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<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.e84187" ID-Pensoft-Pub="2747-8106-96-179" ID-Pensoft-UUID="FE0861D71BB454999EE6637C0D9B2B0C" ID-ZooBank="35B619086E6548B09A177281C2253391" ModsDocID="2747-8106-96-179" checkinTime="1670952513976" checkinUser="pensoft" docAuthor="Gruendel, Joachim, Keupp, Helmut, Lang, Fritz &amp; Nuetzel, Alexander" docDate="2022" docId="B55A4577058A53E6A87D117770034B5F" docLanguage="en" docName="Zitteliana 96: 179-221" docOrigin="Zitteliana 96" docPubDate="2022-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.96.e84187" docTitle="Ptygmatis clio" docType="treatment" docVersion="1" id="FE0861D71BB454999EE6637C0D9B2B0C" lastPageNumber="179" masterDocId="FE0861D71BB454999EE6637C0D9B2B0C" masterDocTitle="Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany)" masterLastPageNumber="221" masterPageNumber="179" pageNumber="179" updateTime="1670952513976" updateUser="pensoft">
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<mods:title>Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gruendel, Joachim</mods:namePart>
<mods:affiliation>Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
<mods:nameIdentifier type="email">joachim.gruendel@lingua-pura.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Keupp, Helmut</mods:namePart>
<mods:affiliation>Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lang, Fritz</mods:namePart>
<mods:affiliation>Drosselweg 16, 96114 Hirschaid, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nuetzel, Alexander</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8852-7688</mods:nameIdentifier>
<mods:affiliation>SNSB-Bayerische Staatssammlung fuer Palaeontologie und Geologie, Richard-Wagner-Str. 10, 80333 Muenchen, Germany &amp; Department of Earth and Environmental Sciences, Paleontology and Geobiology, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zitteliana</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-12</mods:number>
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<mods:detail type="volume">
<mods:number>96</mods:number>
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<mods:start>179</mods:start>
<mods:end>221</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zitteliana.96.e84187</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zitteliana.96.e84187</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:B55A4577058A53E6A87D117770034B5F" httpUri="http://treatment.plazi.org/id/B55A4577058A53E6A87D117770034B5F" lastPageNumber="179" pageId="0" pageNumber="179">
<subSubSection pageId="0" pageNumber="179" type="nomenclature">
<paragraph pageId="0" pageNumber="179">
<taxonomicName LSID="B55A4577-058A-53E6-A87D-117770034B5F" authority="(d'Orbigny, 1852)" baseAuthorityName="d'Orbigny" baseAuthorityYear="1852" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis clio" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="clio">
Ptygmatis clio (
<normalizedToken originalValue="dOrbigny">d'Orbigny</normalizedToken>
, 1852)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="179" type="description">
<paragraph pageId="0" pageNumber="179">
<figureCitation captionStart="Plate 12" captionStartId="F12" captionText="Plate 12. (1 - 3) Ptygmatis clio (d'Orbigny, 1852). (1 - 2) SNSB-BSPG 2021 XV 59, Saal (collection Lang); (1) section of fragment, height 21 mm; (2) detail, height 3.7 mm. (3) SNSB-BSPG 2021 XV 60, Saal (collection Lang), lateral view, height 108 mm. (4) Ptygmatis? polyspira (Quenstedt, 1881 - 1884), holotype, collection University Tuebingen (collection Quenstedt); Nattheim, lateral view, height 32 mm (original to Quenstedt 1881 - 1884, pl. 207, fig. 3. (11 - 12) Bactroptyxis teres (Muenster in Goldfuss, 1844), collection Sauerborn, Nattheim area. (5 - 7) Ptygmatis? ursicina (Thurmann in Thurmann &amp; Etallon, 1861). (5 - 6) SNSB-BSPG 2021 XV 30, Saal (collection Lang); (5) last whorl with aperture, height 4 mm; (6) lateral view, height 9.5 mm. (7) SNSB-BSPG 2021 XV 31, Saal (collection Lang), aperture and ornament of base, width 2.5 mm." figureDoi="10.3897/zitteliana.96.e84187.plate12" httpUri="https://binary.pensoft.net/fig/780390" pageId="0" pageNumber="179">Plate 12: figs 1-3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="179" type="reference_group">
<paragraph pageId="0" pageNumber="179">
<taxonomicName authorityName="d'Orbigny" authorityYear="1850" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea clio" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="clio">*1852 - Nerinea clio</taxonomicName>
<normalizedToken originalValue="dOrbigny">d'Orbigny</normalizedToken>
-
<normalizedToken originalValue="dOrbigny">d'Orbigny</normalizedToken>
: 139, pl. 275, figs 3-5.
</paragraph>
<paragraph pageId="0" pageNumber="179">
<taxonomicName baseAuthorityName="d'Orbigny" baseAuthorityYear="1852" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis clio" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="clio">1886-1888 - Ptygmatis clio</taxonomicName>
<normalizedToken originalValue="dOrbigny">d'Orbigny</normalizedToken>
- Loriol in Loriol and Bourgeat: 84, pl. 7, figs 1-3.
</paragraph>
<paragraph pageId="0" pageNumber="179">
<taxonomicName baseAuthorityName="d'Orbigny" baseAuthorityYear="1852" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis clio" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="clio">1898 - Ptygmatis clio</taxonomicName>
<normalizedToken originalValue="dOrbigny">d'Orbigny</normalizedToken>
- Cossmann: 72, pl. 6, figs 18, 19.
</paragraph>
<paragraph pageId="0" pageNumber="179">
<taxonomicName class="Gastropoda" family="Ptygmatididae" genus="Polyptyxisella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Polyptyxisella clio" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="clio">1997 - Polyptyxisella clio</taxonomicName>
(
<normalizedToken originalValue="dOrbigny">d'Orbigny</normalizedToken>
, 1852) - Fischer and Weber: 55, pl. 12, figs 9, 10.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="179" type="materials_examined">
<paragraph pageId="0" pageNumber="179">Material.</paragraph>
<paragraph pageId="0" pageNumber="179">Two illustrated specimens (SNSB-BSPG 2021 XV 59, 60) and a questionable one, all from Saal, collection Lang.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="179" type="description">
<paragraph pageId="0" pageNumber="179">Description.</paragraph>
<paragraph pageId="0" pageNumber="179">
The largest specimen (Plate
<figureCitation captionStart="Plate 12" captionStartId="F12" captionText="Plate 12. (1 - 3) Ptygmatis clio (d'Orbigny, 1852). (1 - 2) SNSB-BSPG 2021 XV 59, Saal (collection Lang); (1) section of fragment, height 21 mm; (2) detail, height 3.7 mm. (3) SNSB-BSPG 2021 XV 60, Saal (collection Lang), lateral view, height 108 mm. (4) Ptygmatis? polyspira (Quenstedt, 1881 - 1884), holotype, collection University Tuebingen (collection Quenstedt); Nattheim, lateral view, height 32 mm (original to Quenstedt 1881 - 1884, pl. 207, fig. 3. (11 - 12) Bactroptyxis teres (Muenster in Goldfuss, 1844), collection Sauerborn, Nattheim area. (5 - 7) Ptygmatis? ursicina (Thurmann in Thurmann &amp; Etallon, 1861). (5 - 6) SNSB-BSPG 2021 XV 30, Saal (collection Lang); (5) last whorl with aperture, height 4 mm; (6) lateral view, height 9.5 mm. (7) SNSB-BSPG 2021 XV 31, Saal (collection Lang), aperture and ornament of base, width 2.5 mm." figureDoi="10.3897/zitteliana.96.e84187.plate12" httpUri="https://binary.pensoft.net/fig/780390" pageId="0" pageNumber="179">12</figureCitation>
: fig. 3) is 108 mm high. The shell is slender. The whorl face is straight. The last whorl of the largest specimen has a weak, indistinctly delimited subsutural furrow. The sutures are distinct. The whorls lack visible ornament. The basal edge is pronounced and forms an angular transition to the base. The base is incompletely preserved, and is seemingly smooth with a distinct umbilicus. The aperture is not preserved. A columellar section shows the presence of a parietal, two columellar, and one palatal plaits.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="179" type="remarks">
<paragraph pageId="0" pageNumber="179">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="179">
<bibRefCitation author="Fischer, J-C" journalOrPublisher="Masson, Paris" pageId="0" pageNumber="179" refId="B22" refString="Fischer, J-C, Weber, C, 1997. Revision critique de la paleontologie Francaise d'Alcide d'Orbigny. Vol. II: Gastropodes jurassiques. Masson, Paris" title="Revision critique de la paleontologie Francaise d'Alcide d'Orbigny. Vol. II: Gastropodes jurassiques." year="1997">Fischer and Weber (1997)</bibRefCitation>
designated a lectotype (section of a shell,
<bibRefCitation author="Fischer, J-C" journalOrPublisher="Masson, Paris" pageId="0" pageNumber="179" refId="B22" refString="Fischer, J-C, Weber, C, 1997. Revision critique de la paleontologie Francaise d'Alcide d'Orbigny. Vol. II: Gastropodes jurassiques. Masson, Paris" title="Revision critique de la paleontologie Francaise d'Alcide d'Orbigny. Vol. II: Gastropodes jurassiques." year="1997">Fischer and Weber 1997</bibRefCitation>
: pl. 12, fig. 9) that agrees well with the sectioned shell illustrated herein (Plate
<figureCitation captionStart="Plate 12" captionStartId="F12" captionText="Plate 12. (1 - 3) Ptygmatis clio (d'Orbigny, 1852). (1 - 2) SNSB-BSPG 2021 XV 59, Saal (collection Lang); (1) section of fragment, height 21 mm; (2) detail, height 3.7 mm. (3) SNSB-BSPG 2021 XV 60, Saal (collection Lang), lateral view, height 108 mm. (4) Ptygmatis? polyspira (Quenstedt, 1881 - 1884), holotype, collection University Tuebingen (collection Quenstedt); Nattheim, lateral view, height 32 mm (original to Quenstedt 1881 - 1884, pl. 207, fig. 3. (11 - 12) Bactroptyxis teres (Muenster in Goldfuss, 1844), collection Sauerborn, Nattheim area. (5 - 7) Ptygmatis? ursicina (Thurmann in Thurmann &amp; Etallon, 1861). (5 - 6) SNSB-BSPG 2021 XV 30, Saal (collection Lang); (5) last whorl with aperture, height 4 mm; (6) lateral view, height 9.5 mm. (7) SNSB-BSPG 2021 XV 31, Saal (collection Lang), aperture and ornament of base, width 2.5 mm." figureDoi="10.3897/zitteliana.96.e84187.plate12" httpUri="https://binary.pensoft.net/fig/780390" pageId="0" pageNumber="179">12</figureCitation>
: figs 1, 2). The lateral view provided by
<bibRefCitation author="Fischer, J-C" journalOrPublisher="Masson, Paris" pageId="0" pageNumber="179" refId="B22" refString="Fischer, J-C, Weber, C, 1997. Revision critique de la paleontologie Francaise d'Alcide d'Orbigny. Vol. II: Gastropodes jurassiques. Masson, Paris" title="Revision critique de la paleontologie Francaise d'Alcide d'Orbigny. Vol. II: Gastropodes jurassiques." year="1997">Fischer and Weber (1997</bibRefCitation>
: pl. 12, fig. 10) also agrees well with the present material. The specimens illustrated by
<bibRefCitation author="Fischer, J-C" journalOrPublisher="Masson, Paris" pageId="0" pageNumber="179" refId="B22" refString="Fischer, J-C, Weber, C, 1997. Revision critique de la paleontologie Francaise d'Alcide d'Orbigny. Vol. II: Gastropodes jurassiques. Masson, Paris" title="Revision critique de la paleontologie Francaise d'Alcide d'Orbigny. Vol. II: Gastropodes jurassiques." year="1997">Fischer and Weber (1997)</bibRefCitation>
are from the middle Oxfordian of St. Mihiel, but these authors noted that the species is also present in the Kimmeridgian. The illustration given by
<normalizedToken originalValue="dOrbigny">d'Orbigny</normalizedToken>
(1852: pl. 275, figs 3-5) deviates more strongly from our specimens: the shell is slenderer and the whorl face is distinctly concave.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="179" type="relationships">
<paragraph pageId="0" pageNumber="179">Relationships.</paragraph>
<paragraph pageId="0" pageNumber="179">
<taxonomicName baseAuthorityName="Quenstedt" baseAuthorityYear="1858" class="Gastropoda" family="Ptygmatididae" genus="Aptyxiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptyxiella planata" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="planata">
<emphasis italics="true" pageId="0" pageNumber="179">Aptyxiella planata</emphasis>
</taxonomicName>
(Quenstedt) sensu
<bibRefCitation author="Haegele, G" journalOrPublisher="Fossilien, Sonderband" pageId="0" pageNumber="179" pagination="1 - 144" refId="B42" refString="Haegele, G, 1997. Juraschnecken. Fossilien, Sonderband 11: 1 - 144" title="Juraschnecken." volume="11" year="1997">
<normalizedToken originalValue="Hägele">Haegele</normalizedToken>
(1997)
</bibRefCitation>
,
<taxonomicName lsidName="A. quenstedti" pageId="0" pageNumber="179" rank="species" species="quenstedti">
<emphasis italics="true" pageId="0" pageNumber="179">A. quenstedti</emphasis>
</taxonomicName>
Geiger, 1901, and
<taxonomicName lsidName="A. ewaldi" pageId="0" pageNumber="179" rank="species" species="ewaldi">
<emphasis italics="true" pageId="0" pageNumber="179">A. ewaldi</emphasis>
</taxonomicName>
Geiger, 1901 lack plaits.
<taxonomicName class="Gastropoda" family="Eunerineidae" genus="Cossmannea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cossmannea nantuacensis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="nantuacensis">
<emphasis italics="true" pageId="0" pageNumber="179">Cossmannea nantuacensis</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="dOrbigny">d'Orbigny</normalizedToken>
) sensu
<bibRefCitation author="Fischer, J-C" journalOrPublisher="Masson, Paris" pageId="0" pageNumber="179" refId="B22" refString="Fischer, J-C, Weber, C, 1997. Revision critique de la paleontologie Francaise d'Alcide d'Orbigny. Vol. II: Gastropodes jurassiques. Masson, Paris" title="Revision critique de la paleontologie Francaise d'Alcide d'Orbigny. Vol. II: Gastropodes jurassiques." year="1997">Fischer and Weber (1997)</bibRefCitation>
is much larger, lacks an umbilicus and columellar and parietal plaits.
<taxonomicName authorityName="Pchelintsev" authorityYear="1965" class="Gastropoda" family="Nerineidae" genus="Megaptyxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megaptyxis caucasica" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="caucasica">
<emphasis italics="true" pageId="0" pageNumber="179">Megaptyxis caucasica</emphasis>
</taxonomicName>
Pchelintsev, 1965 has a concave whorl face and more oblique sutures.
</paragraph>
</subSubSection>
</treatment>
</document>