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<mods:title id="1E5134494B51065FC9CE723C9234FC6F">Crocodylomorph and dinosaur tracks from the lowermost Jurassic of Le Veillon (western France): ichnotaxonomic revision of the type material (Lapparent collection)</mods:title>
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<emphasis id="3BEEEACEDE007C02FCA697A5DE5CF8C6" bold="true" box="[872,1065,1783,1811]" italics="true" pageId="18" pageNumber="359">Talmontopus tersi</emphasis>
<bibRefCitation id="6D0B4B2DDE007C02FBF497A4DF0AF8C6" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[1082,1407,1783,1810]" pageId="18" pageNumber="359" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat, 1967</bibRefCitation>
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(
<emphasis id="3BEEEACEDE007C02FC1B9646DE01F8E4" box="[981,1140,1817,1843]" italics="true" pageId="18" pageNumber="359">nomen dubium</emphasis>
) (
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A-C)
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<treatmentCitation id="883B10CDDE007C02FCE39608DF3CF8BB" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[813,1353,1876,1900]" page="30" pageId="18" pageNumber="359" year="1967">
<taxonomicName id="CE9A4D5FDE007C02FCE39608DF3CF8BB" authority="Lapparent &amp; Montenat, 1967: 29 - 30" authorityName="&quot; Lapparent &amp; Montenat" authorityPageNumber="29 - 30" authorityYear="1967" box="[813,1353,1876,1900]" family="Anchisauripodidae" genus="Talmontopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="359" phylum="Chordata" rank="species" species="tersi">
<emphasis id="3BEEEACEDE007C02FCE39608D9A4F8BB" box="[813,977,1876,1900]" italics="true" pageId="18" pageNumber="359">Talmontopus tersi</emphasis>
<bibRefCitation id="6D0B4B2DDE007C02FC169609DF3CF8BB" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[984,1353,1876,1900]" pageId="18" pageNumber="359" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat, 1967: 29-30</bibRefCitation>
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, pl. XII.1, fig. 17.
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. — Subjective senior synonym of
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isp.
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. —
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<typeStatus id="D621887EDE007C02FBEA968FDEF1F83C" box="[1060,1156,2003,2027]" pageId="18" pageNumber="359" type="holotype">Holotype</typeStatus>
:
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.
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. 11. — Plaster cast of one footprint from the type trackway of “
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<emphasis id="3BEEEACEDE017C03FD6D9245D924FCFD" box="[675,849,793,810]" italics="true" pageId="19" pageNumber="360">Saltopoides igalensis</emphasis>
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”, that is here invalidated:
<emphasis id="3BEEEACEDE017C03FBEB9245DE32FCFD" bold="true" box="[1061,1095,793,810]" pageId="19" pageNumber="360">A -C</emphasis>
, track ULB-04C01_B, photograph (
<emphasis id="3BEEEACEDE017C03FAA59245DF0DFCFD" bold="true" box="[1387,1400,793,810]" pageId="19" pageNumber="360">A</emphasis>
), DEM and false-colour depth map (
<emphasis id="3BEEEACEDE017C03FEBB926CDBF7FC96" bold="true" box="[373,386,816,833]" pageId="19" pageNumber="360">B</emphasis>
) and interpretative sketch (
<emphasis id="3BEEEACEDE017C03FDAB926CD806FC96" bold="true" box="[613,627,816,833]" pageId="19" pageNumber="360">C</emphasis>
). Scale bars: 5 cm.
</paragraph>
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The track ULB-04C02_A consists of a single, slightly longer than wide (L/W = 1.2),
<quantity id="CE629B39DE017C03FE4192E9DBA1FC18" box="[399,468,949,975]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="360" unit="cm" value="26.0">26 cm</quantity>
long and
<quantity id="CE629B39DE017C03FD8B92E8D8D5FC18" box="[581,672,948,975]" metricMagnitude="-1" metricUnit="m" metricValue="2.15" pageId="19" pageNumber="360" unit="cm" value="21.5">21.5 cm</quantity>
wide tridactyl track (
<figureCitation id="91A12A59DE017C03FEDE9288DB2EFC38" box="[272,347,980,1007]" captionStart="FIG" captionStartId="20.[132,143,703,720]" captionTargetBox="[171,1430,189,675]" captionTargetId="figure-693@20.[171,1017,235,665]" captionTargetPageId="20" captionText="FIG. 12. — Kayentapus isp.: A-C, track ULB-04C02_A (holotype of “Talmontopus tersi” that is here invalidated), photograph (A), DEM and false-colour depth map (B) and interpretative sketch (C). Scale bars: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.11501884" httpUri="https://zenodo.org/record/11501884/files/figure.png" pageId="19" pageNumber="360">Fig. 12</figureCitation>
A-C;
<tableCitation id="44180367DE017C03FE5A9288DB96FC38" box="[404,483,980,1007]" captionStart="TABLE" captionStartId="7.[132,143,220,237]" captionTargetPageId="7" captionText="TABLE 2. — Measured values on Hettangian tridactyl theropod tracks from Le Veillon. Abbreviations: L, length of the trace; W, width of the trace; D, length of the free part of digit III; II-IV, divarication angle between digit II and digit IV (all in cm except II-IV in degrees)." httpUri="http://table.plazi.org/id/5DE56654DE157C17FF4A9180D999FED4" pageId="19" pageNumber="360" tableUuid="5DE56654DE157C17FF4A9180D999FED4">Table 2</tableCitation>
). The angle between II and IV is large (II-IV = 64°). Digit imprints are quite thin and pointed. The traces of digits II and IV are straight and quite similar in length. The trace of digit III is the longest and is slightly curved. The digito-metatarsal base of digit IV is slightly more proximal than that of digit II. The free part of III is quite short (L/D = 2.8). The imprints of pads and claws are only poorly marked.
</paragraph>
</subSubSection>
<subSubSection id="41806557DE017C03FF4A95A8DFDAFB79" pageId="19" pageNumber="360" type="discussion">
<paragraph id="092536DCDE017C03FF4A95A8DA9DFADB" blockId="19.[130,776,1268,2028]" box="[132,232,1268,1294]" pageId="19" pageNumber="360">
<smallCapsWord id="0FC3A000DE017C03FF4A95A8DA9DFADB" baselines="1287,1287" box="[132,232,1268,1294]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="19" pageNumber="360">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="092536DCDE017C03FF4A944FDFDAFB79" blockId="19.[130,776,1268,2028]" lastBlockId="19.[810,1456,885,1198]" pageId="19" pageNumber="360">
Based on the track ULB-04C02_
<bibRefCitation id="6D0B4B2DDE017C03FE249448DAB8FA9A" author="LAPPARENT A. F. DE &amp; MONTENAT C." pageId="19" pageNumber="360" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">A, Lapparent &amp; Montenat (1967)</bibRefCitation>
erected
<taxonomicName id="CE9A4D5FDE017C03FEEE946FDBA5FA9A" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" box="[288,464,1331,1357]" family="Anchisauripodidae" genus="Talmontopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="species" species="tersi">
<emphasis id="3BEEEACEDE017C03FEEE946FDBA5FA9A" box="[288,464,1331,1357]" italics="true" pageId="19" pageNumber="360">Talmontopus tersi</emphasis>
</taxonomicName>
. They justified the creation of “
<taxonomicName id="CE9A4D5FDE017C03FF5E9408DAA0FABA" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" box="[144,213,1364,1389]" family="Anchisauripodidae" genus="Talmontopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="species" species="tersi">
<emphasis id="3BEEEACEDE017C03FF5E9408DAA0FABA" box="[144,213,1364,1389]" italics="true" pageId="19" pageNumber="360">T. tersi</emphasis>
</taxonomicName>
” by a very high T value between thin tracks of digits, and the presence of a putative web mark.We consider the web mark drawn in fig. 17 of
<bibRefCitation id="6D0B4B2DDE017C03FE4094C8D8AFFA7A" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[398,730,1427,1454]" pageId="19" pageNumber="360" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat (1967)</bibRefCitation>
as a sedimentary structure that corresponds to a slight thickening of sediment preserved between the traces of digits (see also plate XII.1 inLapparent &amp; Montenat 1967; ULB-04C02_A). Since ULB-04C02_A shows many characters of
<taxonomicName id="CE9A4D5FDE017C03FD5F974FDB78F99A" authority="Welles, 1971" authorityName="Welles" authorityYear="1971" genus="Kayentapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE017C03FD5F974FD972F9FB" box="[657,775,1555,1580]" italics="true" pageId="19" pageNumber="360">Kayentapus</emphasis>
<bibRefCitation id="6D0B4B2DDE017C03FF4A976EDB78F99A" author="WELLES S. P." box="[132,269,1586,1613]" pageId="19" pageNumber="360" pagination="27 - 38" refId="ref20423" refString="WELLES S. P. 1971. - Dinosaur footprints from the Kayenta Formation of northern Arizona. Plateau 44: 27 - 38." type="journal article" year="1971">Welles, 1971</bibRefCitation>
</taxonomicName>
, we consider “
<taxonomicName id="CE9A4D5FDE017C03FE6F976ED857F99B" authorityName="Lapparent &amp; Montenat" authorityYear="1967" box="[417,546,1586,1612]" family="Anchisauripodidae" genus="Talmontopus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE017C03FE6F976ED857F99B" box="[417,546,1586,1612]" italics="true" pageId="19" pageNumber="360">Talmontopus</emphasis>
</taxonomicName>
” as a subjective senior synonym of this ichnogenus.
<taxonomicName id="CE9A4D5FDE017C03FE7F970FD851F9BB" authorityName="Welles" authorityYear="1971" box="[433,548,1619,1644]" genus="Kayentapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE017C03FE7F970FD851F9BB" box="[433,548,1619,1644]" italics="true" pageId="19" pageNumber="360">Kayentapus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="CE9A4D5FDE017C03FD66970ED970F9BB" authorityName="Hitchcock" authorityYear="1858" box="[680,773,1618,1644]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="19" pageNumber="360" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE017C03FD66970ED970F9BB" box="[680,773,1618,1644]" italics="true" pageId="19" pageNumber="360">Grallator</emphasis>
</taxonomicName>
and
<taxonomicName id="CE9A4D5FDE017C03FF7D972EDB6CF95B" authorityName="Hitchcock" authorityYear="1845" box="[179,281,1650,1676]" family="Grallatoridae" genus="Eubrontes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE017C03FF7D972EDB6CF95B" box="[179,281,1650,1676]" italics="true" pageId="19" pageNumber="360">Eubrontes</emphasis>
</taxonomicName>
in showing larger divarication angle II-IV and a smaller L/W ratio.
<taxonomicName id="CE9A4D5FDE017C03FE9497CFDBBAF97B" authorityName="Welles" authorityYear="1971" box="[346,463,1683,1708]" genus="Kayentapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE017C03FE9497CFDBBAF97B" box="[346,463,1683,1708]" italics="true" pageId="19" pageNumber="360">Kayentapus</emphasis>
</taxonomicName>
was erected by
<bibRefCitation id="6D0B4B2DDE017C03FDBF97CED970F97B" author="WELLES S. P." box="[625,773,1682,1708]" pageId="19" pageNumber="360" pagination="27 - 38" refId="ref20423" refString="WELLES S. P. 1971. - Dinosaur footprints from the Kayenta Formation of northern Arizona. Plateau 44: 27 - 38." type="journal article" year="1971">Welles (1971)</bibRefCitation>
based on the
<typeStatus id="D621887EDE017C03FED697EFDB33F91A" box="[280,326,1715,1741]" pageId="19" pageNumber="360">type</typeStatus>
ichnospecies
<taxonomicName id="CE9A4D5FDE017C03FE1197EFD8BCF91B" authority="Welles, 1971" authorityName="Welles" authorityYear="1971" box="[479,713,1714,1740]" genus="Kayentapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="species" species="hopii">
<emphasis id="3BEEEACEDE017C03FE1197EFD843F91B" box="[479,566,1714,1740]" italics="true" pageId="19" pageNumber="360">K. hopii</emphasis>
<bibRefCitation id="6D0B4B2DDE017C03FDF297EED8BCF91B" author="WELLES S. P." box="[572,713,1714,1740]" pageId="19" pageNumber="360" pagination="27 - 38" refId="ref20423" refString="WELLES S. P. 1971. - Dinosaur footprints from the Kayenta Formation of northern Arizona. Plateau 44: 27 - 38." type="journal article" year="1971">Welles, 1971</bibRefCitation>
</taxonomicName>
from the Early Jurassic Kayenta Formation in
<collectingRegion id="CB5EF83EDE017C03FDF7978ED8E4F93B" box="[569,657,1746,1772]" country="United States of America" name="Arizona" pageId="19" pageNumber="360">Arizona</collectingRegion>
. Two other ichnospecies were later described from Early Jurassic strata of the
<collectingCountry id="718D764CDE017C03FF64964EDB4FF8FB" box="[170,314,1810,1836]" name="United States of America" pageId="19" pageNumber="360">United States</collectingCountry>
and
<collectingCountry id="718D764CDE017C03FEA0964EDBC9F8FB" box="[366,444,1810,1836]" name="Poland" pageId="19" pageNumber="360">Poland</collectingCountry>
:
<taxonomicName id="CE9A4D5FDE017C03FE08964EDACEF89B" authority="(Lull, 1953) Weems, 1987" authorityName="Weems" authorityYear="1987" baseAuthorityName="Lull" baseAuthorityYear="1953" genus="Kayentapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="species" species="minor">
<emphasis id="3BEEEACEDE017C03FE08964ED856F8FC" box="[454,547,1810,1835]" italics="true" pageId="19" pageNumber="360">K. minor</emphasis>
(
<bibRefCitation id="6D0B4B2DDE017C03FDFF964ED8D6F8FB" author="LULL R. S." box="[561,675,1809,1836]" pageId="19" pageNumber="360" refId="ref18568" refString="LULL R. S. 1953. - Triassic life of the Connecticut Valley. State Geological and Natural History Survey, Hartford, 331 p." type="book" year="1953">Lull, 1953</bibRefCitation>
)
<bibRefCitation id="6D0B4B2DDE017C03FD7F964EDACEF89B" author="WEEMS R. E." pageId="19" pageNumber="360" pagination="1 - 79" refId="ref20285" refString="WEEMS R. E. 1987. - A Late Triassic footprint fauna from the Culpeper basin northern Virginia (USA). Transactions of the American Philosophical Society 77: 1 - 79. https: // doi. org / 10.2307 / 1006417" type="journal article" year="1987">Weems, 1987</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="CE9A4D5FDE017C03FF20966ED970F89C" authority="(Gierlinski, 1991) Gierlinski, 1996" authorityName="Gierlinski" authorityYear="1996" baseAuthorityName="Gierlinski" baseAuthorityYear="1991" box="[238,773,1841,1868]" genus="Kayentapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="species" species="soltykovensis">
<emphasis id="3BEEEACEDE017C03FF20966EDBFCF89C" box="[238,393,1841,1867]" italics="true" pageId="19" pageNumber="360">K. soltykovensis</emphasis>
(
<bibRefCitation id="6D0B4B2DDE017C03FE59966DD83CF89B" author="GIERLINSKI G." box="[407,585,1841,1868]" pageId="19" pageNumber="360" pagination="137 - 148" refId="ref17678" refString="GIERLINSKI G. 1991. - New dinosaur ichnotaxa from the Early Jurassic of the Holy Cross Mountains, Poland. Palaeogeography, Palaeoclimatology, Palaeoecology 85: 137 - 148. https: // doi. org / 10.1016 / 0031 - 0182 (91) 90030 - u" type="journal article" year="1991">Gierliński, 1991</bibRefCitation>
)
<bibRefCitation id="6D0B4B2DDE017C03FD98966DD970F89C" author="GIERLINSKI G." box="[598,773,1841,1867]" pageId="19" pageNumber="360" pagination="119 - 128" refId="ref17728" refString="GIERLINSKI G. 1996. - Dinosaur ichnotaxa from the Lower Jurassic of Hungary. Geological Quarterly 40: 119 - 128 [retrieved from https: // gq. pgi. gov. pl / article / view / 8168]." type="journal article" year="1996">Gierliński, 1996</bibRefCitation>
</taxonomicName>
(firstly named
<taxonomicName id="CE9A4D5FDE017C03FED0960DD809F8BC" baseAuthorityName="Gierlinski" baseAuthorityYear="1991" box="[286,636,1873,1900]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="19" pageNumber="360" phylum="Chordata" rank="species" species="soltykovensis" subGenus="Eubrontes">
<emphasis id="3BEEEACEDE017C03FED0960DDB08F8BC" box="[286,381,1873,1899]" italics="true" pageId="19" pageNumber="360">Grallator</emphasis>
(
<emphasis id="3BEEEACEDE017C03FE45960DDB84F8BC" box="[395,497,1873,1899]" italics="true" pageId="19" pageNumber="360">Eubrontes</emphasis>
)
<emphasis id="3BEEEACEDE017C03FE31960DD809F8BC" box="[511,636,1873,1899]" italics="true" pageId="19" pageNumber="360">soltykovensis</emphasis>
</taxonomicName>
in
<bibRefCitation id="6D0B4B2DDE017C03FD52960DDAC9F85C" author="GIERLINSKI G." pageId="19" pageNumber="360" pagination="137 - 148" refId="ref17678" refString="GIERLINSKI G. 1991. - New dinosaur ichnotaxa from the Early Jurassic of the Holy Cross Mountains, Poland. Palaeogeography, Palaeoclimatology, Palaeoecology 85: 137 - 148. https: // doi. org / 10.1016 / 0031 - 0182 (91) 90030 - u" type="journal article" year="1991">Gierliński 1991</bibRefCitation>
and
<bibRefCitation id="6D0B4B2DDE017C03FF3C962DD862F85C" author="GIERLINSKI G. &amp; AHLBERG A." box="[242,535,1905,1931]" pageId="19" pageNumber="360" pagination="99 - 105" refId="ref17771" refString="GIERLINSKI G. &amp; AHLBERG A. 1994. - Late Triassic and Early Jurassic dinosaur footprints in the Hoganas Formation of southern Sweden. Ichnos 3: 99 - 105. https: // doi. org / 10.1080 / 10420949409386377" type="journal article" year="1994">Gierliński &amp; Ahlberg 1994</bibRefCitation>
), respectively (
<bibRefCitation id="6D0B4B2DDE017C03FD77962EDACEF87C" author="WEEMS R. E." pageId="19" pageNumber="360" pagination="1 - 79" refId="ref20285" refString="WEEMS R. E. 1987. - A Late Triassic footprint fauna from the Culpeper basin northern Virginia (USA). Transactions of the American Philosophical Society 77: 1 - 79. https: // doi. org / 10.2307 / 1006417" type="journal article" year="1987">Weems 1987</bibRefCitation>
;
<bibRefCitation id="6D0B4B2DDE017C03FF0996CDDB1EF87C" author="GIERLINSKI G." box="[199,363,1937,1963]" pageId="19" pageNumber="360" pagination="119 - 128" refId="ref17728" refString="GIERLINSKI G. 1996. - Dinosaur ichnotaxa from the Lower Jurassic of Hungary. Geological Quarterly 40: 119 - 128 [retrieved from https: // gq. pgi. gov. pl / article / view / 8168]." type="journal article" year="1996">Gierliński 1996</bibRefCitation>
). Based on its D/W and (L-D)/W ratios (0.43 and 0.78, respectively), the track ULB-04C02_A fully falls within the morphological space of
<taxonomicName id="CE9A4D5FDE017C03FDF8968ED8F9F83C" authorityName="Welles" authorityYear="1971" box="[566,652,2001,2027]" genus="Kayentapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="species" species="hopii">
<emphasis id="3BEEEACEDE017C03FDF8968ED8F9F83C" box="[566,652,2001,2027]" italics="true" pageId="19" pageNumber="360">K. hopii</emphasis>
</taxonomicName>
(
<bibRefCitation id="6D0B4B2DDE017C03FD52968DD916FC58" author="GIERLINSKI G." box="[668,867,885,2027]" pageId="19" pageNumber="360" pagination="119 - 128" refId="ref17728" refString="GIERLINSKI G. 1996. - Dinosaur ichnotaxa from the Lower Jurassic of Hungary. Geological Quarterly 40: 119 - 128 [retrieved from https: // gq. pgi. gov. pl / article / view / 8168]." type="journal article" year="1996">Gierliński 1996</bibRefCitation>
: fig.1 andLockley
<emphasis id="3BEEEACEDE017C03FBE7922ADE2CFC58" box="[1065,1113,885,911]" italics="true" pageId="19" pageNumber="360">et al.</emphasis>
2011: fig. 6B). However,
<taxonomicName id="CE9A4D5FDE017C03FA93922ADFD8FC58" authorityName="Welles" authorityYear="1971" box="[1373,1453,885,911]" genus="Kayentapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="species" species="hopii">
<emphasis id="3BEEEACEDE017C03FA93922ADFD8FC58" box="[1373,1453,885,911]" italics="true" pageId="19" pageNumber="360">K. hopii</emphasis>
</taxonomicName>
is clearly larger (L = 34.0-35.5) than ULB-04C02_A (
<figureCitation id="91A12A59DE017C03FAAD92C9DFEAFC78" box="[1379,1439,917,943]" captionStart="FIG" captionStartId="10.[132,143,1963,1980]" captionTargetBox="[237,1380,197,1889]" captionTargetId="graphics-18@10.[311,1344,1074,1820]" captionTargetPageId="10" captionText="FIG. 4. — Footprints from Le Veillon compared to tridactyl tracks from the Early Jurassic of the Causses Basin (based on data from Demathieu et al. 2002; Moreau et al. 2021) and the Early Jurassic of the eastern United States (based on data from Weems 1992, 2019; Gand et al. 2018): A, bivariate diagram L vs (LD)/D; B, bivariate diagram L vs L/W. L in metres." figureDoi="http://doi.org/10.5281/zenodo.11501858" httpUri="https://zenodo.org/record/11501858/files/figure.png" pageId="19" pageNumber="360">Fig. 4</figureCitation>
). It also differs from ULB-04C02_A in showing traces of digits that are slender and well individualised, and digito-metatarsal pads of the digit IV much more proximal. ULB-04C02_A is close to
<taxonomicName id="CE9A4D5FDE017C03FCB09549D9AEFBF9" authorityName="Weems" authorityYear="1987" baseAuthorityName="Lull" baseAuthorityYear="1953" box="[894,987,1045,1070]" genus="Kayentapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="species" species="minor">
<emphasis id="3BEEEACEDE017C03FCB09549D9AEFBF9" box="[894,987,1045,1070]" italics="true" pageId="19" pageNumber="360">K. minor</emphasis>
</taxonomicName>
(
<figureCitation id="91A12A59DE017C03FC249548DE53FBF9" box="[1002,1062,1044,1070]" captionStart="FIG" captionStartId="10.[132,143,1963,1980]" captionTargetBox="[237,1380,197,1889]" captionTargetId="graphics-18@10.[311,1344,1074,1820]" captionTargetPageId="10" captionText="FIG. 4. — Footprints from Le Veillon compared to tridactyl tracks from the Early Jurassic of the Causses Basin (based on data from Demathieu et al. 2002; Moreau et al. 2021) and the Early Jurassic of the eastern United States (based on data from Weems 1992, 2019; Gand et al. 2018): A, bivariate diagram L vs (LD)/D; B, bivariate diagram L vs L/W. L in metres." figureDoi="http://doi.org/10.5281/zenodo.11501858" httpUri="https://zenodo.org/record/11501858/files/figure.png" pageId="19" pageNumber="360">Fig. 4</figureCitation>
), however, it seems to differ from this ichnospecies in showing a higher D/W ratio (see
<bibRefCitation id="6D0B4B2DDE017C03FA8B9568D910FBB9" author="GIERLINSKI G." pageId="19" pageNumber="360" pagination="119 - 128" refId="ref17728" refString="GIERLINSKI G. 1996. - Dinosaur ichnotaxa from the Lower Jurassic of Hungary. Geological Quarterly 40: 119 - 128 [retrieved from https: // gq. pgi. gov. pl / article / view / 8168]." type="journal article" year="1996">Gierliński 1996</bibRefCitation>
: fig. 1 and
<bibRefCitation id="6D0B4B2DDE017C03FC129508DEDDFBB9" author="LOCKLEY M. G. &amp; GIERLINSKI G. &amp; LUCAS S. G." box="[988,1192,1108,1134]" pageId="19" pageNumber="360" pagination="330 - 336" refId="ref18376" refString="LOCKLEY M. G., GIERLINSKI G. &amp; LUCAS S. G. 2011. - Kayentapus revised: notes on the type material and the importance of this theropod footprint ichnogenus. New Mexico Museum of Natural History and Science Bulletin 53: 330 - 336 (retrieved fromhttps: // nmdigital. unm. edu / digital / collection / bulletins / id / 1423 / rec / 1]." type="journal article" year="2011">
Lockley
<emphasis id="3BEEEACEDE017C03FBF69509DE1FFBB9" box="[1080,1130,1108,1134]" italics="true" pageId="19" pageNumber="360">et al.</emphasis>
2011
</bibRefCitation>
: fig. 6B).
<taxonomicName id="CE9A4D5FDE017C03FADC9509DFDBFBB9" baseAuthorityName="Gierlinski" baseAuthorityYear="1991" box="[1298,1454,1108,1134]" genus="Kayentapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="360" phylum="Chordata" rank="species" species="slotykovensis">
<emphasis id="3BEEEACEDE017C03FADC9509DFDBFBB9" box="[1298,1454,1108,1134]" italics="true" pageId="19" pageNumber="360">K. slotykovensis</emphasis>
</taxonomicName>
differs from ULB-04C02_A in showing a smaller (L-D)/W ratio (see
<bibRefCitation id="6D0B4B2DDE017C03FC9595C8DE70FB79" author="GIERLINSKI G." box="[859,1029,1172,1198]" pageId="19" pageNumber="360" pagination="119 - 128" refId="ref17728" refString="GIERLINSKI G. 1996. - Dinosaur ichnotaxa from the Lower Jurassic of Hungary. Geological Quarterly 40: 119 - 128 [retrieved from https: // gq. pgi. gov. pl / article / view / 8168]." type="journal article" year="1996">Gierliński 1996</bibRefCitation>
: fig. 1 and
<bibRefCitation id="6D0B4B2DDE017C03FBB095C8DF3FFB79" author="LOCKLEY M. G. &amp; GIERLINSKI G. &amp; LUCAS S. G." box="[1150,1354,1172,1198]" pageId="19" pageNumber="360" pagination="330 - 336" refId="ref18376" refString="LOCKLEY M. G., GIERLINSKI G. &amp; LUCAS S. G. 2011. - Kayentapus revised: notes on the type material and the importance of this theropod footprint ichnogenus. New Mexico Museum of Natural History and Science Bulletin 53: 330 - 336 (retrieved fromhttps: // nmdigital. unm. edu / digital / collection / bulletins / id / 1423 / rec / 1]." type="journal article" year="2011">
Lockley
<emphasis id="3BEEEACEDE017C03FB1795C9DF79FB79" box="[1241,1292,1172,1198]" italics="true" pageId="19" pageNumber="360">et al.</emphasis>
2011
</bibRefCitation>
: fig. 6B).
</paragraph>
</subSubSection>
</treatment>
</document>