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<document id="17254CDB8DC8E1AEC39E0DA28A11F216" ID-DOI="10.5281/zenodo.273553" ID-GBIF-Dataset="f09945ae-b957-45f7-99c0-7bbfda0ebf5f" ID-ISSN="1175­5326" ID-Zenodo-Dep="273553" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459787765674" checkinUser="plazi" docAuthor="García, Alfonso N. &amp; Aldrete" docDate="2006" docId="03CB878CA31F4C61FEC8FA08FD09F8BE" docLanguage="en" docName="zt01319p014.pdf" docOrigin="Zootaxa 1319" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Waoraniella erwini García &amp; Aldrete, 2006, n. sp." docType="treatment" docVersion="8" lastPageNumber="11" masterDocId="FFF2FFF4A3164C6BFFC0FF96FFF3FF98" masterDocTitle="New genera of Psocoptera (Insecta), from Mexico, Belize and Ecuador (Psoquillidae, Ptiloneuridae, Lachesillidae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="10" updateTime="1698218827869" updateUser="plazi">
<mods:mods id="592EF1BF2DFC63ECBB1A06099EE1E74D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F8EFFDF16489623BF4558DAFE40A87FF">
<mods:title id="A4360A80C47335CED8654DDBD5920D26">New genera of Psocoptera (Insecta), from Mexico, Belize and Ecuador (Psoquillidae, Ptiloneuridae, Lachesillidae)</mods:title>
</mods:titleInfo>
<mods:name id="0994EB522E503ED3BBCF2B3B2D2D1E90" type="personal">
<mods:role id="0F7E1DDDDDE731B16AB67075820CA90A">
<mods:roleTerm id="5C316EE1CABEE97B7F9A069BDF74260F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D94971EEF31973CCC32DF16CB624E942">García, Alfonso N.</mods:namePart>
</mods:name>
<mods:name id="9F242DCDEBFAAC5E68ED519E43D08E2F" type="personal">
<mods:role id="5D54F129FC21690BF9868399EB4FAF1A">
<mods:roleTerm id="E6DB9AEE1B2CE90B212E8D63FF27A3E3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="01EEC24C6942F4A26582F2982F9F262C">Aldrete</mods:namePart>
</mods:name>
<mods:typeOfResource id="FC6E126816F2DC93FB8B31BC248B5E62">text</mods:typeOfResource>
<mods:relatedItem id="CF60DDFE403752FD93F3A243CA6C0B50" type="host">
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<mods:title id="EEA39B3962F3DB14C44E2D3BADC1D43B">Zootaxa</mods:title>
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<mods:part id="D74FFE488E41A8FB5160BBAD1A4C91F0">
<mods:date id="8C726B7E44D3FF223BD6EB6B496F28AE">2006</mods:date>
<mods:detail id="B339C804739A4317759EFDDB086C29BA" type="volume">
<mods:number id="DDAE3FC193452DC7FB61699794DA5E5D">1319</mods:number>
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<mods:classification id="C23D3871520E04AC9C32CD846017EB93">journal article</mods:classification>
<mods:identifier id="0C9D9C874CCBB1A36E2704D426D4DA8E" type="DOI">10.5281/zenodo.273553</mods:identifier>
<mods:identifier id="6617B757A1D5FB94B4C9AB202E9C67F4" type="GBIF-Dataset">f09945ae-b957-45f7-99c0-7bbfda0ebf5f</mods:identifier>
<mods:identifier id="1ABF80329F01AAB86C4E44F00F6617EA" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="207C666CE68B0F5AF190CEA0ACCAA978" type="Zenodo-Dep">273553</mods:identifier>
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<treatment id="03CB878CA31F4C61FEC8FA08FD09F8BE" ID-DOI="http://doi.org/10.5281/zenodo.6253107" ID-GBIF-Taxon="119349579" ID-Zenodo-Dep="6253107" LSID="urn:lsid:plazi:treatment:03CB878CA31F4C61FEC8FA08FD09F8BE" httpUri="http://treatment.plazi.org/id/03CB878CA31F4C61FEC8FA08FD09F8BE" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="C3786511A31F4C62FEC8FA08FC87F9E3" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BDD369AA31F4C62FEC8FA08FD26FA20" blockId="9.[264,725,1438,1464]" box="[264,725,1438,1464]" pageId="9" pageNumber="10">
<heading id="D09581F6A31F4C62FEC8FA08FD26FA20" bold="true" box="[264,725,1438,1464]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B916EA88A31F4C62FEC8FA08FD26FA20" bold="true" box="[264,725,1438,1464]" pageId="9" pageNumber="10">
<taxonomicName id="4C624D19A31F4C62FEC8FA08FE03FA20" ID-CoL="7G5SN" box="[264,496,1438,1464]" class="Insecta" family="Lachesillidae" genus="Waoraniella" kingdom="Animalia" order="Psocodea" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="erwini" status="sp. nov.">
<emphasis id="B916EA88A31F4C62FEC8FA08FE03FA20" bold="true" box="[264,496,1438,1464]" italics="true" pageId="9" pageNumber="10">Waoraniella erwini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557F3A31F4C62FE37FA08FDC9FA20" box="[503,570,1438,1464]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13592A1FA31F4C62FD89FA08FD3EFA20" box="[585,717,1438,1464]" captionStart="FIGURES 28 – 34" captionStartId="11.[264,382,1622,1646]" captionTargetBox="[317,1292,346,1554]" captionTargetId="figure@11.[283,1308,318,1614]" captionTargetPageId="11" captionText="FIGURES 28 – 34. Waoraniella erwini n. sp. F. 28. Front view of head. 29. Fore­ and hind wings. 30. Subgenital plate. 31. Epiproct and left paraproct. 32. Lacinial apex. 33. Pretarsal claw. 34. Gonapophyses. Scales in mm. Figs. 32 and 33 to common scale." httpUri="https://zenodo.org/record/273557/files/figure.png" pageId="9" pageNumber="10">Fig. 28–34</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD369AA31F4C62FEC8FA7FFC87F9E3" blockId="9.[264,1324,1513,1899]" pageId="9" pageNumber="10">
<emphasis id="B916EA88A31F4C62FEC8FA7FFEA7F99B" bold="true" box="[264,340,1513,1539]" pageId="9" pageNumber="10">Color.</emphasis>
Ground color reddish brown. Compound eyes black, ocelli hyaline, with ochre centripetal crescents. Maxillary palps more pigmented than rest of the body. Antennae and legs pale brown. Wings with a reddish brown hue. Abdomen with transverse, deep reddish brown subcuticular rings, less conspicuous ventrally.
</paragraph>
</subSubSection>
<subSubSection id="C3786511A31F4C61FEF8F91FFBE4FDBE" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph id="8BDD369AA31F4C61FEF8F91FFB96FE4E" blockId="9.[264,1324,1513,1899]" lastBlockId="10.[264,1324,284,1830]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="B916EA88A31F4C62FEF8F91FFE2BF93B" bold="true" box="[312,472,1673,1699]" pageId="9" pageNumber="10">Morphology.</emphasis>
Outer cusp of lacinial apex large, bidentate (
<figureCitation id="13592A1FA31F4C62FBCBF91FFB9BF93B" box="[1035,1128,1673,1699]" captionStart="FIGURES 28 – 34" captionStartId="11.[264,382,1622,1646]" captionTargetBox="[317,1292,346,1554]" captionTargetId="figure@11.[283,1308,318,1614]" captionTargetPageId="11" captionText="FIGURES 28 – 34. Waoraniella erwini n. sp. F. 28. Front view of head. 29. Fore­ and hind wings. 30. Subgenital plate. 31. Epiproct and left paraproct. 32. Lacinial apex. 33. Pretarsal claw. 34. Gonapophyses. Scales in mm. Figs. 32 and 33 to common scale." httpUri="https://zenodo.org/record/273557/files/figure.png" pageId="9" pageNumber="10">Fig. 32</figureCitation>
). Pretarsal claw with a basal seta and a broad pulvillus (
<figureCitation id="13592A1FA31F4C62FD1EF927FCC4F953" box="[734,823,1713,1739]" captionStart="FIGURES 28 – 34" captionStartId="11.[264,382,1622,1646]" captionTargetBox="[317,1292,346,1554]" captionTargetId="figure@11.[283,1308,318,1614]" captionTargetPageId="11" captionText="FIGURES 28 – 34. Waoraniella erwini n. sp. F. 28. Front view of head. 29. Fore­ and hind wings. 30. Subgenital plate. 31. Epiproct and left paraproct. 32. Lacinial apex. 33. Pretarsal claw. 34. Gonapophyses. Scales in mm. Figs. 32 and 33 to common scale." httpUri="https://zenodo.org/record/273557/files/figure.png" pageId="9" pageNumber="10">Fig. 33</figureCitation>
). Forewing pterostigma narrow anteriorly, much wider posteriorly; Rs flexuous, Rs­M joined by a crossvein. Areola postica small (
<figureCitation id="13592A1FA31F4C62FED1F897FE9AF883" box="[273,361,1793,1819]" captionStart="FIGURES 28 – 34" captionStartId="11.[264,382,1622,1646]" captionTargetBox="[317,1292,346,1554]" captionTargetId="figure@11.[283,1308,318,1614]" captionTargetPageId="11" captionText="FIGURES 28 – 34. Waoraniella erwini n. sp. F. 28. Front view of head. 29. Fore­ and hind wings. 30. Subgenital plate. 31. Epiproct and left paraproct. 32. Lacinial apex. 33. Pretarsal claw. 34. Gonapophyses. Scales in mm. Figs. 32 and 33 to common scale." httpUri="https://zenodo.org/record/273557/files/figure.png" pageId="9" pageNumber="10">Fig. 29</figureCitation>
). Hindwing (
<figureCitation id="13592A1FA31F4C62FDC4F897FDAFF883" box="[516,604,1793,1819]" captionStart="FIGURES 28 – 34" captionStartId="11.[264,382,1622,1646]" captionTargetBox="[317,1292,346,1554]" captionTargetId="figure@11.[283,1308,318,1614]" captionTargetPageId="11" captionText="FIGURES 28 – 34. Waoraniella erwini n. sp. F. 28. Front view of head. 29. Fore­ and hind wings. 30. Subgenital plate. 31. Epiproct and left paraproct. 32. Lacinial apex. 33. Pretarsal claw. 34. Gonapophyses. Scales in mm. Figs. 32 and 33 to common scale." httpUri="https://zenodo.org/record/273557/files/figure.png" pageId="9" pageNumber="10">Fig. 29</figureCitation>
). Subgenital plate (
<figureCitation id="13592A1FA31F4C62FC81F897FC64F883" box="[833,919,1793,1819]" captionStart="FIGURES 28 – 34" captionStartId="11.[264,382,1622,1646]" captionTargetBox="[317,1292,346,1554]" captionTargetId="figure@11.[283,1308,318,1614]" captionTargetPageId="11" captionText="FIGURES 28 – 34. Waoraniella erwini n. sp. F. 28. Front view of head. 29. Fore­ and hind wings. 30. Subgenital plate. 31. Epiproct and left paraproct. 32. Lacinial apex. 33. Pretarsal claw. 34. Gonapophyses. Scales in mm. Figs. 32 and 33 to common scale." httpUri="https://zenodo.org/record/273557/files/figure.png" pageId="9" pageNumber="10">Fig. 30</figureCitation>
) setose as illustrated; a pigmented band along sides and posterior margin, this straight. Gonapophyses (
<figureCitation id="13592A1FA31F4C62FB8CF8BFFB5AF8DB" box="[1100,1193,1833,1859]" captionStart="FIGURES 28 – 34" captionStartId="11.[264,382,1622,1646]" captionTargetBox="[317,1292,346,1554]" captionTargetId="figure@11.[283,1308,318,1614]" captionTargetPageId="11" captionText="FIGURES 28 – 34. Waoraniella erwini n. sp. F. 28. Front view of head. 29. Fore­ and hind wings. 30. Subgenital plate. 31. Epiproct and left paraproct. 32. Lacinial apex. 33. Pretarsal claw. 34. Gonapophyses. Scales in mm. Figs. 32 and 33 to common scale." httpUri="https://zenodo.org/record/273557/files/figure.png" pageId="9" pageNumber="10">Fig. 34</figureCitation>
):
<date id="FFDC105AA31F4C62FB01F8BFFB2DF8DB" box="[1217,1246,1833,1859]" pageId="9" pageNumber="10">v1</date>
short, stout, associated to ninth sternum,
<date id="FFDC105AA31F4C62FD70F8C7FD3DF8F3" box="[688,718,1873,1899]" pageId="9" pageNumber="10">v2</date>
an elongate lobe associated with
<date id="FFDC105AA31F4C62FBADF8C7FB7FF8F3" box="[1133,1164,1873,1899]" pageId="9" pageNumber="10">v3</date>
, this wide at base, narrowing distally, with a field of stout setae along outer margin. Paraproct (
<figureCitation id="13592A1FA31C4C61FB06FE8AFAEEFEAE" box="[1222,1309,284,310]" captionStart="FIGURES 28 – 34" captionStartId="11.[264,382,1622,1646]" captionTargetBox="[317,1292,346,1554]" captionTargetId="figure@11.[283,1308,318,1614]" captionTargetPageId="11" captionText="FIGURES 28 – 34. Waoraniella erwini n. sp. F. 28. Front view of head. 29. Fore­ and hind wings. 30. Subgenital plate. 31. Epiproct and left paraproct. 32. Lacinial apex. 33. Pretarsal claw. 34. Gonapophyses. Scales in mm. Figs. 32 and 33 to common scale." httpUri="https://zenodo.org/record/273557/files/figure.png" pageId="10" pageNumber="11">Fig. 31</figureCitation>
), elongate, almost elliptic, with field of setae posteriorly along outer margin, other setae as illustrated; a distinct, bicuspid cone on outer edge, sensory fields with 16
<emphasis id="B916EA88A31C4C61FBA0FEFDFB9CFE1D" bold="true" box="[1120,1135,363,389]" pageId="10" pageNumber="11">–</emphasis>
19 trichobothria issuing from basal rosettes. Epiproct (
<figureCitation id="13592A1FA31C4C61FD02FE02FCE5FE36" box="[706,790,404,430]" captionStart="FIGURES 28 – 34" captionStartId="11.[264,382,1622,1646]" captionTargetBox="[317,1292,346,1554]" captionTargetId="figure@11.[283,1308,318,1614]" captionTargetPageId="11" captionText="FIGURES 28 – 34. Waoraniella erwini n. sp. F. 28. Front view of head. 29. Fore­ and hind wings. 30. Subgenital plate. 31. Epiproct and left paraproct. 32. Lacinial apex. 33. Pretarsal claw. 34. Gonapophyses. Scales in mm. Figs. 32 and 33 to common scale." httpUri="https://zenodo.org/record/273557/files/figure.png" pageId="10" pageNumber="11">Fig. 31</figureCitation>
) straight anteriorly, rounded posteriorly, with a row of five macrosetae along posterior border, other setae as illustrated.
</paragraph>
<paragraph id="8BDD369AA31C4C61FEF8FE75FBE4FDBE" blockId="10.[264,1324,284,1830]" pageId="10" pageNumber="11">
<emphasis id="B916EA88A31C4C61FEF8FE75FE05FE65" bold="true" box="[312,502,483,509]" pageId="10" pageNumber="11">Measurements.</emphasis>
FW: 1659, HW: 1338, F: 421, T: 680, t1: 220, t2: 91, ctt1: 13, Mx4: 136, f1: 238, f2: 151, IO: 360, D: 167, d: 87, IO/D: 2.14, PO: 0.52.
</paragraph>
</subSubSection>
<subSubSection id="C3786511A31C4C61FEF8FDA5FAD9FD5E" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BDD369AA31C4C61FEF8FDA5FAD9FD5E" blockId="10.[264,1324,284,1830]" pageId="10" pageNumber="11">
<emphasis id="B916EA88A31C4C61FEF8FDA5FDE6FDD5" bold="true" box="[312,533,563,589]" pageId="10" pageNumber="11">Material studied.</emphasis>
<typeStatus id="54D98838A31C4C61FDE4FDA2FD62FDD6" box="[548,657,564,590]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
F, 1
<typeStatus id="54D98838A31C4C61FD1FFDA2FCB7FDD6" box="[735,836,564,590]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
F.
<collectingCountry id="F375760AA31C4C61FCB5FDA2FBF4FDD6" box="[885,1031,564,590]" name="Ecuador" pageId="10" pageNumber="11">ECUADOR</collectingCountry>
. Napo. Waorani Ethnic Reserve.
<quantity id="4C9A9B7FA31C4C61FEBEFDCAFE31FDEE" box="[382,450,604,630]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="km" value="1.0">1 km</quantity>
S Onkone Gare Camp.
<specimenCount id="9D64FD13A31C4C61FCC6FDCAFCA2FDEE" box="[774,849,604,630]" pageId="10" pageNumber="11" type="generic">
<quantity id="4C9A9B7FA31C4C61FCC6FDCAFCBEFDEE" box="[774,845,604,630]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="11" unit="m" value="220.0">220m</quantity>
.
</specimenCount>
<date id="FFDC105AA31C4C61FCA1FDCAFC2CFDEE" box="[865,991,604,630]" pageId="10" pageNumber="11" value="1995-02-10">10.II.1995</date>
.
<geoCoordinate id="EE56505DA31C4C61FC33FDCAFB84FDEE" box="[1011,1143,604,630]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="15" value="-0.50277776">0°30’10’’S</geoCoordinate>
:
<geoCoordinate id="EE56505DA31C4C61FB4BFDCAFAD4FDEE" box="[1163,1319,604,630]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="15" value="-76.433334">76°26’00’’W</geoCoordinate>
. Fogging terre firme forest, T. L. Erwin
<emphasis id="B916EA88A31C4C61FCC1FD12FCB1FD05" box="[769,834,644,669]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
The
<typeStatus id="54D98838A31C4C61FC4BFD12FC02FD06" box="[907,1009,644,670]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
will be deposited in the Smithsonian Institution Collection, Washington, D. C. The
<typeStatus id="54D98838A31C4C61FC7FFD3AFBD7FD5E" box="[959,1060,684,710]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
is deposited in
<collectionCode id="ED73AE5FA31C4C61FB1BFD3AFAD4FD5E" box="[1243,1319,684,710]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="10" pageNumber="11">CNIN</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3786511A31C4C61FEF8FD42FCD8FC16" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8BDD369AA31C4C61FEF8FD42FCD8FC16" blockId="10.[264,1324,284,1830]" pageId="10" pageNumber="11">
<emphasis id="B916EA88A31C4C61FEF8FD42FE37FD76" bold="true" box="[312,452,724,750]" pageId="10" pageNumber="11">Etymology.</emphasis>
The genus name is a tribute to the Waorani or Auca “the people of the forest”. The species is dedicated to Dr. Terry L. Erwin, leader of the Smithsonian Institution team that conducted the canopy fogging episodes in the Waorani Reserve, in recognition of his many, ground breaking academic achievements and as a tribute to his expert work in the taxonomy of the
<taxonomicName id="4C624D19A31C4C61FD6DFCE2FCDBFC16" box="[685,808,884,910]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3786511A31C4C61FEF8FC0AFD09F8BE" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BDD369AA31C4C61FEF8FC0AFC4AF935" blockId="10.[264,1324,284,1830]" pageId="10" pageNumber="11">
<emphasis id="B916EA88A31C4C61FEF8FC0AFE42FC2E" bold="true" box="[312,433,924,950]" pageId="10" pageNumber="11">Remarks.</emphasis>
<bibRefCitation id="EFF34B6BA31C4C61FE79FC0AFCFFFC2E" author="Mockford" box="[441,780,924,950]" pageId="10" pageNumber="14" refString="Mockford, E. L. &amp; Sullivan, D. M. (1986) Systematics of the graphocaeciliine psocids with a proposed higher classification of the family Lachesillidae (Psocoptera). Transactions of the American Entomological Society, 112, 1 - 80." type="journal article" year="1986">Mockford &amp; Sullivan (1986)</bibRefCitation>
proposed a higher classification of the family
<taxonomicName id="4C624D19A31C4C61FEC8FC52FE54FC46" box="[264,423,964,990]" class="Insecta" family="Lachesillidae" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Lachesillidae</taxonomicName>
, in which they recognized two subfamilies: Lachesillinae, including
<taxonomicName id="4C624D19A31C4C61FEC8FC7AFE71FB9D" box="[264,386,1004,1029]" class="Insecta" family="Lachesillidae" genus="Lachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FEC8FC7AFE71FB9D" box="[264,386,1004,1029]" italics="true" pageId="10" pageNumber="11">Lachesilla</emphasis>
</taxonomicName>
and
<taxonomicName id="4C624D19A31C4C61FE06FC7AFDDDFB9D" box="[454,558,1004,1029]" class="Insecta" family="Lachesillidae" genus="Nadleria" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FE06FC7AFDDDFB9D" box="[454,558,1004,1029]" italics="true" pageId="10" pageNumber="11">Nadleria</emphasis>
</taxonomicName>
, and Eolachesillinae, with two tribes: Eolachesillini including only
<taxonomicName id="4C624D19A31C4C61FE91FB82FE17FBB5" box="[337,484,1044,1069]" class="Insecta" family="Lachesillidae" genus="Eolachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FE91FB82FE17FBB5" box="[337,484,1044,1069]" italics="true" pageId="10" pageNumber="11">Eolachesilla</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C624D19A31C4C61FDFEFB82FCF7FBB6" box="[574,772,1044,1070]" class="Insecta" family="Lachesillidae" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="tribe" tribe="Graphocaeciliini">Graphocaeciliini</taxonomicName>
, including
<emphasis id="B916EA88A31C4C61FC61FB82FB1AFBCD" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C624D19A31C4C61FC61FB82FB9BFBB5" box="[929,1128,1044,1069]" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Graphocaecilius</taxonomicName>
,
<taxonomicName id="4C624D19A31C4C61FB40FB82FAD5FBB5" box="[1152,1318,1044,1069]" class="Insecta" family="Lachesillidae" genus="Anomopsocus" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Anomopsocus</taxonomicName>
,
<taxonomicName id="4C624D19A31C4C61FEC8FBAAFE5EFBCD" box="[264,429,1084,1109]" class="Insecta" family="Lachesillidae" genus="Antilachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Antilachesilla</taxonomicName>
,
<taxonomicName id="4C624D19A31C4C61FE09FBAAFD9AFBCD" box="[457,617,1084,1109]" class="Insecta" family="Lachesillidae" genus="Prolachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Prolachesilla</taxonomicName>
,
<taxonomicName id="4C624D19A31C4C61FD44FBAAFCCAFBCD" box="[644,825,1084,1109]" class="Insecta" family="Lachesillidae" genus="Nanolachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Nanolachesilla</taxonomicName>
,
<taxonomicName id="4C624D19A31C4C61FC94FBAAFBE6FBCD" box="[852,1045,1084,1109]" class="Insecta" family="Lachesillidae" genus="Tricholachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Tricholachesilla</taxonomicName>
,
<taxonomicName id="4C624D19A31C4C61FBF1FBAAFB16FBCD" box="[1073,1253,1084,1109]" class="Insecta" family="Lachesillidae" genus="Mesolachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Mesolachesilla</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C624D19A31C4C61FEC8FBF2FE40FBE5" box="[264,435,1124,1149]" class="Insecta" family="Lachesillidae" genus="Notolachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FEC8FBF2FE40FBE5" box="[264,435,1124,1149]" italics="true" pageId="10" pageNumber="11">Notolachesilla</emphasis>
</taxonomicName>
.
<taxonomicName id="4C624D19A31C4C61FE01FBF2FDA7FBE5" box="[449,596,1124,1149]" class="Insecta" family="Lachesillidae" genus="Eolachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FE01FBF2FDA7FBE5" box="[449,596,1124,1149]" italics="true" pageId="10" pageNumber="11">Eolachesilla</emphasis>
</taxonomicName>
was transferred to the family
<taxonomicName id="4C624D19A31C4C61FC6FFBF2FBCBFBE6" box="[943,1080,1124,1150]" class="Insecta" family="Elipsocidae" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Elipsocidae</taxonomicName>
by
<bibRefCitation id="EFF34B6BA31C4C61FBA2FBF2FEAAFB3E" author="New" pageId="10" pageNumber="14" refString="New, T. R. &amp; Thornton, I. W. B. (1981) Psocoptera from central and southern Chile. Pacific Insects Monograph, 37, 136 - 178." type="journal article" year="1981">New &amp; Thornton (1981)</bibRefCitation>
, but
<bibRefCitation id="EFF34B6BA31C4C61FE54FB1AFD1AFB3E" author="Mockford" box="[404,745,1164,1190]" pageId="10" pageNumber="14" refString="Mockford, E. L. &amp; Sullivan, D. M. (1986) Systematics of the graphocaeciliine psocids with a proposed higher classification of the family Lachesillidae (Psocoptera). Transactions of the American Entomological Society, 112, 1 - 80." type="journal article" year="1986">Mockford &amp; Sullivan (1986)</bibRefCitation>
decided to leave this genus in the
<taxonomicName id="4C624D19A31C4C61FB47FB1AFAD5FB3E" box="[1159,1318,1164,1190]" class="Insecta" family="Lachesillidae" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Lachesillidae</taxonomicName>
, following
<bibRefCitation id="EFF34B6BA31C4C61FE47FB22FDA5FB56" author="Badonnel" box="[391,598,1204,1230]" pageId="10" pageNumber="14" refString="Badonnel, A. (1967) Psocopteres edaphiques du Chili (2 e note). Biologie de l'Amerique australe, 3, 541 - 585." type="journal article" year="1967">Badonnel (1967)</bibRefCitation>
, and acknowledged that
<taxonomicName id="4C624D19A31C4C61FC50FB22FBDFFB56" box="[912,1068,1204,1230]" class="Insecta" family="Lachesillidae" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Lachesillidae</taxonomicName>
and
<taxonomicName id="4C624D19A31C4C61FBB1FB22FB09FB56" box="[1137,1274,1204,1230]" class="Insecta" family="Elipsocidae" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Elipsocidae</taxonomicName>
are difficult to distinguish.
<bibRefCitation id="EFF34B6BA31C4C61FDE7FB4AFC24FB6E" box="[551,983,1244,1270]" pageId="10" pageNumber="14" refString="Garcia Aldrete, A. N. &amp; Mockford, E. L. (1997) The position of Hemicaecilius (Psocoptera: Lachesillidae) and description of a new species from Venezuela. Entomological News, 108, 102 - 106." type="journal article">García Aldrete &amp; Mockford (1997)</bibRefCitation>
added
<taxonomicName id="4C624D19A31C4C61FBF1FB4AFB2BFB6D" box="[1073,1240,1244,1269]" class="Insecta" family="Lachesillidae" genus="Hemicaecilius" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FBF1FB4AFB2BFB6D" box="[1073,1240,1244,1269]" italics="true" pageId="10" pageNumber="11">Hemicaecilius</emphasis>
</taxonomicName>
to the Lachesillinae, transferring it from
<taxonomicName id="4C624D19A31C4C61FD6BFA92FCC4FA86" box="[683,823,1284,1310]" class="Insecta" family="Elipsocidae" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Elipsocidae</taxonomicName>
, where it had been tentatively placed by
<bibRefCitation id="EFF34B6BA31C4C61FEC8FABAFE23FADE" author="Smithers" box="[264,464,1324,1350]" pageId="10" pageNumber="14" refString="Smithers, C. N. (1972) The classification and phylogeny of the Psocoptera. Australian Museum Memoirs, 14, 1 - 349." type="journal article" year="1972">Smithers (1972</bibRefCitation>
,
<bibRefCitation id="EFF34B6BA31C4C61FE2CFABAFDD8FADE" author="Smithers" box="[492,555,1324,1350]" pageId="10" pageNumber="14" refString="Smithers, C. N. (1990) Key to the families and genera of Psocoptera (Arthropoda: Insecta). Technical Reports of the Australian Museum, 2, 1 - 82." type="journal article" year="1990">1990</bibRefCitation>
). Li Fasheng (1995) described
<taxonomicName id="4C624D19A31C4C61FC2BFABAFB33FADD" box="[1003,1216,1324,1349]" class="Insecta" family="Lachesillidae" genus="Homoeolachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FC2BFABAFB33FADD" box="[1003,1216,1324,1349]" italics="true" pageId="10" pageNumber="11">Homoeolachesilla</emphasis>
</taxonomicName>
in the Lachesillinae, and in 2002 he described
<taxonomicName id="4C624D19A31C4C61FD25FAC2FC54FAF5" box="[741,935,1364,1389]" class="Insecta" family="Lachesillidae" genus="Cyclolachesillus" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FD25FAC2FC54FAF5" box="[741,935,1364,1389]" italics="true" pageId="10" pageNumber="11">Cyclolachesillus</emphasis>
</taxonomicName>
, and erected for it the subfamily
<taxonomicName id="4C624D19A31C4C61FEC8FAEAFE17FA0E" box="[264,484,1404,1430]" class="Insecta" family="Lachesillidae" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subFamily" subFamily="Cyclolachesillinae">Cyclolachesillinae</taxonomicName>
; he also described in the Lachesillinae the genus
<taxonomicName id="4C624D19A31C4C61FBA3FAEAFAD7FA0D" box="[1123,1316,1404,1429]" class="Insecta" family="Lachesillidae" genus="Dicrolachesillus" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FBA3FAEAFAD7FA0D" box="[1123,1316,1404,1429]" italics="true" pageId="10" pageNumber="11">Dicrolachesillus</emphasis>
</taxonomicName>
, subsequently placed in synonymy with
<taxonomicName id="4C624D19A31C4C61FD2FFA32FC9AFA25" box="[751,873,1444,1469]" class="Insecta" family="Lachesillidae" genus="Lachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FD2FFA32FC9AFA25" box="[751,873,1444,1469]" italics="true" pageId="10" pageNumber="11">Lachesilla</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF34B6BA31C4C61FC5EFA32FB95FA26" author="Lienhard" box="[926,1126,1444,1470]" pageId="10" pageNumber="14" refString="Lienhard, C. (2003) Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie, 110, 695 - 721." type="journal article" year="2003">Lienhard (2003)</bibRefCitation>
, and the genera
<emphasis id="B916EA88A31C4C61FEC8FA5AFD50FA7D" box="[264,675,1484,1509]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C624D19A31C4C61FEC8FA5AFE2AFA7D" box="[264,473,1484,1509]" class="Insecta" family="Lachesillidae" genus="Ceratolachesillus" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Ceratolachesillus</taxonomicName>
,
<taxonomicName id="4C624D19A31C4C61FE2DFA5AFD50FA7D" box="[493,675,1484,1509]" class="Insecta" family="Lachesillidae" genus="Zangilachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Zangilachesilla</taxonomicName>
</emphasis>
and
<taxonomicName id="4C624D19A31C4C61FD2EFA5AFC5BFA7D" box="[750,936,1484,1509]" class="Insecta" family="Lachesillidae" genus="Zonolachesillus" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FD2EFA5AFC5BFA7D" box="[750,936,1484,1509]" italics="true" pageId="10" pageNumber="11">Zonolachesillus</emphasis>
</taxonomicName>
. These genera deserve further study, particularly
<taxonomicName id="4C624D19A31C4C61FE33FA62FD46F995" box="[499,693,1524,1549]" class="Insecta" family="Lachesillidae" genus="Cyclolachesillus" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FE33FA62FD46F995" box="[499,693,1524,1549]" italics="true" pageId="10" pageNumber="11">Cyclolachesillus</emphasis>
</taxonomicName>
which, by the illustrations of
<taxonomicName id="4C624D19A31C4C61FB84FA62FE36F9AE" authority="Li Fasheng (2002)" authorityName="Li Fasheng" authorityYear="2002" class="Insecta" family="Lachesillidae" genus="Cyclolachesillus" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ningxiaensis">
<emphasis id="B916EA88A31C4C61FB84FA62FAF1F995" box="[1092,1282,1524,1549]" italics="true" pageId="10" pageNumber="11">C. ningxiaensis</emphasis>
Li Fasheng (2002)
</taxonomicName>
, could probably be an elipsocid. As for the other genera, perhaps their creation needs a broader justification; my opinion is that it would be best to include the species involved in
<taxonomicName id="4C624D19A31C4C61FE3DF9FAFD84F91D" box="[509,631,1644,1669]" class="Insecta" family="Lachesillidae" genus="Lachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FE3DF9FAFD84F91D" box="[509,631,1644,1669]" italics="true" pageId="10" pageNumber="11">Lachesilla</emphasis>
</taxonomicName>
; Li Fasheng himself (1995) described in
<taxonomicName id="4C624D19A31C4C61FBB9F9FAFB00F91D" box="[1145,1267,1644,1669]" class="Insecta" family="Lachesillidae" genus="Lachesilla" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FBB9F9FAFB00F91D" box="[1145,1267,1644,1669]" italics="true" pageId="10" pageNumber="11">Lachesilla</emphasis>
</taxonomicName>
two species that seven years later he placed in
<emphasis id="B916EA88A31C4C61FD37F902FC4AF935" box="[759,953,1684,1709]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C624D19A31C4C61FD37F902FC46F935" box="[759,949,1684,1709]" class="Insecta" family="Lachesillidae" genus="Zonolachesillus" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Zonolachesillus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BDD369AA31C4C61FEF8F92AFD09F8BE" blockId="10.[264,1324,284,1830]" pageId="10" pageNumber="11">
<taxonomicName id="4C624D19A31C4C61FEF8F92AFE3BF94D" box="[312,456,1724,1749]" class="Insecta" family="Lachesillidae" genus="Waoraniella" kingdom="Animalia" order="Psocodea" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA88A31C4C61FEF8F92AFE3BF94D" box="[312,456,1724,1749]" italics="true" pageId="10" pageNumber="11">Waoraniella</emphasis>
</taxonomicName>
presents a distinct autapomorphy (forewing Rs­M joined by a crossvein), that does not permit placing it in any of the two tribes of the subfamily, hence I propose the following new tribe in the Eolachesillinae.
</paragraph>
</subSubSection>
</treatment>
</document>
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