNumber="1">K. nyiyaparli</emphasis>
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</taxonomicName>
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, it is oval and in
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<taxonomicName lsidName="K. kariyarra" pageId="0" pageNumber="1" rank="species" species="kariyarra">
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<emphasis italics="true" pageId="0" pageNumber="1">K. kariyarra</emphasis>
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</taxonomicName>
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it is heart shaped. The lateral lobes of
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<taxonomicName lsidName="K. kariyarra" pageId="0" pageNumber="1" rank="species" species="kariyarra">
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<emphasis italics="true" pageId="0" pageNumber="1">K. kariyarra</emphasis>
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</taxonomicName>
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are pointed at the basomedial margin, and they are not in
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<taxonomicName lsidName="K. nyiyaparli" pageId="0" pageNumber="1" rank="species" species="nyiyaparli">
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<emphasis italics="true" pageId="0" pageNumber="1">K. nyiyaparli</emphasis>
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</taxonomicName>
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. The accessory bulbs are extremely tiny in
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<taxonomicName lsidName="K. kariyarra" pageId="0" pageNumber="1" rank="species" species="kariyarra">
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<emphasis italics="true" pageId="0" pageNumber="1">K. kariyarra</emphasis>
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</taxonomicName>
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(
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3647.3.3" author="Crews, SC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="443 - 469" refId="B11" refString="Crews, SC, 2013. Thirteen new species of the spider genus Karaops (Araneae: Selenopidae) from Western Australia. Zootaxa 3647 (3): 443 - 469, DOI: https://doi.org/10.11646/zootaxa.3647.3.3" title="Thirteen new species of the spider genus Karaops (Araneae: Selenopidae) from Western Australia." url="https://doi.org/10.11646/zootaxa.3647.3.3" volume="3647" year="2013">Crews 2013</bibRefCitation>
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: figs 6, 8).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The female is described in in
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3647.3.3" author="Crews, SC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="443 - 469" refId="B11" refString="Crews, SC, 2013. Thirteen new species of the spider genus Karaops (Araneae: Selenopidae) from Western Australia. Zootaxa 3647 (3): 443 - 469, DOI: https://doi.org/10.11646/zootaxa.3647.3.3" title="Thirteen new species of the spider genus Karaops (Araneae: Selenopidae) from Western Australia." url="https://doi.org/10.11646/zootaxa.3647.3.3" volume="3647" year="2013">Crews (2013)</bibRefCitation>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Male.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species is known only from two nearby localities in the Chichester subregion of the Pilbara, Western Australia (Map
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<figureCitation captionStart="Map 9" captionStartId="F57" captionText="Map 9. Species of the Pilbara-Gascoyne species group A diamonds (juveniles): blue = Erawandoo Hill, orange = Mt. Augustus, red = Nammuldi, yellow = Hillside-Corunna-Glenn Herring, black = Marble Bar-Glacier Valley, pink = Orebody 19, 31, teal = Wheelara North, orange with thick outline = Harding Dam (maybe Karaops ngarluma, see text); Circles: dark blue = Karaops julianneae, dark green = Karaops badgeradda, red = Karaops joehaeneri sp. nov., teal = Karaops karrawarla, blue = Karaops durrantorum sp. nov., brown = Karaops burbidgei, green = Karaops martamarta, purple = Karaops nyangumarta, black = Karaops banyjima, yellow = Karaops morganoconnelli sp. nov., gray = Karaops nyiyaparli, red-brown = Karaops feedtime, bright pink = Karaops nyamal (probably, see text), white = Karaops kariyarra, bright green = Karaops yurlburr, orange = Karaops ngarluma, tan = Karaops jaburrara, peach = Karaops forteyi, teal = new species yet undescribed; small white diamonds are unidentifiable immatures for which no molecular data are available B boxed area expanded for clarity." figureDoi="10.3897/zookeys.1150.93760.map9" httpUri="https://binary.pensoft.net/fig/816437" pageId="0" pageNumber="1">9A, B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="natural history">
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<paragraph pageId="0" pageNumber="1">Natural history.</paragraph>
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<paragraph pageId="0" pageNumber="1">Late March to early April are hot and wet times, transitioning to drier in April. The adult females of several species from the Chichester region have been found in the hotter, wetter times of the year (Suppl. material 2: table S1).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="discussion">
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<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops kariyarra" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kariyarra">
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<emphasis italics="true" pageId="0" pageNumber="1">Karaops kariyarra</emphasis>
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</taxonomicName>
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is only known from two female specimens collected in two separate pitfall traps left out for ~ 10 days in late March-early April. This species is similar to and overlaps with
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<taxonomicName lsidName="K. nyiyaparli" pageId="0" pageNumber="1" rank="species" species="nyiyaparli">
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<emphasis italics="true" pageId="0" pageNumber="1">K. nyiyaparli</emphasis>
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</taxonomicName>
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, but both specimens of
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<taxonomicName lsidName="K. kariyarra" pageId="0" pageNumber="1" rank="species" species="kariyarra">
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<emphasis italics="true" pageId="0" pageNumber="1">K. kariyarra</emphasis>
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</taxonomicName>
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are exactly the same despite being collected at different areas, and these two are both different from
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<taxonomicName lsidName="K. nyiyaparli" pageId="0" pageNumber="1" rank="species" species="nyiyaparli">
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<emphasis italics="true" pageId="0" pageNumber="1">K. nyiyaparli</emphasis>
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</taxonomicName>
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which is widespread; thus, it does not appear to be a variant of
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<taxonomicName lsidName="K. nyiyaparli" pageId="0" pageNumber="1" rank="species" species="nyiyaparli">
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<emphasis italics="true" pageId="0" pageNumber="1">K. nyiyaparli</emphasis>
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</taxonomicName>
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. No molecular data from these specimens were available.
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</paragraph>
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<caption doi="10.3897/zookeys.1150.93760.figure73" httpUri="https://binary.pensoft.net/fig/816462" pageId="0" pageNumber="1" start="Figure 73" startId="F82">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 73.</emphasis>
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Members of the Pilbara-Gascoyne species group
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops nyiyaparli" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nyiyaparli">
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<emphasis italics="true" pageId="0" pageNumber="1">Karaops nyiyaparli</emphasis>
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</taxonomicName>
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, BHP Rail Line (sel_1230, WAM T155606) (Photo: J. DeJong)
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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habitat of
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<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops nyiyaparli" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nyiyaparli">
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<emphasis italics="true" pageId="0" pageNumber="1">Karaops nyiyaparli</emphasis>
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</taxonomicName>
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, Millstream-Chichester National Park
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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habitat of
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<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops nyiyaparli" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nyiyaparli">
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<emphasis italics="true" pageId="0" pageNumber="1">Karaops nyiyaparli</emphasis>
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</taxonomicName>
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along BHP Rail Line
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops nyiyaparli" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nyiyaparli">
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<emphasis italics="true" pageId="0" pageNumber="1">Karaops nyiyaparli</emphasis>
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</taxonomicName>
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, holotype female(WAM T111455)
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops kariyarra" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kariyarra">
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<emphasis italics="true" pageId="0" pageNumber="1">Karaops kariyarra</emphasis>
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</taxonomicName>
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, holotype female, Wodgina Mine (WAM T105208)
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops kariyarra" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kariyarra">
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<emphasis italics="true" pageId="0" pageNumber="1">Karaops kariyarra</emphasis>
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</taxonomicName>
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, adult female, 55 km S of Port Hedland (WAM T106657)
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<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
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<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops feedtime" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="feedtime">
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<emphasis italics="true" pageId="0" pageNumber="1">Karaops feedtime</emphasis>
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</taxonomicName>
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, holotype female, NNW Newman (WAM T111456)
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<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
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<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops forteyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="forteyi">
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<emphasis italics="true" pageId="0" pageNumber="1">Karaops forteyi</emphasis>
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</taxonomicName>
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, holotype female, Cowra Line Camp (WAM T79407)
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<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
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same, paratype male, Cowra Line Camp (WAM T79408).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |