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<document id="36F2DDD3ABD98873A0CC5A23E416027B" ID-CLB-Dataset="298191" ID-DOI="10.5252/geodiversitas2024v46a8" ID-GBIF-Dataset="d0b851c0-a666-45e1-bc92-c28f8eb2992c" ID-ISSN="1638-9395" ID-Zenodo-Dep="11501840" ID-ZooBank="urn:lsid:zoobank.org:pub:431B0382-9073-4676-9EC6-B3CB6A15589D" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1717663391679" checkinUser="plazi" docAuthor="Moreau, Jean-David, Vullo, Romain, Bichr, Elsie, Thomas, Jérôme, Gand, Georges, Gagnaison, Cyril, Barrier, Pascal &amp; Néraudeau, Didier" docDate="2024" docId="813387CADE027C00FF5094CFDF2AF83C" docLanguage="en" docName="geodiversitas.2024.46.8.pdf" docOrigin="Geodiversitas 46 (8)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a8.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Eubrontes veillonensis &quot; Lapparent &amp; Montenat 1967" docType="treatment" docVersion="5" lastPageNumber="357" masterDocId="7D0AFFB2DE127C10FFCE915CDA75FFD7" masterDocTitle="Crocodylomorph and dinosaur tracks from the lowermost Jurassic of Le Veillon (western France): ichnotaxonomic revision of the type material (Lapparent collection)" masterLastPageNumber="366" masterPageNumber="343" pageNumber="357" updateTime="1717754607933" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="8B502C5F5DE6D12F7F1DAC121EE63494">Crocodylomorph and dinosaur tracks from the lowermost Jurassic of Le Veillon (western France): ichnotaxonomic revision of the type material (Lapparent collection)</mods:title>
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<mods:namePart id="FF9EBE4FEB08D9D1F458C4365EFDECBB">Moreau, Jean-David</mods:namePart>
<mods:affiliation id="69398F465475578FD7359835BECF8989">Université Paris-Saclay, CNRS, GEOPS, 91405 Orsay (France) and Biogéosciences, UMR 6282 CNRS, Université Bourgogne, 6 boulevard Gabriel, 21000 Dijon (France)</mods:affiliation>
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<mods:namePart id="EB752CE0563871EFF2D7F04F1DDC3BB8">Vullo, Romain</mods:namePart>
<mods:affiliation id="EDBA0FE9267A43A9D22A0B52D9C448EF">Univ Rennes, CNRS, Géosciences Rennes, UMR 6118, 263 avenue du Général Leclerc, F- 35000 Rennes (France)</mods:affiliation>
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<mods:namePart id="7D9ED151F71FF83EBF905ED465BA3FAE">Thomas, Jérôme</mods:namePart>
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<mods:namePart id="065110A063FBF19DF3B399A79BF9C1FB">Gand, Georges</mods:namePart>
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<mods:affiliation id="CFF00C4052C2D059B2B9C5FCF8217567">Institut Polytechnique UniLaSalle Beauvais, Département Géosciences, Unité Bassins-Réservoirs-Ressources (B 2 R U 2 R 7511), UniLaSalle-Université de Picardie Jules VERNE, 19 rue Pierre Waguet boîte postale 30313, F- 60026 Beauvais cedex (France)</mods:affiliation>
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<taxonomicName id="CE9A4D5FDE027C00FF6494C8D898FA7A" authority="&quot; Lapparent &amp; Montenat, 1967" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" box="[170,749,1427,1454]" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="357" phylum="Chordata" rank="species" species="veillonensis">
<emphasis id="3BEEEACEDE027C00FF6494C8DBE2FA79" bold="true" box="[170,407,1428,1454]" italics="true" pageId="16" pageNumber="357">Eubrontes veillonensis</emphasis>
<bibRefCitation id="6D0B4B2DDE027C00FE6694C8D898FA7A" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[424,749,1427,1454]" pageId="16" pageNumber="357" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat, 1967</bibRefCitation>
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(
<figureCitation id="91A12A59DE027C00FE5E94EFDBB8FA1A" box="[400,461,1459,1485]" captionStart="FIG" captionStartId="17.[132,143,739,756]" captionTargetBox="[142,1417,187,719]" captionTargetId="figure-596@17.[165,993,201,706]" captionTargetPageId="17" captionText="FIG. 9. — Eubrontes giganteus Hitchcock, 1845: A-C, track ULB-04D21_A (plaster cast of the holotype of “E. veillonensis” that is here invalidated), photograph (A), DEM and false-colour depth map (B) and interpretative sketch (C). Scale bars: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.11501876" httpUri="https://zenodo.org/record/11501876/files/figure.png" pageId="16" pageNumber="357">Fig. 9</figureCitation>
A-C)
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<subSubSection id="41806557DE027C00FF4A94B0DB46F9BF" pageId="16" pageNumber="357" type="reference_group">
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<treatmentCitation id="883B10CDDE027C00FF4A94B0D8DBF9D3" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[132,686,1516,1540]" page="27" pageId="16" pageNumber="357" year="1967">
<taxonomicName id="CE9A4D5FDE027C00FF4A94B0D8DBF9D3" authority="Lapparent &amp; Montenat, 1967: 23 - 27" authorityName="&quot; Lapparent &amp; Montenat" authorityPageNumber="23 - 27" authorityYear="1967" box="[132,686,1516,1540]" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="357" phylum="Chordata" rank="species" species="veillonensis">
<emphasis id="3BEEEACEDE027C00FF4A94B0DB3DF9D3" box="[132,328,1516,1540]" italics="true" pageId="16" pageNumber="357">Eubrontes veillonensis</emphasis>
<bibRefCitation id="6D0B4B2DDE027C00FE8394B0D8DBF9D3" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[333,686,1516,1540]" pageId="16" pageNumber="357" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat, 1967: 23-27</bibRefCitation>
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, pl. XI.2- 4, fig. 12.
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. — Subjective junior synonym of
<taxonomicName id="CE9A4D5FDE027C00FD8A976ADB5AF9BF" authority="Hitchcock, 1845" authorityName="Hitchcock" authorityYear="1845" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="357" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="3BEEEACEDE027C00FD8A976AD97DF999" box="[580,776,1590,1614]" italics="true" pageId="16" pageNumber="357">Eubrontes giganteus</emphasis>
<bibRefCitation id="6D0B4B2DDE027C00FF48970CDB5AF9BF" author="HITCHCOCK E." box="[134,303,1616,1640]" pageId="16" pageNumber="357" pagination="23 - 25" refId="ref17918" refString="HITCHCOCK E. 1845. - An attempt to name, classify, and describe the animals that made the fossil footmarks of New England, in Proceedings of the 6 th Meeting American Association of Geologists and Naturalists. Association of American Geologists and Naturalists: 23 - 25." type="book chapter" year="1845">Hitchcock, 1845</bibRefCitation>
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.
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. —
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:
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(plaster cast).
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The track is tridactyl, longer than wide (L/W = 1.3),
<quantity id="CE629B39DE027C00FD0D97AED973F8DB" box="[707,774,1778,1804]" metricMagnitude="-1" metricUnit="m" metricValue="3.4" pageId="16" pageNumber="357" unit="cm" value="34.0">34 cm</quantity>
long and
<quantity id="CE629B39DE027C00FF25964DDB33F8FB" box="[235,326,1809,1836]" metricMagnitude="-1" metricUnit="m" metricValue="2.65" pageId="16" pageNumber="357" unit="cm" value="26.5">26.5 cm</quantity>
wide (
<figureCitation id="91A12A59DE027C00FE5C964EDBA4F8FB" box="[402,465,1810,1836]" captionStart="FIG" captionStartId="17.[132,143,739,756]" captionTargetBox="[142,1417,187,719]" captionTargetId="figure-596@17.[165,993,201,706]" captionTargetPageId="17" captionText="FIG. 9. — Eubrontes giganteus Hitchcock, 1845: A-C, track ULB-04D21_A (plaster cast of the holotype of “E. veillonensis” that is here invalidated), photograph (A), DEM and false-colour depth map (B) and interpretative sketch (C). Scale bars: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.11501876" httpUri="https://zenodo.org/record/11501876/files/figure.png" pageId="16" pageNumber="357">Fig. 9</figureCitation>
A-C;
<tableCitation id="44180367DE027C00FDC0964ED82AF8FB" box="[526,607,1810,1836]" captionStart="TABLE" captionStartId="7.[132,143,220,237]" captionTargetPageId="7" captionText="TABLE 2. — Measured values on Hettangian tridactyl theropod tracks from Le Veillon. Abbreviations: L, length of the trace; W, width of the trace; D, length of the free part of digit III; II-IV, divarication angle between digit II and digit IV (all in cm except II-IV in degrees)." httpUri="http://table.plazi.org/id/5DE56654DE157C17FF4A9180D999FED4" pageId="16" pageNumber="357" tableUuid="5DE56654DE157C17FF4A9180D999FED4">Table 2</tableCitation>
). The divarication angle between digits II and IV is quite large (II-IV = 40°) and D is short (L/D = 3.5). Impressions of digits are particularly wide with well-distinguished and marked pads. The digito-metatarsal pad of digit IV is more proximal than that of digit II. Tracks display elongated and pointed claw marks.
</paragraph>
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<paragraph id="092536DCDE027C00FCE3942FDF2AF83C" blockId="16.[811,1457,1364,2028]" pageId="16" pageNumber="357">
Based on an
<emphasis id="3BEEEACEDE027C00FC7A9428D983FA5A" box="[948,1014,1396,1421]" italics="true" pageId="16" pageNumber="357">in situ</emphasis>
trackway composed of twelve footprints,
<bibRefCitation id="6D0B4B2DDE027C00FCE294C8DE0AFA7A" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[812,1151,1427,1454]" pageId="16" pageNumber="357" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat (1967)</bibRefCitation>
erected (without diagnosis) the ichnospecies “
<taxonomicName id="CE9A4D5FDE027C00FC3794EFDE97FA1A" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" box="[1017,1250,1459,1485]" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="357" phylum="Chordata" rank="species" species="veillonensis">
<emphasis id="3BEEEACEDE027C00FC3794EFDE97FA1A" box="[1017,1250,1459,1485]" italics="true" pageId="16" pageNumber="357">Eubrontes veillonensis</emphasis>
</taxonomicName>
”. ULB-04D21_A (
<figureCitation id="91A12A59DE027C00FCF8948FD903FA3A" box="[822,886,1491,1517]" captionStart="FIG" captionStartId="17.[132,143,739,756]" captionTargetBox="[142,1417,187,719]" captionTargetId="figure-596@17.[165,993,201,706]" captionTargetPageId="17" captionText="FIG. 9. — Eubrontes giganteus Hitchcock, 1845: A-C, track ULB-04D21_A (plaster cast of the holotype of “E. veillonensis” that is here invalidated), photograph (A), DEM and false-colour depth map (B) and interpretative sketch (C). Scale bars: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.11501876" httpUri="https://zenodo.org/record/11501876/files/figure.png" pageId="16" pageNumber="357">Fig. 9</figureCitation>
A-C) is the plaster cast of one of the best-preserved footprints of this trackway. A recent survey in 2022 on the tidal flat from Le Veillon could not relocate the specimen, which was most probably destroyed.
<taxonomicName id="CE9A4D5FDE027C00FBBA976FDFDAF99A" authority="Hitchcock, 1845" authorityName="Hitchcock" authorityYear="1845" box="[1140,1455,1586,1613]" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="357" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="3BEEEACEDE027C00FBBA976FDE85F99B" box="[1140,1264,1587,1612]" italics="true" pageId="16" pageNumber="357">E. giganteus</emphasis>
<bibRefCitation id="6D0B4B2DDE027C00FB36976EDFDAF99A" author="HITCHCOCK E." box="[1272,1455,1586,1613]" pageId="16" pageNumber="357" pagination="23 - 25" refId="ref17918" refString="HITCHCOCK E. 1845. - An attempt to name, classify, and describe the animals that made the fossil footmarks of New England, in Proceedings of the 6 th Meeting American Association of Geologists and Naturalists. Association of American Geologists and Naturalists: 23 - 25." type="book chapter" year="1845">Hitchcock, 1845</bibRefCitation>
</taxonomicName>
was introduced based on tridactyl tracks from the Early Jurassic rocks of the Dinosaur Footprint Reservation in Holyoke, Massachusetts (
<bibRefCitation id="6D0B4B2DDE027C00FC1D97CEDEFCF97B" author="OLSEN P. E. &amp; SMITH J. H. &amp; MC &amp; DONALD N. G." box="[979,1161,1682,1708]" pageId="16" pageNumber="357" pagination="586 - 601" refId="ref19432" refString="OLSEN P. E., SMITH J. H. &amp; MC DONALD N. G. 1998. - Type material of the type species of the classic theropod footprint genera Eubrontes, Anchisauripus and Grallator (Early Jurassic, Hartford and Deerfield basins, Connecticut and Massachusetts, U. S. A.). Journal of Vertebrate Paleontology 18: 586 - 601. https: // www. jstor. org / stable / 4523930" type="journal article" year="1998">
Olsen
<emphasis id="3BEEEACEDE027C00FBD797CFDE3EF97B" box="[1049,1099,1682,1708]" italics="true" pageId="16" pageNumber="357">et al.</emphasis>
1998
</bibRefCitation>
). In
<figureCitation id="91A12A59DE027C00FB7797CEDE83F97B" box="[1209,1270,1682,1708]" captionStart="FIG" captionStartId="10.[132,143,1963,1980]" captionTargetBox="[237,1380,197,1889]" captionTargetId="graphics-18@10.[311,1344,1074,1820]" captionTargetPageId="10" captionText="FIG. 4. — Footprints from Le Veillon compared to tridactyl tracks from the Early Jurassic of the Causses Basin (based on data from Demathieu et al. 2002; Moreau et al. 2021) and the Early Jurassic of the eastern United States (based on data from Weems 1992, 2019; Gand et al. 2018): A, bivariate diagram L vs (LD)/D; B, bivariate diagram L vs L/W. L in metres." figureDoi="http://doi.org/10.5281/zenodo.11501858" httpUri="https://zenodo.org/record/11501858/files/figure.png" pageId="16" pageNumber="357">Fig. 4</figureCitation>
, ULB-04D21_A is compared with Early Jurassic
<taxonomicName id="CE9A4D5FDE027C00FBB897EEDF48F91B" authorityName="Hitchcock" authorityYear="1845" box="[1142,1341,1714,1740]" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="357" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="3BEEEACEDE027C00FBB897EEDF48F91B" box="[1142,1341,1714,1740]" italics="true" pageId="16" pageNumber="357">Eubrontes giganteus</emphasis>
</taxonomicName>
specimens from
<collectingCountry id="718D764CDE027C00FCA8978ED9DBF93B" box="[870,942,1746,1772]" name="France" pageId="16" pageNumber="357">France</collectingCountry>
and from the
<collectingCountry id="718D764CDE027C00FB8B978EDEA2F93B" box="[1093,1239,1746,1772]" name="United States of America" pageId="16" pageNumber="357">United States</collectingCountry>
. This analysis shows that the
<taxonomicName id="CE9A4D5FDE027C00FC4B97AED99EF8DB" authorityName="Hitchcock" authorityYear="1845" box="[901,1003,1778,1804]" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="357" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE027C00FC4B97AED99EF8DB" box="[901,1003,1778,1804]" italics="true" pageId="16" pageNumber="357">Eubrontes</emphasis>
</taxonomicName>
track from Le Veillon fully falls within the morphological range of this ichnospecies.
<taxonomicName id="CE9A4D5FDE027C00FACE964EDF0AF8FC" authorityName="Hitchcock" authorityYear="1845" box="[1280,1407,1810,1835]" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="357" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="3BEEEACEDE027C00FACE964EDF0AF8FC" box="[1280,1407,1810,1835]" italics="true" pageId="16" pageNumber="357">E. giganteus</emphasis>
</taxonomicName>
and “
<taxonomicName id="CE9A4D5FDE027C00FCF6966ED9B0F89C" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" box="[824,965,1841,1867]" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="357" phylum="Chordata" rank="species" species="veillonensis">
<emphasis id="3BEEEACEDE027C00FCF6966ED9B0F89C" box="[824,965,1841,1867]" italics="true" pageId="16" pageNumber="357">E. veillonensis</emphasis>
</taxonomicName>
” show similar shape of footprints with massive impressions of digits II, III and IV bearing well-marked pads and long claw marks as well as the presence of a well-marked digito-metatarsal pad of digit IV more proximal than that of digit II. Based on these observations we consider “
<taxonomicName id="CE9A4D5FDE027C00FA8B96EED92EF83C" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="357" phylum="Chordata" rank="species" species="veillonensis">
<emphasis id="3BEEEACEDE027C00FA8B96EED92EF83C" italics="true" pageId="16" pageNumber="357">E. veillonensis</emphasis>
</taxonomicName>
” as a subjective junior synonym of
<emphasis id="3BEEEACEDE027C00FB15968EDF2AF83C" box="[1243,1375,2002,2027]" italics="true" pageId="16" pageNumber="357">
<taxonomicName id="CE9A4D5FDE027C00FB15968EDF2EF83C" authorityName="Hitchcock" authorityYear="1845" box="[1243,1371,2002,2027]" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="357" phylum="Chordata" rank="species" species="giganteus">E. giganteus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>