494 lines
51 KiB
XML
494 lines
51 KiB
XML
<document id="BFD928DDFA65D39E9FD036B3A43C36C1" ID-DOI="10.11646/zootaxa.4088.4.6" ID-GBIF-Dataset="689b51bb-e0d7-4e6b-b7d4-e89a82dbbf5b" ID-ISSN="1175-5326" ID-Zenodo-Dep="255979" ID-ZooBank="44C19F07-853C-4D0E-A8F6-4B5230FAE927" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457683668170" checkinUser="plazi" docAuthor="Heiss, Ernst" docDate="2016" docId="CF3987BCFFDC021CFF124C4FFEB42685" docLanguage="en" docName="15606-5126-1-PB.pdf.imf" docOrigin="Zootaxa 4088 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helenus hirsutus Champion 1898" docType="treatment" docVersion="11" lastPageNumber="561" masterDocId="3300FFC4FFDE021AFF85480DFF872755" masterDocTitle="RevIew of the NeotropIcal genus Helenus WhIte, 1879, wIth descrIptIon of a new specIes (HemIptera: Heteroptera: AradIdae)" masterLastPageNumber="562" masterPageNumber="555" pageNumber="557" updateTime="1698177951616" updateUser="plazi">
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<mods:titleInfo id="00D0095D0A5D7F70F40FF41EDEC3FEA4">
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<mods:title id="93BE5F52D4CDD61CB461900EEC742B97">RevIew of the NeotropIcal genus Helenus WhIte, 1879, wIth descrIptIon of a new specIes (HemIptera: Heteroptera: AradIdae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B83F43FF2A013D675D275F31C6E9D072" type="personal">
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<mods:roleTerm id="7562BFAB72B86FE6499A35F24E4E7AE0">Author</mods:roleTerm>
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<mods:namePart id="97D3AE7DDCAB726A36D8C383565D6100">Heiss, Ernst</mods:namePart>
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<mods:typeOfResource id="66CB2D17A6886FBED202986754DEC6FC">text</mods:typeOfResource>
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<mods:title id="7D1D6241A65A4AB78387257914BE8F02">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="410B40D59C65D9A0C035164EC7C4A52C">
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<mods:date id="0150B9143678E9158ECCC4AC9EB517B3">2016</mods:date>
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<mods:detail id="2FB0185441BCE0AD1C0B76FFEDCE6335" type="volume">
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<mods:number id="2149EEA96B218A1DD72DBF88D4FFBCD8">4088</mods:number>
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</mods:detail>
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<mods:detail id="AA0A8866066AF9489FF2F7B0F097F460" type="issue">
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<mods:number id="6D3B94CB940987C97D014135F8678A9F">4</mods:number>
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<mods:classification id="26AE946C03980B953EA58B647D159F2D">journal article</mods:classification>
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<mods:identifier id="2A1C73C6055D3FE7F233AB863226EB6F" type="DOI">10.11646/zootaxa.4088.4.6</mods:identifier>
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<mods:identifier id="F6C5DC463448E8F1DDF2319208BD9599" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="CF3987BCFFDC021CFF124C4FFEB42685" ID-DOI="http://doi.org/10.5281/zenodo.6083007" ID-GBIF-Taxon="117222022" ID-Zenodo-Dep="6083007" LSID="urn:lsid:plazi:treatment:CF3987BCFFDC021CFF124C4FFEB42685" httpUri="http://treatment.plazi.org/id/CF3987BCFFDC021CFF124C4FFEB42685" lastPageId="6" lastPageNumber="561" pageId="2" pageNumber="557">
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<subSubSection id="0F8A6521FFDC0218FF124C4FFEDF232B" pageId="2" pageNumber="557" type="nomenclature">
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<paragraph id="472F36AAFFDC0218FF124C4FFDBC2308" blockId="2.[151,571,1090,1150]" box="[151,571,1090,1117]" pageId="2" pageNumber="557">
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<heading id="1C6781C6FFDC0218FF124C4FFDBC2308" bold="true" box="[151,571,1090,1117]" fontSize="11" level="1" pageId="2" pageNumber="557" reason="1">
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<taxonomicName id="80904D29FFDC0218FF124C4FFDBC2308" authority="Champion, 1898" authorityName="Champion" authorityYear="1898" box="[151,571,1090,1117]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="hirsutus">
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<emphasis id="75E4EAB8FFDC0218FF124C4FFDBC2308" bold="true" box="[151,571,1090,1117]" pageId="2" pageNumber="557">
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<emphasis id="75E4EAB8FFDC0218FF124C4FFEE12309" bold="true" box="[151,358,1090,1116]" italics="true" pageId="2" pageNumber="557">Helenus hirsutus</emphasis>
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Champion, 1898
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="472F36AAFFDC0218FF124C68FEDF232B" blockId="2.[151,571,1090,1150]" box="[151,344,1125,1150]" pageId="2" pageNumber="557">
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(
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<figureCitation id="DFAB2A2FFFDC0218FF1A4C68FE8E2328" box="[159,265,1125,1150]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1938,1959]" captionTargetBox="[188,1425,245,1897]" captionTargetId="figure@3.[151,1436,195,1913]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Helenus spp.: 1 — Helenus hesiformis, holotype male, dorsal view; 2 — Helenus hirsutus, lectotype male, dorsal view; 3 — labels attached to holotype of hesiformis; 4 — ditto, reverse side; 5 — labels attached to holotype of hirsutus." httpUri="https://zenodo.org/record/255980/files/figure.png" pageId="2" pageNumber="557">Figs. 2, 5</figureCitation>
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,
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<figureCitation id="DFAB2A2FFFDC0218FE934C6BFEA2232B" box="[278,293,1126,1150]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1702,1723]" captionTargetBox="[158,1410,205,1663]" captionTargetId="figure@4.[152,1435,193,1681]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Helenus spp.: 6 — Helenus wachteli n. sp., holotype male, dorsal view; 7 — Helenus wachteli n. sp., paratype female, dorsal view; 8 – 10. Male pygophore: 8 — hesiformis; 9 — hirsutus; 10 — wachteli n. sp." httpUri="https://zenodo.org/record/255981/files/figure.png" pageId="2" pageNumber="557">9</figureCitation>
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,
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<figureCitation id="DFAB2A2FFFDC0218FEB74C68FEC9232B" box="[306,334,1125,1150]" captionStart="FIGURES 11 — 13" captionStartId="5.[151,264,1300,1321]" captionTargetBox="[278,1308,689,1274]" captionTargetId="figure@5.[270,1316,681,1278]" captionTargetPageId="5" captionText="FIGURES 11 — 13. Left paramere of Hesus spp.: 11 — hesiformis; 12 — hirsutus; 13 — wachteli n. sp." httpUri="https://zenodo.org/record/255982/files/figure.png" pageId="2" pageNumber="557">12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="0F8A6521FFDC0218FF124CA3FEA0217B" pageId="2" pageNumber="557" type="materials_examined">
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<paragraph id="472F36AAFFDC0218FF124CA3FD512267" blockId="2.[151,1437,1197,1870]" pageId="2" pageNumber="557">
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<emphasis id="75E4EAB8FFDC0218FF124CA3FE4A2393" bold="true" box="[151,461,1197,1222]" pageId="2" pageNumber="557">
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<typeStatus id="982B8808FFDC0218FF124CA3FF552390" box="[151,210,1198,1221]" pageId="2" pageNumber="557">Type</typeStatus>
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specimen examined:
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</emphasis>
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Male (
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<figureCitation id="DFAB2A2FFFDC0218FD9A4CA0FDE52393" box="[543,610,1197,1222]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1938,1959]" captionTargetBox="[188,1425,245,1897]" captionTargetId="figure@3.[151,1436,195,1913]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Helenus spp.: 1 — Helenus hesiformis, holotype male, dorsal view; 2 — Helenus hirsutus, lectotype male, dorsal view; 3 — labels attached to holotype of hesiformis; 4 — ditto, reverse side; 5 — labels attached to holotype of hirsutus." httpUri="https://zenodo.org/record/255980/files/figure.png" pageId="2" pageNumber="557">Fig. 2</figureCitation>
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) (glued on card, right antennal segment III+IV, left and right mesotarsus missing) labeled (
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<figureCitation id="DFAB2A2FFFDC0218FEDA4CDCFE2623BC" box="[351,417,1233,1258]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,264,1938,1959]" captionTargetBox="[188,1425,245,1897]" captionTargetId="figure@3.[151,1436,195,1913]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Helenus spp.: 1 — Helenus hesiformis, holotype male, dorsal view; 2 — Helenus hirsutus, lectotype male, dorsal view; 3 — labels attached to holotype of hesiformis; 4 — ditto, reverse side; 5 — labels attached to holotype of hirsutus." httpUri="https://zenodo.org/record/255980/files/figure.png" pageId="2" pageNumber="557">Fig. 5</figureCitation>
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): “
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<typeStatus id="982B8808FFDC0218FE4E4CDCFD8223BF" box="[459,517,1233,1258]" pageId="2" pageNumber="557">Type</typeStatus>
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” (pr., round label with red margin) // “Bugaba /
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<quantity id="80689B4FFFDC0218FBA74CDFFB2D23BF" box="[1058,1194,1233,1258]" metricMagnitude="2" metricUnit="m" metricValue="3.5052" metricValueMax="4.572" metricValueMin="2.4384" pageId="2" pageNumber="557" unit="ft" value="1150.0" valueMax="1500.0" valueMin="800.0">800–1500 ft</quantity>
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/ Champion” (pr) // “ B.C.A. Rhyn.II (pr) /
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<taxonomicName id="80904D29FFDC0218FE0A4CF8FE6C225B" box="[399,491,1269,1294]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">Helenus</taxonomicName>
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(pr) /
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<taxonomicName id="80904D29FFDC0218FDB04CF8FD09225B" box="[565,654,1269,1294]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="hirsutus">hirsutus</taxonomicName>
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(hw) / ♂ Ch “ (hw) // “sp.figured” (pr) // “ cleaned and remounted / E.Heiss
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<date id="332E106AFFDC0218FF714D17FECF2267" box="[244,328,1305,1330]" pageId="2" pageNumber="557" value="2015-10">X 2015</date>
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” (pr) // “ BMNH/ 1705803” (pr) //.
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</paragraph>
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<paragraph id="472F36AAFFDC0218FF424D33FCE92294" blockId="2.[151,1437,1197,1870]" pageId="2" pageNumber="557">
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As indicated by the author (p.75) the description was based on male and female specimens who are considered
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<typeStatus id="982B8808FFDC0218FF124D6FFF7A222C" box="[151,253,1378,1401]" pageId="2" pageNumber="557" type="syntype">syntypes</typeStatus>
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. The male specimen borrowed is also figured (Tab.V, fig. 16) and therefore selected as
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<typeStatus id="982B8808FFDC0218FA914D6CFAFD222F" box="[1300,1402,1377,1402]" pageId="2" pageNumber="557" type="lectotype">lectotype</typeStatus>
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in accordance with article 74B ICZN. It is designated here and labeled: “
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<typeStatus id="982B8808FFDC0218FC234D8BFB9F22CB" box="[934,1048,1414,1438]" pageId="2" pageNumber="557" type="lectotype">Lectotype</typeStatus>
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♂ /
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<taxonomicName id="80904D29FFDC0218FBCD4D88FB2322CB" box="[1096,1188,1413,1438]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">Helenus</taxonomicName>
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/
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<taxonomicName id="80904D29FFDC0218FB3F4D88FA9422CB" box="[1210,1299,1413,1438]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="hirsutus">hirsutus</taxonomicName>
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/ Champion 1898 / des. E.Heiss 2015” (pr, red label) //. Deposited in BMNH.
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</paragraph>
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<paragraph id="472F36AAFFDC0218FF424DC0FEA0217B" blockId="2.[151,1437,1197,1870]" pageId="2" pageNumber="557">
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<emphasis id="75E4EAB8FFDC0218FF424DC0FE3922B3" bold="true" box="[199,446,1485,1510]" pageId="2" pageNumber="557">Additional material.</emphasis>
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<specimenCount id="5196FD23FFDC0218FE4F4DC0FE6A22B3" box="[458,493,1485,1510]" pageId="2" pageNumber="557" type="generic">3♂</specimenCount>
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1♀ “
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<collectingCountry id="3F87763AFFDC0218FDB64DC3FD3422B3" box="[563,691,1485,1510]" name="Costa Rica" pageId="2" pageNumber="557">Costa Rica</collectingCountry>
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/ Carrara Nat. Res. / Westküste /
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<date id="332E106AFFDC0218FBCB4DC3FB3922B3" box="[1102,1214,1486,1510]" pageId="2" pageNumber="557" value="1987-02-01" valueMax="1987-02-08" valueMin="1987-02-01">1–8 II 87</date>
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Heiss” (pr) //;
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<specimenCount id="5196FD23FFDC0218FAFC4DC0FA1B22B3" box="[1401,1436,1485,1510]" pageId="2" pageNumber="557" type="generic">1♂</specimenCount>
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“
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<collectingCountry id="3F87763AFFDC0218FF274DFCFE9C215F" box="[162,283,1521,1546]" name="Nicaragua" pageId="2" pageNumber="557">Nicaragua</collectingCountry>
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: Rio San Juan / Bartola Field Stn.
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<quantity id="80689B4FFFDC0218FD224DFFFC96215F" box="[679,785,1522,1546]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="2" pageNumber="557" unit="m" value="85.0" valueMax="100.0" valueMin="70.0">70–100m</quantity>
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/
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<geoCoordinate id="22A4506DFFDC0218FCAC4DFFFC3F215F" box="[809,952,1522,1546]" direction="north" orientation="latitude" pageId="2" pageNumber="557" precision="15" value="10.9725">10°58’21’’N</geoCoordinate>
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,
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<geoCoordinate id="22A4506DFFDC0218FC434DFFFBDD215F" box="[966,1114,1522,1546]" direction="west" orientation="longitude" pageId="2" pageNumber="557" precision="15" value="-84.33917">84°20’21’’W</geoCoordinate>
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/ leg. Schuh
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<date id="332E106AFFDC0218FB774DFFFA16215F" box="[1266,1425,1521,1546]" pageId="2" pageNumber="557" value="2005-06-07" valueMax="2005-06-11" valueMin="2005-06-07">7–11 VI 2005</date>
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” (pr) // CEHI.
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</paragraph>
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</subSubSection>
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<subSubSection id="0F8A6521FFDC0218FF424E37FEC121CF" pageId="2" pageNumber="557" type="description">
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||
<paragraph id="472F36AAFFDC0218FF424E37FEC121CF" blockId="2.[151,1437,1197,1870]" pageId="2" pageNumber="557">
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||
<emphasis id="75E4EAB8FFDC0218FF424E37FEFA2104" bold="true" box="[199,381,1594,1617]" pageId="2" pageNumber="557">Measurements.</emphasis>
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<typeStatus id="982B8808FFDC0218FE014E37FE7F2104" box="[388,504,1594,1617]" pageId="2" pageNumber="557" type="lectotype">Lectotype</typeStatus>
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. Length
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<quantity id="80689B4FFFDC0218FDDC4E37FD2D2107" box="[601,682,1594,1618]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="557" unit="mm" value="8.2">8.2mm</quantity>
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, head length / width 28/22; pronotum l/w 34/60; length of antennal segments I/II/III/IV = 23/
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<date id="332E106AFFDC0218FE324E50FD9E2123" box="[439,537,1629,1654]" pageId="2" pageNumber="557" value="2013-11-18">13/18/11</date>
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; ratio length of antennae / width of head 2.95; width of abdomen across deltg IV and V
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<quantity id="80689B4FFFDC0218FF644E8FFEC521CF" box="[225,322,1666,1690]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" pageId="2" pageNumber="557" unit="mm" value="3.55">3.55mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="0F8A6521FFDC0218FF424EA8FD5221B7" pageId="2" pageNumber="557" type="distribution">
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<paragraph id="472F36AAFFDC0218FF424EA8FD5221B7" blockId="2.[151,1437,1197,1870]" pageId="2" pageNumber="557">
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<emphasis id="75E4EAB8FFDC0218FF424EA8FEE721EB" bold="true" box="[199,352,1701,1726]" pageId="2" pageNumber="557">Distribution.</emphasis>
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The so far available record data reaching from
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<collectingCountry id="3F87763AFFDC0218FC054EABFC5E21E8" box="[896,985,1702,1725]" name="Panama" pageId="2" pageNumber="557">Panama</collectingCountry>
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to
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<collectingCountry id="3F87763AFFDC0218FC7A4EABFBFD21EB" box="[1023,1146,1701,1726]" name="Costa Rica" pageId="2" pageNumber="557">Costa Rica</collectingCountry>
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and
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||
<collectingCountry id="3F87763AFFDC0218FB364EA8FAA121EB" box="[1203,1318,1701,1726]" name="Nicaragua" pageId="2" pageNumber="557">Nicaragua</collectingCountry>
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indicate a distribution restricted to Central
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<collectingCountry id="3F87763AFFDC0218FD814EC4FDEF21B7" box="[516,616,1737,1762]" name="United States of America" pageId="2" pageNumber="557">America</collectingCountry>
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. (
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<figureCitation id="DFAB2A2FFFDC0218FD064EC4FD4C21B7" box="[643,715,1737,1762]" captionStart="FIGURE 14" captionStartId="6.[151,250,1654,1675]" captionTargetBox="[160,1428,528,1616]" captionTargetId="figure@6.[151,1435,519,1632]" captionTargetPageId="6" captionText="FIGURE 14. Distribution of Helenus species in Northern South and Central America." httpUri="https://zenodo.org/record/255983/files/figure.png" pageId="2" pageNumber="557">Fig.14</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="0F8A6521FFDC021EFF424EE0FEE22008" lastPageId="4" lastPageNumber="559" pageId="2" pageNumber="557" type="discussion">
|
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<paragraph id="472F36AAFFDC0218FF424EE0FCE82018" blockId="2.[151,1437,1197,1870]" pageId="2" pageNumber="557">
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<emphasis id="75E4EAB8FFDC0218FF424EE0FECB2053" bold="true" box="[199,332,1773,1798]" pageId="2" pageNumber="557">Discussion.</emphasis>
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Size, colouration and habitus of
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<taxonomicName id="80904D29FFDC0218FD684EE0FCC02053" box="[749,839,1773,1798]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="hirsutus">
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<emphasis id="75E4EAB8FFDC0218FD684EE0FCC02053" box="[749,839,1773,1798]" italics="true" pageId="2" pageNumber="557">hirsutus</emphasis>
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</taxonomicName>
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is similar to
|
||
<taxonomicName id="80904D29FFDC0218FC7C4EE0FBE92053" box="[1017,1134,1773,1798]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="hesiformis">
|
||
<emphasis id="75E4EAB8FFDC0218FC7C4EE0FBE92053" box="[1017,1134,1773,1798]" italics="true" pageId="2" pageNumber="557">hesiformis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="80904D29FFDC0218FB314EE0FA942053" box="[1204,1299,1773,1798]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="wachteli" status="sp. nov.">
|
||
<emphasis id="75E4EAB8FFDC0218FB314EE0FA942053" box="[1204,1299,1773,1798]" italics="true" pageId="2" pageNumber="557">wachteli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6ED757C3FFDC0218FAA74EE3FAEE2050" box="[1314,1385,1774,1797]" pageId="2" pageNumber="557" rank="species">
|
||
<emphasis id="75E4EAB8FFDC0218FAA74EE3FAEE2050" bold="true" box="[1314,1385,1774,1797]" pageId="2" pageNumber="557">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
but characterized and distinguished from them both by the less pilose body, abdominal margins with produced peangles of deltg II–VI and the shape of pygophore and parameres.
|
||
</paragraph>
|
||
<caption id="13EF6622FFDD0219FF124F9FFAB02092" httpUri="https://zenodo.org/record/255980/files/figure.png" pageId="3" pageNumber="558" targetBox="[188,1425,245,1897]" targetPageId="3">
|
||
<paragraph id="472F36AAFFDD0219FF124F9FFAB02092" blockId="3.[151,1436,1938,1991]" pageId="3" pageNumber="558">
|
||
<emphasis id="75E4EAB8FFDD0219FF124F9FFE9D20F2" bold="true" box="[151,282,1938,1959]" pageId="3" pageNumber="558">FIGURES 1</emphasis>
|
||
–
|
||
<emphasis id="75E4EAB8FFDD0219FEA34F9EFEBE20FD" bold="true" box="[294,313,1939,1960]" pageId="3" pageNumber="558">5.</emphasis>
|
||
<taxonomicName id="80904D29FFDD0219FEBA4F9FFE1520FD" box="[319,402,1938,1960]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="558" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75E4EAB8FFDD0219FEBA4F9FFE1520FD" box="[319,402,1938,1960]" italics="true" pageId="3" pageNumber="558">Helenus</emphasis>
|
||
</taxonomicName>
|
||
spp.:
|
||
<emphasis id="75E4EAB8FFDD0219FE4A4F9FFE5C20F2" bold="true" box="[463,475,1938,1959]" pageId="3" pageNumber="558">1</emphasis>
|
||
—
|
||
<taxonomicName id="80904D29FFDD0219FE704F9FFD3020FD" box="[501,695,1938,1960]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="558" phylum="Arthropoda" rank="species" species="hesiformis">
|
||
<emphasis id="75E4EAB8FFDD0219FE704F9FFD3020FD" box="[501,695,1938,1960]" italics="true" pageId="3" pageNumber="558">Helenus hesiformis</emphasis>
|
||
</taxonomicName>
|
||
, holotype male, dorsal view;
|
||
<emphasis id="75E4EAB8FFDD0219FC644F9FFC6A20F2" bold="true" box="[993,1005,1938,1959]" pageId="3" pageNumber="558">2</emphasis>
|
||
—
|
||
<taxonomicName id="80904D29FFDD0219FB824F9FFB3520FD" box="[1031,1202,1938,1960]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="558" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="75E4EAB8FFDD0219FB824F9FFB3520FD" box="[1031,1202,1938,1960]" italics="true" pageId="3" pageNumber="558">Helenus hirsutus</emphasis>
|
||
</taxonomicName>
|
||
, lectotype male, dorsal view;
|
||
<emphasis id="75E4EAB8FFDD0219FF504FBCFF662093" bold="true" box="[213,225,1969,1990]" pageId="3" pageNumber="558">3</emphasis>
|
||
—labels attached to holotype of
|
||
<taxonomicName id="80904D29FFDD0219FDA94FBCFD122092" box="[556,661,1969,1991]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="558" phylum="Arthropoda" rank="species" species="hesiformis">
|
||
<emphasis id="75E4EAB8FFDD0219FDA94FBCFD122092" box="[556,661,1969,1991]" italics="true" pageId="3" pageNumber="558">hesiformis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="75E4EAB8FFDD0219FD264FBCFD282093" bold="true" box="[675,687,1969,1990]" pageId="3" pageNumber="558">4</emphasis>
|
||
—ditto, reverse side;
|
||
<emphasis id="75E4EAB8FFDD0219FC0C4FBCFC112093" bold="true" box="[905,918,1969,1990]" pageId="3" pageNumber="558">5</emphasis>
|
||
—labels attached to holotype of
|
||
<taxonomicName id="80904D29FFDD0219FB5A4FBCFAB62092" box="[1247,1329,1969,1991]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="558" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="75E4EAB8FFDD0219FB5A4FBCFAB62092" box="[1247,1329,1969,1991]" italics="true" pageId="3" pageNumber="558">hirsutus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="13EF6622FFDA021EFF124EABFBCF218E" httpUri="https://zenodo.org/record/255981/files/figure.png" pageId="4" pageNumber="559" targetBox="[158,1410,205,1663]" targetPageId="4">
|
||
<paragraph id="472F36AAFFDA021EFF124EABFBCF218E" blockId="4.[151,1436,1702,1755]" pageId="4" pageNumber="559">
|
||
<emphasis id="75E4EAB8FFDA021EFF124EABFE9C21E9" bold="true" box="[151,283,1702,1724]" pageId="4" pageNumber="559">FIGURES 6</emphasis>
|
||
–
|
||
<emphasis id="75E4EAB8FFDA021EFEA24EAAFEC021E9" bold="true" box="[295,327,1703,1724]" pageId="4" pageNumber="559">10.</emphasis>
|
||
<taxonomicName id="80904D29FFDA021EFECB4EABFE2621E9" box="[334,417,1702,1724]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="559" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75E4EAB8FFDA021EFECB4EABFE2621E9" box="[334,417,1702,1724]" italics="true" pageId="4" pageNumber="559">Helenus</emphasis>
|
||
</taxonomicName>
|
||
spp.:
|
||
<emphasis id="75E4EAB8FFDA021EFE5B4EAAFE6C21E9" bold="true" box="[478,491,1703,1724]" pageId="4" pageNumber="559">6</emphasis>
|
||
—
|
||
<taxonomicName id="80904D29FFDA021EFD814EABFD3321E9" box="[516,692,1702,1724]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="559" phylum="Arthropoda" rank="species" species="wachteli" status="sp. nov.">
|
||
<emphasis id="75E4EAB8FFDA021EFD814EABFD3321E9" box="[516,692,1702,1724]" italics="true" pageId="4" pageNumber="559">Helenus wachteli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75E4EAB8FFDA021EFD3E4EAAFD7321E9" bold="true" box="[699,756,1703,1724]" pageId="4" pageNumber="559">
|
||
<taxonomicNameLabel id="6ED757C3FFDA021EFD3E4EAAFD7321E9" box="[699,756,1703,1724]" pageId="4" pageNumber="559" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male, dorsal view;
|
||
<emphasis id="75E4EAB8FFDA021EFBA74EAAFBA921E9" bold="true" box="[1058,1070,1703,1724]" pageId="4" pageNumber="559">7</emphasis>
|
||
—
|
||
<taxonomicName id="80904D29FFDA021EFBCD4EABFB7F21E9" box="[1096,1272,1702,1724]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="559" phylum="Arthropoda" rank="species" species="wachteli" status="sp. nov.">
|
||
<emphasis id="75E4EAB8FFDA021EFBCD4EABFB7F21E9" box="[1096,1272,1702,1724]" italics="true" pageId="4" pageNumber="559">Helenus wachteli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75E4EAB8FFDA021EFB7B4EAAFABF21E9" bold="true" box="[1278,1336,1703,1724]" pageId="4" pageNumber="559">
|
||
<taxonomicNameLabel id="6ED757C3FFDA021EFB7B4EAAFABF21E9" box="[1278,1336,1703,1724]" pageId="4" pageNumber="559" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype female, dorsal view;
|
||
<emphasis id="75E4EAB8FFDA021EFEEE4EC8FEFF218F" bold="true" box="[363,376,1733,1754]" pageId="4" pageNumber="559">8</emphasis>
|
||
–
|
||
<emphasis id="75E4EAB8FFDA021EFE014EC8FE1A218F" bold="true" box="[388,413,1733,1754]" pageId="4" pageNumber="559">10</emphasis>
|
||
. Male pygophore:
|
||
<emphasis id="75E4EAB8FFDA021EFDDB4EC8FDED218F" bold="true" box="[606,618,1733,1754]" pageId="4" pageNumber="559">8</emphasis>
|
||
—
|
||
<taxonomicName id="80904D29FFDA021EFD014EC8FD69218E" box="[644,750,1733,1755]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="559" phylum="Arthropoda" rank="species" species="hesiformis">
|
||
<emphasis id="75E4EAB8FFDA021EFD014EC8FD69218E" box="[644,750,1733,1755]" italics="true" pageId="4" pageNumber="559">hesiformis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="75E4EAB8FFDA021EFD7E4EC8FC8F218F" bold="true" box="[763,776,1733,1754]" pageId="4" pageNumber="559">9</emphasis>
|
||
—
|
||
<taxonomicName id="80904D29FFDA021EFCA44EC8FCF4218E" box="[801,883,1733,1755]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="559" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="75E4EAB8FFDA021EFCA44EC8FCF4218E" box="[801,883,1733,1755]" italics="true" pageId="4" pageNumber="559">hirsutus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="75E4EAB8FFDA021EFC054EC8FC1E218F" bold="true" box="[896,921,1733,1754]" pageId="4" pageNumber="559">10</emphasis>
|
||
—
|
||
<taxonomicName id="80904D29FFDA021EFC364EC8FB8E218E" box="[947,1033,1733,1755]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="559" phylum="Arthropoda" rank="species" species="wachteli" status="sp. nov.">
|
||
<emphasis id="75E4EAB8FFDA021EFC364EC8FB8E218E" box="[947,1033,1733,1755]" italics="true" pageId="4" pageNumber="559">wachteli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6ED757C3FFDA021EFB8A4ECBFBCF218E" box="[1039,1096,1734,1755]" pageId="4" pageNumber="559" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="472F36AAFFDA021EFF124F2DFEE22008" blockId="4.[151,430,1824,1885]" pageId="4" pageNumber="559">
|
||
<taxonomicName id="80904D29FFDA021EFF124F2DFEEF206F" box="[151,360,1824,1850]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="559" phylum="Arthropoda" rank="species" species="wachteli" status="sp. nov.">
|
||
<emphasis id="75E4EAB8FFDA021EFF124F2DFEEF206F" bold="true" box="[151,360,1824,1850]" italics="true" pageId="4" pageNumber="559">Helenus wachteli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75E4EAB8FFDA021EFEEA4F2EFE29206F" bold="true" box="[367,430,1827,1850]" pageId="4" pageNumber="559">
|
||
<taxonomicNameLabel id="6ED757C3FFDA021EFEEA4F2EFE29206F" box="[367,430,1827,1850]" pageId="4" pageNumber="559" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="DFAB2A2FFFDA021EFF1A4F49FEB32009" box="[159,308,1860,1885]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1702,1723]" captionTargetBox="[158,1410,205,1663]" captionTargetId="figure@4.[152,1435,193,1681]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Helenus spp.: 6 — Helenus wachteli n. sp., holotype male, dorsal view; 7 — Helenus wachteli n. sp., paratype female, dorsal view; 8 – 10. Male pygophore: 8 — hesiformis; 9 — hirsutus; 10 — wachteli n. sp." httpUri="https://zenodo.org/record/255981/files/figure.png" pageId="4" pageNumber="559">Figs. 6, 7, 10</figureCitation>
|
||
,
|
||
<figureCitation id="DFAB2A2FFFDA021EFEC54F49FEDB2008" box="[320,348,1860,1885]" captionStart="FIGURES 11 — 13" captionStartId="5.[151,264,1300,1321]" captionTargetBox="[278,1308,689,1274]" captionTargetId="figure@5.[270,1316,681,1278]" captionTargetPageId="5" captionText="FIGURES 11 — 13. Left paramere of Hesus spp.: 11 — hesiformis; 12 — hirsutus; 13 — wachteli n. sp." httpUri="https://zenodo.org/record/255982/files/figure.png" pageId="4" pageNumber="559">13</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F8A6521FFDA021FFF124F81FE6125D0" lastPageId="5" lastPageNumber="560" pageId="4" pageNumber="559" type="materials_examined">
|
||
<paragraph id="472F36AAFFDA021EFF124F81FA1B20B9" blockId="4.[151,1437,1932,2029]" pageId="4" pageNumber="559">
|
||
<emphasis id="75E4EAB8FFDA021EFF124F81FE7B20F1" bold="true" box="[151,508,1932,1957]" pageId="4" pageNumber="559">
|
||
Material examined:
|
||
<typeStatus id="982B8808FFDA021EFE0A4F80FE7B20F1" box="[399,508,1933,1956]" pageId="4" pageNumber="559" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
♂ glued on card, labeled: “
|
||
<collectingCountry id="3F87763AFFDA021EFCC44F81FCF120F1" box="[833,886,1932,1956]" name="Peru" pageId="4" pageNumber="559">Peru</collectingCountry>
|
||
Dep. Huanaco / Rio Llullapichis, Estac. / Biol. Panguana
|
||
<quantity id="80689B4FFFDA021EFE8A4FBDFEC8209D" box="[271,335,1968,1992]" metricMagnitude="2" metricUnit="m" metricValue="2.33" pageId="4" pageNumber="559" unit="m" value="233.0">233m</quantity>
|
||
/ 9°
|
||
<geoCoordinate id="22A4506DFFDA021EFE064FBDFE58209D" box="[387,479,1968,1992]" direction="south" orientation="latitude" pageId="4" pageNumber="559" precision="55" value="-36.823">36.823S</geoCoordinate>
|
||
–74°
|
||
<geoCoordinate id="22A4506DFFDA021EFD914FBDFD07209D" box="[532,640,1968,1992]" direction="west" orientation="longitude" pageId="4" pageNumber="559" precision="55" value="-56.127">56,127W</geoCoordinate>
|
||
/
|
||
<date id="332E106AFFDA021EFD1A4FBDFCC4209D" box="[671,835,1968,1992]" pageId="4" pageNumber="559" value="2010-11-01" valueMax="2010-11-24" valueMin="2010-11-01">1–24 XI 2010</date>
|
||
F.Wachtel” (pr) // “
|
||
<typeStatus id="982B8808FFDA021EFBB44FA2FB22209D" box="[1073,1189,1967,1992]" pageId="4" pageNumber="559" type="holotype">Holotypes</typeStatus>
|
||
resulting from / this material are / property of
|
||
<collectingCountry id="3F87763AFFDA021EFE334FD8FE6E20B9" box="[438,489,2005,2028]" name="Peru" pageId="4" pageNumber="559">Peru</collectingCountry>
|
||
and /
|
||
<typeStatus id="982B8808FFDA021EFDAB4FD8FD1C20B9" box="[558,667,2005,2028]" pageId="4" pageNumber="559" type="paratype">Paratypes</typeStatus>
|
||
ZSM - / Collection only! (pr) // “
|
||
<typeStatus id="982B8808FFDA021EFB974FD9FBFB20B8" box="[1042,1148,2004,2029]" pageId="4" pageNumber="559" type="holotype">Holotype</typeStatus>
|
||
♂ /
|
||
<taxonomicName id="80904D29FFDA021EFB2B4FD9FA8D20B8" box="[1198,1290,2004,2029]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="561" phylum="Arthropoda" rank="genus">Helenus</taxonomicName>
|
||
/
|
||
<taxonomicName id="80904D29FFDA021EFA9A4FD9FAF920B8" box="[1311,1406,2004,2029]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="559" phylum="Arthropoda" rank="species" species="wachteli" status="n.">wachteli</taxonomicName>
|
||
<emphasis id="75E4EAB8FFDA021EFA004FD8FA1B20B9" bold="true" box="[1413,1436,2005,2028]" pageId="4" pageNumber="559">
|
||
<taxonomicNameLabel id="6ED757C3FFDA021EFA004FD8FA1B20B9" box="[1413,1436,2005,2028]" pageId="4" pageNumber="559">n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFDB021FFF124895FE242615" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="560">
|
||
<emphasis id="75E4EAB8FFDB021FFF124895FF3F27FA" bold="true" box="[151,184,152,175]" pageId="5" pageNumber="560">sp.</emphasis>
|
||
/ des. E.Heiss 2015” (pr, red label) // deposited at the "Museo de Historia Natural Javier Prado of the University San Marcos in Lima (Universidad Nacional Mayor de San Marcos)".
|
||
<emphasis id="75E4EAB8FFDB021FFC1948B0FB9A2781" bold="true" box="[924,1053,189,212]" pageId="5" pageNumber="560">
|
||
<typeStatus id="982B8808FFDB021FFC1948B0FB9F2781" box="[924,1048,189,212]" pageId="5" pageNumber="560" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
1♀ “
|
||
<collectingCountry id="3F87763AFFDB021FFBE548B0FB112781" box="[1120,1174,189,212]" name="Peru" pageId="5" pageNumber="560">Peru</collectingCountry>
|
||
, Huanaco / Yuyapichis, Panguana /
|
||
<geoCoordinate id="22A4506DFFDB021FFEA548EDFEEF27AD" box="[288,360,224,248]" direction="south" orientation="latitude" pageId="5" pageNumber="560" precision="925" value="-9.616667">9°37’S</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="22A4506DFFDB021FFEFF48EDFE6427AD" box="[378,483,224,248]" direction="west" orientation="longitude" pageId="5" pageNumber="560" precision="925" value="-74.933334">74°56’W</geoCoordinate>
|
||
,
|
||
<quantity id="80689B4FFFDB021FFE7748EDFDB527AD" box="[498,562,224,248]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="560" unit="m" value="220.0">220m</quantity>
|
||
/
|
||
<date id="332E106AFFDB021FFDCB48EDFC9A27AD" box="[590,797,224,248]" pageId="5" pageNumber="560" value="2004-09-21" valueMax="2004-10-04" valueMin="2004-09-21">21 IX–04 X 2004</date>
|
||
/ leg. Schönitzer / Kothe & Breitsameter “ (pr) // also deposited at "Museo de Historia Natural Javier Prado of the University San Marcos in Lima (Universidad Nacional Mayor de San Marcos)"
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFDB021FFF424941FD3E26DD" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="560">
|
||
<specimenCount id="5196FD23FFDB021FFF424941FF6D2630" box="[199,234,332,357]" pageId="5" pageNumber="560" type="generic">1♂</specimenCount>
|
||
“
|
||
<collectingCountry id="3F87763AFFDB021FFE854941FEB02631" box="[256,311,332,356]" name="Peru" pageId="5" pageNumber="560">Peru</collectingCountry>
|
||
, Panguana, Dept. Huanaco / Rio Llullapichis, Nebenfluß des / Rio Pachitea,
|
||
<geoCoordinate id="22A4506DFFDB021FFB474941FA942631" box="[1218,1299,332,356]" direction="south" orientation="latitude" pageId="5" pageNumber="560" precision="925" value="-9.616667">9°37’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22A4506DFFDB021FFAA74941FA0D2631" box="[1314,1418,332,356]" direction="west" orientation="longitude" pageId="5" pageNumber="560" precision="925" value="-74.933334">74°56’W</geoCoordinate>
|
||
/
|
||
<quantity id="80689B4FFFDB021FFF12497DFF5E26DD" box="[151,217,368,392]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="560" unit="m" value="260.0">260m</quantity>
|
||
,
|
||
<date id="332E106AFFDB021FFF60497DFE2F26DD" box="[229,424,368,392]" pageId="5" pageNumber="560" value="2003-04-06" valueMax="2003-04-17" valueMin="2003-04-06">6 IV–17 IV 2003</date>
|
||
, leg. Diller” (pr) // ZSM
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFDB021FFF424999FCD326F9" blockId="5.[151,1437,151,645]" box="[199,852,404,429]" pageId="5" pageNumber="560">
|
||
<specimenCount id="5196FD23FFDB021FFF424999FF6D26F8" box="[199,234,404,429]" pageId="5" pageNumber="560" type="generic">1♂</specimenCount>
|
||
“
|
||
<collectingCountry id="3F87763AFFDB021FFF794999FEB626F9" box="[252,305,404,428]" name="Peru" pageId="5" pageNumber="560">Peru</collectingCountry>
|
||
/ Iquitos Amazonas /
|
||
<date id="332E106AFFDB021FFDA24999FDF826F9" box="[551,639,404,428]" pageId="5" pageNumber="560" value="1977-02-11">11 II 77</date>
|
||
Heiss (pr) // CEHI
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFDB021FFF4249BAFF6126A1" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="560">
|
||
1♀ “
|
||
<collectingCountry id="3F87763AFFDB021FFE8149B5FEBF269A" box="[260,312,440,463]" name="Peru" pageId="5" pageNumber="560">Peru</collectingCountry>
|
||
Dep. Loreto / Porvenir W Iquitos / 65981 / 9569651 / P5, V
|
||
<geoCoordinate id="22A4506DFFDB021FFC5B49B5FB8B2685" box="[990,1036,440,464]" pageId="5" pageNumber="560">
|
||
5
|
||
<date id="332E106AFFDB021FFC7549B5FB8B2685" box="[1008,1036,440,464]" pageId="5" pageNumber="560" value="2011-07-10">10</date>
|
||
</geoCoordinate>
|
||
VII 2011 / Greg Lamarre lg.” (pr) / / CEHI
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFDB021FFF4249F2FDCD2569" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="560">
|
||
<specimenCount id="5196FD23FFDB021FFF4249F2FF6D254D" box="[199,234,511,536]" pageId="5" pageNumber="560" type="generic">1♂</specimenCount>
|
||
“
|
||
<collectingCountry id="3F87763AFFDB021FFE8849F2FEE5254D" box="[269,354,511,536]" name="Bolivia" pageId="5" pageNumber="560">Bolivia</collectingCountry>
|
||
, Cochabamba Prov. /
|
||
<quantity id="80689B4FFFDB021FFDE249F2FD20254D" box="[615,679,511,536]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="560" unit="km" value="20.0">20km</quantity>
|
||
SW Villa Tunari / Rio Avispha:
|
||
<quantity id="80689B4FFFDB021FFBB64A0DFBF4254D" box="[1075,1139,512,536]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="560" unit="m" value="460.0">460m</quantity>
|
||
<date id="332E106AFFDB021FFBFB4A0DFB7F254D" box="[1150,1272,512,536]" pageId="5" pageNumber="560" value="2000-09-06">6 IX 2000</date>
|
||
/
|
||
<geoCoordinate id="22A4506DFFDB021FFA934A0DFA1F254D" box="[1302,1432,512,536]" direction="south" orientation="latitude" pageId="5" pageNumber="560" precision="9" value="-17.024834">17°01.49’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22A4506DFFDB021FFF124A29FEA62569" box="[151,289,548,572]" direction="west" orientation="longitude" pageId="5" pageNumber="560" precision="9" value="-65.527664">65°31.66’W</geoCoordinate>
|
||
/ M.Hauser” (pr) // CEHI.
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFDB021FFF424A4AFE6125D0" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="560">
|
||
All
|
||
<typeStatus id="982B8808FFDB021FFF714A45FEE7250A" box="[244,352,584,607]" pageId="5" pageNumber="560" type="paratype">paratypes</typeStatus>
|
||
are designated and labeled accordingly: “
|
||
<typeStatus id="982B8808FFDB021FFCC34A45FC2E2535" box="[838,937,584,608]" pageId="5" pageNumber="560" type="paratype">Paratype</typeStatus>
|
||
♂ (or ♀) /
|
||
<taxonomicName id="80904D29FFDB021FFBBD4A4AFB132535" box="[1080,1172,583,608]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="561" phylum="Arthropoda" rank="genus">Helenus</taxonomicName>
|
||
/
|
||
<taxonomicName id="80904D29FFDB021FFB2A4A4AFA892535" box="[1199,1294,583,608]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="560" phylum="Arthropoda" rank="species" species="wachteli">wachteli</taxonomicName>
|
||
n.,sp. / des. E.Heiss 2015” (pr., red label).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="13EF6622FFDB021FFF124D19FB15227C" box="[151,1170,1300,1322]" httpUri="https://zenodo.org/record/255982/files/figure.png" pageId="5" pageNumber="560" targetBox="[278,1308,689,1274]" targetPageId="5">
|
||
<paragraph id="472F36AAFFDB021FFF124D19FB15227C" blockId="5.[151,1170,1300,1322]" box="[151,1170,1300,1322]" pageId="5" pageNumber="560">
|
||
<emphasis id="75E4EAB8FFDB021FFF124D19FEA1227C" bold="true" box="[151,294,1300,1321]" pageId="5" pageNumber="560">FIGURES 11</emphasis>
|
||
—
|
||
<emphasis id="75E4EAB8FFDB021FFEBA4D19FED9227C" bold="true" box="[319,350,1300,1321]" pageId="5" pageNumber="560">13.</emphasis>
|
||
Left paramere of
|
||
<taxonomicName id="80904D29FFDB021FFD904D18FDD4227F" box="[533,595,1301,1322]" class="Insecta" family="Aradidae" genus="Hesus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="560" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75E4EAB8FFDB021FFD904D18FDD4227F" box="[533,595,1301,1322]" italics="true" pageId="5" pageNumber="560">Hesus</emphasis>
|
||
</taxonomicName>
|
||
spp.:
|
||
<emphasis id="75E4EAB8FFDB021FFD154D19FD21227C" bold="true" box="[656,678,1300,1321]" pageId="5" pageNumber="560">11</emphasis>
|
||
—
|
||
<taxonomicName id="80904D29FFDB021FFD444D19FCAD227F" box="[705,810,1300,1322]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="560" phylum="Arthropoda" rank="species" species="hesiformis">
|
||
<emphasis id="75E4EAB8FFDB021FFD444D19FCAD227F" box="[705,810,1300,1322]" italics="true" pageId="5" pageNumber="560">hesiformis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="75E4EAB8FFDB021FFCBD4D19FCD6227C" bold="true" box="[824,849,1300,1321]" pageId="5" pageNumber="560">12</emphasis>
|
||
—
|
||
<taxonomicName id="80904D29FFDB021FFCEE4D19FC3A227F" box="[875,957,1300,1322]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="560" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="75E4EAB8FFDB021FFCEE4D19FC3A227F" box="[875,957,1300,1322]" italics="true" pageId="5" pageNumber="560">hirsutus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="75E4EAB8FFDB021FFC4F4D19FC64227C" bold="true" box="[970,995,1300,1321]" pageId="5" pageNumber="560">13</emphasis>
|
||
—
|
||
<taxonomicName id="80904D29FFDB021FFC794D19FBD5227F" box="[1020,1106,1300,1322]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="560" phylum="Arthropoda" rank="species" species="wachteli" status="sp. nov.">
|
||
<emphasis id="75E4EAB8FFDB021FFC794D19FBD5227F" box="[1020,1106,1300,1322]" italics="true" pageId="5" pageNumber="560">wachteli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6ED757C3FFDB021FFBDC4D19FB15227C" box="[1113,1170,1300,1321]" pageId="5" pageNumber="560" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0F8A6521FFDB021FFF424D5CFC6022D8" pageId="5" pageNumber="560" type="diagnosis">
|
||
<paragraph id="472F36AAFFDB021FFF424D5CFC6022D8" blockId="5.[151,1436,1360,2034]" pageId="5" pageNumber="560">
|
||
<emphasis id="75E4EAB8FFDB021FFF424D5CFEC6223F" bold="true" box="[199,321,1361,1386]" pageId="5" pageNumber="560">Diagnosis.</emphasis>
|
||
Shares the rather dense pilosity with
|
||
<taxonomicName id="80904D29FFDB021FFD694D5DFCFC223C" box="[748,891,1360,1385]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="560" phylum="Arthropoda" rank="species" species="hesiformis">
|
||
<emphasis id="75E4EAB8FFDB021FFD694D5DFCFC223C" box="[748,891,1360,1385]" italics="true" pageId="5" pageNumber="560">H.hesiformis</emphasis>
|
||
</taxonomicName>
|
||
but the male genitalia are distinct.
|
||
<taxonomicName id="80904D29FFDB021FFA894D5DFA05223C" box="[1292,1410,1360,1385]" class="Malacostraca" family="Portunidae" genus="Helenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="560" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="75E4EAB8FFDB021FFA894D5DFA05223C" box="[1292,1410,1360,1385]" italics="true" pageId="5" pageNumber="560">H.hirsutus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished by protruding pe-angles of deltg II–VI and different genitalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F8A6521FFDB021CFF424D94FDF52615" lastPageId="6" lastPageNumber="561" pageId="5" pageNumber="560" type="description">
|
||
<paragraph id="472F36AAFFDB021FFF424D94FDE62280" blockId="5.[151,1436,1360,2034]" pageId="5" pageNumber="560">
|
||
<emphasis id="75E4EAB8FFDB021FFF424D94FEDE22E7" bold="true" box="[199,345,1433,1458]" pageId="5" pageNumber="560">Description.</emphasis>
|
||
Body surface with dispersed yellowish double pilosity; colouration reddish-brown, tibiae with a fading yellowish median and apical ring.
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFDB021FFF424DEFFE9E21DF" blockId="5.[151,1436,1360,2034]" pageId="5" pageNumber="560">
|
||
<emphasis id="75E4EAB8FFDB021FFF424DEFFE8922AC" bold="true" box="[199,270,1506,1529]" pageId="5" pageNumber="560">Head.</emphasis>
|
||
Distinctly longer than wide (27/24); genae slightly protruding over clypeus, anterolaterally rounded and diverging; antenniferous lobes subparallel, apices recurved, shorter than clypeus; antennae about
|
||
<date id="332E106AFFDB021FFA854E08FA9B2148" box="[1280,1308,1541,1565]" pageId="5" pageNumber="560">3x</date>
|
||
as long as width of head (71/24); length of antennal segments I/II/III/IV = 26/
|
||
<date id="332E106AFFDB021FFC0C4E24FC6A2117" box="[905,1005,1577,1602]" pageId="5" pageNumber="560" value="2013-12-20">13/20/12</date>
|
||
; eyes oval inserted in head; postocular lobes slightly rounded and converging posteriorly to constricted collar; rostrum as long as head, arising from a slitlike atrium.
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFDB021FFF424E98FC6521A0" blockId="5.[151,1436,1360,2034]" pageId="5" pageNumber="560">
|
||
<emphasis id="75E4EAB8FFDB021FFF424E98FEC021F9" bold="true" box="[199,327,1685,1708]" pageId="5" pageNumber="560">Pronotum.</emphasis>
|
||
About 1.8x as long as wide (63/35); lateral margins sinuate converging anteriorly to the distinct ring-like collar, humeri subparallel; anterior half of disk with 2(1+1) smooth oval depressions flanked by oblique elevations, separated by a deep transverse suture from raised posterior part.
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFDB021FFF424F0CFF7E206B" blockId="5.[151,1436,1360,2034]" pageId="5" pageNumber="560">
|
||
<emphasis id="75E4EAB8FFDB021FFF424F0CFEC1204F" bold="true" box="[199,326,1793,1818]" pageId="5" pageNumber="560">Scutellum.</emphasis>
|
||
Triangular, basal and lateral margins carinate, disk with a longitudinal carina, transversely rugose laterally.
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFDB021FFF424F47FD3920D3" blockId="5.[151,1436,1360,2034]" pageId="5" pageNumber="560">
|
||
<emphasis id="75E4EAB8FFDB021FFF424F47FEC92034" bold="true" box="[199,334,1866,1889]" pageId="5" pageNumber="560">Hemelytra.</emphasis>
|
||
Corium with carinate setose veins nearly reaching posterior margin of deltg III; membrane covering ½ of tergite VII with irregular venation.
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFDB021FFF424F9CFC4820A7" blockId="5.[151,1436,1360,2034]" pageId="5" pageNumber="560">
|
||
<emphasis id="75E4EAB8FFDB021FFF424F9CFEC420FF" bold="true" box="[199,323,1937,1962]" pageId="5" pageNumber="560">Abdomen.</emphasis>
|
||
Tergal plate separated by a carina from deltg II–VII; deltg II+III fused, its fusion line only laterally discernible; lateral margins of deltg II–VI straight, pe-angles rounded and not protruding; spiracles II–V ventral on vltg II–V, far from lateral margin, VI–VIII lateral and visible from above.
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFD8021CFF424895FB2327E5" blockId="6.[151,1437,151,464]" box="[199,1188,151,176]" pageId="6" pageNumber="561">
|
||
<emphasis id="75E4EAB8FFD8021CFF424895FE8127FA" bold="true" box="[199,262,152,175]" pageId="6" pageNumber="561">Legs.</emphasis>
|
||
Femora and tibiae cylindrical, beset with erect setae, claws with thin long pulvilli.
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFD8021CFF4248B0FC2F27AD" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="561">
|
||
<emphasis id="75E4EAB8FFD8021CFF4248B0FEA22781" bold="true" box="[199,293,189,212]" pageId="6" pageNumber="561">Female.</emphasis>
|
||
Basically as male, pe-angles of deltg VI–VII more rounded, posterior margin of tergite VII straight, ptg VIII strongly transverse, segments IX and X produced posteriorly.
|
||
</paragraph>
|
||
<paragraph id="472F36AAFFD8021CFF424908FDF52615" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="561">
|
||
<emphasis id="75E4EAB8FFD8021CFF424908FEFA2649" bold="true" box="[199,381,261,284]" pageId="6" pageNumber="561">Measurements.</emphasis>
|
||
<typeStatus id="982B8808FFD8021CFE004908FE7E2649" box="[389,505,261,284]" pageId="6" pageNumber="561" type="lectotype">Lectotype</typeStatus>
|
||
. Length
|
||
<quantity id="80689B4FFFD8021CFDD94909FD292649" box="[604,686,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="6" pageNumber="561" unit="mm" value="8.9">8.9mm</quantity>
|
||
; length of antennae
|
||
<quantity id="80689B4FFFD8021CFC114909FC722649" box="[916,1013,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" pageId="6" pageNumber="561" unit="mm" value="3.55">3.55mm</quantity>
|
||
; width of abdomen
|
||
<quantity id="80689B4FFFD8021CFB524909FAAD2649" box="[1239,1322,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="561" unit="mm" value="4.1">4.1mm</quantity>
|
||
. Variation of length in
|
||
<typeStatus id="982B8808FFD8021CFE9A4925FE0C266A" box="[287,395,296,319]" pageId="6" pageNumber="561" type="paratype">paratypes</typeStatus>
|
||
(♂, ♀):
|
||
<quantity id="80689B4FFFD8021CFE6E4925FDE92615" box="[491,622,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="8.8" metricValueMin="8.2" pageId="6" pageNumber="561" unit="mm" value="8.5" valueMax="8.8" valueMin="8.2">8.2–8.8mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F8A6521FFD8021CFF424940FB2626DD" pageId="6" pageNumber="561" type="etymology">
|
||
<paragraph id="472F36AAFFD8021CFF424940FB2626DD" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="561">
|
||
<emphasis id="75E4EAB8FFD8021CFF424940FEC92631" bold="true" box="[199,334,333,356]" pageId="6" pageNumber="561">Etymology.</emphasis>
|
||
This unexpected species is dedicated to Franz Wachtel, expert coleopterist and collector of the
|
||
<typeStatus id="982B8808FFD8021CFF124962FF7B26DD" box="[151,252,367,392]" pageId="6" pageNumber="561" type="holotype">holotype</typeStatus>
|
||
, recognizing his repeated generous donations of by-catched
|
||
<taxonomicName id="80904D29FFD8021CFC1A4962FB8126DD" box="[927,1030,367,392]" class="Insecta" family="Aradidae" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="561" phylum="Arthropoda" rank="family">Aradidae</taxonomicName>
|
||
to the author.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F8A6521FFD8021CFF424999FEB42685" pageId="6" pageNumber="561" type="distribution">
|
||
<paragraph id="472F36AAFFD8021CFF424999FEB42685" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="561">
|
||
<emphasis id="75E4EAB8FFD8021CFF424999FEE726F8" bold="true" box="[199,352,404,429]" pageId="6" pageNumber="561">Distribution.</emphasis>
|
||
All records are from an area east of the Andes mountain range in the lowlands of Amazon basin. (Map
|
||
<figureCitation id="DFAB2A2FFFD8021CFF5E49BAFEAC2685" box="[219,299,439,464]" captionStart="FIGURE 14" captionStartId="6.[151,250,1654,1675]" captionTargetBox="[160,1428,528,1616]" captionTargetId="figure@6.[151,1435,519,1632]" captionTargetPageId="6" captionText="FIGURE 14. Distribution of Helenus species in Northern South and Central America." httpUri="https://zenodo.org/record/255983/files/figure.png" pageId="6" pageNumber="561">Fig. 14</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |