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<mods:title id="EEFFD230FC11E834331ECA90C7CA1A82">The Muscidae (Diptera) of Saudi Arabia, descriptions of two new species, new records and updated list of species</mods:title>
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<mods:namePart id="94BBD9FB6AD01D5793ECA7BA1ECFD5D9">Dawah, Hassan A.</mods:namePart>
<mods:affiliation id="1E1D6009EC408F23323293E25B236A79">Centre for Environmental Research and Studies, Jazan University, P. O. Box 2095, Jazan, Kingdom of Saudi Arabia.</mods:affiliation>
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<mods:namePart id="DF90E46C64507B2624A67FF1BB186BDD">Abdullah, Mohammed A.</mods:namePart>
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<mods:affiliation id="5D1D27ED81C0544E3AC24E34AE3AC0EB">Department of Biology, College of Science, King Khalid University, PO Box 9004, Abha- 61413, Kingdom of Saudi Arabia. mohd _ robiya @ hotmail. com; https: // orcid. org / 0000 - 0002 - 3323 - 3623</mods:affiliation>
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<mods:affiliation id="50B2B0BB3A686E91D16B9003C70783FE">National Museum of Wales, Department of Natural Sciences, Entomology Section, Cardiff, CF 10 3 NP, UK. john. deeming @ museumwales. ac. uk; https: // orcid. org / 0000 - 0002 - 6703 - 8180</mods:affiliation>
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<subSubSection id="C35CFAA22C10FFA2FF3A6513FD400465" box="[151,726,1050,1077]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BF9A9292C10FFA2FF3A6513FD400465" blockId="9.[151,1437,1050,1401]" box="[151,726,1050,1077]" pageId="9" pageNumber="10">
<taxonomicName id="4C46D2AA2C10FFA2FF3A6513FDED0464" authority="Speiser, 1924" authorityName="Speiser" authorityYear="1924" box="[151,635,1050,1077]" class="Insecta" family="Anthomyiidae" genus="Musca" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="confiscata" subGenus="Byomya">
<emphasis id="B932753B2C10FFA2FF3A6513FDED0464" bold="true" box="[151,635,1050,1077]" pageId="9" pageNumber="10">
<emphasis id="B932753B2C10FFA2FF3A6513FF730465" bold="true" box="[151,229,1051,1077]" italics="true" pageId="9" pageNumber="10">Musca</emphasis>
(
<emphasis id="B932753B2C10FFA2FF586513FEC40465" bold="true" box="[245,338,1051,1077]" italics="true" pageId="9" pageNumber="10">Byomya</emphasis>
)
<emphasis id="B932753B2C10FFA2FECE6512FE4F0464" bold="true" box="[355,473,1050,1076]" italics="true" pageId="9" pageNumber="10">confiscata</emphasis>
<bibRefCitation id="EFD7D4D82C10FFA2FE4D6512FDED0464" author="Speiser, P." box="[480,635,1050,1076]" pageId="9" pageNumber="10" pagination="90 - 156" refId="ref36309" refString="Speiser, P. (1924) Eine Ubersicht uber die Dipterenfauna Deutsch-Ostafrikas .. In: Beitrage aus der Tierkunde. Herrn Geh. Regierungsrat Prof. Dr. med. et phil. M. Braun aus Anlass seines goldenen medizinischen Doktor-Jubilaums als Festgabe dargebracht von Schulern und Freunden. Grafe &amp; Unzer, Konigsberg (Kaliningrad), pp. 90 - 156." type="book chapter" year="1924">Speiser, 1924</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation id="137DB5AC2C10FFA2FD276512FD5A0465" box="[650,716,1050,1077]" captionStart="FIGURE" captionStartId="10.[151,251,1919,1943]" captionTargetBox="[160,1420,621,1878]" captionTargetId="figure-231@10.[153,1433,596,1896]" captionTargetPageId="10" captionText="FIGURE. 25. Muscidae species habitus: 2) Morellia nilotica (Loew, 1856), Male; 3) Musca (Byomya) biseta Hough, 1898, Male; 4. Musca (Byomya) conducens Walker, 1859, Female; 5. Musca (Byomya) confiscata Speiser, 1924, Male." figureDoi="http://doi.org/10.5281/zenodo.4418252" httpUri="https://zenodo.org/record/4418252/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BF9A9292C10FFA2FF3A6537FDAC0408" blockId="9.[151,1437,1050,1401]" box="[151,570,1086,1112]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB568E072C10FFA2FF3A6537FDAC0408" box="[151,570,1086,1112]" pageId="9" pageNumber="10">
<treatmentCitation id="0AE78F382C10FFA2FF3A6537FDA00408" author="Speiser, P." box="[151,566,1086,1112]" page="104" pageId="9" pageNumber="10" year="1924">
<taxonomicName id="4C46D2AA2C10FFA2FF3A6537FDA00408" authority="Speiser, 1924: 104" authorityName="Speiser" authorityPageNumber="104" authorityYear="1924" box="[151,566,1086,1112]" class="Insecta" family="Anthomyiidae" genus="Musca" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="confiscata">
<emphasis id="B932753B2C10FFA2FF3A6537FECA0408" box="[151,348,1086,1112]" italics="true" pageId="9" pageNumber="10">Musca confiscata</emphasis>
<bibRefCitation id="EFD7D4D82C10FFA2FECE6536FDA00408" author="Speiser, P." box="[355,566,1086,1112]" pageId="9" pageNumber="10" pagination="90 - 156" refId="ref36309" refString="Speiser, P. (1924) Eine Ubersicht uber die Dipterenfauna Deutsch-Ostafrikas .. In: Beitrage aus der Tierkunde. Herrn Geh. Regierungsrat Prof. Dr. med. et phil. M. Braun aus Anlass seines goldenen medizinischen Doktor-Jubilaums als Festgabe dargebracht von Schulern und Freunden. Grafe &amp; Unzer, Konigsberg (Kaliningrad), pp. 90 - 156." type="book chapter" year="1924">Speiser, 1924: 104</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="8BF9A9292C10FFA2FF3A656BFB7F042D" blockId="9.[151,1437,1050,1401]" box="[151,1257,1122,1149]" pageId="9" pageNumber="10">
<taxonomicName id="4C46D2AA2C10FFA2FF3A656BFE6E042C" authority="Speiser 1924" authorityName="Speiser" authorityYear="1924" box="[151,504,1122,1148]" class="Insecta" family="Anthomyiidae" genus="Musca" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="confiscata">
<emphasis id="B932753B2C10FFA2FF3A656BFECA042C" box="[151,348,1122,1148]" italics="true" pageId="9" pageNumber="10">Musca confiscata</emphasis>
<bibRefCitation id="EFD7D4D82C10FFA2FECE656AFE6E042C" author="Speiser, P." box="[355,504,1122,1148]" pageId="9" pageNumber="10" pagination="90 - 156" refId="ref36309" refString="Speiser, P. (1924) Eine Ubersicht uber die Dipterenfauna Deutsch-Ostafrikas .. In: Beitrage aus der Tierkunde. Herrn Geh. Regierungsrat Prof. Dr. med. et phil. M. Braun aus Anlass seines goldenen medizinischen Doktor-Jubilaums als Festgabe dargebracht von Schulern und Freunden. Grafe &amp; Unzer, Konigsberg (Kaliningrad), pp. 90 - 156." type="book chapter" year="1924">Speiser 1924</bibRefCitation>
</taxonomicName>
: Replacement name for
<taxonomicName id="4C46D2AA2C10FFA2FCA5656BFB9A042C" authority="Stein, 1910" authorityName="Stein" authorityYear="1910" box="[776,1036,1122,1148]" class="Insecta" family="Anthomyiidae" genus="Musca" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B932753B2C10FFA2FCA5656BFC10042C" box="[776,902,1122,1148]" italics="true" pageId="9" pageNumber="10">M. fasciata</emphasis>
<bibRefCitation id="EFD7D4D82C10FFA2FC20656AFB9A042C" author="Stein, P." box="[909,1036,1122,1148]" pageId="9" pageNumber="10" pagination="149 - 163" refId="ref36517" refString="Stein, P. (1910) The Percy Sladen Trust expedition to the Indian Ocean in 1905, under the leadership of Mr J. Stanley Gardiner. Vol. 3. No. 1 X. Diptera, Anthomyidae, mit den Gattungen Rhinia und Idiella. Transaction of The Linnaean Society of London, Zoology, Series 2, 14, 149 - 163. https: // doi. org / 10.1111 / j. 1096 - 3642.1910. tb 00528. x" type="journal article" year="1910">Stein, 1910</bibRefCitation>
</taxonomicName>
(junior homonym).
</paragraph>
<paragraph id="8BF9A9292C10FFA2FF6A658EFB030494" blockId="9.[151,1437,1050,1401]" pageId="9" pageNumber="10">
<emphasis id="B932753B2C10FFA2FF6A658EFE5304F0" bold="true" box="[199,453,1158,1184]" pageId="9" pageNumber="10">Specimens examined.</emphasis>
1f,
<materialsCitation id="3B2EA3742C10FFA2FE42658FFD6704F0" ID-GBIF-Occurrence="3017349444" box="[495,753,1158,1185]" collectorName="Al-Tatweer" location="Jazan, Fifa" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Jazan, Fifa">
<collectingRegion id="498267CB2C10FFA2FE42658FFDFC04F0" box="[495,618,1158,1185]" isEnumeration="true" pageId="9" pageNumber="10">Jazan, Fifa</collectingRegion>
,
<collectorName id="26B3CCFF2C10FFA2FDDF658EFD6704F0" box="[626,753,1158,1184]" pageId="9" pageNumber="10">Al-Tatweer</collectorName>
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<materialsCitation id="3B2EA3742C10FFA2FD5A658EFA0104F0" ID-GBIF-Occurrence="3017349328" box="[759,1431,1158,1185]" collectingDate="2015-09-03" collectingDateMax="2015-09-24" collectingDateMin="2015-09-03" collectionCode="CERS" collectorName="H. A. Dawah" location="Centre" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Centre">
<collectingRegion id="498267CB2C10FFA2FD5A658EFCD304F0" box="[759,837,1158,1184]" country="France" name="Centre" pageId="9" pageNumber="10">Centre</collectingRegion>
,
<date id="FFF88FE92C10FFA2FCE2658EFC7304F1" box="[847,997,1158,1185]" pageId="9" pageNumber="10" value="2015-09-03" valueMax="2015-09-24" valueMin="2015-09-03">
<collectingDate id="EFBC76012C10FFA2FCE2658EFC7304F1" box="[847,997,1158,1185]" pageId="9" pageNumber="10" value="2015-09-03" valueMax="2015-09-24" valueMin="2015-09-03">324.ix.2015</collectingDate>
</date>
,
<collectingMethod id="5207D13E2C10FFA2FC42658EFBEA04F1" box="[1007,1148,1158,1185]" pageId="9" pageNumber="10">Malaise trap</collectingMethod>
,
<collectorName id="26B3CCFF2C10FFA2FB2B658EFA8704F0" box="[1158,1297,1158,1184]" pageId="9" pageNumber="10">H.A. Dawah</collectorName>
(
<collectionCode id="ED5731EC2C10FFA2FA8D658EFAFA04F0" box="[1312,1388,1158,1184]" pageId="9" pageNumber="10">CERS</collectionCode>
); 1f
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,
<materialsCitation id="3B2EA3742C10FFA2FF3A65A2FB070494" ID-GBIF-Occurrence="3017349504" box="[151,1169,1194,1221]" collectingDate="2003-06-01" collectingDateMax="2003-06-17" collectingDateMin="2003-06-01" collectionCode="NMWC" collectorName="H. A. Dawah" location="Al-Hudaithy Fruit Farm" municipality="Maraba" pageId="9" pageNumber="10" specimenCount="1" stateProvince="'Asir">
<collectingRegion id="498267CB2C10FFA2FF3A65A2FF5F0494" box="[151,201,1194,1220]" country="Saudi Arabia" name="'Asir" pageId="9" pageNumber="10">Asir</collectingRegion>
,
<collectingMunicipality id="6B9D33532C10FFA2FF7965A2FEBA0494" box="[212,300,1194,1220]" pageId="9" pageNumber="10">Maraba</collectingMunicipality>
,
<location id="8E99FFF22C10FFA2FE9B65A2FDD00495" LSID="urn:lsid:plazi:treatment:03EF183F2C10FFA2FF3A6513FBC70528:8E99FFF22C10FFA2FE9B65A2FDD00495" box="[310,582,1194,1221]" municipality="Maraba" name="Al-Hudaithy Fruit Farm" pageId="9" pageNumber="10" stateProvince="'Asir">Al-Hudaithy Fruit Farm</location>
,
<date id="FFF88FE92C10FFA2FDFC65A2FD710494" box="[593,743,1194,1220]" pageId="9" pageNumber="10" value="2003-06-01" valueMax="2003-06-17" valueMin="2003-06-01">
<collectingDate id="EFBC76012C10FFA2FDFC65A2FD710494" box="[593,743,1194,1220]" pageId="9" pageNumber="10" value="2003-06-01" valueMax="2003-06-17" valueMin="2003-06-01">117.vi.2003</collectingDate>
</date>
,
<collectingMethod id="5207D13E2C10FFA2FD5E65A2FC170495" box="[755,897,1194,1221]" pageId="9" pageNumber="10">Malaise trap</collectingMethod>
,
<collectorName id="26B3CCFF2C10FFA2FC2165A2FB8F0494" box="[908,1049,1194,1220]" pageId="9" pageNumber="10">H.A. Dawah</collectorName>
(
<collectionCode id="ED5731EC2C10FFA2FB8565A2FB1F0494" box="[1064,1161,1194,1220]" country="United Kingdom" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="9" pageNumber="10" type="Museum">NMWC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35CFAA22C10FFA2FF6A65C6FBC70528" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BF9A9292C10FFA2FF6A65C6FCDB0560" blockId="9.[151,1437,1050,1401]" pageId="9" pageNumber="10">
<emphasis id="B932753B2C10FFA2FF6A65C6FEF704B8" bold="true" box="[199,353,1230,1256]" pageId="9" pageNumber="10">Distribution.</emphasis>
The species was previously recorded from
<collectingCountry id="F351E9B92C10FFA2FCF365C6FC6404B8" box="[862,1010,1230,1256]" name="Saudi Arabia" pageId="9" pageNumber="10">Saudi Arabia</collectingCountry>
by
<bibRefCitation id="EFD7D4D82C10FFA2FB8F65C7FB3804B8" author="Pont, A. C." box="[1058,1198,1230,1257]" pageId="9" pageNumber="10" refId="ref35020" refString="Pont, A. C. (1991) A review of the Fanniidae and Muscidae (Diptera) of the Arabian Peninsula. Fauna of Saudi Arabia, 12, 312 - 265." type="book" year="1991">Pont (1991)</bibRefCitation>
;
<bibRefCitation id="EFD7D4D82C10FFA2FB1165C6FF73055C" author="Dawah, H. A. &amp; Abdullah, M. A." pageId="9" pageNumber="10" pagination="373 - 396" refId="ref31297" refString="Dawah, H. A. &amp; Abdullah, M. A. (2009) The Muscidae (Diptera: Brachycera: Muscomorpha) of south-western Saudi Arabia. Fauna of Arabia, 24, 373 - 396." type="journal article" year="2009">Dawah &amp; Abdullah (2009)</bibRefCitation>
;
<bibRefCitation id="EFD7D4D82C10FFA2FF5C65FAFD87055C" author="El-Hawagry, M. S. &amp; Abdel-Dayem, M. S. &amp; Al Dhafer, H. M." box="[241,529,1266,1293]" pageId="9" pageNumber="10" pagination="1499 - 1530" refId="ref32135" refString="El-Hawagry, M. S., Abdel-Dayem, M. S., El-Sonbati &amp; Al Dhafer, H. M. (2017) A preliminary account of the fly fauna in Gard Raydah Nature Reserve, Kingdom of Saudi Arabia, with new records and biogeographical remarks (Diptera: Insecta). Journal of Natural History, 51 (25 - 26), 1499 - 1530. https: // doi. org / 10.1080 / 00222933.2017.1347299" type="journal article" year="2017">
El-Hawagry
<emphasis id="B932753B2C10FFA2FE2F65FBFE22055C" box="[386,436,1266,1292]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. (2017)
</bibRefCitation>
. It was described from
<collectingCountry id="F351E9B92C10FFA2FCB765FAFC03055C" box="[794,917,1266,1292]" name="Seychelles" pageId="9" pageNumber="10">Seychelles</collectingCountry>
. In the Middle East it has been recorded from
<collectingCountry id="F351E9B92C10FFA2FF3A641EFF4D0560" box="[151,219,1302,1328]" name="Oman" pageId="9" pageNumber="10">Oman</collectingCountry>
by
<bibRefCitation id="EFD7D4D82C10FFA2FEAB641FFE1C0560" author="Pont, A. C." box="[262,394,1302,1329]" pageId="9" pageNumber="10" refId="ref35020" refString="Pont, A. C. (1991) A review of the Fanniidae and Muscidae (Diptera) of the Arabian Peninsula. Fauna of Saudi Arabia, 12, 312 - 265." type="book" year="1991">Pont (1991)</bibRefCitation>
and from
<collectingCountry id="F351E9B92C10FFA2FE50641FFDDD0561" box="[509,587,1303,1329]" name="Yemen" pageId="9" pageNumber="10">Yemen</collectingCountry>
by
<bibRefCitation id="EFD7D4D82C10FFA2FDD8641FFCDC0560" author="van Emden, F. I." box="[629,842,1302,1329]" pageId="9" pageNumber="10" pagination="161 - 175" refId="ref37148" refString="van Emden, F. I. (1948) Diptera: Muscidae. In: Expedition to South-West Arabia 1937 - 1938. Vol. I. British Museum (Natural History), London, pp. 161 - 175." type="book chapter" year="1948">van Emden (1948)</bibRefCitation>
.
</paragraph>
<paragraph id="8BF9A9292C10FFA2FF6A6432FBC70528" blockId="9.[151,1437,1050,1401]" pageId="9" pageNumber="10">
<emphasis id="B932753B2C10FFA2FF6A6432FE240504" bold="true" box="[199,434,1338,1364]" pageId="9" pageNumber="10">Biological remarks.</emphasis>
It was recorded as a sweat fly on cattle and buffalo, it also feeds on blood (
<bibRefCitation id="EFD7D4D82C10FFA2FABB6432FA060504" author="Pont, A. C." box="[1302,1424,1338,1364]" pageId="9" pageNumber="10" refId="ref35020" refString="Pont, A. C. (1991) A review of the Fanniidae and Muscidae (Diptera) of the Arabian Peninsula. Fauna of Saudi Arabia, 12, 312 - 265." type="book" year="1991">Pont 1991</bibRefCitation>
). Larvae are only known to develop in cow dung (
<bibRefCitation id="EFD7D4D82C10FFA2FD1B6456FCF10529" author="Skidmore, P." box="[694,871,1374,1401]" pageId="9" pageNumber="10" refId="ref36277" refString="Skidmore, P. (1985) The Biology of the Muscidae of the World. Series Entomologica 29. Dr. W. Junk Publishers, Dordrecht, 550 pp." type="book" year="1985">Skidmore 1985</bibRefCitation>
;
<bibRefCitation id="EFD7D4D82C10FFA2FCDE6456FBD30528" author="Gregor, F. &amp; Rozkosny, R. &amp; Bartak, M. &amp; Vanhara, J." box="[883,1093,1374,1400]" pageId="9" pageNumber="10" pagination="1 - 280" refId="ref32614" refString="Gregor, F., Rozkosny, R., Bartak, M. &amp; Vanhara, J. (2002) The Muscidae (Diptera) of Central Europe. Folia Facultatis Scientiarum Naturalium, Universitatis Masarykianae Brunensis, Biologia, 107, 1 - 280." type="journal article" year="2002">
Gregor
<emphasis id="B932753B2C10FFA2FC646457FB970528" box="[969,1025,1374,1400]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
2002
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>