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<mods:title id="1348C0748DC8DA50F9291B84A526E469">A taxonomic revision and a review of the biology of the North American seed-harvester ant genus Veromessor (Hymenoptera: Formicidae: Myrmicinae)</mods:title>
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<emphasis id="B94EEAF9FFE2FFC87FC5FAF35E01FA36" bold="true" box="[151,639,1407,1434]" pageId="7" pageNumber="8">
Synonymic list of species of
<taxonomicName id="4C3A4D68FFE2FFC87EA1FA0C5E01FA36" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[499,639,1408,1434]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFE2FFC87EA1FA0C5E01FA36" bold="true" box="[499,639,1408,1434]" italics="true" pageId="7" pageNumber="8">Veromessor</emphasis>
</taxonomicName>
</emphasis>
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<subSubSection id="C3206560FFE2FFC87FC5FA4E58C2F827" pageId="7" pageNumber="8" type="synonymic_list">
<paragraph id="8B8536EBFFE2FFC87FC5FA4E595EFA77" blockId="7.[151,1436,1473,1932]" box="[151,1312,1473,1499]" pageId="7" pageNumber="8">
<heading id="D0CD8187FFE2FFC87FC5FA4E595EFA77" box="[151,1312,1473,1499]" fontSize="11" level="2" pageId="7" pageNumber="8" reason="5">
<taxonomicName id="4C3A4D68FFE2FFC87FC5FA4E5DE4FA77" authority="(Mayr, 1886)" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[151,410,1473,1499]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="andrei">
<emphasis id="B94EEAF9FFE2FFC87FC5FA4E5C85FA77" box="[151,251,1473,1499]" italics="true" pageId="7" pageNumber="8">V. andrei</emphasis>
(
<bibRefCitation id="EFAB4B1AFFE2FFC87E5BFA4D5DECFA77" author="Mayr, G." box="[265,402,1473,1499]" pageId="7" pageNumber="8" pagination="419 - 464" refId="ref70555" refString="Mayr, G. (1886) Die Formiciden der Vereinigten Staaten von Nordamerika. Verhandlungen Zoologisch-Botanische Gesellschaft in Wein, 36, 419 - 464." type="journal article" year="1886">Mayr, 1886</bibRefCitation>
)
</taxonomicName>
:
<materialsCitation id="3B523CB6FFE2FFC87EF7FA4D5962FA77" ID-GBIF-Occurrence="3965991440" box="[421,1308,1473,1499]" country="Mexico" location="Baja California" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Baja California">
448 (w.q.m.l.k*).
<collectingCountry id="F32D767BFFE2FFC87D3EFA4D5EBCFA77" box="[620,706,1473,1499]" name="Mexico" pageId="7" pageNumber="8">Mexico</collectingCountry>
(
<collectingRegion id="49FEF809FFE2FFC87D82FA4D5FFCFA77" box="[720,898,1473,1499]" country="Mexico" name="Baja California" pageId="7" pageNumber="8">Baja California</collectingRegion>
), United States (California, Oregon)
</materialsCitation>
.
</heading>
</paragraph>
<paragraph id="8B8536EBFFE2FFC87FC5FA6A5FAAFA53" blockId="7.[151,1436,1473,1932]" box="[151,980,1509,1536]" pageId="7" pageNumber="8">
<taxonomicName id="4C3A4D68FFE2FFC87FC5FA6A5D82F9AC" authority="(Wheeler, 1915)" baseAuthorityName="Wheeler" baseAuthorityYear="1915" box="[151,508,1509,1536]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chamberlini">
<emphasis id="B94EEAF9FFE2FFC87FC5FA6A5D44FA53" box="[151,314,1509,1535]" italics="true" pageId="7" pageNumber="8">V. chamberlini</emphasis>
(
<bibRefCitation id="EFAB4B1AFFE2FFC87E1AFA695D8DF9AC" author="Wheeler, W. M." box="[328,499,1509,1536]" pageId="7" pageNumber="8" pagination="389 - 421" refId="ref73778" refString="Wheeler, W. M. (1915) Some additions to the North American ant-fauna. Bulletin of the American Museum of Natural History, 34, 389 - 421." type="journal article" year="1915">Wheeler, 1915</bibRefCitation>
)
</taxonomicName>
: 410 (w.q.m.l).
<collectingCountry id="F32D767BFFE2FFC87DF9FA695F3FFA53" box="[683,833,1509,1535]" name="United States of America" pageId="7" pageNumber="8">United States</collectingCountry>
(
<collectingRegion id="49FEF809FFE2FFC87C02FA695FB7FA53" box="[848,969,1509,1535]" country="United States of America" name="California" pageId="7" pageNumber="8">California</collectingRegion>
).
</paragraph>
<paragraph id="8B8536EBFFE2FFC87FC5F9865F09F98F" blockId="7.[151,1436,1473,1932]" box="[151,887,1545,1572]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="AB2A11C5FFE2FFC87FC5F9865F09F98F" box="[151,887,1545,1572]" pageId="7" pageNumber="8">
<treatmentCitation id="0A9B10FAFFE2FFC87FC5F9865D9CF98F" author="Smith, M. R." box="[151,482,1545,1572]" page="37" pageId="7" pageNumber="8" year="1956">
<taxonomicName id="4C3A4D68FFE2FFC87FC5F9865D9CF98F" authority="Smith, 1956: 37" authorityName="Smith" authorityPageNumber="37" authorityYear="1956" box="[151,482,1545,1572]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chicoensis">
<emphasis id="B94EEAF9FFE2FFC87FC5F9865D56F98F" box="[151,296,1545,1571]" italics="true" pageId="7" pageNumber="8">V. chicoensis</emphasis>
<bibRefCitation id="EFAB4B1AFFE2FFC87E7CF9855D9CF98F" author="Smith, M. R." box="[302,482,1545,1572]" pageId="7" pageNumber="8" pagination="36 - 38" refId="ref72723" refString="Smith, M. R. (1956) A key to the workers of Veromessor Forel of the United States and the description of a new subspecies. Pan-Pacific Entomologist, 32 (1), 36 - 38." type="journal article" year="1956">Smith, 1956: 37</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(w.q.m).
<collectingCountry id="F32D767BFFE2FFC87D1CF9855E9AF98F" box="[590,740,1545,1571]" name="United States of America" pageId="7" pageNumber="8">United States</collectingCountry>
(
<collectingRegion id="49FEF809FFE2FFC87DA1F9855F12F98F" box="[755,876,1545,1571]" country="United States of America" name="California" pageId="7" pageNumber="8">California</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8536EBFFE2FFC87FC5F9A258FDF9EB" blockId="7.[151,1436,1473,1932]" box="[151,1155,1581,1607]" pageId="7" pageNumber="8">
<taxonomicName id="4C3A4D68FFE2FFC87FC5F9A25DA6F9EB" authority="(Pergande, 1894)" baseAuthorityName="Pergande" baseAuthorityYear="1894" box="[151,472,1581,1607]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="julianus">
<emphasis id="B94EEAF9FFE2FFC87FC5F9A25D73F9EB" box="[151,269,1581,1607]" italics="true" pageId="7" pageNumber="8">V. julianus</emphasis>
(
<bibRefCitation id="EFAB4B1AFFE2FFC87E49F9A15DAEF9EB" author="Pergande, T." box="[283,464,1581,1607]" pageId="7" pageNumber="8" pagination="161 - 165" refId="ref71104" refString="Pergande, T. (1894) Formicidae of Lower California, Mexico. Proceedings of the California Academy of Sciences, 4 (2), 161 - 165." type="journal article" year="1894">Pergande, 1894</bibRefCitation>
)
</taxonomicName>
:
<materialsCitation id="3B523CB6FFE2FFC87EB1F9A15801F9EB" ID-GBIF-Occurrence="3965991465" box="[483,1151,1581,1607]" country="Mexico" location="Baja California" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Baja California">
164 (w.q.m).
<collectingCountry id="F32D767BFFE2FFC87D2BF9A15EB1F9EB" box="[633,719,1581,1607]" name="Mexico" pageId="7" pageNumber="8">Mexico</collectingCountry>
(
<collectingRegion id="49FEF809FFE2FFC87D8FF9A15FF3F9EB" box="[733,909,1581,1607]" country="Mexico" name="Baja California" pageId="7" pageNumber="8">Baja California</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87CCBF9A15808F9EB" box="[921,1142,1581,1607]" country="Mexico" name="Baja California Sur" pageId="7" pageNumber="8">Baja California Sur</collectingRegion>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8536EBFFE2FFC87FC5F9DE5FBBF9C7" blockId="7.[151,1436,1473,1932]" box="[151,965,1617,1644]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="AB2A11C5FFE2FFC87FC5F9DE5FBBF9C7" box="[151,965,1617,1644]" pageId="7" pageNumber="8">
<treatmentCitation id="0A9B10FAFFE2FFC87FC5F9DE5DB2F9C7" author="Smith, M. R." box="[151,460,1617,1644]" page="94" pageId="7" pageNumber="8" year="1951">
<taxonomicName id="4C3A4D68FFE2FFC87FC5F9DE5DB2F9C7" authority="Smith, 1951: 94" authorityName="Smith" authorityPageNumber="94" authorityYear="1951" box="[151,460,1617,1644]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lariversi">
<emphasis id="B94EEAF9FFE2FFC87FC5F9DE5D6CF9C7" box="[151,274,1617,1643]" italics="true" pageId="7" pageNumber="8">V. lariversi</emphasis>
<materialsCitation id="3B523CB6FFE2FFC87E4BF9DD5DB2F9C7" ID-GBIF-Occurrence="3965991482" box="[281,460,1617,1644]" country="United States of America" location="California" pageId="7" pageNumber="8" specimenCount="1" stateProvince="California">
<bibRefCitation id="EFAB4B1AFFE2FFC87E4BF9DD5DB2F9C7" author="Smith, M. R." box="[281,460,1617,1644]" pageId="7" pageNumber="8" pagination="91 - 96" refId="ref72692" refString="Smith, M. R. (1951) Two new ants from western Nevada. Great Basin Naturalist, 11 (3 - 4), 91 - 96." type="journal article" year="1951">Smith, 1951: 94</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(w.q.m).
<collectingCountry id="F32D767BFFE2FFC87D6AF9DD5EB0F9C7" box="[568,718,1617,1643]" name="United States of America" pageId="7" pageNumber="8">United States</collectingCountry>
(
<collectingRegion id="49FEF809FFE2FFC87D8FF9DD5F2DF9C7" box="[733,851,1617,1643]" country="United States of America" name="California" pageId="7" pageNumber="8">California</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87C0CF9DD5FC7F9C7" box="[862,953,1617,1643]" country="United States of America" name="Nevada" pageId="7" pageNumber="8">Nevada</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8536EBFFE2FFC87FC5F9FA5E0BF91F" blockId="7.[151,1436,1473,1932]" pageId="7" pageNumber="8">
<taxonomicName id="4C3A4D68FFE2FFC87FC5F9FA5E72F923" authority="(Andrews, 1916)" baseAuthorityName="Andrews" baseAuthorityYear="1916" box="[151,524,1653,1679]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lobognathus">
<emphasis id="B94EEAF9FFE2FFC87FC5F9FA5D41F923" box="[151,319,1653,1679]" italics="true" pageId="7" pageNumber="8">V. lobognathus</emphasis>
(
<bibRefCitation id="EFAB4B1AFFE2FFC87E02F9F95E7AF923" author="Andrews, H." box="[336,516,1653,1679]" pageId="7" pageNumber="8" pagination="81 - 83" refId="ref66007" refString="Andrews, H. (1916) A new ant of the genus Messor from Colorado. Psyche, 23 (3), 81 - 83." type="journal article" year="1916">Andrews, 1916</bibRefCitation>
)
</taxonomicName>
:
<materialsCitation id="3B523CB6FFE2FFC87D4BF9F95E0FF91F" ID-GBIF-Occurrence="3965991483" country="United States of America" location="Arizona" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Arizona">
82 (w.q.m).
<collectingCountry id="F32D767BFFE2FFC87DF7F9F95F43F923" box="[677,829,1653,1679]" name="United States of America" pageId="7" pageNumber="8">United States</collectingCountry>
(
<collectingRegion id="49FEF809FFE2FFC87C1CF9F95FD3F923" box="[846,941,1653,1679]" country="United States of America" name="Arizona" pageId="7" pageNumber="8">Arizona</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87CE8F9F9585BF923" box="[954,1061,1653,1679]" country="United States of America" name="Colorado" pageId="7" pageNumber="8">Colorado</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87B60F9F9580BF923" box="[1074,1141,1653,1679]" country="United States of America" name="Idaho" pageId="7" pageNumber="8">Idaho</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87BD0F9FA5896F93C" box="[1154,1256,1654,1680]" country="United States of America" name="Montana" pageId="7" pageNumber="8">Montana</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87BA7F9F95933F923" box="[1269,1357,1653,1679]" country="United States of America" name="Nevada" pageId="7" pageNumber="8">Nevada</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87A09F9F95D64F91F" country="United States of America" name="North Dakota" pageId="7" pageNumber="8">North Dakota</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87E77F9155D02F91F" box="[293,380,1689,1715]" country="United States of America" name="Oregon" pageId="7" pageNumber="8">Oregon</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87ED5F9155E5DF91F" box="[391,547,1689,1715]" country="United States of America" name="South Dakota" pageId="7" pageNumber="8">South Dakota</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87D7CF9155E17F91F" box="[558,617,1689,1715]" country="United States of America" name="Utah" pageId="7" pageNumber="8">Utah</collectingRegion>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8536EBFFE2FFC87FC5F9325E63F957" blockId="7.[151,1436,1473,1932]" pageId="7" pageNumber="8">
<taxonomicName id="4C3A4D68FFE2FFC87FC5F9325DB0F97B" authority="(Mayr, 1886)" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[151,462,1725,1751]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis id="B94EEAF9FFE2FFC87FC5F9325D56F97B" box="[151,296,1725,1751]" italics="true" pageId="7" pageNumber="8">V. pergandei</emphasis>
(
<bibRefCitation id="EFAB4B1AFFE2FFC87E68F9315DB8F97B" author="Mayr, G." box="[314,454,1725,1751]" pageId="7" pageNumber="8" pagination="419 - 464" refId="ref70555" refString="Mayr, G. (1886) Die Formiciden der Vereinigten Staaten von Nordamerika. Verhandlungen Zoologisch-Botanische Gesellschaft in Wein, 36, 419 - 464." type="journal article" year="1886">Mayr, 1886</bibRefCitation>
)
</taxonomicName>
:
<materialsCitation id="3B523CB6FFE2FFC87E8FF9315E64F957" ID-GBIF-Occurrence="3965991472" country="Mexico" location="Baja California" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Baja California">
448 (w.q.m).
<collectingCountry id="F32D767BFFE2FFC87D28F9315EAEF97B" box="[634,720,1725,1751]" name="Mexico" pageId="7" pageNumber="8">Mexico</collectingCountry>
(
<collectingRegion id="49FEF809FFE2FFC87DB0F9315FE8F97B" box="[738,918,1725,1751]" country="Mexico" name="Baja California" pageId="7" pageNumber="8">Baja California</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87CF7F93158F7F97B" box="[933,1161,1725,1751]" country="Mexico" name="Baja California Sur" pageId="7" pageNumber="8">Baja California Sur</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87BCAF9315892F97B" box="[1176,1260,1725,1751]" country="Mexico" name="Sonora" pageId="7" pageNumber="8">Sonora</collectingRegion>
), United States (Arizona, California, Nevada)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8536EBFFE2FFC87FC5F88A5D45F8EF" blockId="7.[151,1436,1473,1932]" pageId="7" pageNumber="8">
<taxonomicName id="4C3A4D68FFE2FFC87FC5F88A5F51F8B3" authority="Johnson, Borowiec, Snelling &amp; Cole" authorityName="Johnson, Borowiec, Snelling &amp; Cole" box="[151,815,1797,1823]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pseudolariversi">
<emphasis id="B94EEAF9FFE2FFC87FC5F88A5D16F8B3" box="[151,360,1797,1823]" italics="true" pageId="7" pageNumber="8">V. pseudolariversi</emphasis>
<materialsCitation id="3B523CB6FFE2FFC87E25F8895F51F8B3" ID-GBIF-Occurrence="3965991507" box="[375,815,1797,1823]" country="United States of America" location="California" pageId="7" pageNumber="8" specimenCount="1" stateProvince="California">Johnson, Borowiec, Snelling &amp; Cole</materialsCitation>
</taxonomicName>
(w.q.m).
<collectingCountry id="F32D767BFFE2FFC87CFBF8895839F8B3" box="[937,1095,1797,1823]" name="United States of America" pageId="7" pageNumber="8">United States</collectingCountry>
(
<collectingRegion id="49FEF809FFE2FFC87B0FF88958ADF8B3" box="[1117,1235,1797,1823]" country="United States of America" name="California" pageId="7" pageNumber="8">California</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87BB4F889593FF8B3" box="[1254,1345,1797,1823]" country="United States of America" name="Nevada" pageId="7" pageNumber="8">Nevada</collectingRegion>
).
<emphasis id="B94EEAF9FFE2FFC87A08F8895D45F8EF" bold="true" pageId="7" pageNumber="8">NEW SPECIES</emphasis>
</paragraph>
<paragraph id="8B8536EBFFE2FFC87FC5F8C259F4F8CB" blockId="7.[151,1436,1473,1932]" box="[151,1418,1869,1896]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="AB2A11C5FFE2FFC87FC5F8C259F4F8CB" box="[151,1418,1869,1896]" pageId="7" pageNumber="8">
<treatmentCitation id="0A9B10FAFFE2FFC87FC5F8C25DCCF8CB" author="Cole, A. C." box="[151,434,1869,1895]" page="678" pageId="7" pageNumber="8" year="1963">
<taxonomicName id="4C3A4D68FFE2FFC87FC5F8C25DCCF8CB" authority="Cole, 1963: 678" authorityName="Cole" authorityPageNumber="678" authorityYear="1963" box="[151,434,1869,1895]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="B94EEAF9FFE2FFC87FC5F8C25C89F8CB" box="[151,247,1869,1895]" italics="true" pageId="7" pageNumber="8">V. smithi</emphasis>
<materialsCitation id="3B523CB6FFE2FFC87FACF8C15DCCF8CB" ID-GBIF-Occurrence="3965991444" box="[254,434,1869,1895]" country="United States of America" location="Arizona" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Arizona">
<bibRefCitation id="EFAB4B1AFFE2FFC87FACF8C15DCCF8CB" author="Cole, A. C." box="[254,434,1869,1895]" pageId="7" pageNumber="8" pagination="678 - 682" refId="ref67279" refString="Cole, A. C. (1963) A new species of Veromessor from the Nevada test site and notes on related species (Hymenoptera: Formicidae). Annals of the Entomological Society of America, 56 (5), 678 - 682. https: // doi. org / 10.1093 / aesa / 56.5.678" type="journal article" year="1963">Cole, 1963: 678</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(w.q.m).
<collectingCountry id="F32D767BFFE2FFC87D4CF8C15ECAF8CB" box="[542,692,1869,1895]" name="United States of America" pageId="7" pageNumber="8">United States</collectingCountry>
(
<collectingRegion id="49FEF809FFE2FFC87D90F8C15F5FF8CB" box="[706,801,1869,1895]" country="United States of America" name="Arizona" pageId="7" pageNumber="8">Arizona</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87C7EF8C15FDEF8CB" box="[812,928,1869,1895]" country="United States of America" name="California" pageId="7" pageNumber="8">California</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87CFEF8C15869F8CB" box="[940,1047,1869,1895]" country="United States of America" name="Colorado" pageId="7" pageNumber="8">Colorado</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87B70F8C15804F8CB" box="[1058,1146,1869,1895]" country="United States of America" name="Nevada" pageId="7" pageNumber="8">Nevada</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87BD4F8C25967F8CB" box="[1158,1305,1869,1896]" country="United States of America" name="New Mexico" pageId="7" pageNumber="8">New Mexico</collectingRegion>
,
<collectingRegion id="49FEF809FFE2FFC87A77F8C15900F8CB" box="[1317,1406,1869,1895]" country="United States of America" name="Oregon" pageId="7" pageNumber="8">Oregon</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8536EBFFE2FFC87FC5F8FE58C2F827" blockId="7.[151,1436,1473,1932]" box="[151,1212,1905,1932]" pageId="7" pageNumber="8">
<taxonomicName id="4C3A4D68FFE2FFC87FC5F8FE5DB4F820" authority="(Emery, 1895)" baseAuthorityName="Emery" baseAuthorityYear="1895" box="[151,458,1905,1932]" class="Insecta" family="Formicidae" genus="Stenamma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFE2FFC87FC5F8FE5D63F827" box="[151,285,1905,1931]" italics="true" pageId="7" pageNumber="8">V. stoddardi</emphasis>
(
<bibRefCitation id="EFAB4B1AFFE2FFC87E7EF8FD5DBFF820" author="Emery, C." box="[300,449,1905,1932]" pageId="7" pageNumber="8" pagination="257 - 360" refId="ref67769" refString="Emery, C. (1895) Beitrage zur Kenntniss der nordamerikanischen Ameisenfauna. Zoologische Jahrbucher. Abtheilung fur Systematik, Geographie und Biologie der Tiere, 8, 257 - 360." type="journal article" year="1895">Emery, 1895</bibRefCitation>
)
</taxonomicName>
: 307 (w.q.m).
<collectingCountry id="F32D767BFFE2FFC87D38F8FD5EBEF827" box="[618,704,1905,1931]" name="Mexico" pageId="7" pageNumber="8">Mexico</collectingCountry>
(
<collectingRegion id="49FEF809FFE2FFC87D9CF8FD5FFEF827" box="[718,896,1905,1931]" country="Mexico" name="Baja California" pageId="7" pageNumber="8">Baja California</collectingRegion>
),
<collectingCountry id="F32D767BFFE2FFC87CC1F8FD5857F827" box="[915,1065,1905,1931]" name="United States of America" pageId="7" pageNumber="8">United States</collectingCountry>
(
<collectingRegion id="49FEF809FFE2FFC87B6AF8FD58CFF827" box="[1080,1201,1905,1931]" country="United States of America" name="California" pageId="7" pageNumber="8">California</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3206560FFE2FFC67FC5F8355EEAFF1D" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="multiple">
<paragraph id="8B8536EBFFE2FFC87FC5F8355F48F87F" blockId="7.[151,822,1977,2004]" box="[151,822,1977,2004]" pageId="7" pageNumber="8">* w = worker, q = queen, m = male, l = larva, k = karyotype</paragraph>
<caption id="DF456663FFEDFFC77FC5F9645D98F83C" ID-DOI="http://doi.org/10.5281/zenodo.7318831" ID-Zenodo-Dep="7318831" httpUri="https://zenodo.org/record/7318831/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1768,1792]" targetBox="[378,1208,181,1742]" targetPageId="8">
<paragraph id="8B8536EBFFEDFFC77FC5F9645D98F83C" blockId="8.[151,1437,1768,1936]" pageId="8" pageNumber="9">
<emphasis id="B94EEAF9FFEDFFC77FC5F9645D66F8AC" bold="true" box="[151,280,1768,1792]" pageId="8" pageNumber="9">FIGURE 1.</emphasis>
Photographs of the metasternal process on males of
<taxonomicName id="4C3A4D68FFEDFFC77C0FF9655FACF8AC" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[861,978,1769,1792]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFEDFFC77C0FF9655FACF8AC" box="[861,978,1769,1792]" italics="true" pageId="8" pageNumber="9">Veromessor</emphasis>
</taxonomicName>
: (A)
<taxonomicName id="4C3A4D68FFEDFFC77B46F965580DF8AC" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[1044,1139,1768,1792]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="andrei">
<emphasis id="B94EEAF9FFEDFFC77B46F965580DF8AC" box="[1044,1139,1768,1792]" italics="true" pageId="8" pageNumber="9">V. andrei</emphasis>
</taxonomicName>
(CASENT4010823), (B)
<taxonomicName id="4C3A4D68FFEDFFC77AD8F9655D6FF888" baseAuthorityName="Wheeler" baseAuthorityYear="1915" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="chamberlini">
<emphasis id="B94EEAF9FFEDFFC77AD8F9655D6FF888" italics="true" pageId="8" pageNumber="9">V. chamberlini</emphasis>
</taxonomicName>
(UCRC_ENT00500152), (C)
<taxonomicName id="4C3A4D68FFEDFFC77D6CF8815EC0F888" authorityName="Smith" authorityYear="1956" box="[574,702,1804,1828]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="chicoensis">
<emphasis id="B94EEAF9FFEDFFC77D6CF8815EC0F888" box="[574,702,1804,1828]" italics="true" pageId="8" pageNumber="9">V.chicoensis</emphasis>
</taxonomicName>
(CASENT0869853), (D)
<taxonomicName id="4C3A4D68FFEDFFC77CEDF8815859F888" baseAuthorityName="Pergande" baseAuthorityYear="1894" box="[959,1063,1804,1828]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="julianus">
<emphasis id="B94EEAF9FFEDFFC77CEDF8815859F888" box="[959,1063,1804,1828]" italics="true" pageId="8" pageNumber="9">V.julianus</emphasis>
</taxonomicName>
(LACMENT359792), (E)
<taxonomicName id="4C3A4D68FFEDFFC77A62F88159E2F888" box="[1328,1436,1804,1828]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lariversi">
<emphasis id="B94EEAF9FFEDFFC77A62F88159E2F888" box="[1328,1436,1804,1828]" italics="true" pageId="8" pageNumber="9">V.lariversi</emphasis>
</taxonomicName>
(CASENT0761204), (F)
<taxonomicName id="4C3A4D68FFEDFFC77EC9F8BD5E4AF8E4" baseAuthorityName="Andrews" baseAuthorityYear="1916" box="[411,564,1840,1864]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lobognathus">
<emphasis id="B94EEAF9FFEDFFC77EC9F8BD5E4AF8E4" box="[411,564,1840,1864]" italics="true" pageId="8" pageNumber="9">V. lobognathus</emphasis>
</taxonomicName>
(LACMENT363986), (G)
<taxonomicName id="4C3A4D68FFEDFFC77C02F8BD5FACF8E4" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[848,978,1840,1864]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis id="B94EEAF9FFEDFFC77C02F8BD5FACF8E4" box="[848,978,1840,1864]" italics="true" pageId="8" pageNumber="9">V. pergandei</emphasis>
</taxonomicName>
(CASENT0869850), (H)
<taxonomicName id="4C3A4D68FFEDFFC77BB1F8BD59E3F8E4" box="[1251,1437,1840,1864]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pseudolariversi">
<emphasis id="B94EEAF9FFEDFFC77BB1F8BD59E3F8E4" box="[1251,1437,1840,1864]" italics="true" pageId="8" pageNumber="9">V. pseudolariversi</emphasis>
</taxonomicName>
(CASENT0869851), (I)
<taxonomicName id="4C3A4D68FFEDFFC77ECAF8D95D8DF8C0" authorityName="Cole" authorityYear="1963" box="[408,499,1876,1900]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="B94EEAF9FFEDFFC77ECAF8D95D8DF8C0" box="[408,499,1876,1900]" italics="true" pageId="8" pageNumber="9">V. smithi</emphasis>
</taxonomicName>
(LACMENT364071), and (J)
<taxonomicName id="4C3A4D68FFEDFFC77C68F8D95FC9F8C0" baseAuthorityName="Emery" baseAuthorityYear="1895" box="[826,951,1876,1900]" class="Insecta" family="Formicidae" genus="Stenamma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFEDFFC77C68F8D95FC9F8C0" box="[826,951,1876,1900]" italics="true" pageId="8" pageNumber="9">V. stoddardi</emphasis>
</taxonomicName>
(LACMENT364102). Photographs by Robert Johnson from www.AntWeb.org.
</paragraph>
</caption>
<paragraph id="8B8536EBFFECFFC67FC5FF1B5EEAFF1D" blockId="9.[151,660,151,177]" box="[151,660,151,177]" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FF1B5EEAFF1D" bold="true" box="[151,660,151,177]" pageId="9" pageNumber="10">
Key to workers for species of
<taxonomicName id="4C3A4D68FFECFFC67D5AFF1B5EEAFF1D" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[520,660,151,177]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFECFFC67D5AFF1B5EEAFF1D" bold="true" box="[520,660,151,177]" italics="true" pageId="9" pageNumber="10">Veromessor</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3206560FFECFFC67FC5FF5059E2F9DE" pageId="9" pageNumber="10" type="key">
<key id="FFAB61DAFFECFFC67FC5FF5059E2F9DE" pageId="9" pageNumber="10">
<keyStep id="30CE2E4EFFECFFC67FC5FF5059E2FECA" pageId="9" pageNumber="10">
<paragraph id="8B8536EBFFECFFC67FC5FF5059E2FEA2" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FF5059E2FEA2" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FF505CDDFF5D" bold="true" box="[151,163,220,241]" pageId="9" pageNumber="10">1</emphasis>
Psammophore well developed, consisting of numerous, long, J-shaped hairs arranged in a distinct row along lateroventral margin of head (
<figureCitation id="13012A6EFFECFFC67ED3FF755D98FEA2" box="[385,486,249,271]" captionStart="FIGURE 6" captionStartId="21.[151,250,1642,1666]" captionTargetBox="[151,1436,181,1618]" captionTargetId="figure-18@21.[151,1436,181,1618]" captionTargetPageId="21" captionText="FIGURE 6. Photographs of the four categories of increasing psammophore development for species of Veromessor based on number and distribution of long J-shaped hairs on the ventral surface of the head capsule (= hypostomal region) (see text). Photograph of: (A) V. chicoensis (CASENT0923125)—J-shaped hairs mostly absent with scattered straight or evenly curved hairs, (B) V. andrei (CASENT0923140)—J-shaped hairs present but not arranged in a distinct row, usually mixed with straight or evenly curved hairs, (C) V. chamberlini (CASENT0761101) and (D) V. smithi (CASENT0923131)—J-shaped hairs present, arranged in a V-shaped row which does not reach the posterior part of the lateroventral margin of head capsule, usually mixed with straight or evenly curved hairs, and (E) V. pergandei (CASENT0923124)—J-shaped hairs present, many long J-shaped hairs arranged in a distinct row around the outer margin of the ventral region of the head capsule. Photographs by Wade Lee from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.7318842" httpUri="https://zenodo.org/record/7318842/files/figure.png" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67ED3FF755D98FEA2" bold="true" box="[385,486,249,271]" pageId="9" pageNumber="10">Figure 6E</emphasis>
</figureCitation>
)............................................................................
<emphasis id="B94EEAF9FFECFFC67AC2FF7559E2FEA2" bold="true" box="[1424,1436,249,270]" pageId="9" pageNumber="10">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFECFFC67FC5FE9B59E2FECA" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FE9B59E2FECA" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FE9B5CE1FE80" bold="true" box="[151,159,279,300]" pageId="9" pageNumber="10">-</emphasis>
Psammophore poorly developed, consisting of numerous, short to medium length hairs scattered across ventral surface of head; long J-shaped hairs lacking, or if present J-shaped hairs arranged in a V-shaped row not reaching posterior part of the lateroventral margin of head capsule, usually mixed with straight or evenly curved hairs (
<figureCitation id="13012A6EFFECFFC67B6EFEDD58B7FECA" box="[1084,1225,337,359]" captionStart="FIGURE 6" captionStartId="21.[151,250,1642,1666]" captionTargetBox="[151,1436,181,1618]" captionTargetId="figure-18@21.[151,1436,181,1618]" captionTargetPageId="21" captionText="FIGURE 6. Photographs of the four categories of increasing psammophore development for species of Veromessor based on number and distribution of long J-shaped hairs on the ventral surface of the head capsule (= hypostomal region) (see text). Photograph of: (A) V. chicoensis (CASENT0923125)—J-shaped hairs mostly absent with scattered straight or evenly curved hairs, (B) V. andrei (CASENT0923140)—J-shaped hairs present but not arranged in a distinct row, usually mixed with straight or evenly curved hairs, (C) V. chamberlini (CASENT0761101) and (D) V. smithi (CASENT0923131)—J-shaped hairs present, arranged in a V-shaped row which does not reach the posterior part of the lateroventral margin of head capsule, usually mixed with straight or evenly curved hairs, and (E) V. pergandei (CASENT0923124)—J-shaped hairs present, many long J-shaped hairs arranged in a distinct row around the outer margin of the ventral region of the head capsule. Photographs by Wade Lee from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.7318842" httpUri="https://zenodo.org/record/7318842/files/figure.png" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67B6EFEDD58B7FECA" bold="true" box="[1084,1225,337,359]" pageId="9" pageNumber="10">Figures 6AD</emphasis>
</figureCitation>
)...............
<emphasis id="B94EEAF9FFECFFC67AC2FEDD59E2FECA" bold="true" box="[1424,1436,337,358]" pageId="9" pageNumber="10">8</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFECFFC67FC5FEE359E2FE70" pageId="9" pageNumber="10">
<paragraph id="8B8536EBFFECFFC67FC5FEE359E3FE0E" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FEE359E3FE0E" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FEE35CDDFE28" bold="true" box="[151,163,367,388]" pageId="9" pageNumber="10">2</emphasis>
Anteromedial margin of clypeus with a small, blunt tooth or projection; in profile, eye narrowed below with a distinct ventral angle; body dark brownish-black to black..........................................................
<taxonomicName id="4C3A4D68FFECFFC67A68FE0059E3FE0E" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[1338,1437,396,418]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis id="B94EEAF9FFECFFC67A68FE0059E3FE0E" bold="true" box="[1338,1437,396,418]" italics="true" pageId="9" pageNumber="10">pergandei</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8B8536EBFFECFFC67FC5FE2659E2FE70" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FE2659E2FE70" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FE265CE1FE13" bold="true" box="[151,159,426,447]" pageId="9" pageNumber="10">-</emphasis>
Anteromedial margin of clypeus entire or concave, but lacking a small medial tooth or projection; in profile, eye rounded below; body usually concolorous yellowish or orangish-brown to rust colored, but never dark brownish-black to black....
<emphasis id="B94EEAF9FFECFFC67AC2FE4B59E2FE70" bold="true" box="[1424,1436,455,476]" pageId="9" pageNumber="10">3</emphasis>
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<keyStep id="30CE2E4EFFECFFC67FC5FE6859E2FDBA" pageId="9" pageNumber="10">
<paragraph id="8B8536EBFFECFFC67FC5FE6859E2FE55" blockId="9.[151,1437,220,1650]" box="[151,1436,484,506]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FE6859E2FE55" box="[151,1436,484,506]" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FE685CDDFE55" bold="true" box="[151,163,484,505]" pageId="9" pageNumber="10">3</emphasis>
Propodeal spines short, length &lt;0.5× maximum diameter of the eye (MOD)......................................
<emphasis id="B94EEAF9FFECFFC67AC2FE6859E2FE55" bold="true" box="[1424,1436,484,505]" pageId="9" pageNumber="10">4</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFECFFC67FC5FD8E59E2FDBA" blockId="9.[151,1437,220,1650]" box="[151,1436,513,535]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FD8E59E2FDBA" box="[151,1436,513,535]" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FD8E5CE1FDBB" bold="true" box="[151,159,514,535]" pageId="9" pageNumber="10">-</emphasis>
Propodeal spines long, length&gt; 1.0× MOD................................................................
<emphasis id="B94EEAF9FFECFFC67AC2FD8D59E2FDBA" bold="true" box="[1424,1436,513,534]" pageId="9" pageNumber="10">6</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFECFFC67FC5FD9359E2FD20" pageId="9" pageNumber="10">
<paragraph id="8B8536EBFFECFFC67FC5FD9359E2FDFE" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FD9359E2FDFE" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FD935CDDFD98" bold="true" box="[151,163,543,564]" pageId="9" pageNumber="10">4</emphasis>
Larger (HW =
<quantity id="4CC29B0EFFECFFC67E22FD935D81FD98" box="[368,511,543,565]" metricMagnitude="-3" metricUnit="m" metricValue="1.47" metricValueMax="1.61" metricValueMin="1.33" pageId="9" pageNumber="10" unit="mm" value="1.47" valueMax="1.61" valueMin="1.33">1.331.61 mm</quantity>
); maximum basal width of scape greater than maximum preapical width; body concolorous dark yellowish-orange or orangish-brown to rust colored; mandible with 8 teeth, rarely 9...........................
<taxonomicName id="4C3A4D68FFECFFC67A0DFDB059E2FDFE" authorityName="Cole" authorityYear="1963" box="[1375,1436,572,594]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="B94EEAF9FFECFFC67A0DFDB059E2FDFE" bold="true" box="[1375,1436,572,594]" italics="true" pageId="9" pageNumber="10">smithi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFECFFC67FC5FDD659E2FD20" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FDD659E2FD20" pageId="9" pageNumber="10">
- Smaller (HW =
<quantity id="4CC29B0EFFECFFC67E2CFDD65E70FDC3" box="[382,526,602,623]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.37" metricValueMin="0.93" pageId="9" pageNumber="10" unit="mm" value="1.15" valueMax="1.37" valueMin="0.93">0.931.37 mm</quantity>
); maximum basal width of scape less than maximum preapical width; body light yellowish to yellowish-orange or yellowish-red, gaster often slightly darker; mandibles with 7 teeth..............................
<emphasis id="B94EEAF9FFECFFC67AC2FDFB59E2FD20" bold="true" box="[1424,1436,631,652]" pageId="9" pageNumber="10">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFECFFC67FC5FD1859E2FC90" pageId="9" pageNumber="10">
<paragraph id="8B8536EBFFECFFC67FC5FD1859E3FCAE" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FD1859E3FCAE" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FD185CDDFD05" bold="true" box="[151,163,660,681]" pageId="9" pageNumber="10">5</emphasis>
Anteroventral margin of postpetiole interrupted by a transverse ridge, in profile appearing as a minute process, margin weakly concave anterior to process; in dorsal view, pronotum with weak, discontinuous to continuous, wavy to irregular transverse rugae; dorsum of mesonotum with one to few weak, wavy to irregular, usually discontinuous longitudinal rugae..................................................................................................
<taxonomicName id="4C3A4D68FFECFFC67A54FD6059E3FCAE" box="[1286,1437,748,770]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pseudolariversi">
<emphasis id="B94EEAF9FFECFFC67A54FD6059E3FCAE" bold="true" box="[1286,1437,748,770]" italics="true" pageId="9" pageNumber="10">pseudolariversi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFECFFC67FC5FC8659E2FC90" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FC8659E2FC90" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FC865CE1FCB3" bold="true" box="[151,159,778,799]" pageId="9" pageNumber="10">-</emphasis>
Anteroventral margin of postpetiole continuous, in profile lacking a minute process; in dorsal view, pronotum and mesonotum arugose, weakly to strongly coriarious to punctulate-granulate...........................................
<taxonomicName id="4C3A4D68FFECFFC67A19FCAA59E2FC90" box="[1355,1436,806,828]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lariversi">
<emphasis id="B94EEAF9FFECFFC67A19FCAA59E2FC90" bold="true" box="[1355,1436,806,828]" italics="true" pageId="9" pageNumber="10">lariversi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFECFFC67FC5FCC859E2FBA5" pageId="9" pageNumber="10">
<paragraph id="8B8536EBFFECFFC67FC5FCC859E2FC38" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FCC859E2FC38" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FCC85CDDFCF5" bold="true" box="[151,163,836,857]" pageId="9" pageNumber="10">6</emphasis>
Base of scape weakly widened; maximum basal width of scape less than maximum preapical width; cephalic dorsum with fine, closely spaced, regular to weakly wavy, longitudinal rugae, interrugae moderately to strongly shining; metasternal process about twice as long as high, apex nearly flat..........................................................
<taxonomicName id="4C3A4D68FFECFFC67A19FCF259E2FC38" baseAuthorityName="Pergande" baseAuthorityYear="1894" box="[1355,1436,894,916]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="julianus">
<emphasis id="B94EEAF9FFECFFC67A19FCF259E2FC38" bold="true" box="[1355,1436,894,916]" italics="true" pageId="9" pageNumber="10">julianus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFECFFC67FC5FC1159E2FBA5" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FC1159E2FBA5" pageId="9" pageNumber="10">
- Base of scape distinctly widened; maximum basal width of scape greater than maximum preapical width; cephalic dorsum with prominent widely spaced, mostly regular longitudinal rugae, interrugae shining
<emphasis id="B94EEAF9FFECFFC67CB2FC365F8BFC63" bold="true" box="[992,1013,954,975]" italics="true" pageId="9" pageNumber="10">or</emphasis>
if rugae closely spaced, then rugae strongly irregular to nearly rugoreticulate, interrugae strongly granulate, dull; metasternal process as long as to slightly longer than high, apex bluntly rounded..................................................................................
<emphasis id="B94EEAF9FFECFFC67AC2FC7859E2FBA5" bold="true" box="[1424,1436,1012,1033]" pageId="9" pageNumber="10">7</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFECFFC67FC5FB9E5D23FA83" pageId="9" pageNumber="10">
<paragraph id="8B8536EBFFECFFC67FC5FB9E59E2FBD3" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FB9E59E2FBD3" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FB9E5CDDFB8B" bold="true" box="[151,163,1042,1063]" pageId="9" pageNumber="10">7</emphasis>
Propodeal spines longer, length about 2.0× the distance between their bases; cephalic dorsum with prominent, widely spaced, mostly regular longitudinal rugae, interrugae shining; sides of pronotum with wavy to regular rugae, interrugae smooth and strongly shining; posterior margin of frontal lobes notably constricted; in frontal view, base of scape asymmetrical with ventral portion of anterior edge greatly expanded compared to dorsal edge (
<figureCitation id="13012A6EFFECFFC67C3DFBE55F9CFBD2" box="[879,994,1129,1151]" captionStart="FIGURE 12" captionStartId="29.[151,250,1849,1873]" captionTargetBox="[209,1376,181,1825]" captionTargetId="figure-18@29.[209,1379,181,1825]" captionTargetPageId="29" captionText="FIGURE 12. Photographs of dorsal surface of scape: (A) Veromessor chamberlini (LACMENT359743)—anterior edge of scape curving inward distad of base (red arrow), posterior edge of scape flat to weakly convex at base (black arrow), narrowing distad of base; widened area at base of scape much longer than width at base, and (B) V. lobognathus (LACMENT363932)—anterior edge of scape flat at base (red arrow), posterior edge of scape narrowing from base (black arrow); widened area at base of scape shorter than width at base. Photographs by Michele Esposito from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.7318854" httpUri="https://zenodo.org/record/7318854/files/figure.png" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67C3DFBE55F9CFBD2" bold="true" box="[879,994,1129,1151]" pageId="9" pageNumber="10">Figure 12A</emphasis>
</figureCitation>
).........................
<taxonomicName id="4C3A4D68FFECFFC67A76FBE559E2FBD3" ID-CoL="427ZH" baseAuthorityName="Wheeler" baseAuthorityYear="1915" box="[1316,1436,1129,1151]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="chamberlini">
<emphasis id="B94EEAF9FFECFFC67A76FBE559E2FBD3" bold="true" box="[1316,1436,1129,1151]" italics="true" pageId="9" pageNumber="10">chamberlini</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFECFFC67FC5FB0B5D23FA83" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FB0B5D23FA83" pageId="9" pageNumber="10">
- Propodeal spines shorter, length only slightly greater than to rarely 1.5× the distance between their bases; cephalic dorsum with closely spaced, strongly irregular rugae to nearly rugoreticulate, interrugae strongly granulate, dull; sides of pronotum with weakly to strongly irregular rugae to weakly rugoreticulate, interrugae moderately to strongly granulate, dull to weakly shining; posterior margin of frontal lobes not constricted; in frontal view, base of scape mostly symmetrical with ventral and dorsal portions of anterior edge similar in size or with dorsal edge expanded more than ventral edge (
<figureCitation id="13012A6EFFECFFC67BFFFB705961FABD" box="[1197,1311,1276,1298]" captionStart="FIGURE 12" captionStartId="29.[151,250,1849,1873]" captionTargetBox="[209,1376,181,1825]" captionTargetId="figure-18@29.[209,1379,181,1825]" captionTargetPageId="29" captionText="FIGURE 12. Photographs of dorsal surface of scape: (A) Veromessor chamberlini (LACMENT359743)—anterior edge of scape curving inward distad of base (red arrow), posterior edge of scape flat to weakly convex at base (black arrow), narrowing distad of base; widened area at base of scape much longer than width at base, and (B) V. lobognathus (LACMENT363932)—anterior edge of scape flat at base (red arrow), posterior edge of scape narrowing from base (black arrow); widened area at base of scape shorter than width at base. Photographs by Michele Esposito from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.7318854" httpUri="https://zenodo.org/record/7318854/files/figure.png" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67BFFFB705961FABD" bold="true" box="[1197,1311,1276,1298]" pageId="9" pageNumber="10">Figure 12B</emphasis>
</figureCitation>
).........
<taxonomicName id="4C3A4D68FFECFFC67FB0FA955D23FA83" baseAuthorityName="Andrews" baseAuthorityYear="1916" box="[226,349,1305,1327]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lobognathus">
<emphasis id="B94EEAF9FFECFFC67FB0FA955D23FA83" bold="true" box="[226,349,1305,1327]" italics="true" pageId="9" pageNumber="10">lobognathus</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFECFFC67FC5FABB59E2FA50" pageId="9" pageNumber="10">
<paragraph id="8B8536EBFFECFFC67FC5FABB59E2FA2B" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FABB59E2FA2B" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5FABB5CDDFAE0" bold="true" box="[151,163,1335,1356]" pageId="9" pageNumber="10">8</emphasis>
Propodeal spines long, in dorsal view, length&gt; 2.5× as long as distance between their bases and&gt; 2.5× MOD; hairs on scape variable in length, suberect to erect, longest hairs about 0.30.4× MOD; anterodorsal margin of pronotum with one to several coarse, irregular, transverse rugae; workers mostly monomorphic..........................................
<taxonomicName id="4C3A4D68FFECFFC67A0EFAFD59E2FA2B" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[1372,1436,1393,1415]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="andrei">
<emphasis id="B94EEAF9FFECFFC67A0EFAFD59E2FA2B" bold="true" box="[1372,1436,1393,1415]" italics="true" pageId="9" pageNumber="10">andrei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFECFFC67FC5FA0359E2FA50" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5FA0359E2FA50" pageId="9" pageNumber="10">
- Propodeal spines short, in dorsal view, length less than to about as long as distance between their bases and less than to about as long as MOD; hairs on scape mostly uniform in length, subdecumbent to suberect, longest hairs about 0.10.2× MOD; anterodorsal margin of pronotum with weak, wavy to regular longitudinal or oblique rugae, rugae rarely transverse; workers polymorphic.........................................................................................
<emphasis id="B94EEAF9FFECFFC67AC2FA6B59E2FA50" bold="true" box="[1424,1436,1511,1532]" pageId="9" pageNumber="10">9</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFECFFC67FC5F98859E2F9DE" pageId="9" pageNumber="10">
<paragraph id="8B8536EBFFECFFC67FC5F98859E3F99B" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5F98859E3F99B" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5F9885CDDF9B5" bold="true" box="[151,163,1540,1561]" pageId="9" pageNumber="10">9</emphasis>
In dorsal view, rugae on mesonotum similar to those on pronotum, interrugae weakly dull to shining; antennal scape narrowest near midlength...............................................................................
<taxonomicName id="4C3A4D68FFECFFC67A64F9AD59E3F99B" authorityName="Smith" authorityYear="1956" box="[1334,1437,1569,1591]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="chicoensis">
<emphasis id="B94EEAF9FFECFFC67A64F9AD59E3F99B" bold="true" box="[1334,1437,1569,1591]" italics="true" pageId="9" pageNumber="10">chicoensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFECFFC67FC5F9B359E2F9DE" blockId="9.[151,1437,220,1650]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5F9B359E2F9DE" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5F9B35CE1F9F8" bold="true" box="[151,159,1599,1620]" pageId="9" pageNumber="10">-</emphasis>
In dorsal view, rugae on mesonotum distinctly weaker than those on pronotum, interrugae shining; antennal scape narrowest immediately distad of basal bend.................................................................
<taxonomicName id="4C3A4D68FFECFFC67A12F9D059E2F9DE" baseAuthorityName="Emery" baseAuthorityYear="1895" box="[1344,1436,1628,1650]" class="Insecta" family="Formicidae" genus="Stenamma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFECFFC67A12F9D059E2F9DE" bold="true" box="[1344,1436,1628,1650]" italics="true" pageId="9" pageNumber="10">stoddardi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="C3206560FFECFFC67FC5F9495EFAF973" box="[151,644,1732,1759]" pageId="9" pageNumber="10" type="multiple">
<paragraph id="8B8536EBFFECFFC67FC5F9495EFAF973" blockId="9.[151,644,1732,1759]" box="[151,644,1732,1759]" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5F9495EFAF973" bold="true" box="[151,644,1732,1759]" pageId="9" pageNumber="10">
Key to queens for species of
<taxonomicName id="4C3A4D68FFECFFC67EAAF9495EFAF973" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[504,644,1733,1759]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFECFFC67EAAF9495EFAF973" bold="true" box="[504,644,1733,1759]" italics="true" pageId="9" pageNumber="10">Veromessor</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3206560FFECFFC57FC5F88559E2FBD9" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="key">
<key id="FFAB61DAFFECFFC57FC5F88559E2FBD9" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<keyStep id="30CE2E4EFFECFFC67FC5F88559E2F8DA" pageId="9" pageNumber="10">
<paragraph id="8B8536EBFFECFFC67FC5F88559E2F890" blockId="9.[151,1437,1801,2029]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5F88559E2F890" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5F8855CDDF8B2" bold="true" box="[151,163,1801,1822]" pageId="9" pageNumber="10">1</emphasis>
Psammophore well developed, consisting of numerous, long, J-shaped hairs arranged in a distinct row along lateroventral margin of head.......................................................................................
<emphasis id="B94EEAF9FFECFFC67AC2F8AB59E2F890" bold="true" box="[1424,1436,1831,1852]" pageId="9" pageNumber="10">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFECFFC67FC5F8C859E2F8DA" blockId="9.[151,1437,1801,2029]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5F8C859E2F8DA" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5F8C85CE1F8F5" bold="true" box="[151,159,1860,1881]" pageId="9" pageNumber="10">-</emphasis>
Psammophore poorly developed, consisting of numerous, short to medium length hairs scattered across ventral surface of head; long J-shaped hairs sparse or lacking.....................................................................
<emphasis id="B94EEAF9FFECFFC67AC2F8ED59E2F8DA" bold="true" box="[1424,1436,1889,1910]" pageId="9" pageNumber="10">8</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFECFFC57FC5F8F359E2FF4B" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<paragraph id="8B8536EBFFECFFC67FC5F8F359E3F863" blockId="9.[151,1437,1801,2029]" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC67FC5F8F359E3F863" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5F8F35CDDF838" bold="true" box="[151,163,1919,1940]" pageId="9" pageNumber="10">2</emphasis>
Anteromedial margin of clypeus with a small, blunt tooth or projection; in posterodorsal view, medial and posteromedial surface of petiolar node usually weakly to moderately depressed or concave; concolorous blackish to black; propodeal spines short and triangular, length much less than width at base and length less than distance between their bases......
<taxonomicName id="4C3A4D68FFECFFC67A68F83559E3F863" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[1338,1437,1977,1999]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis id="B94EEAF9FFECFFC67A68F83559E3F863" bold="true" box="[1338,1437,1977,1999]" italics="true" pageId="9" pageNumber="10">pergandei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFECFFC57FC5F85B59E2FF4B" blockId="9.[151,1437,1801,2029]" lastBlockId="10.[151,1437,151,1141]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<keyLead id="30CB95DEFFECFFC57FC5F85B59E2FF4B" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="B94EEAF9FFECFFC67FC5F85B5CE1F840" bold="true" box="[151,159,2007,2028]" pageId="9" pageNumber="10">-</emphasis>
Anterior margin of clypeus entire, lacking a small medial tooth or projection; in posterodorsal view, medial and posteromedial portion of petiolar node usually evenly rounded; concolorous orangish or orangish-brown to reddish-brown or bicolored with head and mesosoma orange and gaster blackish
<emphasis id="B94EEAF9FFEFFFC57DFCFF395EBDFF66" bold="true" box="[686,707,181,202]" italics="true" pageId="10" pageNumber="11">or</emphasis>
if concolorous dark brownish-black, then propodeal spines elongate, length greater than width at base and length similar to distance between their bases......................................
<emphasis id="B94EEAF9FFEFFFC57AC2FF5E59E2FF4B" bold="true" box="[1424,1436,210,231]" pageId="10" pageNumber="11">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5FF6359E2FE8E" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5FF6359E2FEA8" blockId="10.[151,1437,151,1141]" box="[151,1436,239,261]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FF6359E2FEA8" box="[151,1436,239,261]" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FF635CDDFEA8" bold="true" box="[151,163,239,260]" pageId="10" pageNumber="11">3</emphasis>
Maximum basal width of scape greater than maximum preapical width..........................................
<emphasis id="B94EEAF9FFEFFFC57AC2FF6359E2FEA8" bold="true" box="[1424,1436,239,260]" pageId="10" pageNumber="11">4</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5FE8159E2FE8E" blockId="10.[151,1437,151,1141]" box="[151,1436,269,291]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FE8159E2FE8E" box="[151,1436,269,291]" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FE815CE1FE8E" bold="true" box="[151,159,269,290]" pageId="10" pageNumber="11">-</emphasis>
Maximum basal width of scape less than to similar to maximum preapical width...................................
<emphasis id="B94EEAF9FFEFFFC57AC2FE8159E2FE8E" bold="true" box="[1424,1436,269,290]" pageId="10" pageNumber="11">6</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5FEA659E2FED6" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5FEA659E2FEF1" blockId="10.[151,1437,151,1141]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FEA659E2FEF1" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FEA65CDDFE93" bold="true" box="[151,163,298,319]" pageId="10" pageNumber="11">4</emphasis>
Rugae on cephalic dorsum becoming faint to disappearing above level of ocelli, lacking on posterior margin; in profile, rugae absent posterior to eyes...........................................................................
<taxonomicName id="4C3A4D68FFEFFFC57A0DFECB59E2FEF1" authorityName="Cole" authorityYear="1963" box="[1375,1436,327,349]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="B94EEAF9FFEFFFC57A0DFECB59E2FEF1" bold="true" box="[1375,1436,327,349]" italics="true" pageId="10" pageNumber="11">smithi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5FEE959E2FED6" blockId="10.[151,1437,151,1141]" box="[151,1436,357,379]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FEE959E2FED6" box="[151,1436,357,379]" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FEE95CE1FED6" bold="true" box="[151,159,357,378]" pageId="10" pageNumber="11">-</emphasis>
Rugae on cephalic dorsum continuing to posterior margin; in profile, rugae present posterior to eyes...................
<emphasis id="B94EEAF9FFEFFFC57AC2FEE959E2FED6" bold="true" box="[1424,1436,357,378]" pageId="10" pageNumber="11">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5FE0E59E3FE5C" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5FE0E59E2FE19" blockId="10.[151,1437,151,1141]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FE0E59E2FE19" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FE0E5CDDFE3B" bold="true" box="[151,163,386,407]" pageId="10" pageNumber="11">5</emphasis>
Posterior margin of frontal lobes notably constricted; in frontal view, base of scape asymmetrical with ventral portion of anterior edge greatly expanded compared to dorsal edge.............................................
<taxonomicName id="4C3A4D68FFEFFFC57A76FE1359E2FE19" ID-CoL="427ZH" baseAuthorityName="Wheeler" baseAuthorityYear="1915" box="[1316,1436,415,437]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="chamberlini">
<emphasis id="B94EEAF9FFEFFFC57A76FE1359E2FE19" bold="true" box="[1316,1436,415,437]" italics="true" pageId="10" pageNumber="11">chamberlini</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5FE3159E3FE5C" blockId="10.[151,1437,151,1141]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FE3159E3FE5C" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FE315CE1FE7E" bold="true" box="[151,159,445,466]" pageId="10" pageNumber="11">-</emphasis>
Posterior margin of frontal lobes not notably constricted; in frontal view, base of scape mostly symmetrical with ventral and dorsal portions of anterior edge similar in size or with dorsal edge expanded more than ventral edge.........
<taxonomicName id="4C3A4D68FFEFFFC57A70FE5659E3FE5C" baseAuthorityName="Andrews" baseAuthorityYear="1916" box="[1314,1437,474,496]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lobognathus">
<emphasis id="B94EEAF9FFEFFFC57A70FE5659E3FE5C" bold="true" box="[1314,1437,474,496]" italics="true" pageId="10" pageNumber="11">lobognathus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5FE7B59E2FD2E" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5FE7B59E2FD86" blockId="10.[151,1437,151,1141]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FE7B59E2FD86" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FE7B5CDDFDA0" bold="true" box="[151,163,503,524]" pageId="10" pageNumber="11">6</emphasis>
Rugae on cephalic dorsum continue onto posterior margin; propodeal spines elongate, length greater than width at base and length similar to distance between their bases; restricted to the
<collectingRegion id="49FEF809FFEFFFC57C6CFD995FAFFD87" box="[830,977,533,555]" country="Mexico" name="Baja California" pageId="10" pageNumber="11">Baja California</collectingRegion>
peninsula of
<collectingCountry id="F32D767BFFEFFFC57B01FD9958E2FD87" box="[1107,1180,533,555]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
.............
<taxonomicName id="4C3A4D68FFEFFFC57A19FD9859E2FD86" baseAuthorityName="Pergande" baseAuthorityYear="1894" box="[1355,1436,532,554]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="julianus">
<emphasis id="B94EEAF9FFEFFFC57A19FD9859E2FD86" bold="true" box="[1355,1436,532,554]" italics="true" pageId="10" pageNumber="11">julianus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5FDBE59E2FD2E" blockId="10.[151,1437,151,1141]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FDBE59E2FD2E" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FDBE5CE1FDEB" bold="true" box="[151,159,562,583]" pageId="10" pageNumber="11">-</emphasis>
Rugae on cephalic dorsum fade and disappear posterad, posterior margins smooth and shining; propodeal spines triangular, length less than width at base and length much less than distance between their bases; only known from
<collectingRegion id="49FEF809FFEFFFC57A5CFDC35911FDC9" box="[1294,1391,591,613]" country="United States of America" name="California" pageId="10" pageNumber="11">California</collectingRegion>
and
<collectingRegion id="49FEF809FFEFFFC57FB0FDE15D55FD2F" box="[226,299,621,643]" country="United States of America" name="Nevada" pageId="10" pageNumber="11">Nevada</collectingRegion>
.............................................................................................
<emphasis id="B94EEAF9FFEFFFC57AC2FDE159E2FD2E" bold="true" box="[1424,1436,621,642]" pageId="10" pageNumber="11">7</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5FD0659E2FC9E" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5FD0659E3FD76" blockId="10.[151,1437,151,1141]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FD0659E3FD76" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FD065CDDFD33" bold="true" box="[151,163,650,671]" pageId="10" pageNumber="11">7</emphasis>
In profile, anteroventral margin of postpetiole interrupted by a minute process, margin weakly concave anterior to process; smaller (HW =
<quantity id="4CC29B0EFFEFFFC57E2AFD2B5E78FD11" box="[376,518,679,701]" metricMagnitude="-3" metricUnit="m" metricValue="1.285" metricValueMax="1.36" metricValueMin="1.21" pageId="10" pageNumber="11" unit="mm" value="1.285" valueMax="1.36" valueMin="1.21">1.211.36 mm</quantity>
; ML =
<quantity id="4CC29B0EFFEFFFC57D1EFD2B5EA5FD11" box="[588,731,679,701]" metricMagnitude="-3" metricUnit="m" metricValue="1.995" metricValueMax="2.04" metricValueMin="1.95" pageId="10" pageNumber="11" unit="mm" value="1.995" valueMax="2.04" valueMin="1.95">1.952.04 mm</quantity>
); longitudinal rugae present over most to all of katepisternum...............................................................................................
<taxonomicName id="4C3A4D68FFEFFFC57A54FD4859E3FD76" box="[1286,1437,708,730]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pseudolariversi">
<emphasis id="B94EEAF9FFEFFFC57A54FD4859E3FD76" bold="true" box="[1286,1437,708,730]" italics="true" pageId="10" pageNumber="11">pseudolariversi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5FD6E59E2FC9E" blockId="10.[151,1437,151,1141]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FD6E59E2FC9E" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FD6E5CE1FD5B" bold="true" box="[151,159,738,759]" pageId="10" pageNumber="11">-</emphasis>
In profile, anteroventral margin of postpetiole continuous, lacking a minute process; larger (HW =
<quantity id="4CC29B0EFFEFFFC57B96FD6E592BFD54" box="[1220,1365,738,760]" metricMagnitude="-3" metricUnit="m" metricValue="1.575" metricValueMax="1.62" metricValueMin="1.53" pageId="10" pageNumber="11" unit="mm" value="1.575" valueMax="1.62" valueMin="1.53">1.531.62 mm</quantity>
; ML =
<quantity id="4CC29B0EFFEFFFC57FB0FC8C5D0FFCB9" box="[226,369,768,789]" metricMagnitude="-3" metricUnit="m" metricValue="2.88" metricValueMax="2.98" metricValueMin="2.78" pageId="10" pageNumber="11" unit="mm" value="2.88" valueMax="2.98" valueMin="2.78">2.782.98 mm</quantity>
); longitudinal rugae present on anterior and posterior portion of katepisternum, central portion lacking rugae..............................................................................................
<taxonomicName id="4C3A4D68FFEFFFC57A19FC9059E2FC9E" box="[1355,1436,796,818]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lariversi">
<emphasis id="B94EEAF9FFEFFFC57A19FC9059E2FC9E" bold="true" box="[1355,1436,796,818]" italics="true" pageId="10" pageNumber="11">lariversi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5FCB659E2FC68" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5FCB659E2FCC1" blockId="10.[151,1437,151,1141]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FCB659E2FCC1" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FCB65CDDFCE3" bold="true" box="[151,163,826,847]" pageId="10" pageNumber="11">8</emphasis>
Dorsal base of scape with a well developed, horizontal flange; mesoscutellum with numerous longitudinal rugae; medial lobe of clypeus with several coarse longitudinal rugae; hairs on scape variable in length, longest hairs suberect to erect...
<taxonomicName id="4C3A4D68FFEFFFC57A0EFCDB59E2FCC1" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[1372,1436,855,877]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="andrei">
<emphasis id="B94EEAF9FFEFFFC57A0EFCDB59E2FCC1" bold="true" box="[1372,1436,855,877]" italics="true" pageId="10" pageNumber="11">andrei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5FCF959E2FC68" blockId="10.[151,1437,151,1141]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FCF959E2FC68" pageId="10" pageNumber="11">
- Dorsal base of scape may be broadened but lacking a horizontal flange; mesoscutellum arugose, smooth and shining; medial lobe of clypeus smooth and shining, lacking rugae, or rugae occasionally near anterior margin; hairs on scape similar in length, all hairs decumbent...................................................................................
<emphasis id="B94EEAF9FFEFFFC57AC2FC2359E2FC68" bold="true" box="[1424,1436,943,964]" pageId="10" pageNumber="11">9</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5FC4159E2FBD9" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5FC4159E3FBB1" blockId="10.[151,1437,151,1141]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FC4159E3FBB1" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FC415CDDFC4E" bold="true" box="[151,163,973,994]" pageId="10" pageNumber="11">9</emphasis>
Smaller (HW
<quantity id="4CC29B0EFFEFFFC57E35FC415D8AFC4E" box="[359,500,973,995]" metricMagnitude="-3" metricUnit="m" metricValue="1.4849999999999999" metricValueMax="1.58" metricValueMin="1.39" pageId="10" pageNumber="11" unit="mm" value="1.4849999999999999" valueMax="1.58" valueMin="1.39">1.391.58 mm</quantity>
; ML =
<quantity id="4CC29B0EFFEFFFC57D6AFC415EB9FC4E" box="[568,711,973,995]" metricMagnitude="-3" metricUnit="m" metricValue="2.1799999999999997" metricValueMax="2.26" metricValueMin="2.1" pageId="10" pageNumber="11" unit="mm" value="2.1799999999999997" valueMax="2.26" valueMin="2.1">2.102.26 mm</quantity>
); katepisternum with longitudinal rugae; hairs on first gastral tergum longer, length of longest hairs approaching to equaling MOD; rugae on cephalic dorsum continue onto posterior margin.....................................................................................................
<taxonomicName id="4C3A4D68FFEFFFC57A64FB8B59E3FBB1" authorityName="Smith" authorityYear="1956" box="[1334,1437,1031,1053]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="chicoensis">
<emphasis id="B94EEAF9FFEFFFC57A64FB8B59E3FBB1" bold="true" box="[1334,1437,1031,1053]" italics="true" pageId="10" pageNumber="11">chicoensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5FBA959E2FBD9" blockId="10.[151,1437,151,1141]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FBA959E2FBD9" pageId="10" pageNumber="11">
- Larger (HW =
<quantity id="4CC29B0EFFEFFFC57E21FBA95E7CFB96" box="[371,514,1061,1083]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" metricValueMax="1.75" metricValueMin="1.61" pageId="10" pageNumber="11" unit="mm" value="1.68" valueMax="1.75" valueMin="1.61">1.611.75 mm</quantity>
; ML =
<quantity id="4CC29B0EFFEFFFC57D19FBA95EA5FB96" box="[587,731,1061,1082]" metricMagnitude="-3" metricUnit="m" metricValue="2.87" metricValueMax="2.97" metricValueMin="2.77" pageId="10" pageNumber="11" unit="mm" value="2.87" valueMax="2.97" valueMin="2.77">2.772.97 mm</quantity>
); katepisternum mostly smooth and shining; hairs on first gastral tergum shorter, length of longest hairs &lt;0.40.5× MOD; rugae on cephalic dorsum fade to disappear on posterior margin.....................................................................................................
<taxonomicName id="4C3A4D68FFEFFFC57A12FBD359E2FBD9" baseAuthorityName="Emery" baseAuthorityYear="1895" box="[1344,1436,1119,1141]" class="Insecta" family="Formicidae" genus="Stenamma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFEFFFC57A12FBD359E2FBD9" bold="true" box="[1344,1436,1119,1141]" italics="true" pageId="10" pageNumber="11">stoddardi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="C3206560FFEFFFC57FC5FB445E08FB4E" box="[151,630,1224,1250]" pageId="10" pageNumber="11" type="multiple">
<paragraph id="8B8536EBFFEFFFC57FC5FB445E08FB4E" blockId="10.[151,630,1224,1250]" box="[151,630,1224,1250]" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FB445E08FB4E" bold="true" box="[151,630,1224,1250]" pageId="10" pageNumber="11">
Key to males for species of
<taxonomicName id="4C3A4D68FFEFFFC57EB8FB445E08FB4E" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[490,630,1224,1250]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFEFFFC57EB8FB445E08FB4E" bold="true" box="[490,630,1224,1250]" italics="true" pageId="10" pageNumber="11">Veromessor</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3206560FFEFFFC47FC5FA8159E2FEA9" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="key">
<key id="FFAB61DAFFEFFFC47FC5FA8159E2FEA9" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<keyStep id="30CE2E4EFFEFFFC57FC5FA8159E2FA93" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5FA8159E2FA8E" blockId="10.[151,1437,1293,2019]" box="[151,1436,1293,1315]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FA8159E2FA8E" box="[151,1436,1293,1315]" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FA815CDDFA8E" bold="true" box="[151,163,1293,1314]" pageId="10" pageNumber="11">1</emphasis>
Dorsum of mesoscutum smooth and shining, rugae and/or striae lacking.........................................
<emphasis id="B94EEAF9FFEFFFC57AC2FA8159E2FA8E" bold="true" box="[1424,1436,1293,1314]" pageId="10" pageNumber="11">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5FAA659E2FA93" blockId="10.[151,1437,1293,2019]" box="[151,1436,1322,1344]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FAA659E2FA93" box="[151,1436,1322,1344]" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FAA65CE1FA93" bold="true" box="[151,159,1322,1343]" pageId="10" pageNumber="11">-</emphasis>
Rugae and/or striae cover some to most of dorsum of mesoscutum..............................................
<emphasis id="B94EEAF9FFEFFFC57AC2FAA659E2FA93" bold="true" box="[1424,1436,1322,1343]" pageId="10" pageNumber="11">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5FACB59E2FA19" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5FACB59E2FAD6" blockId="10.[151,1437,1293,2019]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FACB59E2FAD6" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FACB5CDDFAF0" bold="true" box="[151,163,1351,1372]" pageId="10" pageNumber="11">2</emphasis>
Mandibles with 23 teeth basad of preapical tooth; in profile; metasternal process triangular, acuminate (
<figureCitation id="13012A6EFFEFFFC57B82FACB594BFAF0" box="[1232,1333,1351,1373]" captionStart="FIGURE 1" captionStartId="8.[151,250,1768,1792]" captionTargetBox="[378,1208,181,1742]" captionTargetId="figure-26@8.[378,1208,181,1744]" captionTargetPageId="8" captionText="FIGURE 1. Photographs of the metasternal process on males of Veromessor: (A) V. andrei (CASENT4010823), (B) V. chamberlini (UCRC_ENT00500152), (C)V.chicoensis (CASENT0869853), (D) V.julianus (LACMENT359792),(E)V.lariversi (CASENT0761204), (F) V. lobognathus (LACMENT363986), (G) V. pergandei (CASENT0869850), (H) V. pseudolariversi (CASENT0869851), (I) V. smithi (LACMENT364071), and (J) V. stoddardi (LACMENT364102). Photographs by Robert Johnson from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.7318831" httpUri="https://zenodo.org/record/7318831/files/figure.png" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57B82FACB594BFAF0" bold="true" box="[1232,1333,1351,1373]" pageId="10" pageNumber="11">Figure 1E</emphasis>
</figureCitation>
)....................................................................................................
<taxonomicName id="4C3A4D68FFEFFFC57A19FAE859E2FAD6" box="[1355,1436,1380,1402]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lariversi">
<emphasis id="B94EEAF9FFEFFFC57A19FAE859E2FAD6" bold="true" box="[1355,1436,1380,1402]" italics="true" pageId="10" pageNumber="11">lariversi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5FA0E59E2FA19" blockId="10.[151,1437,1293,2019]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FA0E59E2FA19" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FA0E5CE1FA3B" bold="true" box="[151,159,1410,1431]" pageId="10" pageNumber="11">-</emphasis>
Mandibles with 12 minute denticles or teeth basad of preapical tooth; metasternal process elongate, slender, with a withered look (
<figureCitation id="13012A6EFFEFFFC57E49FA135D07FA18" box="[283,377,1439,1461]" captionStart="FIGURE 1" captionStartId="8.[151,250,1768,1792]" captionTargetBox="[378,1208,181,1742]" captionTargetId="figure-26@8.[378,1208,181,1744]" captionTargetPageId="8" captionText="FIGURE 1. Photographs of the metasternal process on males of Veromessor: (A) V. andrei (CASENT4010823), (B) V. chamberlini (UCRC_ENT00500152), (C)V.chicoensis (CASENT0869853), (D) V.julianus (LACMENT359792),(E)V.lariversi (CASENT0761204), (F) V. lobognathus (LACMENT363986), (G) V. pergandei (CASENT0869850), (H) V. pseudolariversi (CASENT0869851), (I) V. smithi (LACMENT364071), and (J) V. stoddardi (LACMENT364102). Photographs by Robert Johnson from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.7318831" httpUri="https://zenodo.org/record/7318831/files/figure.png" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57E49FA135D07FA18" bold="true" box="[283,377,1439,1461]" pageId="10" pageNumber="11">Figure 1I</emphasis>
</figureCitation>
).................................................................................
<taxonomicName id="4C3A4D68FFEFFFC57A0DFA1359E2FA19" authorityName="Cole" authorityYear="1963" box="[1375,1436,1439,1461]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="B94EEAF9FFEFFFC57A0DFA1359E2FA19" bold="true" box="[1375,1436,1439,1461]" italics="true" pageId="10" pageNumber="11">smithi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5FA3159E2F986" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5FA3159E3FA5C" blockId="10.[151,1437,1293,2019]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FA3159E3FA5C" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FA315CDDFA7E" bold="true" box="[151,163,1469,1490]" pageId="10" pageNumber="11">3</emphasis>
Psammophore consists of numerous, long, J-shaped hairs along lateroventral margin of head
<emphasis id="B94EEAF9FFEFFFC57B08FA305801FA7E" bold="true" box="[1114,1151,1468,1490]" italics="true" pageId="10" pageNumber="11">and</emphasis>
larger (HW =
<quantity id="4CC29B0EFFEFFFC57A59FA3159E6FA7E" box="[1291,1432,1469,1491]" metricMagnitude="-3" metricUnit="m" metricValue="1.345" metricValueMax="1.46" metricValueMin="1.23" pageId="10" pageNumber="11" unit="mm" value="1.345" valueMax="1.46" valueMin="1.23">1.231.46 mm</quantity>
; PPW =
<quantity id="4CC29B0EFFEFFFC57E7EFA565DC4FA5C" box="[300,442,1498,1520]" metricMagnitude="-3" metricUnit="m" metricValue="1.135" metricValueMax="1.29" metricValueMin="0.98" pageId="10" pageNumber="11" unit="mm" value="1.135" valueMax="1.29" valueMin="0.98">0.981.29 mm</quantity>
; EL = 0.500.59)...........................................................
<taxonomicName id="4C3A4D68FFEFFFC57A68FA5659E3FA5C" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[1338,1437,1498,1520]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis id="B94EEAF9FFEFFFC57A68FA5659E3FA5C" bold="true" box="[1338,1437,1498,1520]" italics="true" pageId="10" pageNumber="11">pergandei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5FA7459E2F986" blockId="10.[151,1437,1293,2019]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5FA7459E2F986" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5FA745CE1F9A1" bold="true" box="[151,159,1528,1549]" pageId="10" pageNumber="11">-</emphasis>
Psammophore consists of numerous hairs scattered across ventral surface of head
<emphasis id="B94EEAF9FFEFFFC57CA4FA7B5849F9A1" bold="true" box="[1014,1079,1527,1549]" italics="true" pageId="10" pageNumber="11">and/or</emphasis>
smaller (HW = 0.681.00 mm; PPW =
<quantity id="4CC29B0EFFEFFFC57FA7F9995DFDF986" box="[245,387,1557,1578]" metricMagnitude="-4" metricUnit="m" metricValue="5.35" metricValueMax="7.0" metricValueMin="3.7" pageId="10" pageNumber="11" unit="mm" value="0.5349999999999999" valueMax="0.7" valueMin="0.37">0.370.70 mm</quantity>
; EL =
<quantity id="4CC29B0EFFEFFFC57E91F9995E2EF986" box="[451,592,1557,1578]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="4.1" metricValueMin="3.3" pageId="10" pageNumber="11" unit="mm" value="0.37" valueMax="0.41" valueMin="0.33">0.330.41 mm</quantity>
)....................................................................
<emphasis id="B94EEAF9FFEFFFC57AC2F99959E2F986" bold="true" box="[1424,1436,1557,1578]" pageId="10" pageNumber="11">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5F9BE59E2F933" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5F9BE59E2F9C9" blockId="10.[151,1437,1293,2019]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5F9BE59E2F9C9" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5F9BE5CDDF9EB" bold="true" box="[151,163,1586,1607]" pageId="10" pageNumber="11">4</emphasis>
Occurring east of the Sierra
<collectingRegion id="49FEF809FFEFFFC57EAEF9BE5E3BF9E4" box="[508,581,1586,1608]" country="United States of America" name="Nevada" pageId="10" pageNumber="11">Nevada</collectingRegion>
and north of San Gabriel Mountains of
<collectingRegion id="49FEF809FFEFFFC57C8AF9BE5847F9E4" box="[984,1081,1586,1608]" country="United States of America" name="California" pageId="10" pageNumber="11">California</collectingRegion>
and east into
<collectingRegion id="49FEF809FFEFFFC57B9AF9BE596AF9E4" box="[1224,1300,1586,1608]" country="United States of America" name="Nevada" pageId="10" pageNumber="11">Nevada</collectingRegion>
,
<collectingRegion id="49FEF809FFEFFFC57A4CF9BE5910F9E4" box="[1310,1390,1586,1608]" country="United States of America" name="Arizona" pageId="10" pageNumber="11">Arizona</collectingRegion>
, and further east..........................................................................................
<emphasis id="B94EEAF9FFEFFFC57AC2F9DC59E2F9C9" bold="true" box="[1424,1436,1616,1637]" pageId="10" pageNumber="11">5</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5F9E159E2F933" blockId="10.[151,1437,1293,2019]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5F9E159E2F933" pageId="10" pageNumber="11">
- Occurring in southwestern Oregon, California west of the Sierra Nevada, southwestern California, and the
<collectingRegion id="49FEF809FFEFFFC57A55F9E159E2F92F" box="[1287,1436,1645,1667]" country="Mexico" name="Baja California" pageId="10" pageNumber="11">Baja California</collectingRegion>
peninsula,
<collectingCountry id="F32D767BFFEFFFC57E18F9065DEDF90C" box="[330,403,1674,1696]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
....................................................................................
<emphasis id="B94EEAF9FFEFFFC57AC2F90659E2F933" bold="true" box="[1424,1436,1674,1695]" pageId="10" pageNumber="11">6</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5F92459E3F8B9" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5F92459E3F976" blockId="10.[151,1437,1293,2019]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5F92459E3F976" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5F9245CDDF911" bold="true" box="[151,163,1704,1725]" pageId="10" pageNumber="11">5</emphasis>
Body blackish to black; in profile, anteroventral margin of postpetiole interrupted by small transverse ridge that appears as a minute process; anepisternum and katepisternum rugose..........................................
<taxonomicName id="4C3A4D68FFEFFFC57A54F94859E3F976" box="[1286,1437,1732,1754]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pseudolariversi">
<emphasis id="B94EEAF9FFEFFFC57A54F94859E3F976" bold="true" box="[1286,1437,1732,1754]" italics="true" pageId="10" pageNumber="11">pseudolariversi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5F96E59E3F8B9" blockId="10.[151,1437,1293,2019]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5F96E59E3F8B9" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5F96E5CE1F95B" bold="true" box="[151,159,1762,1783]" pageId="10" pageNumber="11">-</emphasis>
Body light brown to brown; in profile, anteroventral margin of postpetiole continuous, lacking a minute process; anepisternum and katepisternum mostly smooth and shining to coriarious.........................................
<taxonomicName id="4C3A4D68FFEFFFC57A70F97359E3F8B9" baseAuthorityName="Andrews" baseAuthorityYear="1916" box="[1314,1437,1791,1813]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lobognathus">
<emphasis id="B94EEAF9FFEFFFC57A70F97359E3F8B9" bold="true" box="[1314,1437,1791,1813]" italics="true" pageId="10" pageNumber="11">lobognathus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5F89159E2F8E3" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5F89159E2F89E" blockId="10.[151,1437,1293,2019]" box="[151,1436,1821,1843]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5F89159E2F89E" box="[151,1436,1821,1843]" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5F8915CDDF89E" bold="true" box="[151,163,1821,1842]" pageId="10" pageNumber="11">6</emphasis>
In profile, propodeum evenly rounded, lacking tubercles, denticles, or enlarged rugae...............................
<emphasis id="B94EEAF9FFEFFFC57AC2F89159E2F89E" bold="true" box="[1424,1436,1821,1842]" pageId="10" pageNumber="11">7</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5F8B659E2F8E3" blockId="10.[151,1437,1293,2019]" box="[151,1436,1850,1872]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5F8B659E2F8E3" box="[151,1436,1850,1872]" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5F8B65CE1F8E3" bold="true" box="[151,159,1850,1871]" pageId="10" pageNumber="11">-</emphasis>
In profile, propodeum not evenly rounded, profile interrupted by denticles, tubercles, or enlarged rugae................
<emphasis id="B94EEAF9FFEFFFC57AC2F8B659E2F8E3" bold="true" box="[1424,1436,1850,1871]" pageId="10" pageNumber="11">8</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5F8D459E2F826" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5F8D459E2F8C1" blockId="10.[151,1437,1293,2019]" box="[151,1436,1879,1901]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5F8D459E2F8C1" box="[151,1436,1879,1901]" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5F8D45CDDF8C1" bold="true" box="[151,163,1880,1901]" pageId="10" pageNumber="11">7</emphasis>
Restricted to the
<collectingRegion id="49FEF809FFEFFFC57ED7F8DB5E66F8C1" box="[389,536,1879,1901]" country="Mexico" name="Baja California" pageId="10" pageNumber="11">Baja California</collectingRegion>
peninsula of
<collectingCountry id="F32D767BFFEFFFC57DC8F8DB5E9DF8C1" box="[666,739,1879,1901]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
..................................................
<taxonomicName id="4C3A4D68FFEFFFC57A19F8DB59E2F8C1" baseAuthorityName="Pergande" baseAuthorityYear="1894" box="[1355,1436,1879,1901]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="julianus">
<emphasis id="B94EEAF9FFEFFFC57A19F8DB59E2F8C1" bold="true" box="[1355,1436,1879,1901]" italics="true" pageId="10" pageNumber="11">julianus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5F8F959E2F826" blockId="10.[151,1437,1293,2019]" box="[151,1436,1908,1931]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5F8F959E2F826" box="[151,1436,1908,1931]" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5F8F95CE1F826" bold="true" box="[151,159,1909,1930]" pageId="10" pageNumber="11">-</emphasis>
Restricted to
<collectingRegion id="49FEF809FFEFFFC57E30F8F95DBDF827" box="[354,451,1909,1931]" country="United States of America" name="California" pageId="10" pageNumber="11">California</collectingRegion>
south and west of San Gabriel Mountains.....................................
<taxonomicName id="4C3A4D68FFEFFFC57A76F8F859E2F826" ID-CoL="427ZH" baseAuthorityName="Wheeler" baseAuthorityYear="1915" box="[1316,1436,1908,1930]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="chamberlini">
<emphasis id="B94EEAF9FFEFFFC57A76F8F859E2F826" bold="true" box="[1316,1436,1908,1930]" italics="true" pageId="10" pageNumber="11">chamberlini</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC57FC5F81E59E2F868" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC57FC5F81E59E2F804" blockId="10.[151,1437,1293,2019]" box="[151,1436,1938,1960]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5F81E59E2F804" box="[151,1436,1938,1960]" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5F81E5CDDF80B" bold="true" box="[151,163,1938,1959]" pageId="10" pageNumber="11">8</emphasis>
Anepisternum rugose.............................................................................
<taxonomicName id="4C3A4D68FFEFFFC57A0EF81E59E2F804" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[1372,1436,1938,1960]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="andrei">
<emphasis id="B94EEAF9FFEFFFC57A0EF81E59E2F804" bold="true" box="[1372,1436,1938,1960]" italics="true" pageId="10" pageNumber="11">andrei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEFFFC57FC5F83C59E2F868" blockId="10.[151,1437,1293,2019]" box="[151,1436,1967,1989]" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC57FC5F83C59E2F868" box="[151,1436,1967,1989]" pageId="10" pageNumber="11">
- Anepisternum contiguously punctate-granulate to mostly smooth to roughened or with faint rugae.....................
<emphasis id="B94EEAF9FFEFFFC57AC2F82359E2F868" bold="true" box="[1424,1436,1967,1988]" pageId="10" pageNumber="11">9</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2E4EFFEFFFC47FC5F84159E2FEA9" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<paragraph id="8B8536EBFFEFFFC47FC5F84159E3FF66" blockId="10.[151,1437,1293,2019]" lastBlockId="11.[151,1437,151,261]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<keyLead id="30CB95DEFFEFFFC47FC5F84159E3FF66" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="B94EEAF9FFEFFFC57FC5F8415CDDF84E" bold="true" box="[151,163,1997,2018]" pageId="10" pageNumber="11">9</emphasis>
Anepisternum and katepisternum contiguously punctulate-granulate, anepisternum occasionally with scattered rugae; sides of propodeum contiguously punctulate-granulate with scattered, weak longitudinal rugae; in profile, area posterior to eyes and posterior corners strongly punctulate-granulate with several weak longitudinal rugae.......................
<taxonomicName id="4C3A4D68FFEEFFC47A64FF3859E3FF66" authorityName="Smith" authorityYear="1956" box="[1334,1437,180,202]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="chicoensis">
<emphasis id="B94EEAF9FFEEFFC47A64FF3859E3FF66" bold="true" box="[1334,1437,180,202]" italics="true" pageId="11" pageNumber="12">chicoensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8536EBFFEEFFC47FC5FF5E59E2FEA9" blockId="11.[151,1437,151,261]" pageId="11" pageNumber="12">
<keyLead id="30CB95DEFFEEFFC47FC5FF5E59E2FEA9" pageId="11" pageNumber="12">
<emphasis id="B94EEAF9FFEEFFC47FC5FF5E5CE1FF4B" bold="true" box="[151,159,210,231]" pageId="11" pageNumber="12">-</emphasis>
Anepisternum mostly smooth to roughened or with faint rugae; katepisternum roughened or coriarious; in profile, area posterior to eyes and posterior corners with prominent longitudinal rugae, interrugae moderately punctulate-granulate.....
<taxonomicName id="4C3A4D68FFEEFFC47A12FF6359E2FEA9" baseAuthorityName="Emery" baseAuthorityYear="1895" box="[1344,1436,239,261]" class="Insecta" family="Formicidae" genus="Stenamma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFEEFFC47A12FF6359E2FEA9" bold="true" box="[1344,1436,239,261]" italics="true" pageId="11" pageNumber="12">stoddardi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>