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<document id="390FAC6B36D45CE4BC83FD80123E82AD" ID-DOI="10.11646/zootaxa.3691.2.4" ID-GBIF-Dataset="0d7e6fd5-6762-4709-9d21-865617a12794" ID-ISSN="1175-5326" ID-Zenodo-Dep="222829" ID-ZooBank="8EEA9F53-4F57-4C02-8FCB-2F6A67FC34FC" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460650533101" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Rahayu, Dwi Listyo" docDate="2013" docId="03EF9F137B35941340BDFCECCCAFE3CF" docLanguage="en" docName="zt03691p252.pdf" docOrigin="Zootaxa 3691 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lophopagurus (Lophopagurus) minimus Komai &amp; Rahayu, 2013, n. sp." docType="treatment" docVersion="8" lastPageNumber="246" masterDocId="FFD6E76B7B349415402AFFB3CC40E06B" masterDocTitle="A new species of the pagurid hermit crab genus Lophopagurus McLaughlin, 1981 (Crustacea: Decapoda: Anomura) from French Polynesia, and additional record of L. indonesiensis McLaughlin, 1997 from the Philippines" masterLastPageNumber="252" masterPageNumber="240" pageNumber="241" updateTime="1698337886706" updateUser="plazi">
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<mods:title id="4458F992623D1E4A0441FB4C801DA7D8">A new species of the pagurid hermit crab genus Lophopagurus McLaughlin, 1981 (Crustacea: Decapoda: Anomura) from French Polynesia, and additional record of L. indonesiensis McLaughlin, 1997 from the Philippines</mods:title>
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<mods:namePart id="AF15C29CB46DB63DE85B475656FFBB07">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="83AD6F1C7996C2AB23C7B6DD4B7A220B">Rahayu, Dwi Listyo</mods:namePart>
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<mods:title id="F15DA27E9A96F1A480FA44E3B27EEED8">Zootaxa</mods:title>
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<mods:date id="78FF089573591550334DEBC0A2CB9049">2013</mods:date>
<mods:detail id="29AC1995EB719835AC28646BCA9476C8" type="volume">
<mods:number id="DADBE2C4F1C53CEB018D53DA45BA97E8">3691</mods:number>
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<mods:identifier id="0CC2B8C393DC494B497FC0815A1F1D1D" type="DOI">10.11646/zootaxa.3691.2.4</mods:identifier>
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<treatment id="03EF9F137B35941340BDFCECCCAFE3CF" ID-DOI="http://doi.org/10.5281/zenodo.6153527" ID-GBIF-Taxon="119572782" ID-Zenodo-Dep="6153527" LSID="urn:lsid:plazi:treatment:03EF9F137B35941340BDFCECCCAFE3CF" httpUri="http://treatment.plazi.org/id/03EF9F137B35941340BDFCECCCAFE3CF" lastPageId="6" lastPageNumber="246" pageId="1" pageNumber="241">
<subSubSection id="C35C7D8E7B35941440BDFCECCD51E3F1" pageId="1" pageNumber="241" type="nomenclature">
<paragraph id="8BF92E057B35941440BDFCECCE8FE312" blockId="1.[151,719,863,922]" box="[151,719,863,889]" pageId="1" pageNumber="241">
<heading id="D0B199697B35941440BDFCECCE8FE312" bold="true" box="[151,719,863,889]" fontSize="11" level="1" pageId="1" pageNumber="241" reason="1">
<emphasis id="B932F2177B35941440BDFCECCE8FE312" bold="true" box="[151,719,863,889]" pageId="1" pageNumber="241">
<taxonomicName id="4C4655867B35941440BDFCECCEC6E312" box="[151,646,863,889]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="241" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov." subGenus="Lophopagurus">
<emphasis id="B932F2177B35941440BDFCECCD09E312" bold="true" box="[151,329,863,889]" italics="true" pageId="1" pageNumber="241">Lophopagurus</emphasis>
(
<emphasis id="B932F2177B3594144173FCECCE4BE312" bold="true" box="[345,523,863,889]" italics="true" pageId="1" pageNumber="241">Lophopagurus</emphasis>
)
<emphasis id="B932F2177B3594144236FCECCEC6E312" bold="true" box="[540,646,863,889]" italics="true" pageId="1" pageNumber="241">minimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2014F6C7B35941442A7FCECCE8FE312" box="[653,719,863,889]" pageId="1" pageNumber="241" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF92E057B35941440BDFC31CD51E3F1" blockId="1.[151,719,863,922]" box="[151,273,898,922]" pageId="1" pageNumber="241">
(
<figureCitation id="137D32807B35941440B5FC31CD49E3F1" box="[159,265,898,922]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1884,1907]" captionStartId-1="4.[151,250,1609,1632]" captionStartId-2="5.[151,250,1798,1821]" captionTargetBox-0="[196,1383,203,1854]" captionTargetBox-1="[164,1427,197,1583]" captionTargetBox-2="[163,1432,205,1769]" captionTargetId-0="figure@3.[183,1415,193,1857]" captionTargetId-1="figure@4.[151,1436,193,1588]" captionTargetId-2="figure@5.[151,1436,193,1777]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." captionText-1="FIGURE 2. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, right cheliped, mesial view; B, same, lateral view; C, left cheliped, mesial view; D, same, lateral view. Scale bar: 0.5 mm." captionText-2="FIGURE 3. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, right second pereopod, lateral view; B, same, dactylus, mesial view; C, left third pereopod, lateral view; D, same, dactylus, mesial view. Scale bar: 0.5 mm." httpUri-0="https://zenodo.org/record/222830/files/figure.png" httpUri-1="https://zenodo.org/record/222831/files/figure.png" httpUri-2="https://zenodo.org/record/222832/files/figure.png" pageId="1" pageNumber="241">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35C7D8E7B35941440BDFC79C801E425" pageId="1" pageNumber="241" type="materials_examined">
<paragraph id="8BF92E057B35941440BDFC79CE1BE441" blockId="1.[151,1436,970,2002]" pageId="1" pageNumber="241">
<emphasis id="B932F2177B35941440BDFC79CD39E388" bold="true" box="[151,377,970,995]" pageId="1" pageNumber="241">Material examined</emphasis>
.
<typeStatus id="54FD90A77B35941441A2FC79CDB5E389" box="[392,501,970,994]" pageId="1" pageNumber="241" type="holotype">Holotype</typeStatus>
: ovigerous female (sl
<quantity id="4CBE83E07B35941442C7FC79CF08E389" box="[749,840,970,994]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="241" unit="mm" value="1.3">1.3 mm</quantity>
), off Motu, Maeherehonae, Rangiroa Atoll, Tuamoto Archipelago, Frenth Polynesia,
<geoCoordinate id="EE7248C27B3594144229FC5DCEC5E46D" box="[515,645,1006,1030]" direction="south" orientation="latitude" pageId="1" pageNumber="241" precision="9" value="-14.154834">14°09.29S</geoCoordinate>
,
<geoCoordinate id="EE7248C27B35941442BEFC5DCF6CE46D" box="[660,812,1006,1030]" direction="west" orientation="longitude" pageId="1" pageNumber="241" precision="9" value="-147.14317">147°08.59W</geoCoordinate>
,
<quantity id="4CBE83E07B3594144310FC5DCFD4E46D" box="[826,916,1006,1030]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="1" pageNumber="241" unit="m" value="9.0" valueMax="12.0" valueMin="6.0">612 m</quantity>
, outer reef slope, under rocks,
<date id="FFF808C57B359414452AFC5DCC92E441" pageId="1" pageNumber="241" value="2001-11-10">10 November 2001</date>
, coll. Gustav Paulay, FMNH 1858.
</paragraph>
<paragraph id="8BF92E057B35941440EDFB84C801E425" blockId="1.[151,1436,970,2002]" box="[199,1089,1078,1103]" pageId="1" pageNumber="241">
<typeStatus id="54FD90A77B35941440EDFB84CD6DE424" box="[199,301,1079,1103]" pageId="1" pageNumber="241" type="paratype">Paratype</typeStatus>
: 1 ovigerous female (sl
<quantity id="4CBE83E07B3594144212FB85CED2E425" box="[568,658,1078,1102]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="241" unit="mm" value="1.2">1.2 mm</quantity>
), same data as
<typeStatus id="54FD90A77B3594144313FB85CFDEE425" box="[825,926,1078,1102]" pageId="1" pageNumber="241" type="holotype">holotype</typeStatus>
, FMNH 1813.
</paragraph>
</subSubSection>
<subSubSection id="C35C7D8E7B35941140EDFBE9CE6BE71A" lastPageId="4" lastPageNumber="244" pageId="1" pageNumber="241" type="description">
<paragraph id="8BF92E057B35941440EDFBE9C83FE4B5" blockId="1.[151,1436,970,2002]" pageId="1" pageNumber="241">
<emphasis id="B932F2177B35941440EDFBE9CD19E418" bold="true" box="[199,345,1114,1139]" pageId="1" pageNumber="241">Description.</emphasis>
Shield (
<figureCitation id="137D32807B3594144197FBE9CDBFE419" box="[445,511,1114,1138]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="1" pageNumber="241">Fig. 1</figureCitation>
A) approximately as long as wide; anterior margin between rostrum and lateral projections slightly concave; lateral margins strongly convex; dorsal surface with few tufts of short setae laterally; paragastric grooves delimited only posteriorly. Rostrum broadly triangular, extending beyond lateral projections, terminating in small spine. Lateral projections obsolete, each with 1 submarginal spinule.
</paragraph>
<paragraph id="8BF92E057B35941440EDFB59CC93E5F9" blockId="1.[151,1436,970,2002]" pageId="1" pageNumber="241">
Ocular peduncles (
<figureCitation id="137D32807B35941441B0FB59CD99E569" box="[410,473,1258,1282]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="1" pageNumber="241">Fig. 1</figureCitation>
A) about 0.7 times as long as shield, moderately stout, faintly constricted proximal to base of corneas, somewhat inflated basally, each with longitudinal row of tufts of short setae on dorsal surface in distal half; cornea slightly dilated; basal width distinctly greater than corneal width. Ocular acicles narrowly subtriangular, with submarginal spinule distally; moderately separated basally. Interocular lobe visible in dorsal view.
</paragraph>
<paragraph id="8BF92E057B35941440EDFA2DCD47E649" blockId="1.[151,1436,970,2002]" pageId="1" pageNumber="241">
Antennular peduncles (
<figureCitation id="137D32807B35941441E6FA2DCE4AE5DD" box="[460,522,1438,1462]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="1" pageNumber="241">Fig. 1</figureCitation>
A) overreaching distal corneal margins by about 0.3 length of ultimate segments. Ultimate segment with prominent tuft of long setae on dorsolateral distal angle, dorsal surface almost glabrous. Basal segment with moderately inflated statocyst lobe bearing prominent spine laterally; ventrodistal angle slightly produced.
</paragraph>
<paragraph id="8BF92E057B35941440EDF99DC957E775" blockId="1.[151,1436,970,2002]" pageId="1" pageNumber="241">
Antennal peduncles (
<figureCitation id="137D32807B3594144192F99DCDB8E62D" box="[440,504,1582,1606]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="1" pageNumber="241">Fig. 1</figureCitation>
A) slightly falling short of distal corneal margins. Fifth and fourth segments with few stiff setae on lateral and mesial margins. Third segment with tiny spine at ventromesial distal angle. Second segment with dorsolateral distal angle produced to strong spine reaching midlength of fourth segment; mesial margin with 1 additional spinule subterminally; dorsomesial distal angle with small spine. First segment unarmed on lateral surface; ventrodistal margin strongly produced. Antennal acicle reaching corneal base, terminating in tiny spine, with row of moderately long setae dorsomesially. Antennal flagellum about 4 times as long as shield, overreaching extended right cheliped; each article with 13 short setae and 1 long seta every 2 articles.
</paragraph>
<paragraph id="8BF92E057B35941440EDF899CEF4E7E1" blockId="1.[151,1436,970,2002]" pageId="1" pageNumber="241">
Third maxilliped (
<figureCitation id="137D32807B359414418EF899CDADE729" box="[420,493,1834,1858]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="1" pageNumber="241">Fig. 1</figureCitation>
B) moderately stout. Ischium (
<figureCitation id="137D32807B3594144342F899CFEFE729" box="[872,943,1834,1858]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="1" pageNumber="241">Fig. 1</figureCitation>
C) bearing well-developed crista dentata consisting of row of corneous teeth increasing in size proximally and 1 accessory tooth. Merus and carpus unarmed. Exopod reaching midlength of carpus.
</paragraph>
<paragraph id="8BF92E057B35941740EDF825CF25E37F" blockId="1.[151,1436,970,2002]" lastBlockId="2.[151,1437,151,2012]" lastPageId="2" lastPageNumber="242" pageId="1" pageNumber="241">
Chelipeds appreciably unequal and dissimilar. Right cheliped (
<figureCitation id="137D32807B35941443B5F825CFADE7C5" box="[927,1005,1942,1966]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="1" pageNumber="241">Figs. 1</figureCitation>
D, E; 2A, B) with chela subovate in dorsal view, with mesial margin of palm nearly straight and lateral margin arcuate. Dactylus subequal in length to palm, slightly curved ventrally; dorsal surface convex transversely, smooth, without median ridge; dorsomesial or mesial margin not delimited, mesial face with some tiny tubercles along midline proximally; surfaces with scattered moderately short setae; cutting edge with row of obtuse calcareous teeth, terminating in small calcareous claw. Palm slightly shorter than carpus; dorsal surface gently convex, with sparse granules and scattered short to long setae; dorsolateral margin sharply delimited by tuberculate carina extending onto fixed finger; dorsomesial margin delimited by double row of low, rounded tubercles; lateral surface smooth, with scattered short setae; mesial surface also smooth, with few long setae; ventral surface convex, with few long setae. Fixed finger broader than dactylus; cutting edge with 2 blunt calcareous teeth in proximal 0.4, smooth in distal 0.6, terminating in calcareous claw. Carpus longer than merus, distinctly widened distally; dorsomesial margin with row of 4 moderately strong spines increasing in size distally and tufts of long setae; dorsal surface smooth, with few long setae; dorsolateral margin not clearly delimited, with 1 or 2 small tubercles and tufts of long setae; mesial surface with 2 short transverse ridges bearing row of long setae in proximal half and 1 small submarginal spine adjacent to distal angle, and also with scattered long setae; lateral surface smooth, with several short to long setae dorsally; ventral surface with anterior half forming concavity to accommodate carpus when cheliped flexed, and posterior half convex, with several long setae. Merus unarmed on dorsodistal margin; dorsal surface also unarmed, glabrous; lateral surface smooth, with some long setae proximoventrally, ventrolateral margin with 2 spines distally; mesial surface smooth, glabrous, ventromesial margin unarmed, with row of numerous long setae; ventral surface slightly concave transversely, with several long setae. Ischium unarmed.
</paragraph>
<paragraph id="8BF92E057B36941740EDFCACCEDDE513" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="242">
Left cheliped (
<figureCitation id="137D32807B3694174146FCACCDF7E35C" box="[364,439,799,823]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="2" pageNumber="242">Figs. 1</figureCitation>
F; 2C, D) with chela elongate subovate in general outline. Dactylus 1.9 times as long as palm, surfaces unarmed, bearing scattered tufts of short to long setae; cutting edge with row of closely set corneous teeth, terminating in small corneous claw. Palm short, about half length of carpus, triangular in cross-section; prominently elevated dorsal midline with row of small spines or tubercles extending onto midlength of fixed finger; lateral margin delimited by row of blunt small tubercles, extending onto midlength of fixed finger; dorsomesial margin not delimited; mesial surface smooth, with scattered long setae; ventral surface slightly convex, with scattered long setae. Cutting edge of fixed finger with row of corneous teeth and few tiny, sharp calcareous teeth in distal half. Carpus slightly widened distally, subequal in length to merus; dorsolateral margin with row of 3 prominent spines in distal half, dorsomesial margin with row of 2 spines in distal 0.3, both margins with tufts of long stiff setae; dorsodistal margin with 1 additional minute spine laterally; lateral surface with few tufts of long setae dorsally, otherwise glabrous, ventrolateral margin denticulate; mesial surface smooth, with long setae in distal half and 2 short transverse ridges bearing setae in posterior half dorsally, ventromesial margin strongly convex, unarmed; anterior half of ventral surface forming concavity, posterior half with several long setae. Merus unarmed on dorsodistal margin; dorsal surface almost glabrous; lateral surface glabrous, nearly straight with row of 3 prominent spines and 1 or 2 low protuberances; mesial surface also smooth, with few setae ventrally, ventromesial margin with 2 small spines proximally; ventral surface with several long setae. Ischium unarmed, with short to long setae dorsally and ventrally.
</paragraph>
<paragraph id="8BF92E057B36941740EDFA37C842E743" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="242">
Ambulatory legs (
<figureCitation id="137D32807B36941741BEFA37CD96E5F7" box="[404,470,1412,1436]" captionStart="FIGURE 3" captionStartId="5.[151,250,1798,1821]" captionTargetBox="[163,1432,205,1769]" captionTargetId="figure@5.[151,1436,193,1777]" captionTargetPageId="5" captionText="FIGURE 3. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, right second pereopod, lateral view; B, same, dactylus, mesial view; C, left third pereopod, lateral view; D, same, dactylus, mesial view. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/222832/files/figure.png" pageId="2" pageNumber="242">Fig. 3</figureCitation>
AD) generally similar from right to left, moderately slender. Dactyli slightly curved ventrally in lateral view, nearly straight in dorsal view, about 1.1 times as long as propodi, 6.57.0 times as long as wide; dorsal margins each with sparse row of tufts of moderately short setae; lateral and mesial surfaces without sulcus or groove, mesial surfaces unarmed (second) (
<figureCitation id="137D32807B36941742CCFA43CF67E663" box="[742,807,1520,1544]" captionStart="FIGURE 3" captionStartId="5.[151,250,1798,1821]" captionTargetBox="[163,1432,205,1769]" captionTargetId="figure@5.[151,1436,193,1777]" captionTargetPageId="5" captionText="FIGURE 3. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, right second pereopod, lateral view; B, same, dactylus, mesial view; C, left third pereopod, lateral view; D, same, dactylus, mesial view. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/222832/files/figure.png" pageId="2" pageNumber="242">Fig. 3</figureCitation>
B) or 2 minute spinules adjacent to dorsal margin (third) (
<figureCitation id="137D32807B36941740B5F9A7CCA1E647" box="[159,225,1556,1580]" captionStart="FIGURE 3" captionStartId="5.[151,250,1798,1821]" captionTargetBox="[163,1432,205,1769]" captionTargetId="figure@5.[151,1436,193,1777]" captionTargetPageId="5" captionText="FIGURE 3. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, right second pereopod, lateral view; B, same, dactylus, mesial view; C, left third pereopod, lateral view; D, same, dactylus, mesial view. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/222832/files/figure.png" pageId="2" pageNumber="242">Fig. 3</figureCitation>
D); ventral margins each with row of 6 or 7 slender corneous spines. Propodi with row of tufts of long setae on dorsal margin, 4.34.5 times as long as wide; lateral and mesial faces with few short setae (second) or glabrous (third); ventral margins with row of moderately long setae (second) or with very short setae (third); ventrodistal margins each with pair of corneous spines. Carpi each with small dorsodistal spine (second) or unarmed (third); dorsal margins each with row of tufts of long setae; ventral surfaces with few setae. Meri each with row of moderately long setae on dorsal margins; lateral and mesial faces smooth; ventral margins each with tufts of long setae (setae longer in second than in third), second with ventrolateral distal spine and 1 ventral spine located at distal one-third, third unarmed. Female with unpaired left gonopore (
<figureCitation id="137D32807B36941743B1F8A3CF9BE743" box="[923,987,1808,1832]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="2" pageNumber="242">Fig. 1</figureCitation>
H).
</paragraph>
<paragraph id="8BF92E057B36941740EDF887C8F1E71B" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="242">
Fourth pereopods (
<figureCitation id="137D32807B369417418AF887CDA1E727" box="[416,481,1844,1868]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="2" pageNumber="242">Fig. 1</figureCitation>
G) semichelate. Dactyli each with row of relatively long corneous teeth on ventral margins; no preungual process. Propodi with rasp consisting of single row of corneous scales.
</paragraph>
<paragraph id="8BF92E057B36941740EDF8CFCDBBE7B7" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="242">
Sixth thoracic sternite (
<figureCitation id="137D32807B36941741E1F8CFCE49E7FF" box="[459,521,1916,1940]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="2" pageNumber="242">Fig. 1</figureCitation>
H) with anterior lobe subrectangular, slightly skewed to left; sparse short setae on anterior margin. Eighth thoracic sternite (
<figureCitation id="137D32807B369417425DF813CEF9E7D3" box="[631,697,1952,1976]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="2" pageNumber="242">Fig. 1</figureCitation>
E) with two narrowly spaced rounded lobes (concealed by first pleopods), with few short setae.
</paragraph>
<caption id="DF397E8D7B37941640BDF8EFC92BE786" httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="3" pageNumber="243" targetBox="[196,1383,203,1854]" targetPageId="3">
<paragraph id="8BF92E057B37941640BDF8EFC92BE786" blockId="3.[151,1436,1884,2029]" pageId="3" pageNumber="243">
<emphasis id="B932F2177B37941640BDF8EFCD4DE71A" bold="true" box="[151,269,1884,1907]" pageId="3" pageNumber="243">FIGURE 1</emphasis>
.
<taxonomicName id="4C4655867B3794164136F8EFCEF9E719" box="[284,697,1884,1906]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov." subGenus="Lophopagurus">
<emphasis id="B932F2177B3794164136F8EFCDF0E719" box="[284,432,1884,1906]" italics="true" pageId="3" pageNumber="243">Lophopagurus</emphasis>
(
<emphasis id="B932F2177B3794164195F8EFCE13E719" box="[447,595,1884,1906]" italics="true" pageId="3" pageNumber="243">Lophopagurus</emphasis>
)
<emphasis id="B932F2177B3794164249F8EECEF9E719" box="[611,697,1885,1906]" italics="true" pageId="3" pageNumber="243">minimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2014F6C7B37941642EBF8EECEBBE719" box="[705,763,1885,1906]" pageId="3" pageNumber="243" rank="species">
<emphasis id="B932F2177B37941642EBF8EECEBBE719" bold="true" box="[705,763,1885,1906]" pageId="3" pageNumber="243">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="DF397E8D7B30941140BDF9FAC973E615" httpUri="https://zenodo.org/record/222831/files/figure.png" pageId="4" pageNumber="244" targetBox="[164,1427,197,1583]" targetPageId="4">
<paragraph id="8BF92E057B30941140BDF9FAC973E615" blockId="4.[151,1436,1609,1662]" pageId="4" pageNumber="244">
<emphasis id="B932F2177B30941140BDF9FACD4FE634" bold="true" box="[151,271,1609,1632]" pageId="4" pageNumber="244">FIGURE 2</emphasis>
.
<taxonomicName id="4C4655867B3094114137F9F9CEFCE60B" box="[285,700,1610,1632]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="244" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov." subGenus="Lophopagurus">
<emphasis id="B932F2177B3094114137F9F9CDF1E60B" box="[285,433,1610,1632]" italics="true" pageId="4" pageNumber="244">Lophopagurus</emphasis>
(
<emphasis id="B932F2177B30941141E8F9F9CE16E60B" box="[450,598,1610,1632]" italics="true" pageId="4" pageNumber="244">Lophopagurus</emphasis>
)
<emphasis id="B932F2177B309411424CF9F8CEFCE60B" box="[614,700,1611,1632]" italics="true" pageId="4" pageNumber="244">minimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2014F6C7B30941142EEF9F9CF40E634" box="[708,768,1610,1631]" pageId="4" pageNumber="244" rank="species">
<emphasis id="B932F2177B30941142EEF9F9CF40E634" bold="true" box="[708,768,1610,1631]" pageId="4" pageNumber="244">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, right cheliped, mesial view; B, same, lateral view; C, left cheliped, mesial view; D, same, lateral view. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BF92E057B30941140EDF915CD0DE725" blockId="4.[151,1436,1702,1943]" pageId="4" pageNumber="244">
Pleon dextrally twisted. Female with paired first pleopods modified as gonopods (
<figureCitation id="137D32807B3094114470F915C8DAE6D5" box="[1114,1178,1702,1726]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="4" pageNumber="244">
Fig.
<date id="FFF808C57B30941144A4F915C8DAE6D5" box="[1166,1178,1702,1726]" pageId="4" pageNumber="244">1</date>
</figureCitation>
I), protopod and ramus distinctly articulated; unpaired, left second to fifth pleopods present, second and third subequally biramous, fourth and fifth not reduced in size, uniramous. Uropods strongly asymmetrical. Telson (
<figureCitation id="137D32807B3094114479F95DC8D6E76D" box="[1107,1174,1774,1798]" captionStart="FIGURE 1" captionStartId="3.[151,250,1884,1907]" captionTargetBox="[196,1383,203,1854]" captionTargetId="figure@3.[183,1415,193,1857]" captionTargetPageId="3" captionText="FIGURE 1. Lophopagurus (Lophopagurus) minimus n. sp., holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, shield and cephalic appendages, dorsal view; B, left third maxilliped, lateral view; C, same, ischium, ventral view; D, chela of right cheliped, dorsal view; E, carpus of right cheliped, dorsal view; F, chela and carpus of left cheliped, dorsal view; G, left fourth pereopod, lateral view; H, coxae of third pereopods and sixth thoracic sternite, ventral view; I, coxae of fifth pereopods and paired first pleopods, ventral view; J. telson, dorsal view; inset, terminal margins, posterodorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/222830/files/figure.png" pageId="4" pageNumber="244">Fig. 1</figureCitation>
J) with shallow lateral indentations; terminal margins slightly oblique, each with 3 or 4 spinules; posterolateral margins forming thin chitinous plates.
</paragraph>
<paragraph id="8BF92E057B30941140EDF8EACE6BE71A" blockId="4.[151,1436,1702,1943]" box="[199,555,1881,1906]" pageId="4" pageNumber="244">
<emphasis id="B932F2177B30941140EDF8EACDD2E719" bold="true" box="[199,402,1881,1906]" pageId="4" pageNumber="244">Coloration in life</emphasis>
. Not Known.
</paragraph>
</subSubSection>
<subSubSection id="C35C7D8E7B30941140EDF8CEC8D3E7FD" box="[199,1171,1917,1943]" pageId="4" pageNumber="244" type="distribution">
<paragraph id="8BF92E057B30941140EDF8CEC8D3E7FD" blockId="4.[151,1436,1702,1943]" box="[199,1171,1917,1943]" pageId="4" pageNumber="244">
<emphasis id="B932F2177B30941140EDF8CECD19E7FD" bold="true" box="[199,345,1917,1942]" pageId="4" pageNumber="244">Distribution</emphasis>
. Known only from the
<typeStatus id="54FD90A77B309411424AF8CCCED0E7FC" box="[608,656,1919,1943]" pageId="4" pageNumber="244">type</typeStatus>
locality, Tuamotu Archipelago,
<quantity id="4CBE83E07B30941143D0F8CDC80FE7FD" box="[1018,1103,1918,1942]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="4" pageNumber="244" unit="m" value="9.0" valueMax="12.0" valueMin="6.0">612 m</quantity>
deep.
</paragraph>
</subSubSection>
<caption id="DF397E8D7B31941040BDF8B5CDD3E731" httpUri="https://zenodo.org/record/222832/files/figure.png" pageId="5" pageNumber="245" targetBox="[163,1432,205,1769]" targetPageId="5">
<paragraph id="8BF92E057B31941040BDF8B5CDD3E731" blockId="5.[151,1436,1798,1882]" pageId="5" pageNumber="245">
<emphasis id="B932F2177B31941040BDF8B5CD4FE777" bold="true" box="[151,271,1798,1821]" pageId="5" pageNumber="245">FIGURE 3</emphasis>
.
<taxonomicName id="4C4655867B3194104137F8B4CEFCE776" box="[285,700,1799,1821]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="245" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov." subGenus="Lophopagurus">
<emphasis id="B932F2177B3194104137F8B4CDF1E776" box="[285,433,1799,1821]" italics="true" pageId="5" pageNumber="245">Lophopagurus</emphasis>
(
<emphasis id="B932F2177B31941041E8F8B4CE16E776" box="[450,598,1799,1821]" italics="true" pageId="5" pageNumber="245">Lophopagurus</emphasis>
)
<emphasis id="B932F2177B319410424CF8BBCEFCE776" box="[614,700,1800,1821]" italics="true" pageId="5" pageNumber="245">minimus</emphasis>
</taxonomicName>
<emphasis id="B932F2177B31941042EEF8B4CF40E777" bold="true" box="[708,768,1799,1820]" pageId="5" pageNumber="245">
<taxonomicNameLabel id="A2014F6C7B31941042EEF8B4CF40E777" box="[708,768,1799,1820]" pageId="5" pageNumber="245" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female (sl 1.3 mm), FMNH 1858. A, right second pereopod, lateral view; B, same, dactylus, mesial view; C, left third pereopod, lateral view; D, same, dactylus, mesial view. Scale bar: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C35C7D8E7B31941340EDF831C8E1E337" lastPageId="6" lastPageNumber="246" pageId="5" pageNumber="245" type="discussion">
<paragraph id="8BF92E057B31941340EDF831CCA1E2EF" blockId="5.[151,1436,1921,2017]" lastBlockId="6.[151,1436,151,932]" lastPageId="6" lastPageNumber="246" pageId="5" pageNumber="245">
<emphasis id="B932F2177B31941040EDF831CD74E7F1" bold="true" box="[199,308,1922,1946]" pageId="5" pageNumber="245">Remarks</emphasis>
. Although no male specimens are available for study, the present new species is assigned to the genus
<taxonomicName id="4C4655867B31941040CAF816CDC3E7D6" box="[224,387,1957,1981]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="B932F2177B31941040CAF816CDC3E7D6" box="[224,387,1957,1981]" italics="true" pageId="5" pageNumber="245">Lophopagurus</emphasis>
</taxonomicName>
with little hesitation because of the possession of 11 pairs of biserial gills, the occurrence of paired first pleopods in females and the similarities in the general structure of both chelipeds.
<taxonomicName id="4C4655867B31941344D3F87ACDF3E0DB" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" lastPageId="6" lastPageNumber="246" order="Decapoda" pageId="5" pageNumber="245" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov." subGenus="Lophopagurus">
<emphasis id="B932F2177B31941044D3F87AC9DCE78A" box="[1273,1436,1993,2017]" italics="true" pageId="5" pageNumber="245">Lophopagurus</emphasis>
(
<emphasis id="B932F2177B329413408AFF2BCD04E0DB" box="[160,324,152,176]" italics="true" pageId="6" pageNumber="246">Lophopagurus</emphasis>
)
<emphasis id="B932F2177B329413417EFF2ACDF3E0DB" box="[340,435,153,176]" italics="true" pageId="6" pageNumber="246">minimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2014F6C7B3294134191FF2BCDBBE0DB" box="[443,507,152,176]" pageId="6" pageNumber="246" rank="species">
<emphasis id="B932F2177B3294134191FF2BCDBBE0DB" bold="true" box="[443,507,152,176]" pageId="6" pageNumber="246">n. sp.</emphasis>
</taxonomicNameLabel>
appears closest to
<emphasis id="B932F2177B32941342F8FF2ACEA1E0DB" box="[722,737,153,176]" italics="true" pageId="6" pageNumber="246">L</emphasis>
. (
<emphasis id="B932F2177B32941342D0FF2ACF49E0DB" box="[762,777,153,176]" italics="true" pageId="6" pageNumber="246">L</emphasis>
.)
<taxonomicName id="4C4655867B329413430BFF2ACF24E0DB" box="[801,868,153,176]" class="Malacostraca" family="Paguridae" genus="Pylopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B932F2177B329413430BFF2ACF24E0DB" box="[801,868,153,176]" italics="true" pageId="6" pageNumber="246">nanus</emphasis>
</taxonomicName>
known from southeastern
<collectingCountry id="F3516E957B32941344B8FF24C8B9E0C4" box="[1170,1273,151,175]" name="Australia" pageId="6" pageNumber="246">Australia</collectingCountry>
and Tasmania in carpi of second pereopods bearing only a dorsodistal spine and the flat lateral surface of the dactylus of the left third pereopod. However, the new species is easily distinguished from the latter species by many morphological characters (cf. McLaughlin &amp; Gunn 1994). The ocular peduncles are basally inflated in
<taxonomicName id="4C4655867B3294134447FEB6C8AAE177" box="[1133,1258,261,284]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B932F2177B3294134447FEB6C8AAE177" box="[1133,1258,261,284]" italics="true" pageId="6" pageNumber="246">L. minimus</emphasis>
</taxonomicName>
<emphasis id="B932F2177B32941344DBFEB7C96FE177" bold="true" box="[1265,1327,260,284]" pageId="6" pageNumber="246">
<taxonomicNameLabel id="A2014F6C7B32941344DBFEB7C96FE177" box="[1265,1327,260,284]" pageId="6" pageNumber="246" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but such inflation is not seen in the ocular peduncles of
<taxonomicName id="4C4655867B32941342B5FE9ACEBFE12B" box="[671,767,297,320]" class="Malacostraca" family="Paguridae" genus="Pylopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B932F2177B32941342B5FE9ACEBFE12B" box="[671,767,297,320]" italics="true" pageId="6" pageNumber="246">L. nanus</emphasis>
</taxonomicName>
. The dactylus of the right cheliped is unarmed on the dorsal surface in
<taxonomicName id="4C4655867B3294134120FEFECDC6E10F" box="[266,390,333,356]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B932F2177B3294134120FEFECDC6E10F" box="[266,390,333,356]" italics="true" pageId="6" pageNumber="246">L. minimus</emphasis>
</taxonomicName>
<emphasis id="B932F2177B32941341A6FEFFCD8AE10F" bold="true" box="[396,458,332,356]" pageId="6" pageNumber="246">
<taxonomicNameLabel id="A2014F6C7B32941341A6FEFFCD8AE10F" box="[396,458,332,356]" pageId="6" pageNumber="246" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, whereas there is a single or double row of tubercles or protuberances along the midline in
<taxonomicName id="4C4655867B329413409EFEC2CD56E1E3" box="[180,278,369,392]" class="Malacostraca" family="Paguridae" genus="Pylopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B932F2177B329413409EFEC2CD56E1E3" box="[180,278,369,392]" italics="true" pageId="6" pageNumber="246">L. nanus</emphasis>
</taxonomicName>
; the dorsomesial margin is not delimited in
<taxonomicName id="4C4655867B3294134326FEC2CFC9E1E3" box="[780,905,369,392]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B932F2177B3294134326FEC2CFC9E1E3" box="[780,905,369,392]" italics="true" pageId="6" pageNumber="246">L. minimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2014F6C7B32941343BBFEC3CF90E1E3" box="[913,976,368,392]" pageId="6" pageNumber="246" rank="species">
<emphasis id="B932F2177B32941343BBFEC3CF90E1E3" bold="true" box="[913,976,368,392]" pageId="6" pageNumber="246">n. sp.</emphasis>
</taxonomicNameLabel>
, rather than sharply delimited by row of small tubercles in
<taxonomicName id="4C4655867B329413414FFE26CD86E1C7" box="[357,454,405,428]" class="Malacostraca" family="Paguridae" genus="Pylopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B932F2177B329413414FFE26CD86E1C7" box="[357,454,405,428]" italics="true" pageId="6" pageNumber="246">L. nanus</emphasis>
</taxonomicName>
. The dorsal surface of the right palm bears only sparse, minute tubercles in
<taxonomicName id="4C4655867B3294134535FE26C9DCE1C7" box="[1311,1436,405,428]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B932F2177B3294134535FE26C9DCE1C7" box="[1311,1436,405,428]" italics="true" pageId="6" pageNumber="246">L. minimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2014F6C7B32941340BDFE0BCC98E1BB" box="[151,216,440,464]" pageId="6" pageNumber="246" rank="species">
<emphasis id="B932F2177B32941340BDFE0BCC98E1BB" bold="true" box="[151,216,440,464]" pageId="6" pageNumber="246">n. sp.</emphasis>
</taxonomicNameLabel>
, whereas it is armed with a longitudinal row of small tubercles laterad to the midline as well as scattered minute tubercles in
<taxonomicName id="4C4655867B3294134159FE6ECD92E19F" box="[371,466,477,500]" class="Malacostraca" family="Paguridae" genus="Pylopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B932F2177B3294134159FE6ECD92E19F" box="[371,466,477,500]" italics="true" pageId="6" pageNumber="246">L. nanus</emphasis>
</taxonomicName>
. The carpus of the right cheliped has only one dorsomesial row of spines in
<taxonomicName id="4C4655867B329413450AFE6EC9DCE19F" box="[1312,1436,477,500]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B932F2177B329413450AFE6EC9DCE19F" box="[1312,1436,477,500]" italics="true" pageId="6" pageNumber="246">L. minimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2014F6C7B32941340BDFDB3CC96E273" box="[151,214,512,536]" pageId="6" pageNumber="246" rank="species">
<emphasis id="B932F2177B32941340BDFDB3CC96E273" bold="true" box="[151,214,512,536]" pageId="6" pageNumber="246">n. sp.</emphasis>
</taxonomicNameLabel>
, rather than two dorsomesial rows of spines in
<taxonomicName id="4C4655867B32941342C6FDB2CF0DE273" box="[748,845,513,536]" class="Malacostraca" family="Paguridae" genus="Pylopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B932F2177B32941342C6FDB2CF0DE273" box="[748,845,513,536]" italics="true" pageId="6" pageNumber="246">L. nanus</emphasis>
</taxonomicName>
. The propodi of the ambulatory legs are each armed only with a pair of corneous spines on the distoventral margin in
<taxonomicName id="4C4655867B329413435AFD96CFACE257" box="[880,1004,549,572]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B932F2177B329413435AFD96CFACE257" box="[880,1004,549,572]" italics="true" pageId="6" pageNumber="246">L. minimus</emphasis>
</taxonomicName>
, but there is a row of minute corneous spines on ventral margins of each propodus, in addition to two or three spines on the distoventral margin, in
<taxonomicName id="4C4655867B32941345AFFDFACC9AE2EF" class="Malacostraca" family="Paguridae" genus="Pylopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B932F2177B32941345AFFDFACC9AE2EF" italics="true" pageId="6" pageNumber="246">L. nanus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF92E057B32941340EDFD23C8E1E337" blockId="6.[151,1436,151,932]" pageId="6" pageNumber="246">
<taxonomicName id="4C4655867B32941340EDFD23CD92E2C3" box="[199,466,656,680]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B932F2177B32941340EDFD23CD92E2C3" box="[199,466,656,680]" italics="true" pageId="6" pageNumber="246">Lophopagurus minimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2014F6C7B32941341F1FD23CE5BE2C3" box="[475,539,656,680]" pageId="6" pageNumber="246" rank="species">
<emphasis id="B932F2177B32941341F1FD23CE5BE2C3" bold="true" box="[475,539,656,680]" pageId="6" pageNumber="246">n. sp.</emphasis>
</taxonomicNameLabel>
is the smallest representative of the genus, measuring only
<quantity id="4CBE83E07B32941344E0FD3CC913E2CC" box="[1226,1363,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="6" pageNumber="246" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.21.3 mm</quantity>
in the shield length in ovigerous females. De Saint Laurent &amp; McLaughlin (2000) ranked
<emphasis id="B932F2177B329413446DFD06C816E2A7" box="[1095,1110,693,716]" italics="true" pageId="6" pageNumber="246">L</emphasis>
. (
<emphasis id="B932F2177B3294134445FD06C83EE2A7" box="[1135,1150,693,716]" italics="true" pageId="6" pageNumber="246">L</emphasis>
.)
<taxonomicName id="4C4655867B32941344BDFD07C8AFE2A7" box="[1175,1263,692,716]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B932F2177B32941344BDFD07C8AFE2A7" box="[1175,1263,692,716]" italics="true" pageId="6" pageNumber="246">pumilus</emphasis>
</taxonomicName>
as the smallest species (measuring
<quantity id="4CBE83E07B3294134159FD64CDB9E284" box="[371,505,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.9" metricValueMin="1.4" pageId="6" pageNumber="246" unit="mm" value="2.65" valueMax="3.9" valueMin="1.4">1.43.9 mm</quantity>
in the shield length in ovigerous females) in the genus known at the time. This new species also represents the first representative of the subgenus
<taxonomicName id="4C4655867B329413434FFD4FC849E37F" box="[869,1033,764,788]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="genus">
<emphasis id="B932F2177B329413434FFD4FC849E37F" box="[869,1033,764,788]" italics="true" pageId="6" pageNumber="246">Lophopagurus</emphasis>
</taxonomicName>
occurring in tropical waters. Other seven known species in the subgenus are distributed in
<collectingCountry id="F3516E957B329413430CFC92CFFFE35C" box="[806,959,799,824]" name="New Zealand" pageId="6" pageNumber="246">New Zealand</collectingCountry>
(
<taxonomicName id="4C4655867B32941343FEFC92C814E353" box="[980,1108,801,824]" class="Malacostraca" family="Paguridae" genus="Pylopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="crenatus">
<emphasis id="B932F2177B32941343FEFC92C814E353" box="[980,1108,801,824]" italics="true" pageId="6" pageNumber="246">L. crenatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4655867B329413444CFC92C88EE353" box="[1126,1230,800,824]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="B932F2177B329413444CFC92C88EE353" box="[1126,1230,800,824]" italics="true" pageId="6" pageNumber="246">L. foresti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4655867B32941344CAFC92C933E353" box="[1248,1395,800,824]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="nodulosus">
<emphasis id="B932F2177B32941344CAFC92C933E353" box="[1248,1395,800,824]" italics="true" pageId="6" pageNumber="246">L. nodulosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4655867B32941345AFFC92CD4AE337" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="lacertosus">
<emphasis id="B932F2177B32941345AFFC92CD4AE337" italics="true" pageId="6" pageNumber="246">L. lacertosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4655867B3294134132FCF6CDCEE337" box="[280,398,836,860]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B932F2177B3294134132FCF6CDCEE337" box="[280,398,836,860]" italics="true" pageId="6" pageNumber="246">L. pumilus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4655867B32941341EEFCF6CE16E337" box="[452,598,836,860]" class="Malacostraca" family="Paguridae" genus="Pylopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis id="B932F2177B32941341EEFCF6CE16E337" box="[452,598,836,860]" italics="true" pageId="6" pageNumber="246">L. thompsoni</emphasis>
</taxonomicName>
) or southeastern
<collectingCountry id="F3516E957B3294134332FCF7CF3FE337" box="[792,895,836,860]" name="Australia" pageId="6" pageNumber="246">Australia</collectingCountry>
and Tasmania (
<taxonomicName id="4C4655867B329413441BFCF6C8D1E337" box="[1073,1169,837,860]" class="Malacostraca" family="Paguridae" genus="Pylopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B932F2177B329413441BFCF6C8D1E337" box="[1073,1169,837,860]" italics="true" pageId="6" pageNumber="246">L. nanus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35C7D8E7B32941340EDFCDBCCAFE3CF" pageId="6" pageNumber="246" type="etymology">
<paragraph id="8BF92E057B32941340EDFCDBCCAFE3CF" blockId="6.[151,1436,151,932]" pageId="6" pageNumber="246">
<emphasis id="B932F2177B32941340EDFCDBCD06E3EB" bold="true" box="[199,326,872,896]" pageId="6" pageNumber="246">Etymology</emphasis>
. The specific name from the Latin
<taxonomicName id="4C4655867B32941342C3FCDACF08E3EB" box="[745,840,873,896]" class="Malacostraca" family="Paguridae" genus="Lophopagurus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B932F2177B32941342C3FCDACF08E3EB" box="[745,840,873,896]" italics="true" pageId="6" pageNumber="246">minimus</emphasis>
</taxonomicName>
, meaning smallest, reflecting the small size of this species.
</paragraph>
</subSubSection>
</treatment>
</document>