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<document id="6D612775EF3A0F1CF9CAF45F289569A5" ID-CLB-Dataset="31508" ID-DOI="10.11646/zootaxa.4079.4.3" ID-GBIF-Dataset="9e67a942-2592-41d5-a7e2-b7771e20ba9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050863" ID-ZooBank="4F1CC4B7-2A6F-4BB6-8F05-00650BDB001A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510811401326" checkinUser="plazi" docAuthor="Cranston, Peter S., Martin, Jon &amp; Spies, Martin" docDate="2016" docId="03CB87E7FF995258FF23FD25FCE0677B" docLanguage="en" docName="zootaxa.4079.4.3.pdf" docOrigin="Zootaxa 4079 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polypedilum quasinubifer Cranston &amp; Martin &amp; Spies 2016, sp. n." docType="treatment" docUuid="70DEF8F7-C4A2-445B-8550-B8417B29393F" docUuidSource="ZooBank" docVersion="8" lastPageNumber="441" masterDocId="FFF2FF9FFF925254FFB4FFE8FFDD6547" masterDocTitle="Cryptic species in the nuisance midge Polypedilum nubifer (Skuse) (Diptera: Chironomidae) and the status of Tripedilum Kieffer" masterLastPageNumber="447" masterPageNumber="429" pageNumber="440" updateTime="1698493448602" updateUser="ExternalLinkService">
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<mods:title id="6486D558290DFDE9109CCA1FAFC06AF9">Cryptic species in the nuisance midge Polypedilum nubifer (Skuse) (Diptera: Chironomidae) and the status of Tripedilum Kieffer</mods:title>
</mods:titleInfo>
<mods:name id="A2E742B2082D81E6119367BEED0A3222" type="personal">
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<mods:namePart id="8BB2280962CABD4DB545B68A356B9917">Cranston, Peter S.</mods:namePart>
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<mods:namePart id="2A1378A193AE956ADA9CECEA2D3EAFC1">Martin, Jon</mods:namePart>
</mods:name>
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<mods:namePart id="0FC870B1AA2D01A34A995DB580E17423">Spies, Martin</mods:namePart>
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<mods:title id="4583B477285B875D3E79B02FF3FE0C55">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4FF99A4A29E348E504F4C4829A38D08D">
<mods:date id="757C7AC428FECE71E6B199DEE23C80DA">2016</mods:date>
<mods:detail id="5E3E2F9AD18CEEE7521930023421D8D3" type="pubDate">
<mods:number id="37B9FFBD7C2D62B2D28B0B67861F852D">2016-02-15</mods:number>
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<mods:number id="ECD621010D2C41CEAA8A4FF82A5486D6">4079</mods:number>
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<mods:identifier id="7D746248F1026D3107CF7C59A3CC8ECB" type="DOI">10.11646/zootaxa.4079.4.3</mods:identifier>
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<treatment id="03CB87E7FF995258FF23FD25FCE0677B" ID-DOI="http://doi.org/10.5281/zenodo.6075641" ID-GBIF-Taxon="137307355" ID-Zenodo-Dep="6075641" LSID="urn:lsid:zoobank.org:act:70DEF8F7-C4A2-445B-8550-B8417B29393F" httpUri="http://treatment.plazi.org/id/03CB87E7FF995258FF23FD25FCE0677B" lastPageId="12" lastPageNumber="441" pageId="11" pageNumber="440">
<subSubSection id="C378657AFF99525FFF23FD25FD80664E" pageId="11" pageNumber="440" type="nomenclature">
<paragraph id="8BDD36F1FF99525FFF23FD25FD4B67A0" blockId="11.[151,954,717,813]" box="[151,662,717,743]" pageId="11" pageNumber="440">
<heading id="D095819DFF99525FFF23FD25FD4B67A0" bold="true" box="[151,662,717,743]" fontSize="11" level="1" pageId="11" pageNumber="440" reason="1">
<emphasis id="B916EAE3FF99525FFF23FD25FD4B67A0" bold="true" box="[151,662,717,743]" pageId="11" pageNumber="440">
<taxonomicName id="4C624D72FF99525FFF23FD25FD9167A0" ID-CoL="9HNW3" authority="Cranston" authorityName="Cranston &amp; Martin &amp; Spies" authorityYear="2016" box="[151,588,717,743]" class="Insecta" family="Chironomidae" genus="Polypedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="440" phylum="Arthropoda" rank="species" species="quasinubifer" status="sp. nov.">
<emphasis id="B916EAE3FF99525FFF23FD25FE0D67A0" bold="true" box="[151,464,717,743]" italics="true" pageId="11" pageNumber="440">Polypedilum quasinubifer</emphasis>
Cranston
</taxonomicName>
<taxonomicNameLabel id="A2255798FF99525FFDE0FD25FD4B67A0" box="[596,662,717,743]" pageId="11" pageNumber="440" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD36F1FF99525FFF23FD18FD80664E" blockId="11.[151,954,717,813]" box="[151,605,752,777]" pageId="11" pageNumber="440">
(
<figureCitation id="13592A74FF99525FFF2BFD18FEE6664F" box="[159,315,752,777]" captionStart="FIGURE 1" captionStartId="6.[151,250,1822,1844]" captionTargetBox="[262,1320,359,1789]" captionTargetId="figure@6.[249,1337,339,1798]" captionTargetPageId="6" captionText="FIGURE 1. Polypedilum spp., adults. A. Wing of female P. quasinubifer sp. n. B, C. Apex of fore tibia of: B. P. nubifer (Skuse), C. P. quasinubifer. D. Posterior abdomen of P. paranubifer sp. n. E G. Male genitalia of: E. P. nubife r (left: dorsal, right: stylised ventral plus apodemes in segment IX); F. P. quasinubifer (dorsal, left side only); G. P. paranubifer (T IX dorsomedial setae, anal point, superior volsella). H. Female genitalia of P. nubifer (ventral, parts of some paired structures omitted)." httpUri="https://zenodo.org/record/1051646/files/figure.png" pageId="11" pageNumber="440">Figs 1A, C, F</figureCitation>
; 2C, F; 3C; 4B, D, F; 5B)
</paragraph>
</subSubSection>
<subSubSection id="C378657AFF99525FFF23FCFCFC67666A" box="[151,954,788,813]" pageId="11" pageNumber="440" type="description">
<paragraph id="8BDD36F1FF99525FFF23FCFCFC67666A" blockId="11.[151,954,717,813]" box="[151,954,788,813]" pageId="11" pageNumber="440">
urn:lsid:zoobank.org:act:70DEF8F7-C4A2-445B-8550-B8417
<accessionNumber id="9431AB12FF99525FFCFBFCFCFC77666A" box="[847,938,788,813]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/B29393" pageId="11" pageNumber="440" type="EnaNcbi">B29393</accessionNumber>
F
</paragraph>
</subSubSection>
<subSubSection id="C378657AFF99525FFF23FCBDFEC46005" pageId="11" pageNumber="440" type="materials_examined">
<paragraph id="8BDD36F1FF99525FFF23FCBDFCB066D5" blockId="11.[151,1437,853,2034]" pageId="11" pageNumber="440">
<materialsCitation id="3B0A3CACFF99525FFF23FCBDFCB066D5" ID-GBIF-Occurrence="1700783278" collectingDate="1995-05-13" collectionCode="ANIC" country="Australia" county="Le" latitude="-18.35" location="Lawn Hill National Park" longLatPrecision="1275" longitude="138.15" municipality="Pe" pageId="11" pageNumber="440" specimenCount="1" specimenCount-male="1" stateProvince="Queensland" typeStatus="holotype">
<emphasis id="B916EAE3FF99525FFF23FCBDFE126629" bold="true" box="[151,463,853,878]" pageId="11" pageNumber="440">
<typeStatus id="54D98853FF99525FFF23FCBDFF0F6629" box="[151,210,853,878]" pageId="11" pageNumber="440">Type</typeStatus>
material.
<typeStatus id="54D98853FF99525FFED6FCBDFE126629" box="[354,463,853,878]" pageId="11" pageNumber="440" type="holotype">Holotype</typeStatus>
</emphasis>
,
<collectingCounty id="62BC4E7DFF99525FFE53FCBEFDDB6629" box="[487,518,854,878]" pageId="11" pageNumber="440">Le</collectingCounty>
/
<collectingMunicipality id="6BB9AC8BFF99525FFDBBFCBEFDF06629" box="[527,557,854,878]" pageId="11" pageNumber="440">Pe</collectingMunicipality>
/
<specimenCount id="9D64FD78FF99525FFD82FCBEFD946629" box="[566,585,854,878]" pageId="11" pageNumber="440" type="male"></specimenCount>
,
<collectingCountry id="F3757661FF99525FFDEBFCBEFCDF6629" box="[607,770,854,878]" name="Australia" pageId="11" pageNumber="440">AUSTRALIA</collectingCountry>
:
<collectingRegion id="49A6F813FF99525FFCA3FCBDFC436629" box="[791,926,853,878]" country="Australia" name="Queensland" pageId="11" pageNumber="440">Queensland</collectingRegion>
,
<location id="8EBD602AFF99525FFC01FCBEFB306629" LSID="urn:lsid:plazi:treatment:03CB87E7FF995258FF23FD25FCE0677B:8EBD602AFF99525FFC01FCBEFB306629" box="[949,1261,853,878]" country="Australia" county="Le" latitude="-18.35" longLatPrecision="1275" longitude="138.15" municipality="Pe" name="Lawn Hill National Park" pageId="11" pageNumber="440" stateProvince="Queensland">Lawn Hill National Park</location>
,
<location id="8EBD602AFF99525FFAB7FCBDFA456629" LSID="urn:lsid:plazi:treatment:03CB87E7FF995258FF23FD25FCE0677B:8EBD602AFF99525FFAB7FCBDFA456629" box="[1283,1432,853,878]" country="Australia" county="Le" latitude="-18.35" longLatPrecision="1275" longitude="138.15" municipality="Pe" name="Musselbrook" pageId="11" pageNumber="440" stateProvince="Queensland">Musselbrook</location>
,
<location id="8EBD602AFF99525FFF23FC91FEE966D5" LSID="urn:lsid:plazi:treatment:03CB87E7FF995258FF23FD25FCE0677B:8EBD602AFF99525FFF23FC91FEE966D5" box="[151,308,889,914]" country="Australia" county="Le" latitude="-18.35" longLatPrecision="1275" longitude="138.15" municipality="Pe" name="Amphitheatre" pageId="11" pageNumber="440" stateProvince="Queensland">Amphitheatre</location>
,
<geoCoordinate id="EE565036FF99525FFE8BFC92FE4666D5" box="[319,411,890,914]" direction="south" orientation="latitude" pageId="11" pageNumber="440" precision="925" value="-18.35">18°21S</geoCoordinate>
<geoCoordinate id="EE565036FF99525FFE16FC92FDD666D5" box="[418,523,890,914]" direction="east" orientation="longitude" pageId="11" pageNumber="440" precision="925" value="138.15">138°09'E</geoCoordinate>
,
<collectingDate id="EF98E9D9FF99525FFDA3FC91FD5B66D5" box="[535,646,889,914]" pageId="11" pageNumber="440" value="1995-05-13">
<date id="FFDC1031FF99525FFDA3FC91FD5B66D5" box="[535,646,889,914]" pageId="11" pageNumber="440" value="1995-05-13">13.v.1995</date>
</collectingDate>
(
<location id="8EBD602AFF99525FFD21FC92FD2066D6" LSID="urn:lsid:plazi:treatment:03CB87E7FF995258FF23FD25FCE0677B:8EBD602AFF99525FFD21FC92FD2066D6" box="[661,765,890,913]" country="Australia" county="Le" latitude="-18.35" longLatPrecision="1275" longitude="138.15" municipality="Pe" name="Cranston" pageId="11" pageNumber="440" stateProvince="Queensland">
<emphasis id="B916EAE3FF99525FFD21FC92FD2066D6" box="[661,765,890,913]" italics="true" pageId="11" pageNumber="440">Cranston</emphasis>
</location>
) (
<collectionCode id="ED73AE34FF99525FFCA2FC92FCBC66D5" box="[790,865,890,914]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="440">ANIC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BDD36F1FF99525FFF73FC75FCC2669D" blockId="11.[151,1437,853,2034]" pageId="11" pageNumber="440">
<typeStatus id="54D98853FF99525FFF73FC75FEE266F1" box="[199,319,925,950]" pageId="11" pageNumber="440" type="paratype">
<emphasis id="B916EAE3FF99525FFF73FC75FEE266F1" bold="true" box="[199,319,925,950]" pageId="11" pageNumber="440">Paratypes</emphasis>
</typeStatus>
, Le/Pe/
<specimenCount id="9D64FD78FF99525FFE14FC75FE6E66F2" box="[416,435,925,949]" pageId="11" pageNumber="440" type="male"></specimenCount>
, Le/Pe/
<specimenCount id="9D64FD78FF99525FFDA2FC75FDF966F2" box="[534,548,925,949]" pageId="11" pageNumber="440" type="female"></specimenCount>
, as
<typeStatus id="54D98853FF99525FFDEDFC75FD1866F2" box="[601,709,925,949]" pageId="11" pageNumber="440" type="holotype">Holotype</typeStatus>
; Pe,
<collectingRegion id="49A6F813FF99525FFCB0FC75FC5666F2" box="[772,907,925,949]" country="Australia" name="Queensland" pageId="11" pageNumber="440">Queensland</collectingRegion>
, Lawn Hill N.P., Musselbrook, Border Pool,
<geoCoordinate id="EE565036FF99525FFF23FC29FEC6669D" box="[151,283,961,986]" direction="south" orientation="latitude" pageId="11" pageNumber="440" precision="15" value="-18.614723">18°3653”S</geoCoordinate>
137°5925”'E,
<date id="FFDC1031FF99525FFE7DFC29FDE5669D" box="[457,568,961,986]" pageId="11" pageNumber="440" value="1995-05-14">14.v.1995</date>
(
<emphasis id="B916EAE3FF99525FFDFCFC2AFD6D669E" box="[584,688,962,985]" italics="true" pageId="11" pageNumber="440">Cranston</emphasis>
) (
<collectionCode id="ED73AE34FF99525FFD7CFC2AFCCE669D" box="[712,787,962,986]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="440">ANIC</collectionCode>
).
</paragraph>
<paragraph id="8BDD36F1FF99525FFF73FC0DFBC26101" blockId="11.[151,1437,853,2034]" pageId="11" pageNumber="440">
<materialsCitation id="3B0A3CACFF99525FFF73FC0DFBC16101" ID-GBIF-Occurrence="1700783298" collectingDate="2013-09-11" collectionCode="ANIC, RMBR-MV" country="Singapore" county="Bedok Reservoir" latitude="1.3463889" location="Other" longLatPrecision="21" longitude="103.92528" municipality="Upper Seletar Reservoir" pageId="11" pageNumber="440" specimenCount="1">
<emphasis id="B916EAE3FF99525FFF73FC0DFE2266B9" bold="true" box="[199,511,997,1022]" pageId="11" pageNumber="440">
<location id="8EBD602AFF99525FFF73FC0DFED266B9" LSID="urn:lsid:plazi:treatment:03CB87E7FF995258FF23FD25FCE0677B:8EBD602AFF99525FFF73FC0DFED266B9" box="[199,271,997,1022]" country="Singapore" county="Bedok Reservoir" latitude="1.3463889" longLatPrecision="21" longitude="103.92528" municipality="Upper Seletar Reservoir" name="Other" pageId="11" pageNumber="440">Other</location>
material examined
</emphasis>
.
<collectingCountry id="F3757661FF99525FFDA6FC0EFD6966B9" box="[530,692,998,1022]" name="Singapore" pageId="11" pageNumber="440">SINGAPORE</collectingCountry>
:
<collectingMunicipality id="6BB9AC8BFF99525FFD77FC0EFC0066B9" box="[707,989,997,1022]" pageId="11" pageNumber="440">Upper Seletar Reservoir</collectingMunicipality>
,
<geoCoordinate id="EE565036FF99525FFC5AFC0EFBBA66B9" box="[1006,1127,998,1022]" direction="north" orientation="latitude" pageId="11" pageNumber="440" precision="15" value="1.4027778">1°2410&quot;N</geoCoordinate>
<geoCoordinate id="EE565036FF99525FFBC0FC0EFAD866B9" box="[1140,1285,998,1022]" direction="east" orientation="longitude" pageId="11" pageNumber="440" precision="15" value="103.8075">103°48'27&quot;E</geoCoordinate>
,
<collectingDate id="EF98E9D9FF99525FFAA2FC0DFE546165" isEnumeration="true" pageId="11" pageNumber="440">
<date id="FFDC1031FF99525FFAA2FC0DFA4566B9" box="[1302,1432,997,1022]" pageId="11" pageNumber="440" value="2012-07-29">29.vii.2012</date>
,
<date id="FFDC1031FF99525FFF23FBE1FEC46165" box="[151,281,1033,1058]" pageId="11" pageNumber="440" value="2013-07-16">16.vii.2013</date>
,
<date id="FFDC1031FF99525FFE93FBE2FE546165" box="[295,393,1034,1058]" pageId="11" pageNumber="440" value="2013-10-08">8.x.2013</date>
</collectingDate>
(
<emphasis id="B916EAE3FF99525FFE2FFBE2FDD26166" box="[411,527,1034,1057]" italics="true" pageId="11" pageNumber="440">NUS team</emphasis>
) (all life stages);
<collectingCounty id="62BC4E7DFF99525FFD6BFBE1FC7C6165" box="[735,929,1033,1058]" pageId="11" pageNumber="440">Bedok Reservoir</collectingCounty>
,
<geoCoordinate id="EE565036FF99525FFC1AFBE2FBF46165" box="[942,1065,1034,1058]" direction="north" orientation="latitude" pageId="11" pageNumber="440" precision="15" value="1.3463889">1°2047”N</geoCoordinate>
<geoCoordinate id="EE565036FF99525FFB86FBE1FB146165" box="[1074,1225,1033,1058]" direction="east" orientation="longitude" pageId="11" pageNumber="440" precision="15" value="103.92528">103°5531”E</geoCoordinate>
,
<collectingDate id="EF98E9D9FF99525FFB63FBE1FA926165" box="[1239,1359,1033,1058]" pageId="11" pageNumber="440" value="2013-09-11">
<date id="FFDC1031FF99525FFB63FBE1FA926165" box="[1239,1359,1033,1058]" pageId="11" pageNumber="440" value="2013-09-11">11.ix.2013</date>
</collectingDate>
, (
<emphasis id="B916EAE3FF99525FFAD3FBE2FF0B6102" italics="true" pageId="11" pageNumber="440">NUS team)</emphasis>
(all life stages) (some of each stage to
<collectionCode id="ED73AE34FF99525FFD38FBC6FD0E6101" box="[652,723,1070,1094]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="440">ANIC</collectionCode>
, remainder to
<collectionCode id="ED73AE34FF99525FFCC0FBC6FBC96101" box="[884,1044,1070,1094]" pageId="11" pageNumber="440">RMBR—MV</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDD36F1FF99525FFF73FBBAFEC46005" blockId="11.[151,1437,853,2034]" pageId="11" pageNumber="440">
<materialsCitation id="3B0A3CACFF99525FFF73FBBAFEAE61CA" ID-GBIF-Occurrence="1700783258" collectingDate="2004-05-15" collectorName="Amphur Doi Saket &amp; Nam Kuang" country="Thailand" latitude="19.0" location="Chiang Mai" longLatPrecision="1273" longitude="99.28333" pageId="11" pageNumber="440" specimenCount="554" stateProvince="Chiang Mai">
<collectingCountry id="F3757661FF99525FFF73FBBAFE81612D" box="[199,348,1106,1130]" name="Thailand" pageId="11" pageNumber="440">THAILAND</collectingCountry>
: single L from each of:
<collectingRegion id="49A6F813FF99525FFD3BFBB9FCC3612D" box="[655,798,1105,1130]" country="Thailand" name="Chiang Mai" pageId="11" pageNumber="440">Chiang Mai</collectingRegion>
,
<collectorName id="26975327FF99525FFC84FBB9FBCE612D" box="[816,1043,1105,1130]" pageId="11" pageNumber="440">Amphur Doi Saket</collectorName>
,
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15.v.
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,
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L-689 (
<emphasis id="B916EAE3FF99525FFE57FB9DFD2761CA" box="[483,762,1141,1166]" italics="true" pageId="11" pageNumber="440">Vitheepradit &amp; Prommi</emphasis>
);
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,
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,
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<date id="FFDC1031FF99525FFB38FB9DFB2361C9" box="[1164,1278,1141,1166]" pageId="11" pageNumber="440" value="2004-05-15">15.v.2004</date>
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(
<emphasis id="B916EAE3FF99525FFAA2FB9DFA4F61C9" box="[1302,1426,1141,1166]" italics="true" pageId="11" pageNumber="440">DuCharme</emphasis>
)
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, Amphur Khao Wong, Tad Tong Waterfall,
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,
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<quantity id="4C9A9B14FF99525FFC7EFB72FBC561F6" box="[970,1048,1178,1202]" metricMagnitude="2" metricUnit="m" metricValue="2.04" pageId="11" pageNumber="440" unit="m" value="204.0">204 m</quantity>
</elevation>
.
</specimenCount>
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, TH62 (
<emphasis id="B916EAE3FF99525FFBC8FB71FADB61F5" box="[1148,1286,1177,1202]" italics="true" pageId="11" pageNumber="440">Vitheepradit</emphasis>
); Petchabun, Khao Khor N.P., stream from Tamtip Waterfall, 16°39.594N 101°07.901E,
<quantity id="4C9A9B14FF99525FFB9DFB56FBA76192" box="[1065,1146,1214,1238]" metricMagnitude="2" metricUnit="m" metricValue="1.8199999999999998" pageId="11" pageNumber="440" unit="m" value="182.0">182 m</quantity>
.,
<date id="FFDC1031FF99525FFB24FB56FADD6191" box="[1168,1280,1214,1238]" pageId="11" pageNumber="440" value="2004-05-11">11.v.2004</date>
, L-678, TH5 (
<emphasis id="B916EAE3FF99525FFF14FB09FE7361BE" box="[160,430,1249,1274]" italics="true" pageId="11" pageNumber="440">Vitheepradit &amp; Prommi</emphasis>
);
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<collectingRegion id="49A6F813FF99525FFE7DFB09FD8661BD" box="[457,603,1249,1274]" country="Thailand" name="Chaiyaphum" pageId="11" pageNumber="440">Chaiyaphum</collectingRegion>
,
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,
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,
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,
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<elevation id="004FD1C2FF99525FFB6DFB09FAF461BE" box="[1241,1321,1249,1274]" metricMagnitude="2" metricUnit="m" metricValue="5.89" pageId="11" pageNumber="440" unit="m" value="589.0">
<quantity id="4C9A9B14FF99525FFB6DFB09FAF461BE" box="[1241,1321,1249,1274]" metricMagnitude="2" metricUnit="m" metricValue="5.89" pageId="11" pageNumber="440" unit="m" value="589.0">589 m</quantity>
</elevation>
.
</specimenCount>
,
<collectingDate id="EF98E9D9FF99525FFA88FB0AFA4161BD" box="[1340,1436,1250,1274]" pageId="11" pageNumber="440" value="2004-05-02">
<date id="FFDC1031FF99525FFA88FB0AFA4161BD" box="[1340,1436,1250,1274]" pageId="11" pageNumber="440" value="2004-05-02">2.v.2004</date>
</collectingDate>
(
<emphasis id="B916EAE3FF99525FFF14FAEDFEC16059" box="[160,284,1285,1310]" italics="true" pageId="11" pageNumber="440">DuCharme</emphasis>
)
</materialsCitation>
;
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<collectingRegion id="49A6F813FF99525FFE80FAEDFE186059" box="[308,453,1285,1310]" country="Thailand" name="Prachin Buri" pageId="11" pageNumber="440">Prachin Buri</collectingRegion>
,
<collectorName id="26975327FF99525FFE64FAEDFDAF605A" box="[464,626,1285,1310]" pageId="11" pageNumber="440">Khao Yai N.P.</collectorName>
,
<collectorName id="26975327FF99525FFDC9FAEEFCBD6059" box="[637,864,1285,1310]" pageId="11" pageNumber="440">Kaeng Wang Thong</collectorName>
,
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<geoCoordinate id="EE565036FF99525FFBB0FAEEFB7D6059" box="[1028,1184,1286,1310]" direction="east" orientation="longitude" pageId="11" pageNumber="440" precision="1" value="101.72366">101°43.420E</geoCoordinate>
(
<emphasis id="B916EAE3FF99525FFB04FAEEFA4E6059" box="[1200,1427,1285,1310]" italics="true" pageId="11" pageNumber="440">Sites &amp; Vitheepradit</emphasis>
) (all
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C378657AFF995258FF73FAA5FCE0677B" lastPageId="12" lastPageNumber="441" pageId="11" pageNumber="440" type="description">
<paragraph id="8BDD36F1FF99525FFF73FAA5FDC663B5" blockId="11.[151,1437,853,2034]" pageId="11" pageNumber="440">
<emphasis id="B916EAE3FF99525FFF73FAA5FE846021" bold="true" box="[199,345,1357,1382]" pageId="11" pageNumber="440">Description.</emphasis>
Adults (
<figureCitation id="13592A74FF99525FFE0EFAA5FD8A6022" box="[442,599,1357,1382]" captionStart="FIGURE 1" captionStartId="6.[151,250,1822,1844]" captionTargetBox="[262,1320,359,1789]" captionTargetId="figure@6.[249,1337,339,1798]" captionTargetPageId="6" captionText="FIGURE 1. Polypedilum spp., adults. A. Wing of female P. quasinubifer sp. n. B, C. Apex of fore tibia of: B. P. nubifer (Skuse), C. P. quasinubifer. D. Posterior abdomen of P. paranubifer sp. n. E G. Male genitalia of: E. P. nubife r (left: dorsal, right: stylised ventral plus apodemes in segment IX); F. P. quasinubifer (dorsal, left side only); G. P. paranubifer (T IX dorsomedial setae, anal point, superior volsella). H. Female genitalia of P. nubifer (ventral, parts of some paired structures omitted)." httpUri="https://zenodo.org/record/1051646/files/figure.png" pageId="11" pageNumber="440">Figs 1A, C, F</figureCitation>
,
<figureCitation id="13592A74FF99525FFDD7FAA6FD5B6021" box="[611,646,1358,1382]" captionStart="FIGURE 2" captionStartId="7.[151,250,1007,1029]" captionTargetBox="[348,1257,196,979]" captionTargetId="figure@7.[335,1275,193,984]" captionTargetPageId="7" captionText="FIGURE 2. Polypedilum spp. A C. Female genitalia, gonapophysis VIII of: A. P. nubifer (Skuse); B. P. paranubifer; C. P. quasinubifer; DmL—dorsomesal lobe, VlL—ventrolateral lobe. D F. Pupae, posterolateral corner of abdominal segment VIII (ventral) of: D. P. nubifer; E. P. paranubifer, F. P. quasinubifer; l — 4 th (posterior-most) lateral seta. Scale bars = 100 µm." httpUri="https://zenodo.org/record/1051648/files/figure.png" pageId="11" pageNumber="440">2C</figureCitation>
,
<figureCitation id="13592A74FF99525FFD27FAA6FD6B6021" box="[659,694,1358,1382]" captionStart="FIGURE 3" captionStartId="8.[151,250,1785,1807]" captionTargetBox="[250,1315,201,1730]" captionTargetId="figure@8.[235,1327,193,1742]" captionTargetPageId="8" captionText="FIGURE 3. Polypedilum spp., pupae. A. Cephalic tubercles, frontal setae of P. nubifer (Skuse). B, C. Abdomen (dorsal) of: B. P. nubifer, C. P. quasinubifer sp. n. D, E. Posterolateral corner of abdominal segment VIII (ventral) of: D. P. nubifer; E. P. paranubifer sp. n." httpUri="https://zenodo.org/record/1051650/files/figure.png" pageId="11" pageNumber="440">3B</figureCitation>
). For mensural features see
<tableCitation id="C6E0034AFF99525FFC42FAA5FB976021" box="[1014,1098,1357,1382]" captionStart="TABLE 1" captionStartId="3.[125,213,741,764]" captionText="TABLE 1. Mensural features of Polypedilum spp. adults. n / a = not visible; n / s not stated (by author)." pageId="11" pageNumber="440">Table 1</tableCitation>
. Frontal tubercles very small, 810 µm long. Foretibial apex triangularly tapering without apical spine (
<figureCitation id="13592A74FF99525FFC65FA99FBF560CD" box="[977,1064,1393,1418]" captionStart="FIGURE 1" captionStartId="6.[151,250,1822,1844]" captionTargetBox="[262,1320,359,1789]" captionTargetId="figure@6.[249,1337,339,1798]" captionTargetPageId="6" captionText="FIGURE 1. Polypedilum spp., adults. A. Wing of female P. quasinubifer sp. n. B, C. Apex of fore tibia of: B. P. nubifer (Skuse), C. P. quasinubifer. D. Posterior abdomen of P. paranubifer sp. n. E G. Male genitalia of: E. P. nubife r (left: dorsal, right: stylised ventral plus apodemes in segment IX); F. P. quasinubifer (dorsal, left side only); G. P. paranubifer (T IX dorsomedial setae, anal point, superior volsella). H. Female genitalia of P. nubifer (ventral, parts of some paired structures omitted)." httpUri="https://zenodo.org/record/1051646/files/figure.png" pageId="11" pageNumber="440">Fig. 1C</figureCitation>
). Pulvillus (
<figureCitation id="13592A74FF99525FFB05FA99FADB60CD" box="[1201,1286,1393,1418]" captionStart="FIGURE 5" captionStartId="10.[151,250,1100,1122]" captionTargetBox="[350,1233,201,1059]" captionTargetId="figure@10.[340,1247,193,1079]" captionTargetPageId="10" captionText="FIGURE 5. Polypedilum spp., adult pulvilli: of A. P. nubifer (Skuse), B. P. quasinubifer sp. n. C. P. longiforceps Kieffer, D. P. lobiferum Freeman." httpUri="https://zenodo.org/record/1051654/files/figure.png" pageId="11" pageNumber="440">Fig. 5B</figureCitation>
) divided in 2 parts; outer plumose but not forming pad, extending to mid-claw; inner part simple, unbranched, subequal in length to outer; empodium sparsely plumose, extending beyond pulvillus apex. Wing pattern very faint on (teneral)
<typeStatus id="54D98853FF99525FFF23FA36FF1A60B1" box="[151,199,1502,1526]" pageId="11" pageNumber="440">type</typeStatus>
specimens, variably to strongly developed on those from
<collectingCountry id="F3757661FF99525FFCE3FA35FC1460B1" box="[855,969,1501,1526]" name="Singapore" pageId="11" pageNumber="440">Singapore</collectingCountry>
(
<figureCitation id="13592A74FF99525FFC6CFA35FBEF60B1" box="[984,1074,1501,1526]" captionStart="FIGURE 1" captionStartId="6.[151,250,1822,1844]" captionTargetBox="[262,1320,359,1789]" captionTargetId="figure@6.[249,1337,339,1798]" captionTargetPageId="6" captionText="FIGURE 1. Polypedilum spp., adults. A. Wing of female P. quasinubifer sp. n. B, C. Apex of fore tibia of: B. P. nubifer (Skuse), C. P. quasinubifer. D. Posterior abdomen of P. paranubifer sp. n. E G. Male genitalia of: E. P. nubife r (left: dorsal, right: stylised ventral plus apodemes in segment IX); F. P. quasinubifer (dorsal, left side only); G. P. paranubifer (T IX dorsomedial setae, anal point, superior volsella). H. Female genitalia of P. nubifer (ventral, parts of some paired structures omitted)." httpUri="https://zenodo.org/record/1051646/files/figure.png" pageId="11" pageNumber="440">Fig. 1A</figureCitation>
). Male T VIII waisted (tapering anteriorly); TIX with cluster of setae not lying within delimited paler area; anal point narrow. Male genitalia (
<figureCitation id="13592A74FF99525FFADAF9E9FF696379" captionStart="FIGURE 1" captionStartId="6.[151,250,1822,1844]" captionTargetBox="[262,1320,359,1789]" captionTargetId="figure@6.[249,1337,339,1798]" captionTargetPageId="6" captionText="FIGURE 1. Polypedilum spp., adults. A. Wing of female P. quasinubifer sp. n. B, C. Apex of fore tibia of: B. P. nubifer (Skuse), C. P. quasinubifer. D. Posterior abdomen of P. paranubifer sp. n. E G. Male genitalia of: E. P. nubife r (left: dorsal, right: stylised ventral plus apodemes in segment IX); F. P. quasinubifer (dorsal, left side only); G. P. paranubifer (T IX dorsomedial setae, anal point, superior volsella). H. Female genitalia of P. nubifer (ventral, parts of some paired structures omitted)." httpUri="https://zenodo.org/record/1051646/files/figure.png" pageId="11" pageNumber="440">Fig. 1F</figureCitation>
) with conventional gonocoxite and slender gonostylus tapering to blunt apex. Superior volsella curved, shorter and stouter than in
<taxonomicName id="4C624D72FF99525FFECCF9A2FE3B6325" box="[376,486,1609,1634]" class="Insecta" family="Chironomidae" genus="Chironomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="440" phylum="Arthropoda" rank="species" species="nubifer">
<emphasis id="B916EAE3FF99525FFECCF9A2FE3B6325" box="[376,486,1609,1634]" italics="true" pageId="11" pageNumber="440">P. nubifer</emphasis>
</taxonomicName>
, bearing dorsolateral seta inserted at about 2/3 length of volsella. Inferior volsella slender, with apical seta arising from weakly bilobed apex, scarcely longer than any other in sparse cluster of setae. Female genitalia: gonocoxite IX with 711 setae. Lobes of gonapophysis VIII (
<figureCitation id="13592A74FF99525FFB95F979FBA163ED" box="[1057,1148,1681,1706]" captionStart="FIGURE 2" captionStartId="7.[151,250,1007,1029]" captionTargetBox="[348,1257,196,979]" captionTargetId="figure@7.[335,1275,193,984]" captionTargetPageId="7" captionText="FIGURE 2. Polypedilum spp. A C. Female genitalia, gonapophysis VIII of: A. P. nubifer (Skuse); B. P. paranubifer; C. P. quasinubifer; DmL—dorsomesal lobe, VlL—ventrolateral lobe. D F. Pupae, posterolateral corner of abdominal segment VIII (ventral) of: D. P. nubifer; E. P. paranubifer, F. P. quasinubifer; l — 4 th (posterior-most) lateral seta. Scale bars = 100 µm." httpUri="https://zenodo.org/record/1051648/files/figure.png" pageId="11" pageNumber="440">Fig. 2C</figureCitation>
): ventrolateral lobe squat (1520 µm) bearing short setae; dorsomedial lobe no more than a rounded microtrichiose contour without setae, c.
<date id="FFDC1031FF99525FFF23F932FF1563B6" box="[151,200,1754,1778]" pageId="11" pageNumber="440" value="1990-10">90 x</date>
125 µm, rounded rectangular.
</paragraph>
<paragraph id="8BDD36F1FF99525FFF73F916FAF762AA" blockId="11.[151,1437,853,2034]" pageId="11" pageNumber="440">
Pupa (
<figureCitation id="13592A74FF99525FFEA7F915FEAD6251" box="[275,368,1789,1814]" captionStart="FIGURE 2" captionStartId="7.[151,250,1007,1029]" captionTargetBox="[348,1257,196,979]" captionTargetId="figure@7.[335,1275,193,984]" captionTargetPageId="7" captionText="FIGURE 2. Polypedilum spp. A C. Female genitalia, gonapophysis VIII of: A. P. nubifer (Skuse); B. P. paranubifer; C. P. quasinubifer; DmL—dorsomesal lobe, VlL—ventrolateral lobe. D F. Pupae, posterolateral corner of abdominal segment VIII (ventral) of: D. P. nubifer; E. P. paranubifer, F. P. quasinubifer; l — 4 th (posterior-most) lateral seta. Scale bars = 100 µm." httpUri="https://zenodo.org/record/1051648/files/figure.png" pageId="11" pageNumber="440">Figs 2F</figureCitation>
,
<figureCitation id="13592A74FF99525FFE35F916FE786251" box="[385,421,1790,1814]" captionStart="FIGURE 3" captionStartId="8.[151,250,1785,1807]" captionTargetBox="[250,1315,201,1730]" captionTargetId="figure@8.[235,1327,193,1742]" captionTargetPageId="8" captionText="FIGURE 3. Polypedilum spp., pupae. A. Cephalic tubercles, frontal setae of P. nubifer (Skuse). B, C. Abdomen (dorsal) of: B. P. nubifer, C. P. quasinubifer sp. n. D, E. Posterolateral corner of abdominal segment VIII (ventral) of: D. P. nubifer; E. P. paranubifer sp. n." httpUri="https://zenodo.org/record/1051650/files/figure.png" pageId="11" pageNumber="440">3C</figureCitation>
). Exuviae 5.0
<quantity id="4C9A9B14FF99525FFDEDF915FD696252" box="[601,692,1789,1814]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="11" pageNumber="440" unit="mm" value="5.5">5.5 mm</quantity>
long, pale with yellow highlights and yellow-brown comb on abdominal segment VIII. Cephalic tubercles triangular, 2535 µm long and about equally wide at base, bearing 40 55 µm long frontal seta inserted basally on tubercle. Thoracic horn plumose, arising from oval, golden basal ring. Thorax mid-dorsally with dense cluster of small tubercles. Abdominal tergite spine/spinule pattern and setation as in
<figureCitation id="13592A74FF99525FFF00F865FED162E1" box="[180,268,1933,1958]" captionStart="FIGURE 3" captionStartId="8.[151,250,1785,1807]" captionTargetBox="[250,1315,201,1730]" captionTargetId="figure@8.[235,1327,193,1742]" captionTargetPageId="8" captionText="FIGURE 3. Polypedilum spp., pupae. A. Cephalic tubercles, frontal setae of P. nubifer (Skuse). B, C. Abdomen (dorsal) of: B. P. nubifer, C. P. quasinubifer sp. n. D, E. Posterolateral corner of abdominal segment VIII (ventral) of: D. P. nubifer; E. P. paranubifer sp. n." httpUri="https://zenodo.org/record/1051650/files/figure.png" pageId="11" pageNumber="440">Fig. 3C</figureCitation>
. Hook row on segment II c. 4550% of tergite width, with c. 3042 hooklets. Caudolateral spur on VIII (
<figureCitation id="13592A74FF99525FFF2BF859FF2A628D" box="[159,247,1969,1994]" captionStart="FIGURE 2" captionStartId="7.[151,250,1007,1029]" captionTargetBox="[348,1257,196,979]" captionTargetId="figure@7.[335,1275,193,984]" captionTargetPageId="7" captionText="FIGURE 2. Polypedilum spp. A C. Female genitalia, gonapophysis VIII of: A. P. nubifer (Skuse); B. P. paranubifer; C. P. quasinubifer; DmL—dorsomesal lobe, VlL—ventrolateral lobe. D F. Pupae, posterolateral corner of abdominal segment VIII (ventral) of: D. P. nubifer; E. P. paranubifer, F. P. quasinubifer; l — 4 th (posterior-most) lateral seta. Scale bars = 100 µm." httpUri="https://zenodo.org/record/1051648/files/figure.png" pageId="11" pageNumber="440">Figs 2F</figureCitation>
,
<figureCitation id="13592A74FF99525FFEB1F85AFEF8628D" box="[261,293,1970,1994]" captionStart="FIGURE 3" captionStartId="8.[151,250,1785,1807]" captionTargetBox="[250,1315,201,1730]" captionTargetId="figure@8.[235,1327,193,1742]" captionTargetPageId="8" captionText="FIGURE 3. Polypedilum spp., pupae. A. Cephalic tubercles, frontal setae of P. nubifer (Skuse). B, C. Abdomen (dorsal) of: B. P. nubifer, C. P. quasinubifer sp. n. D, E. Posterolateral corner of abdominal segment VIII (ventral) of: D. P. nubifer; E. P. paranubifer sp. n." httpUri="https://zenodo.org/record/1051650/files/figure.png" pageId="11" pageNumber="440">3E</figureCitation>
) well-developed, with dominant (apical-most) spine and few significantly smaller subsidiary spines not extending to L4 setal base. Anal lobe fringe evenly uniserial, with 2330 taeniae, without dorsal seta.
</paragraph>
<paragraph id="8BDD36F1FF9E5258FF73FF70FD1B6407" blockId="12.[151,1437,151,573]" pageId="12" pageNumber="441">
Larva (
<figureCitation id="13592A74FF9E5258FEA3FF7FFE6D65F7" box="[279,432,151,176]" captionStart="FIGURE 4" captionStartId="9.[151,250,1973,1995]" captionTargetBox="[236,1332,1010,1913]" captionTargetId="figure@9.[229,1340,999,1924]" captionTargetPageId="9" captionText="FIGURE 4. Polypedilum spp., larvae. A, B. Antenna of: A. P. nubifer (Skuse), B. P. quasinubifer sp. n. C, D. Mentum of: C. P. nubifer, D. P. quasinubifer. E, F. Lateral end of ventromental plate of: E. P. nubifer, F. P. quasinubifer." httpUri="https://zenodo.org/record/1051652/files/figure.png" pageId="12" pageNumber="441">Fig. 4B, D, F</figureCitation>
). For mensural features see
<tableCitation id="C6E0034AFF9E5258FD5DFF7FFCE265F7" box="[745,831,151,176]" captionStart="TABLE 2" captionStartId="4.[120,208,740,763]" captionText="TABLE 2. Mensural features of Polypedilum spp. larvae. n / a = not visible; n / s = not stated (by author)." pageId="12" pageNumber="441">Table 2</tableCitation>
. Head yellow, with pale postmentum and narrow dark occipital margin; head total length, postmentum, mentum (
<figureCitation id="13592A74FF9E5258FC8AFF54FC446593" box="[830,921,188,213]" captionStart="FIGURE 4" captionStartId="9.[151,250,1973,1995]" captionTargetBox="[236,1332,1010,1913]" captionTargetId="figure@9.[229,1340,999,1924]" captionTargetPageId="9" captionText="FIGURE 4. Polypedilum spp., larvae. A, B. Antenna of: A. P. nubifer (Skuse), B. P. quasinubifer sp. n. C, D. Mentum of: C. P. nubifer, D. P. quasinubifer. E, F. Lateral end of ventromental plate of: E. P. nubifer, F. P. quasinubifer." httpUri="https://zenodo.org/record/1051652/files/figure.png" pageId="12" pageNumber="441">Fig. 4D</figureCitation>
) and mandible c. 3040% shorter than in
<taxonomicName id="4C624D72FF9E5258FA33FF55FF3B65BF" class="Insecta" family="Chironomidae" genus="Chironomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="441" phylum="Arthropoda" rank="species" species="nubifer">
<emphasis id="B916EAE3FF9E5258FA33FF55FF3B65BF" italics="true" pageId="12" pageNumber="441">P. nubifer</emphasis>
</taxonomicName>
. Antennal segment lengths shorter but ratios similar: Antennal blade relatively longer, reaching antennal apex (
<figureCitation id="13592A74FF9E5258FF68FEECFEE9645B" box="[220,308,260,285]" captionStart="FIGURE 4" captionStartId="9.[151,250,1973,1995]" captionTargetBox="[236,1332,1010,1913]" captionTargetId="figure@9.[229,1340,999,1924]" captionTargetPageId="9" captionText="FIGURE 4. Polypedilum spp., larvae. A, B. Antenna of: A. P. nubifer (Skuse), B. P. quasinubifer sp. n. C, D. Mentum of: C. P. nubifer, D. P. quasinubifer. E, F. Lateral end of ventromental plate of: E. P. nubifer, F. P. quasinubifer." httpUri="https://zenodo.org/record/1051652/files/figure.png" pageId="12" pageNumber="441">Fig. 4B</figureCitation>
). Microsculpture of ventromental plate (
<figureCitation id="13592A74FF9E5258FD48FEECFC8F645B" box="[764,850,260,285]" captionStart="FIGURE 4" captionStartId="9.[151,250,1973,1995]" captionTargetBox="[236,1332,1010,1913]" captionTargetId="figure@9.[229,1340,999,1924]" captionTargetPageId="9" captionText="FIGURE 4. Polypedilum spp., larvae. A, B. Antenna of: A. P. nubifer (Skuse), B. P. quasinubifer sp. n. C, D. Mentum of: C. P. nubifer, D. P. quasinubifer. E, F. Lateral end of ventromental plate of: E. P. nubifer, F. P. quasinubifer." httpUri="https://zenodo.org/record/1051652/files/figure.png" pageId="12" pageNumber="441">Fig. 4F</figureCitation>
): with sparser striae, terminating anteriorly on outer (lateralmost) corner of plate in 34 shallow lobes.
</paragraph>
<paragraph id="8BDD36F1FF9E5258FF73FEA4FCE0677B" blockId="12.[151,1437,151,573]" pageId="12" pageNumber="441">
<emphasis id="B916EAE3FF9E5258FF73FEA4FECC6422" bold="true" box="[199,273,332,357]" pageId="12" pageNumber="441">Notes.</emphasis>
<taxonomicName id="4C624D72FF9E5258FEA9FEA4FD986422" authorityName="Cranston &amp; Martin &amp; Spies" authorityYear="2016" box="[285,581,332,357]" class="Insecta" family="Chironomidae" genus="Polypedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="441" phylum="Arthropoda" rank="species" species="quasinubifer">
<emphasis id="B916EAE3FF9E5258FEA9FEA4FD986422" box="[285,581,332,357]" italics="true" pageId="12" pageNumber="441">Polypedilum quasinubifer</emphasis>
</taxonomicName>
is smaller in all measurements (e.g. wing length c. 40% of
<emphasis id="B916EAE3FF9E5258FA96FEA5FEC164CF" italics="true" pageId="12" pageNumber="441">
<taxonomicName id="4C624D72FF9E5258FA96FEA5FA4D6422" box="[1314,1424,332,357]" class="Insecta" family="Chironomidae" genus="Chironomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="441" phylum="Arthropoda" rank="species" species="nubifer">P. nubifer</taxonomicName>
/
<taxonomicName id="4C624D72FF9E5258FF23FE87FEC164CF" authorityName="Cranston &amp; Martin &amp; Spies" authorityYear="2016" box="[151,284,367,392]" class="Insecta" family="Chironomidae" genus="Polypedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="441" phylum="Arthropoda" rank="species" species="paranubifer">paranubifer</taxonomicName>
</emphasis>
, has fewer setae, and differs significantly in ratios, notably the fore leg ratio of 1.92.2 (in both sexes). The pupa is smaller and paler than those of
<emphasis id="B916EAE3FF9E5258FD2DFE7DFC4264EA" box="[665,927,404,429]" italics="true" pageId="12" pageNumber="441">
<taxonomicName id="4C624D72FF9E5258FD2DFE7DFCD964EA" box="[665,772,404,429]" class="Insecta" family="Chironomidae" genus="Chironomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="441" phylum="Arthropoda" rank="species" species="nubifer">P. nubifer</taxonomicName>
/
<taxonomicName id="4C624D72FF9E5258FCADFE7CFC4264EA" authorityName="Cranston &amp; Martin &amp; Spies" authorityYear="2016" box="[793,927,404,429]" class="Insecta" family="Chironomidae" genus="Polypedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="441" phylum="Arthropoda" rank="species" species="paranubifer">paranubifer</taxonomicName>
</emphasis>
), with smaller cephalic tubercles but longer frontal setae arising nearer the tubercle base. The thorax has a dorsal cluster of small tubercles (rather than aligned rows of larger tubercles). The hook row is narrower (no more than 50% of tergite width) and with fewer hooklets. The 'comb' on segment VIII is paler, smaller, with a more dominant major spine and fewer subsidiary spines. The anal lobe has fewer taeniae (&lt;30) in a uniform uniserial row.
</paragraph>
</subSubSection>
</treatment>
</document>