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<document id="5BB6B481573B5B844072F7BBA4A78052" ID-DOI="10.5852/ejt.2019.526" ID-GBIF-Dataset="2da65c0a-9c48-43f5-ada3-e088de28dec1" ID-ISSN="2118-9773" ID-Zenodo-Dep="3152217" ID-ZooBank="3FFC527D-E9CA-4D9F-BF59-076FA60171AC" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1558535825802" checkinUser="plazi" docAuthor="Nurinsiyah, Ayu Savitri, Neiber, Marco T. &amp; Hausdorf, Bernhard" docDate="2019" docId="03B1DB4C8B6C4E5B4C64FD82387DFCD7" docLanguage="en" docName="ej-526_nurinsiyah_neiber_hausdorf.pdf.imf" docOrigin="European Journal of Taxonomy 526" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria menorehensis Nurinsiyah &amp; Neiber &amp; Hausdorf 2019, sp. nov." docType="treatment" docUuid="E09F6255-8F7A-49E7-A3BC-E5B795EE2015" docUuidSource="ZooBank" docVersion="16" lastPageNumber="67" masterDocId="FF88A3348B2D4E194E35FFFC3C0DFFDB" masterDocTitle="Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java" masterLastPageNumber="73" masterPageNumber="1" pageNumber="66" updateTime="1716320095690" updateUser="carolina">
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<mods:title id="F2CEE098D0FD45A406A0E30CF416CDF5">Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java</mods:title>
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<mods:namePart id="C3A81F94BD3C5919E54A48765CFAC3DE">Nurinsiyah, Ayu Savitri</mods:namePart>
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<mods:affiliation id="F73136FEE682EDBD12D9A216E959A965">Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia.</mods:affiliation>
<mods:nameIdentifier id="70CB5248FFBC78A3CB651DA22EE1FC21" type="email">ayu_nurinsiyah@yahoo.com &amp; ayus002@lipi.go.id</mods:nameIdentifier>
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<mods:namePart id="210291E16E4ED25B27AA65AC9DFDB8EA">Neiber, Marco T.</mods:namePart>
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<mods:affiliation id="BD844C2DB6C60ECBFB323F5772083A7E">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier id="C0786BC9882B818FB64CA9D9A5F74980" type="email">mneiber@hotmail.de</mods:nameIdentifier>
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<mods:namePart id="9AD999AFAA8EA5148CB3E9B74BDEF135">Hausdorf, Bernhard</mods:namePart>
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<mods:affiliation id="69F67DAAF6DB1C412B134E01CEC98AA9">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
<mods:nameIdentifier id="D9C8AAD9645FD83298D4E83E396FFC1D" type="email">hausdorf@zoologie.uni-hamburg.de</mods:nameIdentifier>
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<mods:title id="6113FC03938959EB4A4A3C7C3FFE9FE4">European Journal of Taxonomy</mods:title>
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<mods:date id="B6EC8FF49761E8717DACC217BF81201B">2019</mods:date>
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<treatment id="03B1DB4C8B6C4E5B4C64FD82387DFCD7" ID-DOI="http://doi.org/10.5281/zenodo.5587354" ID-GBIF-Taxon="157239578" ID-Zenodo-Dep="5587354" LSID="urn:lsid:zoobank.org:act:E09F6255-8F7A-49E7-A3BC-E5B795EE2015" httpUri="http://treatment.plazi.org/id/03B1DB4C8B6C4E5B4C64FD82387DFCD7" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
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<paragraph id="8BA76A5A8B6C4E584C64FD823FEEFD42" blockId="65.[369,1218,638,736]" box="[593,995,638,665]" pageId="65" pageNumber="66">
<heading id="D0EFDD368B6C4E584C64FD823FEEFD42" box="[593,995,638,665]" centered="true" fontSize="11" level="2" pageId="65" pageNumber="66" reason="2">
<taxonomicName id="4C1811D98B6C4E584C64FD823F8CFD42" ID-CoL="7RTHN" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[593,897,638,665]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="65" pageNumber="66" phylum="Mollusca" rank="species" species="menorehensis" status="sp. nov.">
<emphasis id="B96CB6488B6C4E584C64FD823F8CFD42" bold="true" box="[593,897,638,665]" italics="true" pageId="65" pageNumber="66">Landouria menorehensis</emphasis>
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<taxonomicNameLabel id="A25F0B338B6C4E584DBDFD833FEEFD42" box="[904,995,639,665]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="C30239D18B6C4E584F44FD5D3FC2FD04" pageId="65" pageNumber="66" type="description">
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urn:lsid:zoobank.org:act:
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<paragraph id="8BA76A5A8B6C4E584C50FD393FC2FD04" blockId="65.[369,1218,638,736]" box="[613,975,709,736]" pageId="65" pageNumber="66">
<figureCitation id="132376DF8B6C4E584C50FD393EB2FD04" box="[613,703,709,735]" captionStart="Figs 29–47" captionStartId="29.[189,241,1366,1392]" captionTargetBox="[189,1399,649,1337]" captionTargetId="figure@29.[189,1399,649,1337]" captionTargetPageId="29" captionText="Figs 29–47. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm." httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="65" pageNumber="66">Figs 47</figureCitation>
,
<figureCitation id="132376DF8B6C4E584CFEFD393EE6FD3B" box="[715,747,709,736]" captionStart="Figs 52–55" captionStartId="33.[189,241,1717,1743]" captionTargetBox="[189,1398,265,1993]" captionTargetId="figure@34.[189,1398,265,1993]" captionTargetPageId="34" captionText="Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867)." httpUri="https://zenodo.org/record/3152265/files/figure.png" pageId="65" pageNumber="66">55</figureCitation>
,
<figureCitation id="132376DF8B6C4E584CC2FD393F1AFD04" box="[759,791,709,735]" captionStart="Figs 63–71" captionStartId="48.[189,241,1668,1694]" captionTargetBox="[189,1394,515,1612]" captionTargetId="figure@48.[189,1398,510,1629]" captionTargetPageId="48" captionText="Figs 63–71. Genitalia of species of Landouria from Java. 63. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 64. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 65. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 66. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 67. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 68. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 69. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 70. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 71. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152282/files/figure.png" pageId="65" pageNumber="66">71</figureCitation>
,
<figureCitation id="132376DF8B6C4E584D16FD393F4EFD04" box="[803,835,709,735]" captionStart="Figs 85–88" captionStartId="62.[189,241,1859,1885]" captionTargetBox="[232,1355,265,1830]" captionTargetId="figure@62.[232,1355,265,1830]" captionTargetPageId="62" captionText="Figs 85–88. Shells of species of Landouria from Java. 85. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 86. L. dharmai sp. nov., East Java, Tulungagung, Popoh (holotype, MZB 21272). 87. L. davini Dharma, 2015, Central Java, Baturraden, Pancuran Tujuh, Gunung Slamet (holotype, MZB 17063). 88. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152308/files/figure.png" pageId="65" pageNumber="66">88</figureCitation>
;
<tableCitation id="C69A5FE18B6C4E584D7AFD393FC2FD04" box="[847,975,709,735]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="65" pageNumber="66" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B6C4E584E88FCFA3FB1FCA9" pageId="65" pageNumber="66" type="diagnosis">
<paragraph id="8BA76A5A8B6C4E584E88FCFA3D3BFCFB" blockId="65.[189,310,774,800]" box="[189,310,774,800]" pageId="65" pageNumber="66">
<heading id="D0EFDD368B6C4E584E88FCFA3D3BFCFB" bold="true" box="[189,310,774,800]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="B96CB6488B6C4E584E88FCFA3D3BFCFB" bold="true" box="[189,310,774,800]" pageId="65" pageNumber="66">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B6C4E584E88FCC93FB1FCA9" blockId="65.[189,1398,821,883]" pageId="65" pageNumber="66">
<taxonomicName id="4C1811D98B6C4E584E88FCC93DEDFC94" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,480,821,847]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="65" pageNumber="66" phylum="Mollusca" rank="species" species="menorehensis" status="sp. nov.">
<emphasis id="B96CB6488B6C4E584E88FCC93DEDFC94" box="[189,480,821,847]" italics="true" pageId="65" pageNumber="66">Landouria menorehensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B6C4E584FD3FCC93E4DFC94" box="[486,576,821,847]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by a whitish, globose shell with a distinct brown band above the rounded periphery and a moderately narrow umbilicus.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B6C4E584E88FC6638D8FC38" box="[189,1237,922,995]" pageId="65" pageNumber="66" type="etymology">
<paragraph id="8BA76A5A8B6C4E584E88FC663D49FC6F" blockId="65.[189,324,922,948]" box="[189,324,922,948]" pageId="65" pageNumber="66">
<heading id="D0EFDD368B6C4E584E88FC663D49FC6F" bold="true" box="[189,324,922,948]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="B96CB6488B6C4E584E88FC663D49FC6F" bold="true" box="[189,324,922,948]" pageId="65" pageNumber="66">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B6C4E584E88FC3438D8FC38" blockId="65.[189,1237,968,995]" box="[189,1237,968,995]" pageId="65" pageNumber="66">
<taxonomicName id="4C1811D98B6C4E584E88FC343DEDFC39" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,480,968,994]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="65" pageNumber="66" phylum="Mollusca" rank="species" species="menorehensis" status="sp. nov.">
<emphasis id="B96CB6488B6C4E584E88FC343DEDFC39" box="[189,480,968,994]" italics="true" pageId="65" pageNumber="66">Landouria menorehensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B6C4E584FDDFC353E4FFC38" box="[488,578,969,995]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
is named after its occurrence in the Menoreh karst area.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B6C4E584E88FBF53D68F91C" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="8BA76A5A8B6C4E584E88FBF53DA7FBF8" blockId="65.[189,426,1033,1059]" box="[189,426,1033,1059]" pageId="65" pageNumber="66">
<heading id="D0EFDD368B6C4E584E88FBF53DA7FBF8" bold="true" box="[189,426,1033,1059]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="B96CB6488B6C4E584E88FBF53DA7FBF8" bold="true" box="[189,426,1033,1059]" pageId="65" pageNumber="66">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B7060078B6C4E584EDFFBC43D58FB67" ID-GBIF-Occurrence="2243137312" collectingDate="2013-01-08" collectionCode="MZB" collectorName="A. Nurinsiyah" country="Indonesia" elevation="687" latitude="-7.747778" location="Sibolong" longLatPrecision="21" longitude="110.136665" municipality="Kulonprogo" pageId="65" pageNumber="66" specimenCode="19201" specimenCount="1" stateProvince="Yogyakarta" typeStatus="holotype">
<paragraph id="8BA76A5A8B6C4E584EDFFBC43D51FB89" blockId="65.[189,1399,1080,1212]" box="[234,348,1080,1106]" pageId="65" pageNumber="66">
<typeStatus id="54A3D4F88B6C4E584EDFFBC43D51FB89" box="[234,348,1080,1106]" pageId="65" pageNumber="66" type="holotype">
<emphasis id="B96CB6488B6C4E584EDFFBC43D51FB89" bold="true" box="[234,348,1080,1106]" pageId="65" pageNumber="66">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BA76A5A8B6C4E584E88FBA73D58FB67" blockId="65.[189,1399,1080,1212]" pageId="65" pageNumber="66">
<collectingCountry id="F30F2ACA8B6C4E584E88FBA73D56FBAE" box="[189,347,1115,1141]" name="Indonesia" pageId="65" pageNumber="66">INDONESIA</collectingCountry>
• holotype (det. anat.);
<collectingRegion id="49DCA4B88B6C4E584CBEFBA73F19FBAE" box="[651,788,1115,1141]" country="Indonesia" name="Yogyakarta" pageId="65" pageNumber="66">Yogyakarta</collectingRegion>
,
<collectingMunicipality id="6BC3F0208B6C4E584D10FBA73FBBFBAE" box="[805,950,1115,1141]" pageId="65" pageNumber="66">Kulonprogo</collectingMunicipality>
,
<location id="8EC73C818B6C4E584DF2FBA73839FBAE" LSID="urn:lsid:plazi:treatment:03B1DB4C8B6C4E5B4C64FD82387DFCD7:8EC73C818B6C4E584DF2FBA73839FBAE" box="[967,1076,1115,1141]" country="Indonesia" latitude="-7.747778" longLatPrecision="21" longitude="110.136665" municipality="Kulonprogo" name="Sibolong" pageId="65" pageNumber="66" stateProvince="Yogyakarta">Sibolong</location>
, agroforestry;
<geoCoordinate id="EE2C0C9D8B6C4E584ADFFBA5397CFBAE" box="[1258,1393,1113,1141]" degrees="7" direction="south" minutes="44" orientation="latitude" pageId="65" pageNumber="66" precision="15" seconds="52" value="-7.747778">7°44′52″ S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B6C4E584E88FB813D50FB42" box="[189,349,1149,1177]" degrees="110" direction="east" minutes="08" orientation="longitude" pageId="65" pageNumber="66" precision="15" seconds="12" value="110.136665">110°08′12″ E</geoCoordinate>
;
<elevation id="00358D698B6C4E584F5FFB823DF9FB43" box="[362,500,1150,1177]" metricMagnitude="2" metricUnit="m" metricValue="6.87" pageId="65" pageNumber="66" unit="m" value="687.0">
<quantity id="4CE0C7BF8B6C4E584F5FFB823DB9FB42" box="[362,436,1150,1177]" metricMagnitude="2" metricUnit="m" metricValue="6.87" pageId="65" pageNumber="66" unit="m" value="687.0">687 m</quantity>
a.s.l.
</elevation>
;
<collectingDate id="EFE2B5728B6C4E584C34FB823E86FB43" box="[513,651,1150,1177]" pageId="65" pageNumber="66" value="2013-01-08">
<date id="FFA64C9A8B6C4E584C34FB823E86FB43" box="[513,651,1150,1177]" pageId="65" pageNumber="66" value="2013-01-08">8 Jan. 2013</date>
</collectingDate>
;
<collectorName id="26ED0F8C8B6C4E584CA0FB833F36FB43" box="[661,827,1150,1177]" pageId="65" pageNumber="66">A. Nurinsiyah</collectorName>
leg.; measurements: D =
<quantity id="4CE0C7BF8B6C4E584A5CFB8238D8FB42" box="[1129,1237,1150,1177]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="65" pageNumber="66" unit="mm" value="13.4">13.4 mm</quantity>
, H =
<quantity id="4CE0C7BF8B6C4E584B20FB82397FFB42" box="[1301,1394,1150,1177]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="65" pageNumber="66" unit="mm" value="8.6">8.6 mm</quantity>
;
<collectionCode id="ED09F29F8B6C4E584E88FB5E3CF0FB67" box="[189,253,1186,1212]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="65" pageNumber="66" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B6C4E584F31FB5E3D5CFB67" box="[260,337,1186,1212]" pageId="65" pageNumber="66">19201</specimenCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="3B7060078B6C4E584EDFFB1F3836FA9F" ID-GBIF-Occurrence="2243137385" collectedFrom="agroforestry" collectionCode="ZMH" country="Indonesia" county="Kulonprogo" elevation="721" latitude="-7.75" location="Sibolong Hill" longLatPrecision="21" longitude="110.11667" municipality="Sibolong" pageId="65" pageNumber="66" specimenCode="ZMH 133441" specimenCount="1" stateProvince="Central Java" typeStatus="paratype">
<paragraph id="8BA76A5A8B6C4E584EDFFB1F3D6AFB26" blockId="65.[189,1399,1251,1735]" box="[234,359,1251,1277]" pageId="65" pageNumber="66">
<typeStatus id="54A3D4F88B6C4E584EDFFB1F3D6AFB26" box="[234,359,1251,1277]" pageId="65" pageNumber="66" type="paratype">
<emphasis id="B96CB6488B6C4E584EDFFB1F3D6AFB26" bold="true" box="[234,359,1251,1277]" pageId="65" pageNumber="66">Paratypes</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BA76A5A8B6C4E584E88FAFA3D68F91C" blockId="65.[189,1399,1251,1735]" pageId="65" pageNumber="66">
<collectingCountry id="F30F2ACA8B6C4E584E88FAFA3D56FAFB" box="[189,347,1286,1312]" name="Indonesia" pageId="65" pageNumber="66">INDONESIA</collectingCountry>
–
<emphasis id="B96CB6488B6C4E584FB5FAFA3EC5FAFB" bold="true" box="[384,712,1286,1312]" pageId="65" pageNumber="66">
<collectingRegion id="49DCA4B88B6C4E584FB5FAFA3E2AFAFB" box="[384,551,1286,1312]" country="Indonesia" name="Jawa Tengah" pageId="65" pageNumber="66">Central Java</collectingRegion>
,
<collectingRegion id="49DCA4B88B6C4E584C02FAFA3EC5FAFB" box="[567,712,1286,1312]" country="Indonesia" name="Yogyakarta" pageId="65" pageNumber="66">Yogyakarta</collectingRegion>
</emphasis>
•
<specimenCount id="9D1EA1D38B6C4E584CDDFAFA3F4DFAFA" box="[744,832,1286,1313]" count="1" pageId="65" pageNumber="66" type="generic">1 spec.</specimenCount>
;
<collectingCounty id="62C612D68B6C4E584D65FAFA3FECFAFB" box="[848,993,1286,1312]" pageId="65" pageNumber="66">Kulonprogo</collectingCounty>
,
<collectingMunicipality id="6BC3F0208B6C4E584DC5FAFA3850FAFB" box="[1008,1117,1286,1312]" pageId="65" pageNumber="66">Sibolong</collectingMunicipality>
,
<collectedFrom id="380429A78B6C4E584A59FAFA38F6FAFB" box="[1132,1275,1286,1312]" pageId="65" pageNumber="66">agroforestry</collectedFrom>
along the way to
<location id="8EC73C818B6C4E584F21FAD53DBAFA98" LSID="urn:lsid:plazi:treatment:03B1DB4C8B6C4E5B4C64FD82387DFCD7:8EC73C818B6C4E584F21FAD53DBAFA98" box="[276,439,1321,1347]" country="Indonesia" county="Kulonprogo" latitude="-7.75" longLatPrecision="21" longitude="110.11667" municipality="Sibolong" name="Sibolong Hill" pageId="65" pageNumber="66" stateProvince="Central Java">Sibolong Hill</location>
;
<geoCoordinate id="EE2C0C9D8B6C4E584FF1FAD43E48FA9F" box="[452,581,1320,1348]" degrees="7" direction="south" minutes="45" orientation="latitude" pageId="65" pageNumber="66" precision="15" seconds="00" value="-7.75">7°45′00″ S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B6C4E584C66FAD43EF9FA9F" box="[595,756,1320,1348]" degrees="110" direction="east" minutes="07" orientation="longitude" pageId="65" pageNumber="66" precision="15" seconds="00" value="110.11667">110°07′00″ E</geoCoordinate>
;
<elevation id="00358D698B6C4E584D34FAD63F81FA98" box="[769,908,1321,1348]" metricMagnitude="2" metricUnit="m" metricValue="7.209999999999999" pageId="65" pageNumber="66" unit="m" value="721.0">
<quantity id="4CE0C7BF8B6C4E584D34FAD63F41FA9F" box="[769,844,1322,1348]" metricMagnitude="2" metricUnit="m" metricValue="7.209999999999999" pageId="65" pageNumber="66" unit="m" value="721.0">721 m</quantity>
a.s.l.
</elevation>
;
<specimenCode id="DBBEC2218B6C4E584DAFFAD63836FA9F" box="[922,1083,1322,1348]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="65" pageNumber="66" type="Museum">ZMH 133441</specimenCode>
•
<materialsCitation id="3B7060078B6C4E584A60FAD63DEBFABC" ID-GBIF-Occurrence="2243137536" collectionCode="ZMH" country="Indonesia" county="Kulonprogo" elevation="721" latitude="-7.75" location="Sibolong Hill" longLatPrecision="21" longitude="110.11667" municipality="Sibolong" pageId="65" pageNumber="66" specimenCode="ZMH 133442" specimenCount="2" stateProvince="Yogyakarta" typeStatus="paratype">
<specimenCount id="9D1EA1D38B6C4E584A60FAD638A7FA9F" box="[1109,1194,1322,1348]" count="2" pageId="65" pageNumber="66" type="generic">2 spec.</specimenCount>
; same data as for preceding;
<specimenCode id="DBBEC2218B6C4E584F76FAB13DEBFABC" box="[323,486,1357,1383]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="65" pageNumber="66" type="Museum">ZMH 133442</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B7060078B6C4E584C31FAB1386EFABC" ID-GBIF-Occurrence="2243137492" box="[516,1123,1357,1383]" collectionCode="ZMH" country="Indonesia" county="Kulonprogo" elevation="721" latitude="-7.75" location="Sibolong Hill" longLatPrecision="21" longitude="110.11667" municipality="Sibolong" pageId="65" pageNumber="66" specimenCode="ZMH 133443" specimenCount="1" stateProvince="Yogyakarta" typeStatus="paratype">
<specimenCount id="9D1EA1D38B6C4E584C31FAB13E56FABC" box="[516,603,1357,1383]" count="1" pageId="65" pageNumber="66" type="generic">1 spec.</specimenCount>
; same data as for preceding;
<specimenCode id="DBBEC2218B6C4E584D8AFAB1386EFABC" box="[959,1123,1357,1383]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="65" pageNumber="66" type="Museum">ZMH 133443</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B7060078B6C4E584AB4FAB138F4FA51" ID-GBIF-Occurrence="2243137443" collectedFrom="agroforestry" collectionCode="MZB" country="Indonesia" elevation="716" latitude="-7.745556" location="Sibolong" longLatPrecision="21" longitude="110.13389" municipality="Kulonprogo" pageId="65" pageNumber="66" specimenCode="17686" specimenCount="12" stateProvince="Yogyakarta" typeStatus="paratype">
<specimenCount id="9D1EA1D38B6C4E584AB4FAB138E9FABC" box="[1153,1252,1357,1383]" count="12" pageId="65" pageNumber="66" type="generic">12 spec.</specimenCount>
(det. anat.);
<collectingMunicipality id="6BC3F0208B6C4E584E88FA8C3D43FA51" box="[189,334,1392,1418]" pageId="65" pageNumber="66">Kulonprogo</collectingMunicipality>
,
<location id="8EC73C818B6C4E584F68FA8C3DC7FA51" LSID="urn:lsid:plazi:treatment:03B1DB4C8B6C4E5B4C64FD82387DFCD7:8EC73C818B6C4E584F68FA8C3DC7FA51" box="[349,458,1392,1418]" country="Indonesia" latitude="-7.745556" longLatPrecision="21" longitude="110.13389" municipality="Kulonprogo" name="Sibolong" pageId="65" pageNumber="66" stateProvince="Yogyakarta">Sibolong</location>
,
<collectedFrom id="380429A78B6C4E584FECFA8C3E61FA51" box="[473,620,1392,1418]" pageId="65" pageNumber="66">agroforestry</collectedFrom>
;
<geoCoordinate id="EE2C0C9D8B6C4E584C4EFA923F0DFA51" box="[635,768,1390,1418]" degrees="7" direction="south" minutes="44" orientation="latitude" pageId="65" pageNumber="66" precision="15" seconds="44" value="-7.745556">7°44′44″ S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B6C4E584D25FA923FB9FA51" box="[784,948,1390,1418]" degrees="110" direction="east" minutes="08" orientation="longitude" pageId="65" pageNumber="66" precision="15" seconds="02" value="110.13389">110°08′02″ E</geoCoordinate>
;
<elevation id="00358D698B6C4E584DF1FA8C3858FA51" box="[964,1109,1392,1418]" metricMagnitude="2" metricUnit="m" metricValue="7.159999999999999" pageId="65" pageNumber="66" unit="m" value="716.0">
<quantity id="4CE0C7BF8B6C4E584DF1FA8C381FFA51" box="[964,1042,1392,1418]" metricMagnitude="2" metricUnit="m" metricValue="7.159999999999999" pageId="65" pageNumber="66" unit="m" value="716.0">716 m</quantity>
a.s.l.
</elevation>
;
<collectionCode id="ED09F29F8B6C4E584A53FA8C38ABFA51" box="[1126,1190,1392,1418]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="65" pageNumber="66" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B6C4E584A85FA8C38F4FA51" box="[1200,1273,1392,1418]" pageId="65" pageNumber="66">17686</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B7060078B6C4E584B2CFA8C3E91FA76" ID-GBIF-Occurrence="2243137399" collectionCode="MZB" country="Indonesia" elevation="716" latitude="-7.745556" location="Sibolong" longLatPrecision="21" longitude="110.13389" municipality="Kulonprogo" pageId="65" pageNumber="66" specimenCode="17702" specimenCount="9" stateProvince="Yogyakarta" typeStatus="paratype">
<specimenCount id="9D1EA1D38B6C4E584B2CFA8C397FFA51" box="[1305,1394,1392,1418]" count="9" pageId="65" pageNumber="66" type="generic">9 spec.</specimenCount>
; same data as for preceding;
<collectionCode id="ED09F29F8B6C4E584C3EFA683E46FA75" box="[523,587,1428,1454]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="65" pageNumber="66" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B6C4E584C66FA6F3E91FA76" box="[595,668,1427,1453]" pageId="65" pageNumber="66">17702</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B7060078B6C4E584C82FA6F3904FA76" ID-GBIF-Occurrence="2243137431" box="[695,1289,1427,1454]" collectionCode="ZMH" country="Indonesia" elevation="716" latitude="-7.745556" location="Sibolong" longLatPrecision="21" longitude="110.13389" municipality="Kulonprogo" pageId="65" pageNumber="66" specimenCode="ZMH 133311" specimenCount="5" stateProvince="Yogyakarta" typeStatus="paratype">
<specimenCount id="9D1EA1D38B6C4E584C82FA6F3F00FA75" box="[695,781,1427,1454]" count="5" pageId="65" pageNumber="66" type="generic">5 spec.</specimenCount>
; same data as for preceding;
<specimenCode id="DBBEC2218B6C4E584A5DFA683904FA76" box="[1128,1289,1427,1454]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="65" pageNumber="66" type="Museum">ZMH 133311</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B7060078B6C4E584B11FA6F3D0BFA2F" ID-GBIF-Occurrence="2243137323" collectedFrom="snakefruit plantation" collectionCode="MZB" country="Indonesia" elevation="716" latitude="-7.745556" location="Sibolong" longLatPrecision="21" longitude="110.13389" municipality="Kulonprogo" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Yogyakarta" typeStatus="paratype">
<specimenCount id="9D1EA1D38B6C4E584B11FA6F397AFA75" box="[1316,1399,1427,1454]" count="1" pageId="65" pageNumber="66" type="generic">1 spec.</specimenCount>
(det. anat.);
<collectingMunicipality id="6BC3F0208B6C4E584F7EFA4A3DD1FA0B" box="[331,476,1462,1488]" pageId="65" pageNumber="66">Kulonprogo</collectingMunicipality>
,
<location id="8EC73C818B6C4E584FDDFA4A3E58FA0B" LSID="urn:lsid:plazi:treatment:03B1DB4C8B6C4E5B4C64FD82387DFCD7:8EC73C818B6C4E584FDDFA4A3E58FA0B" box="[488,597,1462,1488]" country="Indonesia" latitude="-7.745556" longLatPrecision="21" longitude="110.13389" municipality="Kulonprogo" name="Sibolong" pageId="65" pageNumber="66" stateProvince="Yogyakarta">Sibolong</location>
,
<collectedFrom id="380429A78B6C4E584C54FA4A3F5AFA0B" box="[609,855,1462,1488]" pageId="65" pageNumber="66">snakefruit plantation</collectedFrom>
;
<geoCoordinate id="EE2C0C9D8B6C4E584D56FA483FE9FA0B" box="[867,996,1460,1488]" degrees="7" direction="south" minutes="44" orientation="latitude" pageId="65" pageNumber="66" precision="15" seconds="44" value="-7.745556">7°44′44″ S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B6C4E584DC4FA48389FFA0A" box="[1009,1170,1460,1489]" degrees="110" direction="east" minutes="08" orientation="longitude" pageId="65" pageNumber="66" precision="15" seconds="02" value="110.13389">110°08′02″ E</geoCoordinate>
;
<elevation id="00358D698B6C4E584AAAFA4A3924FA0B" box="[1183,1321,1462,1489]" metricMagnitude="2" metricUnit="m" metricValue="7.159999999999999" pageId="65" pageNumber="66" unit="m" value="716.0">
<quantity id="4CE0C7BF8B6C4E584AAAFA4A38E4FA0A" box="[1183,1257,1462,1489]" metricMagnitude="2" metricUnit="m" metricValue="7.159999999999999" pageId="65" pageNumber="66" unit="m" value="716.0">716 m</quantity>
a.s.l.
</elevation>
;
<collectionCode id="ED09F29F8B6C4E584B02FA4B397AFA0A" box="[1335,1399,1463,1489]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="65" pageNumber="66" type="Museum">MZB</collectionCode>
17724
</materialsCitation>
•
<materialsCitation id="3B7060078B6C4E584F14FA263F6FFA2F" ID-GBIF-Occurrence="2243137514" box="[289,866,1497,1524]" collectionCode="MZB" country="Indonesia" elevation="716" latitude="-7.745556" location="Sibolong" longLatPrecision="21" longitude="110.13389" municipality="Kulonprogo" pageId="65" pageNumber="66" specimenCode="17725" specimenCount="1" stateProvince="Yogyakarta" typeStatus="paratype">
<specimenCount id="9D1EA1D38B6C4E584F14FA263D7AFA2F" box="[289,375,1498,1524]" count="1" pageId="65" pageNumber="66" type="generic">1 spec.</specimenCount>
; same data as for preceding;
<collectionCode id="ED09F29F8B6C4E584CE4FA263F1CFA2F" box="[721,785,1498,1524]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="65" pageNumber="66" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B6C4E584D2CFA253F6FFA2F" box="[793,866,1497,1524]" pageId="65" pageNumber="66">17725</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B7060078B6C4E584D48FA26388EF9CD" ID-GBIF-Occurrence="2243137315" collectedFrom="shrubs" collectionCode="MZB" country="Indonesia" county="Kulonprogo" elevation="721" latitude="-7.7472224" location="near Kiskendo Cave" longLatPrecision="21" longitude="110.13583" municipality="Sibolong" pageId="65" pageNumber="66" specimenCode="17644" specimenCount="2" stateProvince="Yogyakarta" typeStatus="paratype">
<specimenCount id="9D1EA1D38B6C4E584D48FA263FDDFA2F" box="[893,976,1498,1524]" count="2" pageId="65" pageNumber="66" type="generic">2 spec.</specimenCount>
(det. anat.);
<collectingCounty id="62C612D68B6C4E584A5DFA2538F4FA28" box="[1128,1273,1497,1523]" pageId="65" pageNumber="66">Kulonprogo</collectingCounty>
,
<collectingMunicipality id="6BC3F0208B6C4E584B30FA25397FFA28" box="[1285,1394,1497,1523]" pageId="65" pageNumber="66">Sibolong</collectingMunicipality>
,
<location id="8EC73C818B6C4E584E88FA013DA2F9CC" LSID="urn:lsid:plazi:treatment:03B1DB4C8B6C4E5B4C64FD82387DFCD7:8EC73C818B6C4E584E88FA013DA2F9CC" box="[189,431,1533,1559]" country="Indonesia" county="Kulonprogo" latitude="-7.7472224" longLatPrecision="21" longitude="110.13583" municipality="Sibolong" name="near Kiskendo Cave" pageId="65" pageNumber="66" stateProvince="Yogyakarta">near Kiskendo Cave</location>
,
<collectedFrom id="380429A78B6C4E584F89FA013E01F9CC" box="[444,524,1533,1559]" pageId="65" pageNumber="66">shrubs</collectedFrom>
;
<geoCoordinate id="EE2C0C9D8B6C4E584C2CFA073E96F9CC" box="[537,667,1531,1559]" degrees="7" direction="south" minutes="44" orientation="latitude" pageId="65" pageNumber="66" precision="15" seconds="50" value="-7.7472224">7°44′50″ S</geoCoordinate>
,
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; same data as for preceding;
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; same data as for holotype;
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19202
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<specimenCount id="9D1EA1D38B6C4E584AB6F9DC38DAF9E1" box="[1155,1239,1568,1594]" count="3" pageId="65" pageNumber="66" type="generic">3 spec.</specimenCount>
; same data as for holotype;
<specimenCode id="DBBEC2218B6C4E584F6AF9B83E0CF986" box="[351,513,1603,1630]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="65" pageNumber="66" type="Museum">ZMH 133439</specimenCode>
</materialsCitation>
•
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; same data as for holotype;
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a.s.l.
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; same data as for preceding;
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<specimenCount id="9D1EA1D38B6C4E584FCBF9763E59F97F" box="[510,596,1674,1700]" count="2" pageId="65" pageNumber="66" type="generic">2 spec.</specimenCount>
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<collectingMunicipality id="6BC3F0208B6C4E584C54F9753EFFF978" box="[609,754,1673,1699]" pageId="65" pageNumber="66">Kulonprogo</collectingMunicipality>
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,
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.
</paragraph>
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</subSubSection>
<subSubSection id="C30239D18B6C4E5B4E88F9123842FE23" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="description">
<paragraph id="8BA76A5A8B6C4E584E88F9123D43F8D3" blockId="65.[189,334,1774,1800]" box="[189,334,1774,1800]" pageId="65" pageNumber="66">
<heading id="D0EFDD368B6C4E584E88F9123D43F8D3" bold="true" box="[189,334,1774,1800]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="B96CB6488B6C4E584E88F9123D43F8D3" bold="true" box="[189,334,1774,1800]" pageId="65" pageNumber="66">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B6C4E584E88F8E13916F83D" blockId="65.[189,1399,1820,2023]" pageId="65" pageNumber="66">
SHELL (
<figureCitation id="132376DF8B6C4E584F2FF8E03D75F8EC" box="[282,376,1820,1847]" captionStart="Figs 85–88" captionStartId="62.[189,241,1859,1885]" captionTargetBox="[232,1355,265,1830]" captionTargetId="figure@62.[232,1355,265,1830]" captionTargetPageId="62" captionText="Figs 85–88. Shells of species of Landouria from Java. 85. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 86. L. dharmai sp. nov., East Java, Tulungagung, Popoh (holotype, MZB 21272). 87. L. davini Dharma, 2015, Central Java, Baturraden, Pancuran Tujuh, Gunung Slamet (holotype, MZB 17063). 88. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152308/files/figure.png" pageId="65" pageNumber="66">Fig. 88</figureCitation>
;
<tableCitation id="C69A5FE18B6C4E584FB9F8E03E1AF8EC" box="[396,535,1820,1847]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="65" pageNumber="66" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
). Slightly depressed conical-globose, with 5.25–5.75 convex whorls; protoconch smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; without scaly processes; whitish with a distinct brown band above periphery; body whorl rounded; aperture almost circular; upper insertion of peristome distinctly descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, moderately narrow, comprising 16–24% of shell diameter, hardly obscured by columellar edge.
</paragraph>
<paragraph id="8BA76A5A8B6F4E5B4E88FEF73842FE23" blockId="66.[189,1399,267,505]" pageId="66" pageNumber="67">
GENITALIA (
<figureCitation id="132376DF8B6F4E5B4F0AFEF73D96FEFE" box="[319,411,267,293]" captionStart="Figs 29–47" captionStartId="29.[189,241,1366,1392]" captionTargetBox="[189,1399,649,1337]" captionTargetId="figure@29.[189,1399,649,1337]" captionTargetPageId="29" captionText="Figs 29–47. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm." httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="66" pageNumber="67">Figs 47</figureCitation>
,
<figureCitation id="132376DF8B6F4E5B4F93FEF73DCBFEFE" box="[422,454,267,293]" captionStart="Figs 63–71" captionStartId="48.[189,241,1668,1694]" captionTargetBox="[189,1394,515,1612]" captionTargetId="figure@48.[189,1398,510,1629]" captionTargetPageId="48" captionText="Figs 63–71. Genitalia of species of Landouria from Java. 63. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 64. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 65. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 66. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 67. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 68. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 69. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 70. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 71. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152282/files/figure.png" pageId="66" pageNumber="67">71</figureCitation>
;
<tableCitation id="C69A5FE18B6F4E5B4FE7FEF73E27FEFE" box="[466,554,267,293]" captionStart="Table 4" captionStartId="22.[188,257,551,579]" captionTargetBox="[205,1929,661,1732]" captionTargetPageId="22" captionText="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="66" pageNumber="67" tableUuid="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Table 4</tableCitation>
). Atrium short; penis short, distal part cylindrical, proximal part slightly dilated, without penis coecum; membranaceous sheath around penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to distal part of short epiphallus; flagellum long, broadest in its middle part, almost smooth, without node, tapering towards its proximal end; vas deferens narrow and long; vagina long; oviduct short; peduncle of bursa copulatrix subdivided into a short, broad distal part and a narrow and long proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B6F4E5B4E88FDD438F5FD4F" pageId="66" pageNumber="67" type="discussion">
<paragraph id="8BA76A5A8B6F4E5B4E88FDD43D22FD99" blockId="66.[189,303,552,578]" box="[189,303,552,578]" pageId="66" pageNumber="67">
<heading id="D0EFDD368B6F4E5B4E88FDD43D22FD99" bold="true" box="[189,303,552,578]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="B96CB6488B6F4E5B4E88FDD43D22FD99" bold="true" box="[189,303,552,578]" pageId="66" pageNumber="67">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B6F4E5B4E88FDAA38F5FD4F" blockId="66.[189,1398,598,660]" pageId="66" pageNumber="67">
<taxonomicName id="4C1811D98B6F4E5B4E88FDAA3DECFDAB" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,481,598,624]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="menorehensis" status="sp. nov.">
<emphasis id="B96CB6488B6F4E5B4E88FDAA3DECFDAB" box="[189,481,598,624]" italics="true" pageId="66" pageNumber="67">Landouria menorehensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B6F4E5B4FDFFDAB3E4BFDAA" box="[490,582,599,625]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
shares with
<taxonomicName id="4C1811D98B6F4E5B4CE9FDAB3F7CFDAB" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[732,881,598,624]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="pakidulan" status="sp. nov.">
<emphasis id="B96CB6488B6F4E5B4CE9FDAB3F7CFDAB" box="[732,881,598,624]" italics="true" pageId="66" pageNumber="67">L. pakidulan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B6F4E5B4D4CFDAB3FD8FDAA" box="[889,981,599,625]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1811D98B6F4E5B4A25FDAB38A1FDAB" authorityName="in Dharma" authorityYear="2015" box="[1040,1196,598,624]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="ciliocincta">
<emphasis id="B96CB6488B6F4E5B4A25FDAB38A1FDAB" box="[1040,1196,598,624]" italics="true" pageId="66" pageNumber="67">L. ciliocincta</emphasis>
</taxonomicName>
the whitish shell with a brown band at the periphery. For differences, see remarks under
<taxonomicName id="4C1811D98B6F4E5B4A34FD863898FD48" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1025,1173,633,659]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="pakidulan" status="sp. nov.">
<emphasis id="B96CB6488B6F4E5B4A34FD863898FD48" box="[1025,1173,633,659]" italics="true" pageId="66" pageNumber="67">L. pakidulan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B6F4E5B4AA8FD8638F5FD4F" box="[1181,1272,634,660]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B6F4E5B4E88FD3E387DFCD7" box="[189,1136,706,780]" pageId="66" pageNumber="67" type="distribution">
<paragraph id="8BA76A5A8B6F4E5B4E88FD3E3D5BFD07" blockId="66.[189,342,706,732]" box="[189,342,706,732]" pageId="66" pageNumber="67">
<heading id="D0EFDD368B6F4E5B4E88FD3E3D5BFD07" bold="true" box="[189,342,706,732]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="B96CB6488B6F4E5B4E88FD3E3D5BFD07" bold="true" box="[189,342,706,732]" pageId="66" pageNumber="67">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B6F4E5B4E88FD0D387DFCD7" blockId="66.[189,1136,753,780]" box="[189,1136,753,780]" pageId="66" pageNumber="67">
<taxonomicName id="4C1811D98B6F4E5B4E88FD0D3DEDFCD0" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,480,753,779]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="menorehensis" status="sp. nov.">
<emphasis id="B96CB6488B6F4E5B4E88FD0D3DEDFCD0" box="[189,480,753,779]" italics="true" pageId="66" pageNumber="67">Landouria menorehensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B6F4E5B4FDDFD0D3E4FFCD0" box="[488,578,753,779]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
is endemic to the Menoreh karst area (
<figureCitation id="132376DF8B6F4E5B4A3EFD0D3861FCD7" box="[1035,1132,753,780]" captionStart="Figs 52–55" captionStartId="33.[189,241,1717,1743]" captionTargetBox="[189,1398,265,1993]" captionTargetId="figure@34.[189,1398,265,1993]" captionTargetPageId="34" captionText="Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867)." httpUri="https://zenodo.org/record/3152265/files/figure.png" pageId="66" pageNumber="67">Fig. 55)</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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