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<mods:title id="3F29A934A440A52E5D27EE6D0B9757BD">A taxonomic revision and a review of the biology of the North American seed-harvester ant genus Veromessor (Hymenoptera: Formicidae: Myrmicinae)</mods:title>
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<mods:namePart id="6CAAF9D2052D660643E42E38F959EED9">Snelling, Roy R.</mods:namePart>
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<paragraph id="8B8536EBFFBEFF947FC5FF1B5DE5FF1E" blockId="91.[151,423,151,250]" box="[151,411,151,178]" pageId="91" pageNumber="92">
<heading id="D0CD8187FFBEFF947FC5FF1B5DE5FF1E" bold="true" box="[151,411,151,178]" fontSize="11" level="2" pageId="91" pageNumber="92" reason="4">
<taxonomicName id="4C3A4D68FFBEFF947FC5FF1B5DE5FF1E" baseAuthorityName="Emery" baseAuthorityYear="1895" box="[151,411,151,178]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFBEFF947FC5FF1B5DE5FF1E" bold="true" box="[151,411,151,178]" italics="true" pageId="91" pageNumber="92">Veromessor stoddardi</emphasis>
</taxonomicName>
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<subSubSection id="C3206560FFBEFF947FC5FF375DD9FF56" pageId="91" pageNumber="92" type="description">
<paragraph id="8B8536EBFFBEFF947FC5FF375D0CFF7A" blockId="91.[151,423,151,250]" box="[151,370,187,214]" pageId="91" pageNumber="92">
(
<figureCitation id="13012A6EFFBEFF947FCDFF375D68FF79" box="[159,278,187,213]" captionStart="FIGURE 1" captionStartId="8.[151,250,1768,1792]" captionTargetBox="[378,1208,181,1742]" captionTargetId="figure-26@8.[378,1208,181,1744]" captionTargetPageId="8" captionText="FIGURE 1. Photographs of the metasternal process on males of Veromessor: (A) V. andrei (CASENT4010823), (B) V. chamberlini (UCRC_ENT00500152), (C)V.chicoensis (CASENT0869853), (D) V.julianus (LACMENT359792),(E)V.lariversi (CASENT0761204), (F) V. lobognathus (LACMENT363986), (G) V. pergandei (CASENT0869850), (H) V. pseudolariversi (CASENT0869851), (I) V. smithi (LACMENT364071), and (J) V. stoddardi (LACMENT364102). Photographs by Robert Johnson from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.7318831" httpUri="https://zenodo.org/record/7318831/files/figure.png" pageId="91" pageNumber="92">Figures 1J</figureCitation>
,
<figureCitation id="13012A6EFFBEFF947E70FF375D14FF7A" box="[290,362,187,214]" captionStart-0="FIGURE 48" captionStart-1="FIGURE 49" captionStart-2="FIGURE 50" captionStart-3="FIGURE 51" captionStartId-0="93.[151,250,1943,1967]" captionStartId-1="94.[151,250,1943,1967]" captionStartId-2="95.[151,250,1944,1968]" captionStartId-3="96.[151,250,1943,1967]" captionTargetBox-0="[404,1184,181,1920]" captionTargetBox-1="[420,1168,181,1919]" captionTargetBox-2="[404,1184,181,1920]" captionTargetBox-3="[405,1180,181,1920]" captionTargetId-0="figure-18@93.[404,1184,181,1920]" captionTargetId-1="figure-26@94.[420,1168,181,1919]" captionTargetId-2="figure-18@95.[404,1184,181,1920]" captionTargetId-3="figure-26@96.[405,1182,181,1920]" captionTargetPageId-0="93" captionTargetPageId-1="94" captionTargetPageId-2="95" captionTargetPageId-3="96" captionText-0="FIGURE 48. Photograph of Veromessor stoddardi minor worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT0922825). Photographs by Michele Esposito from www.AntWeb.org." captionText-1="FIGURE 49. Photograph of Veromessor stoddardi major worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT0922824). Photographs by Michele Esposito from www.AntWeb.org." captionText-2="FIGURE 50. Photograph of Veromessor stoddardi alate queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (LACMENT370303). Photographs by Wade Lee from www.AntWeb.org." captionText-3="FIGURE 51. Photograph of Veromessor stoddardi male: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (LACMENT364104). Photographs by Michele Esposito from www.AntWeb.org." figureDoi-0="http://doi.org/10.5281/zenodo.7318938" figureDoi-1="http://doi.org/10.5281/zenodo.7318940" figureDoi-2="http://doi.org/10.5281/zenodo.7318942" figureDoi-3="http://doi.org/10.5281/zenodo.7318944" httpUri-0="https://zenodo.org/record/7318938/files/figure.png" httpUri-1="https://zenodo.org/record/7318940/files/figure.png" httpUri-2="https://zenodo.org/record/7318942/files/figure.png" httpUri-3="https://zenodo.org/record/7318944/files/figure.png" pageId="91" pageNumber="92">4851</figureCitation>
)
</paragraph>
<paragraph id="8B8536EBFFBEFF947FC5FF535DD9FF56" blockId="91.[151,423,151,250]" box="[151,423,223,250]" pageId="91" pageNumber="92">
Distribution—
<figureCitation id="13012A6EFFBEFF947E69FF535DD9FF56" box="[315,423,223,250]" captionStart="FIGURE 52" captionStartId="97.[151,250,1652,1676]" captionTargetBox="[151,1434,633,1628]" captionTargetId="figure-213@97.[151,1435,633,1629]" captionTargetPageId="97" captionText="FIGURE 52. Geographic distribution of Veromessor stoddardi. The larger black circle denotes the type locality." figureDoi="http://doi.org/10.5281/zenodo.7318946" httpUri="https://zenodo.org/record/7318946/files/figure.png" pageId="91" pageNumber="92">Figure 52</figureCitation>
</paragraph>
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<treatmentCitation id="0A9B10FAFFBEFF947FC5FEA85EE4FE90" author="Emery, C." box="[151,666,291,316]" page="307" pageId="91" pageNumber="92" year="1895">
<taxonomicName id="4C3A4D68FFBEFF947FC5FEA85EE4FE90" authority="Emery, 1895: 307" authorityName="Emery" authorityPageNumber="307" authorityYear="1895" box="[151,666,291,316]" class="Insecta" family="Formicidae" genus="Stenamma" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="stoddardi" subGenus="Messor">
<emphasis id="B94EEAF9FFBEFF947FC5FEA85D7FFE97" box="[151,257,292,315]" italics="true" pageId="91" pageNumber="92">Stenamma</emphasis>
(
<emphasis id="B94EEAF9FFBEFF947E46FEA85D21FE97" box="[276,351,292,315]" italics="true" pageId="91" pageNumber="92">Messor</emphasis>
)
<emphasis id="B94EEAF9FFBEFF947E20FEA85DACFE90" box="[370,466,292,316]" italics="true" pageId="91" pageNumber="92">stoddardi</emphasis>
<bibRefCitation id="EFAB4B1AFFBEFF947E8EFEA85EE4FE90" author="Emery, C." box="[476,666,291,316]" pageId="91" pageNumber="92" pagination="257 - 360" refId="ref67769" refString="Emery, C. (1895) Beitrage zur Kenntniss der nordamerikanischen Ameisenfauna. Zoologische Jahrbucher. Abtheilung fur Systematik, Geographie und Biologie der Tiere, 8, 257 - 360." type="journal article" year="1895">Emery, 1895: 307</bibRefCitation>
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(worker).
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examined:
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[
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],
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[
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,
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#55466,
<collectingCountry id="F32D767BFFBEFF947E70FECF5D9AFEF6" box="[290,484,322,346]" name="United States of America" pageId="91" pageNumber="92">UNITED STATES</collectingCountry>
,
<collectingRegion id="49FEF809FFBEFF947EA2FECE5E25FEF6" box="[496,603,322,346]" country="United States of America" name="California" pageId="91" pageNumber="92">California</collectingRegion>
: Riverside County, San Jacinto (Hyatt?). See also
<bibRefCitation id="EFAB4B1AFFBEFF947B3BFECE59E3FEF6" author="Wheeler, W. M. &amp; Creighton, W. S." box="[1129,1437,322,346]" pageId="91" pageNumber="92" pagination="341 - 387" refId="ref73843" refString="Wheeler, W. M. &amp; Creighton, W. S. (1934) A study of the ant genera Novomessor and Veromessor. Proceedings of the American Academy of Arts and Sciences, 69 (9), 341 - 387. https: // doi. org / 10.2307 / 20023057" type="journal article" year="1934">Wheeler and Creighton 1934:</bibRefCitation>
385, plate II, fig. 1. USNM worker here designated
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<typeStatus id="54818849FFBEFF947D81FEED5F11FED5" box="[723,879,353,377]" pageId="91" pageNumber="92" type="lectotype">LECTOTYPE</typeStatus>
</emphasis>
[USNMENT00922798].
</paragraph>
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<taxonomicName id="4C3A4D68FFBEFF947FC5FE0C5DA7FE34" authority="(Emery)" baseAuthorityName="Emery" box="[151,473,384,408]" class="Insecta" family="Formicidae" genus="Novomessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFBEFF947FC5FE0C5D05FE34" box="[151,379,384,408]" italics="true" pageId="91" pageNumber="92">Novomessor stoddardi</emphasis>
(Emery)
</taxonomicName>
;
<bibRefCitation id="EFAB4B1AFFBEFF947EB1FE0C5EF5FE34" author="Emery, C." box="[483,651,383,408]" pageId="91" pageNumber="92" pagination="67 - 75" refId="ref67802" refString="Emery, C. (1915) Definizione del genere Aphaenogaster e partizione di esso in sottogeneri. Parapheidole e Novomessor nn. gg. Rendiconti delle Sessioni della Reale Accademia delle Scienze dell'Istituto di Bologna. Classe di Scienze Fisiche, New Series, 19, 67 - 75." type="journal article" year="1915">Emery, 1915: 73</bibRefCitation>
(first combination in
<taxonomicName id="4C3A4D68FFBEFF947C3AFE0C5F9BFE3B" baseAuthorityName="Donisthorpe" baseAuthorityYear="1943" box="[872,997,384,407]" class="Insecta" family="Formicidae" genus="Novomessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFBEFF947C3AFE0C5F9BFE3B" box="[872,997,384,407]" italics="true" pageId="91" pageNumber="92">Novomessor</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B8536EBFFBEFF947FC5FE13596BFE1A" blockId="91.[151,1437,291,622]" box="[151,1301,414,438]" pageId="91" pageNumber="92">
<taxonomicName id="4C3A4D68FFBEFF947FC5FE135E1BFE1A" authority="(Emery)" baseAuthorityName="Emery" box="[151,613,414,438]" class="Insecta" family="Formicidae" genus="Novomessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="stoddardi" subGenus="Veromessor">
<emphasis id="B94EEAF9FFBEFF947FC5FE135D6AFE1A" box="[151,276,415,438]" italics="true" pageId="91" pageNumber="92">Novomessor</emphasis>
(
<emphasis id="B94EEAF9FFBEFF947E71FE135DE6FE1A" box="[291,408,415,438]" italics="true" pageId="91" pageNumber="92">Veromessor</emphasis>
)
<emphasis id="B94EEAF9FFBEFF947EF5FE125E79FE1A" box="[423,519,414,438]" italics="true" pageId="91" pageNumber="92">stoddardi</emphasis>
(Emery)
</taxonomicName>
;
<bibRefCitation id="EFAB4B1AFFBEFF947D22FE125F66FE1A" author="Forel, A." box="[624,792,414,438]" pageId="91" pageNumber="92" pagination="229 - 253" refId="ref68064" refString="Forel, A. (1917) Cadre synoptique actuel de la faune universelle des fourmis. Bulletin de la Societe Vaudoise des Sciences Naturelles, 51, 229 - 253." type="journal article" year="1917">Forel, 1917: 235</bibRefCitation>
(first combination in
<taxonomicName id="4C3A4D68FFBEFF947CA7FE13580CFE1A" baseAuthorityName="Donisthorpe" baseAuthorityYear="1943" box="[1013,1138,415,438]" class="Insecta" family="Formicidae" genus="Novomessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFBEFF947CA7FE13580CFE1A" box="[1013,1138,415,438]" italics="true" pageId="91" pageNumber="92">Novomessor</emphasis>
</taxonomicName>
([
<taxonomicName id="4C3A4D68FFBEFF947BDBFE135880FE1A" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[1161,1278,415,438]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFBEFF947BDBFE135880FE1A" box="[1161,1278,415,438]" italics="true" pageId="91" pageNumber="92">Veromessor</emphasis>
</taxonomicName>
]).
</paragraph>
<paragraph id="8B8536EBFFBEFF947FC5FE3258E4FE79" blockId="91.[151,1437,291,622]" box="[151,1178,445,469]" pageId="91" pageNumber="92">
<taxonomicName id="4C3A4D68FFBEFF947FC5FE325DAFFE79" authority="(Emery)" baseAuthorityName="Emery" box="[151,465,445,469]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFBEFF947FC5FE325D0DFE79" box="[151,371,445,469]" italics="true" pageId="91" pageNumber="92">Veromessor stoddardi</emphasis>
(Emery)
</taxonomicName>
;
<bibRefCitation id="EFAB4B1AFFBEFF947E89FE315F44FE79" author="Wheeler, W. M. &amp; Creighton, W. S." box="[475,826,445,469]" pageId="91" pageNumber="92" pagination="341 - 387" refId="ref73843" refString="Wheeler, W. M. &amp; Creighton, W. S. (1934) A study of the ant genera Novomessor and Veromessor. Proceedings of the American Academy of Arts and Sciences, 69 (9), 341 - 387. https: // doi. org / 10.2307 / 20023057" type="journal article" year="1934">Wheeler and Creighton, 1934: 385</bibRefCitation>
(first combination in
<taxonomicName id="4C3A4D68FFBEFF947B45FE3258F2FE79" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[1047,1164,446,469]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFBEFF947B45FE3258F2FE79" box="[1047,1164,446,469]" italics="true" pageId="91" pageNumber="92">Veromessor</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B8536EBFFBEFF947FC5FE505C85FDBE" blockId="91.[151,1437,291,622]" pageId="91" pageNumber="92">
<taxonomicName id="4C3A4D68FFBEFF947FC5FE505E0FFE58" authority="(Emery)" baseAuthorityName="Emery" box="[151,625,476,500]" class="Insecta" family="Formicidae" genus="Novomessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="stoddardi" subGenus="Veromessor">
<emphasis id="B94EEAF9FFBEFF947FC5FE505D6AFE5F" box="[151,276,476,499]" italics="true" pageId="91" pageNumber="92">Novomessor</emphasis>
(
<emphasis id="B94EEAF9FFBEFF947E75FE505DE2FE5F" box="[295,412,476,499]" italics="true" pageId="91" pageNumber="92">Veromessor</emphasis>
)
<emphasis id="B94EEAF9FFBEFF947EFDFE505E71FE58" box="[431,527,476,500]" italics="true" pageId="91" pageNumber="92">stoddardi</emphasis>
(Emery)
</taxonomicName>
;
<bibRefCitation id="EFAB4B1AFFBEFF947DD2FE505F24FE5F" author="Enzmann, J." box="[640,858,475,500]" pageId="91" pageNumber="92" pagination="147 - 152" refId="ref67881" refString="Enzmann, J. (1947) New forms of Aphaenogaster and Novomessor. Journal of the New York Entomological Society, 55 (2), 147 - 152." type="journal article" year="1947">Enzmann, 1947: 152</bibRefCitation>
(revived combination in
<taxonomicName id="4C3A4D68FFBEFF947B38FE505899FE5F" baseAuthorityName="Donisthorpe" baseAuthorityYear="1943" box="[1130,1255,476,499]" class="Insecta" family="Formicidae" genus="Novomessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFBEFF947B38FE505899FE5F" box="[1130,1255,476,499]" italics="true" pageId="91" pageNumber="92">Novomessor</emphasis>
</taxonomicName>
[
<taxonomicName id="4C3A4D68FFBEFF947BA8FE505911FE5F" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[1274,1391,476,499]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFBEFF947BA8FE505911FE5F" box="[1274,1391,476,499]" italics="true" pageId="91" pageNumber="92">Veromessor</emphasis>
</taxonomicName>
], in key).
</paragraph>
<paragraph id="8B8536EBFFBEFF947FC5FD965854FD9D" blockId="91.[151,1437,291,622]" box="[151,1066,537,561]" pageId="91" pageNumber="92">
<taxonomicName id="4C3A4D68FFBEFF947FC5FD965DBAFD9D" authority="(Mayr)" baseAuthorityName="Mayr" box="[151,452,537,561]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFBEFF947FC5FD965D0DFD9D" box="[151,371,537,561]" italics="true" pageId="91" pageNumber="92">Veromessor stoddardi</emphasis>
(Mayr)
</taxonomicName>
;
<bibRefCitation id="EFAB4B1AFFBEFF947E9CFD955ED8FD9D" author="Creighton, W. S." box="[462,678,537,561]" pageId="91" pageNumber="92" pagination="1 - 585" refId="ref67372" refString="Creighton, W. S. (1950) The ants of North America. Bulletin of the Museum of Comparative Zoology, 104, 1 - 585." type="journal article" year="1950">Creighton, 1950: 157</bibRefCitation>
(revived combination in
<taxonomicName id="4C3A4D68FFBEFF947CF5FD965862FD9D" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[935,1052,538,561]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFBEFF947CF5FD965862FD9D" box="[935,1052,538,561]" italics="true" pageId="91" pageNumber="92">Veromessor</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B8536EBFFBEFF947FC5FDB45FE0FDFC" blockId="91.[151,1437,291,622]" box="[151,926,568,592]" pageId="91" pageNumber="92">
<taxonomicName id="4C3A4D68FFBEFF947FC5FDB45DD8FDFC" ID-CoL="4283H" authority="(Emery)" baseAuthorityName="Emery" baseAuthorityYear="1895" box="[151,422,568,592]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFBEFF947FC5FDB45D36FDFC" box="[151,328,568,592]" italics="true" pageId="91" pageNumber="92">Messor stoddardi</emphasis>
(Emery)
</taxonomicName>
;
<bibRefCitation id="EFAB4B1AFFBEFF947EE2FDB45E16FDFC" author="Bolton, B." box="[432,616,568,592]" pageId="91" pageNumber="92" pagination="307 - 370" refId="ref66249" refString="Bolton, B. (1982) Afrotropical species of the myrmicine ant genera Cardiocondyla, Leptothorax, Melissotarsus, Messor, and Cataulcus (Formicidae). Bulletin of the British Museum (Natural History) Entomology, 45 (4), 307 - 370." type="journal article" year="1982">Bolton, 1982: 341</bibRefCitation>
(first combination in
<taxonomicName id="4C3A4D68FFBEFF947C17FDB45FEEFDE3" ID-CoL="5Q45" authorityName="Forel" authorityYear="1890" box="[837,912,568,591]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFBEFF947C17FDB45FEEFDE3" box="[837,912,568,591]" italics="true" pageId="91" pageNumber="92">Messor</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B8536EBFFBEFF947FC5FDDB58CEFDC2" blockId="91.[151,1437,291,622]" box="[151,1200,598,622]" pageId="91" pageNumber="92">
<taxonomicName id="4C3A4D68FFBEFF947FC5FDDB5F52FDC2" authority="Ward, Brady, Fisher, and Schultz, 2015: 13" authorityName="Ward, Brady, Fisher, and Schultz" authorityPageNumber="13" authorityYear="2015" box="[151,812,598,622]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFBEFF947FC5FDDB5D0DFDC2" box="[151,371,598,622]" italics="true" pageId="91" pageNumber="92">Veromessor stoddardi</emphasis>
<bibRefCitation id="EFAB4B1AFFBEFF947E2BFDDA5F52FDC2" author="Ward, P. S. &amp; Brady, S. G. &amp; Fisher, B. L. &amp; Schultz, T. R." box="[377,812,598,622]" pageId="91" pageNumber="92" pagination="61 - 81" refId="ref73206" refString="Ward, P. S., Brady, S. G., Fisher, B. L. &amp; Schultz, T. R. (2015) The evolution of myrmicine ants: phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Systematic Entomology, 40 (1), 61 - 81. https: // doi. org / 10.1111 / syen. 12090" type="journal article" year="2015">Ward, Brady, Fisher, and Schultz, 2015: 13</bibRefCitation>
</taxonomicName>
(revived combination in
<taxonomicName id="4C3A4D68FFBEFF947B7FFDDB58DCFDC2" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[1069,1186,599,622]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FFBEFF947B7FFDDB58DCFDC2" box="[1069,1186,599,622]" italics="true" pageId="91" pageNumber="92">Veromessor</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B8536EBFFBEFF947FC5FD115E63FB03" blockId="91.[151,1437,668,2027]" pageId="91" pageNumber="92">
<emphasis id="B94EEAF9FFBEFF947FC5FD115D0CFD1A" bold="true" box="[151,370,668,695]" pageId="91" pageNumber="92">Worker diagnosis.</emphasis>
This species is uniquely characterized by the following combination of features: (1) head and mesosoma orangish-red to orangish-brown, gaster blackish-orange to blackish, (2) medial lobe of clypeus lacking medial groove but with several lateral longitudinal rugae, medial lobe not thick and protuberant in profile, not elevated above lateral lobes in frontal view, (3) mandibles with 8 teeth, (4) antennal scape narrowest immediately distad of basal bend, (5) MOD distinctly less than OMD, OI &lt;25.0, (6) cephalic dorsum moderately shining between fine, widely spaced, longitudinal rugae; antennal fossa lacking concentric rugae, (7) psammophore poorly developed; ventral surface of head capsule with scattered straight or evenly curved hairs, but J-shaped hairs mostly absent, (8) dorsum of pronotum with fine, wavy to weakly irregular, longitudinal rugae; sides of pronotum with longitudinal rugae that fade to disappear posterad, posterior one-third often granulate; mesonotum mostly smooth and shining, lacking rugae or with longitudinal rugae that are distinctly weaker than those on dorsum of pronotum; mesopleura with rugae that traverse longitudinally to posterodorsally, interrugae weakly coriarious to moderately granulate, (9) propodeal spines short, acuminate, length less than distance between their bases; infraspinal facet and propodeal declivity rugose or weakly to strongly coriarious, weakly to moderately shining, and (10) metasternal process large, narrowed laterally, slightly higher than long with angulate apex; anterior and posterior surfaces decline steeply; partly translucent (
<figureCitation id="13012A6EFFBEFF947E3BFB185E6FFB03" box="[361,529,1172,1199]" captionStart-0="FIGURE 48" captionStart-1="FIGURE 49" captionStartId-0="93.[151,250,1943,1967]" captionStartId-1="94.[151,250,1943,1967]" captionTargetBox-0="[404,1184,181,1920]" captionTargetBox-1="[420,1168,181,1919]" captionTargetId-0="figure-18@93.[404,1184,181,1920]" captionTargetId-1="figure-26@94.[420,1168,181,1919]" captionTargetPageId-0="93" captionTargetPageId-1="94" captionText-0="FIGURE 48. Photograph of Veromessor stoddardi minor worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT0922825). Photographs by Michele Esposito from www.AntWeb.org." captionText-1="FIGURE 49. Photograph of Veromessor stoddardi major worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT0922824). Photographs by Michele Esposito from www.AntWeb.org." figureDoi-0="http://doi.org/10.5281/zenodo.7318938" figureDoi-1="http://doi.org/10.5281/zenodo.7318940" httpUri-0="https://zenodo.org/record/7318938/files/figure.png" httpUri-1="https://zenodo.org/record/7318940/files/figure.png" pageId="91" pageNumber="92">Figures 4849</figureCitation>
).
</paragraph>
<paragraph id="8B8536EBFFBEFF947F95FB355E0CFA93" blockId="91.[151,1437,668,2027]" pageId="91" pageNumber="92">
<emphasis id="B94EEAF9FFBEFF947F95FB355D08FB7F" bold="true" box="[199,374,1209,1235]" pageId="91" pageNumber="92">Measurements</emphasis>
.
<typeStatus id="54818849FFBEFF947ED6FB345D95FB7E" box="[388,491,1208,1234]" pageId="91" pageNumber="92" type="lectotype">lectotype</typeStatus>
(
<emphasis id="B94EEAF9FFBEFF947EA9FB355E77FB7E" box="[507,521,1209,1234]" italics="true" pageId="91" pageNumber="92">n</emphasis>
= 66). HL 1.72 (1.031.80); HW 1.80 (0.931.83); MOD 0.33 (0.210.33); OMD 0.44 (0.230.44); SL 1.22 (0.831.45); PNW 1.02 (0.621.11); HFL 1.63 (0.921.71); ML 2.20 (1.272.20); PW 0.34 (0.190.38); PPW 0.50 (0.260.52). Indices: SI 67.68 (66.85100.69); CI 104.65 (87.07109.20); OI 18.33 (16.0924.75); HFI 90.56 (68.03114.85).
</paragraph>
<paragraph id="8B8536EBFFBEFF947F95FAC45F7EF8F7" blockId="91.[151,1437,668,2027]" pageId="91" pageNumber="92">
<emphasis id="B94EEAF9FFBEFF947F95FAC45DEDFACE" bold="true" box="[199,403,1352,1378]" pageId="91" pageNumber="92">Queen diagnosis.</emphasis>
This caste is diagnosed by the following combination of features: (1) head dark orangishbrown to reddish-brown, gaster slightly lighter; pronotum, dorsal portion of anepisternum, mesoscutellum, propodeum dark orangish-brown; rest of mesosoma dark blackish-orange to blackish, (2) medial lobe of clypeus lacking medial groove but with several lateral longitudinal rugae, (3) mandibles with 8 teeth, (4) dorsal surface of base of scape slightly flattened; maximum basal width of scape similar to maximum preapical width, (5) MOD slightly greater than OMD, (6) cephalic dorsum with fine, wavy to irregular longitudinal rugae that fade to disappear near posterior margin; interrugae moderately shining, (7) psammophore poorly developed, (8) sides of pronotum weakly to moderately punctate between fine longitudinal rugae; mesoscutum and mesoscutellum smooth and shining with scattered piligerous punctures; anepisternum shining, weakly punctate between fine longitudinal rugae, both rugae and punctuation weak to absent ventrad; katepisternum mostly shining and superficially roughened with short, longitudinal striae anterad and posterad, (9) sides of propodeum weakly shining and densely punctate between longitudinal and oblique rugae; propodeal spines triangular, acuminate, distinctly shorter than distance between their bases; infraspinal facet shining, superficially punctate above, propodeal declivity smooth and shining, and (10) metasternal process large, narrowed laterally, slightly higher than long with angulate apex; anterior and posterior surfaces decline steeply; partly translucent (
<figureCitation id="13012A6EFFBEFF947DD0F8CC5E8AF8F7" box="[642,756,1856,1883]" captionStart="FIGURE 50" captionStartId="95.[151,250,1944,1968]" captionTargetBox="[404,1184,181,1920]" captionTargetId="figure-18@95.[404,1184,181,1920]" captionTargetPageId="95" captionText="FIGURE 50. Photograph of Veromessor stoddardi alate queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (LACMENT370303). Photographs by Wade Lee from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.7318942" httpUri="https://zenodo.org/record/7318942/files/figure.png" pageId="91" pageNumber="92">Figure 50</figureCitation>
).
</paragraph>
<paragraph id="8B8536EBFFBEFF947F95F8E95E76F86B" blockId="91.[151,1437,668,2027]" pageId="91" pageNumber="92">
<emphasis id="B94EEAF9FFBEFF947F95F8E95D08F8D3" bold="true" box="[199,374,1893,1919]" pageId="91" pageNumber="92">Measurements</emphasis>
. (
<emphasis id="B94EEAF9FFBEFF947EC0F8E95DDEF8D2" box="[402,416,1893,1918]" italics="true" pageId="91" pageNumber="92">n</emphasis>
= 6). HL 1.561.72; HW 1.611.75; MOD 0.400.44; OMD 0.340.41; SL 1.201.32; HFL 1.611.69; ML 2.772.97; PW 0.390.45; PPW 0.610.67. Indices: SI 74.0776.97; CI 96.99110.76; OI 24.1226.71; HFI 93.71101.24.
</paragraph>
<paragraph id="8B8536EBFFBEFF937F95F85D5FD2FE01" blockId="91.[151,1437,668,2027]" lastBlockId="92.[151,1437,151,2043]" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92">
<emphasis id="B94EEAF9FFBEFF947F95F85D5DF8F846" bold="true" box="[199,390,2000,2027]" pageId="91" pageNumber="92">Male diagnosis.</emphasis>
This caste is diagnosed by the following combination of features: (11) blackish brown, appendages light brown, (12) medial lobe of clypeus abruptly descendant distad, disc weakly shining with coarse oblique to longitudinal rugae, (13) mandibles with 34 teeth basad of preapical tooth, (14) anterior ocellus above level of top of eyes, (15) anepisternum weakly shining, finely punctate between closely spaced, moderately coarse, longitudinal rugae; katepisternum shinier, disc weakly roughened, posterad with moderately coarse oblique to longitudinal rugae, (16) propodeum densely punctate between coarse, mostly longitudinal rugae, interrugae weakly shining; propodeal spines triangular, acuminate, length less than distance between their bases, (17) metasternal process distinctly higher than long, apex narrowly rounded, and (18) subpetiolar process short, elongate-triangular to spiniform or digitiform, apex acuminate to rounded (
<figureCitation id="13012A6EFFB9FF937C51FE1F5F0AFE01" box="[771,884,403,429]" captionStart="FIGURE 1" captionStartId="8.[151,250,1768,1792]" captionTargetBox="[378,1208,181,1742]" captionTargetId="figure-26@8.[378,1208,181,1744]" captionTargetPageId="8" captionText="FIGURE 1. Photographs of the metasternal process on males of Veromessor: (A) V. andrei (CASENT4010823), (B) V. chamberlini (UCRC_ENT00500152), (C)V.chicoensis (CASENT0869853), (D) V.julianus (LACMENT359792),(E)V.lariversi (CASENT0761204), (F) V. lobognathus (LACMENT363986), (G) V. pergandei (CASENT0869850), (H) V. pseudolariversi (CASENT0869851), (I) V. smithi (LACMENT364071), and (J) V. stoddardi (LACMENT364102). Photographs by Robert Johnson from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.7318831" httpUri="https://zenodo.org/record/7318831/files/figure.png" pageId="92" pageNumber="93">Figures1J</figureCitation>
,
<figureCitation id="13012A6EFFB9FF937C2DFE1E5FE0FE01" box="[895,926,402,429]" captionStart="FIGURE 51" captionStartId="96.[151,250,1943,1967]" captionTargetBox="[405,1180,181,1920]" captionTargetId="figure-26@96.[405,1182,181,1920]" captionTargetPageId="96" captionText="FIGURE 51. Photograph of Veromessor stoddardi male: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (LACMENT364104). Photographs by Michele Esposito from www.AntWeb.org." figureDoi="http://doi.org/10.5281/zenodo.7318944" httpUri="https://zenodo.org/record/7318944/files/figure.png" pageId="92" pageNumber="93">51</figureCitation>
).
</paragraph>
<paragraph id="8B8536EBFFB9FF937F95FE3B585EFDB5" blockId="92.[151,1437,151,2043]" pageId="92" pageNumber="93">
<emphasis id="B94EEAF9FFB9FF937F95FE3B5D08FE7D" bold="true" box="[199,374,439,465]" pageId="92" pageNumber="93">Measurements</emphasis>
. (
<emphasis id="B94EEAF9FFB9FF937EC0FE3B5DDEFE7C" box="[402,416,439,464]" italics="true" pageId="92" pageNumber="93">n</emphasis>
= 2). HL 1.151.19; HW 1.121.15; MOD 0.450.46; OMD 0.110.14; SL 0.460.52; HFL 1.962.07; ML 3.073.12; PW 0.460.49; PPW 0.830.90;
<collectingRegion id="49FEF809FFB9FF937C3BFE575FD8FE59" box="[873,934,475,501]" country="New Zealand" name="West Coast" pageId="92" pageNumber="93">AOD</collectingRegion>
0.120.13; IOD 0.320.32; OOD 0.400.42. Indices: SI 41.0745.22; CI 94.12100.00; OI 40.0040.18; HFI 175.00180.00.
</paragraph>
<paragraph id="8B8536EBFFB9FF937F95FDAE585BFB0D" blockId="92.[151,1437,151,2043]" pageId="92" pageNumber="93">
<emphasis id="B94EEAF9FFB9FF937F95FDAE5E4CFD90" bold="true" box="[199,562,546,572]" pageId="92" pageNumber="93">Additional material examined.</emphasis>
<materialsCitation id="3B523CB6FFB9FF937D6AFDAE5DC6FDCD" ID-GBIF-Occurrence="3965991329" collectingDate="1993-02-10" collectionCode="RAJC, UCDC" country="Mexico" location="El Progreso" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<emphasis id="B94EEAF9FFB9FF937D6AFDAE5F0CFD91" bold="true" box="[568,882,546,573]" pageId="92" pageNumber="93">
<collectingCountry id="F32D767BFFB9FF937D6AFDAE5ECDFD90" box="[568,691,546,572]" name="Mexico" pageId="92" pageNumber="93">MEXICO</collectingCountry>
:
<emphasis id="B94EEAF9FFB9FF937DEFFDAF5F0CFD91" bold="true" box="[701,882,546,573]" italics="true" pageId="92" pageNumber="93">
<collectingRegion id="49FEF809FFB9FF937DEFFDAF5F0CFD91" box="[701,882,546,573]" country="Mexico" name="Baja California" pageId="92" pageNumber="93">Baja California</collectingRegion>
</emphasis>
</emphasis>
:
<locationDeviation id="9A372FC9FFB9FF937C2DFDAE580CFD91" box="[895,1138,546,573]" location="Catavina" pageId="92" pageNumber="93">
<quantity id="4CC29B0EFFB9FF937C2DFDAE5FAAFD91" box="[895,980,546,573]" metricMagnitude="4" metricUnit="m" metricValue="6.1959744" pageId="92" pageNumber="93" unit="mi" value="38.5">38.5 mi</quantity>
NW Catavina
</locationDeviation>
at
<quantity id="4CC29B0EFFB9FF937BC3FDAF58BDFD91" box="[1169,1219,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="92" pageNumber="93" unit="mi" value="1.0">1 mi</quantity>
N
<location id="8EE56030FFB9FF937BB1FDAF5916FD91" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937BB1FDAF5916FD91" box="[1251,1384,547,573]" country="Mexico" name="El Progreso" pageId="92" pageNumber="93" stateProvince="Baja California">El Progreso</location>
,
<date id="FF84102BFFB9FF937A20FDAF5C87FDCD" pageId="92" pageNumber="93" value="1993-02-10">
<collectingDate id="EFC0E9C3FFB9FF937A20FDAF5C87FDCD" pageId="92" pageNumber="93" value="1993-02-10">Feb 10, 1993</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937E5AFDCB5D2FFDCD" box="[264,337,583,609]" pageId="92" pageNumber="93">RAJC</collectionCode>
;
<collectionCode id="ED2BAE2EFFB9FF937E0FFDCB5DCEFDCD" box="[349,432,583,609]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="92" pageNumber="93" type="Museum">UCDC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937E96FDCB58AEFDCD" ID-GBIF-Occurrence="3965991435" box="[452,1232,583,609]" collectingDate="1998-03-11" collectionCode="RAJC" country="Mexico" location="San Borja" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
Sierra
<location id="8EE56030FFB9FF937D5CFDCB5E03FDCD" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937D5CFDCB5E03FDCD" box="[526,637,583,609]" country="Mexico" name="San Borja" pageId="92" pageNumber="93" stateProvince="Baja California">San Borja</location>
at
<locationDeviation id="9A372FC9FFB9FF937DF2FDCB5F02FDCD" box="[672,892,583,609]" location="San Borja" pageId="92" pageNumber="93">
<quantity id="4CC29B0EFFB9FF937DF2FDCB5E96FDCD" box="[672,744,583,609]" metricMagnitude="4" metricUnit="m" metricValue="1.59325056" pageId="92" pageNumber="93" unit="mi" value="9.9">9.9 mi</quantity>
N San Borja
</locationDeviation>
, 1970,
<date id="FF84102BFFB9FF937C85FDCB5810FDCD" box="[983,1134,583,609]" pageId="92" pageNumber="93" value="1998-03-11">
<collectingDate id="EFC0E9C3FFB9FF937C85FDCB5810FDCD" box="[983,1134,583,609]" pageId="92" pageNumber="93" value="1998-03-11">Mar 11, 1998</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937B2CFDCB58B7FDCD" box="[1150,1225,583,609]" pageId="92" pageNumber="93">RAJC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937B8EFDCB5F1FFD29" ID-GBIF-Occurrence="3965991432" collectingDate="2001-03-25" collectionCode="CIDA, NHMW, RAJC, UCDC" country="Mexico" location="Meling Ranch" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<quantity id="4CC29B0EFFB9FF937B8EFDCB595AFDCD" box="[1244,1316,583,609]" metricMagnitude="3" metricUnit="m" metricValue="7.0811136" pageId="92" pageNumber="93" unit="mi" value="4.4">4.4 mi</quantity>
W
<location id="8EE56030FFB9FF937A19FDCB5C9FFD29" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937A19FDCB5C9FFD29" country="Mexico" name="Meling Ranch" pageId="92" pageNumber="93" stateProvince="Baja California">Meling Ranch</location>
, 2450,
<date id="FF84102BFFB9FF937E12FDE75DA3FD29" box="[320,477,618,645]" pageId="92" pageNumber="93" value="2001-03-25">
<collectingDate id="EFC0E9C3FFB9FF937E12FDE75DA3FD29" box="[320,477,618,645]" pageId="92" pageNumber="93" value="2001-03-25">Mar 25, 2001</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937EBCFDE75E49FD29" box="[494,567,619,645]" country="USA" httpUri="http://grbio.org/cool/z54r-t2qg" name="Albertson College, Museum of Natural History" pageId="92" pageNumber="93">CIDA</collectionCode>
;
<collectionCode id="ED2BAE2EFFB9FF937D17FDE75EDDFD29" box="[581,675,619,645]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="92" pageNumber="93" type="Museum">NHMW</collectionCode>
;
<collectionCode id="ED2BAE2EFFB9FF937DE2FDE75E86FD29" box="[688,760,619,645]" pageId="92" pageNumber="93">RAJC</collectionCode>
;
<collectionCode id="ED2BAE2EFFB9FF937C54FDE75F27FD29" box="[774,857,619,645]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="92" pageNumber="93" type="Museum">UCDC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937C3CFDE75D04FD05" ID-GBIF-Occurrence="3965991510" collectorName="R. Snelling" country="Mexico" location="Meling Ranch" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<location id="8EE56030FFB9FF937C3CFDE7586CFD29" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937C3CFDE7586CFD29" box="[878,1042,619,645]" country="Mexico" name="Meling Ranch" pageId="92" pageNumber="93" stateProvince="Baja California">Meling Ranch</location>
, 2200, no date (not examined;
<collectorName id="26CF533DFFB9FF937AD0FDE75C89FD05" pageId="92" pageNumber="93">R. Snelling</collectorName>
, pers. obs.)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937ED4FD035839FD05" ID-GBIF-Occurrence="3965991449" box="[390,1095,654,681]" collectingDate="1952-05-14" collectionCode="LACM, UAIC" country="Mexico" location="Mision Descanso" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<quantity id="4CC29B0EFFB9FF937ED4FD035DC7FD05" box="[390,441,655,681]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="92" pageNumber="93" unit="mi" value="4.0">4 mi</quantity>
S
<location id="8EE56030FFB9FF937E84FD035EE5FD05" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937E84FD035EE5FD05" box="[470,667,655,681]" country="Mexico" name="Mision Descanso" pageId="92" pageNumber="93" stateProvince="Baja California">Misión Descanso</location>
, 700,
<date id="FF84102BFFB9FF937DBAFD035FFBFD05" box="[744,901,654,681]" pageId="92" pageNumber="93" value="1952-05-14">
<collectingDate id="EFC0E9C3FFB9FF937DBAFD035FFBFD05" box="[744,901,654,681]" pageId="92" pageNumber="93" value="1952-05-14">May 14, 1952</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937CC6FD035F94FD05" box="[916,1002,655,681]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="92" pageNumber="93" type="Museum">LACM</collectionCode>
;
<collectionCode id="ED2BAE2EFFB9FF937CA7FD035840FD05" box="[1013,1086,655,681]" country="0" httpUri="http://grbio.org/cool/tizb-nd41" name="University of Alabama, Ichthyological Collection" pageId="92" pageNumber="93">UAIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937B00FD035DCBFD61" ID-GBIF-Occurrence="3965991419" collectingDate="1938-06-14" collectionCode="CASC, LACM, USNM" country="Mexico" location="Rosarito Beach" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<location id="8EE56030FFB9FF937B00FD03597FFD05" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937B00FD03597FFD05" box="[1106,1281,655,681]" country="Mexico" name="Rosarito Beach" pageId="92" pageNumber="93" stateProvince="Baja California">Rosarito Beach</location>
,
<date id="FF84102BFFB9FF937A5EFD0359E2FD05" box="[1292,1436,655,681]" pageId="92" pageNumber="93" value="1938-06-14">
<collectingDate id="EFC0E9C3FFB9FF937A5EFD0359E2FD05" box="[1292,1436,655,681]" pageId="92" pageNumber="93" value="1938-06-14">Jun 14, 1938</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937FCDFD3F5C93FD61" box="[159,237,691,717]" pageId="92" pageNumber="93">CASC</collectionCode>
;
<collectionCode id="ED2BAE2EFFB9FF937FAAFD3F5D33FD61" box="[248,333,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="92" pageNumber="93" type="Museum">LACM</collectionCode>
;
<collectionCode id="ED2BAE2EFFB9FF937E05FD3F5DD3FD61" box="[343,429,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="92" pageNumber="93" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937EEDFD3F5831FD61" ID-GBIF-Occurrence="3965991383" box="[447,1103,690,717]" collectingDate="1952-05-15" collectionCode="LACM, USNM" country="Mexico" location="Ensenada" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<quantity id="4CC29B0EFFB9FF937EEDFD3F5E7EFD61" box="[447,512,691,717]" metricMagnitude="4" metricUnit="m" metricValue="3.0577536" pageId="92" pageNumber="93" unit="mi" value="19.0">19 mi</quantity>
N
<location id="8EE56030FFB9FF937D72FD3F5EF0FD61" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937D72FD3F5EF0FD61" box="[544,654,691,717]" country="Mexico" name="Ensenada" pageId="92" pageNumber="93" stateProvince="Baja California">Ensenada</location>
, 1100,
<date id="FF84102BFFB9FF937DB5FD3F5FFCFD61" box="[743,898,690,717]" pageId="92" pageNumber="93" value="1952-05-15">
<collectingDate id="EFC0E9C3FFB9FF937DB5FD3F5FFCFD61" box="[743,898,690,717]" pageId="92" pageNumber="93" value="1952-05-15">May 15, 1952</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937CC2FD3F5F98FD61" box="[912,998,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="92" pageNumber="93" type="Museum">LACM</collectionCode>
;
<collectionCode id="ED2BAE2EFFB9FF937CA3FD3F5839FD61" box="[1009,1095,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="92" pageNumber="93" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937B0BFD3F5D16FD5D" ID-GBIF-Occurrence="3965991466" collectingDate="1952-05-14" collectionCode="USNM" country="Mexico" location="La Mision" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<quantity id="4CC29B0EFFB9FF937B0BFD3F58F2FD61" box="[1113,1164,691,717]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="92" pageNumber="93" unit="mi" value="4.0">4 mi</quantity>
S
<location id="8EE56030FFB9FF937BF5FD3F5963FD61" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937BF5FD3F5963FD61" box="[1191,1309,691,717]" country="Mexico" name="La Mision" pageId="92" pageNumber="93" stateProvince="Baja California">La Misión</location>
, 700,
<date id="FF84102BFFB9FF937A3BFD3F5C84FD5D" pageId="92" pageNumber="93" value="1952-05-14">
<collectingDate id="EFC0E9C3FFB9FF937A3BFD3F5C84FD5D" pageId="92" pageNumber="93" value="1952-05-14">May 14, 1952</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937E5BFD5B5D1EFD5D" box="[265,352,727,753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="92" pageNumber="93" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937E26FD5B5C8FFCB9" ID-GBIF-Occurrence="3965991506" collectingDate="1994-08-26" collectorName="M. Bennett" country="Mexico" elevation="260" location="Rancho Canadas" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<location id="8EE56030FFB9FF937E26FD5B5E4EFD5D" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937E26FD5B5E4EFD5D" box="[372,560,727,753]" country="Mexico" name="Rancho Canadas" pageId="92" pageNumber="93" stateProvince="Baja California">Rancho Cañadas</location>
at
<locationDeviation id="9A372FC9FFB9FF937D01FD5B5F56FD5D" box="[595,808,727,753]" location="Camalu" pageId="92" pageNumber="93">
<quantity id="4CC29B0EFFB9FF937D01FD5B5EE4FD5D" box="[595,666,727,753]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="92" pageNumber="93" unit="km" value="13.0">13 km</quantity>
NE Camalú
</locationDeviation>
,
<quantity id="4CC29B0EFFB9FF937C66FD5B5F03FD5D" box="[820,893,727,753]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="92" pageNumber="93" unit="m" value="260.0">
<elevation id="0017D1D8FFB9FF937C66FD5B5F03FD5D" box="[820,893,727,753]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="92" pageNumber="93" unit="m" value="260.0">260 m</elevation>
</quantity>
,
<date id="FF84102BFFB9FF937CDAFD5B585CFD5D" box="[904,1058,727,753]" pageId="92" pageNumber="93" value="1994-08-26">
<collectingDate id="EFC0E9C3FFB9FF937CDAFD5B585CFD5D" box="[904,1058,727,753]" pageId="92" pageNumber="93" value="1994-08-26">Aug 26, 1994</collectingDate>
</date>
(not examined;
<collectorName id="26CF533DFFB9FF937B88FD5B5925FD5D" box="[1242,1371,727,753]" pageId="92" pageNumber="93">M. Bennett</collectorName>
, pers. comm.)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937FAFFD775E9AFCB9" ID-GBIF-Occurrence="3965991473" box="[253,740,763,789]" collectingDate="1979-07-14" collectionCode="UCDC" country="Mexico" location="San Telmo" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<quantity id="4CC29B0EFFB9FF937FAFFD775D38FCB9" box="[253,326,763,789]" metricMagnitude="3" metricUnit="m" metricValue="4.5061632" pageId="92" pageNumber="93" unit="mi" value="2.8">2.8 mi</quantity>
E
<location id="8EE56030FFB9FF937E34FD775D9FFCB9" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937E34FD775D9FFCB9" box="[358,481,763,789]" country="Mexico" name="San Telmo" pageId="92" pageNumber="93" stateProvince="Baja California">San Telmo</location>
,
<date id="FF84102BFFB9FF937EBFFD775E06FCB9" box="[493,632,763,789]" pageId="92" pageNumber="93" value="1979-07-14">
<collectingDate id="EFC0E9C3FFB9FF937EBFFD775E06FCB9" box="[493,632,763,789]" pageId="92" pageNumber="93" value="1979-07-14">Jul 14, 1979</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937DDAFD775EA2FCB9" box="[648,732,763,789]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="92" pageNumber="93" type="Museum">UCDC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937DA2FD775DCCFC95" ID-GBIF-Occurrence="3965991352" collectingDate="1998-04-07" collectionCode="UCDC" collectorName="Angeles" country="Mexico" elevation="480" location="Bahia" municipality="Road" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<locationDeviation id="9A372FC9FFB9FF937DA2FD775FA4FCB9" box="[752,986,763,789]" location="Jct Hwy" pageId="92" pageNumber="93">
<quantity id="4CC29B0EFFB9FF937DA2FD775F49FCB9" box="[752,823,763,789]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="92" pageNumber="93" unit="km" value="14.0">14 km</quantity>
ENE Jct Hwy
</locationDeviation>
1 at
<collectingMunicipality id="6BE1AC91FFB9FF937B46FD775831FCB9" box="[1044,1103,763,789]" pageId="92" pageNumber="93">Road</collectingMunicipality>
to
<location id="8EE56030FFB9FF937B26FD7758CBFCB9" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937B26FD7758CBFCB9" box="[1140,1205,763,789]" country="Mexico" municipality="Road" name="Bahia" pageId="92" pageNumber="93" stateProvince="Baja California">Bahía</location>
de los
<collectorName id="26CF533DFFB9FF937A5AFD705919FCBB" box="[1288,1383,764,791]" pageId="92" pageNumber="93">Ángeles</collectorName>
,
<quantity id="4CC29B0EFFB9FF937A20FD775CD1FC95" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="92" pageNumber="93" unit="m" value="480.0">
<elevation id="0017D1D8FFB9FF937A20FD775CD1FC95" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="92" pageNumber="93" unit="m" value="480.0">480 m</elevation>
</quantity>
,
<date id="FF84102BFFB9FF937FE9FC935D3BFC95" box="[187,325,799,825]" pageId="92" pageNumber="93" value="1998-04-07">
<collectingDate id="EFC0E9C3FFB9FF937FE9FC935D3BFC95" box="[187,325,799,825]" pageId="92" pageNumber="93" value="1998-04-07">Apr 7, 1998</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937E04FC935DD4FC95" box="[342,426,799,825]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="92" pageNumber="93" type="Museum">UCDC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937EEDFC935808FC95" ID-GBIF-Occurrence="3965991410" box="[447,1142,798,825]" collectingDate="1999-05-09" collectionCode="UCDC" country="Mexico" location="The Oaks" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<location id="8EE56030FFB9FF937E9BFC935E42FC95" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937E9BFC935E42FC95" box="[457,572,799,825]" country="Mexico" name="The Oaks" pageId="92" pageNumber="93" stateProvince="Baja California">The Oaks</location>
” at
<locationDeviation id="9A372FC9FFB9FF937D39FC925F15FC95" box="[619,875,798,825]" location="Santo Thomas" pageId="92" pageNumber="93">
<quantity id="4CC29B0EFFB9FF937D39FC925EDEFC95" box="[619,672,798,825]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="92" pageNumber="93" unit="mi" value="5.0">5 mi</quantity>
N Santo Thomas
</locationDeviation>
,
<date id="FF84102BFFB9FF937C2AFC935877FC95" box="[888,1033,799,825]" pageId="92" pageNumber="93" value="1999-05-09">
<collectingDate id="EFC0E9C3FFB9FF937C2AFC935877FC95" box="[888,1033,799,825]" pageId="92" pageNumber="93" value="1999-05-09">May 9, 1999</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937B48FC935810FC95" box="[1050,1134,799,825]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="92" pageNumber="93" type="Museum">UCDC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937BD1FC935EF6FCF1" ID-GBIF-Occurrence="3965991346" collectorName="Snelling" country="Mexico" location="10 mi E San Quintin" pageId="92" pageNumber="93" specimenCount="1" stateProvince="Baja California">
<location id="8EE56030FFB9FF937BD1FC93590DFC95" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937BD1FC93590DFC95" box="[1155,1395,799,825]" country="Mexico" name="10 mi E San Quintin" pageId="92" pageNumber="93" stateProvince="Baja California">
<quantity id="4CC29B0EFFB9FF937BD1FC9358B8FC95" box="[1155,1222,799,825]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="92" pageNumber="93" unit="mi" value="10.0">10 mi</quantity>
E San Quintin
</location>
, no date (not examined; R.
<collectorName id="26CF533DFFB9FF937EF1FCCF5E7DFCF1" box="[419,515,835,861]" pageId="92" pageNumber="93">Snelling</collectorName>
, pers. obs.)
</materialsCitation>
.
<materialsCitation id="3B523CB6FFB9FF937DC6FCCF5E1CFC2D" ID-GBIF-Occurrence="3965991318" collectingDate="1941-04-25" collectionCode="LACM" collectorName="FS Hastings Natural History Reserve" country="United States of America" county="Monterey Co." location="Monterey Co." pageId="92" pageNumber="93" specimenCount="1" stateProvince="California">
<emphasis id="B94EEAF9FFB9FF937DC6FCCF587DFCF1" bold="true" box="[660,1027,834,861]" pageId="92" pageNumber="93">
<collectingCountry id="F32D767BFFB9FF937DC6FCCF5F0BFCF0" box="[660,885,834,861]" name="United States of America" pageId="92" pageNumber="93">UNITED STATES</collectingCountry>
:
<emphasis id="B94EEAF9FFB9FF937CD0FCCE5F84FCF1" bold="true" box="[898,1018,834,861]" italics="true" pageId="92" pageNumber="93">
<collectingRegion id="49FEF809FFB9FF937CD0FCCE5F84FCF1" box="[898,1018,834,861]" country="United States of America" name="California" pageId="92" pageNumber="93">California</collectingRegion>
</emphasis>
:
</emphasis>
<collectingCounty id="62E44E67FFB9FF937B5EFCCF58D8FCF1" box="[1036,1190,835,861]" pageId="92" pageNumber="93">
<emphasis id="B94EEAF9FFB9FF937B5EFCCF58DEFCF1" box="[1036,1184,835,861]" italics="true" pageId="92" pageNumber="93">Monterey Co</emphasis>
.
</collectingCounty>
:
<collectorName id="26CF533DFFB9FF937BE4FCCF5D2EFC2D" pageId="92" pageNumber="93">FS Hastings Natural History Reserve</collectorName>
,
<date id="FF84102BFFB9FF937E09FCEB5D8CFC2D" box="[347,498,870,897]" pageId="92" pageNumber="93" value="1941-04-25">
<collectingDate id="EFC0E9C3FFB9FF937E09FCEB5D8CFC2D" box="[347,498,870,897]" pageId="92" pageNumber="93" value="1941-04-25">Apr 25, 1941</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937D50FCEB5E27FC2D" box="[514,601,871,897]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="92" pageNumber="93" type="Museum">LACM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937D3CFCEB5D13FC09" ID-GBIF-Occurrence="3965991447" collectingDate="1963-06" collectionCode="USNM" collectorName="R. Snelling" country="United States of America" location="Paraiso Springs" pageId="92" pageNumber="93" specimenCount="1" stateProvince="California">
<location id="8EE56030FFB9FF937D3CFCEB5F5FFC2D" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937D3CFCEB5F5FFC2D" box="[622,801,871,897]" country="United States of America" name="Paraiso Springs" pageId="92" pageNumber="93" stateProvince="California">Paraiso Springs</location>
, no date (not examined;
<collectorName id="26CF533DFFB9FF937B69FCEB58C3FC2D" box="[1083,1213,871,897]" pageId="92" pageNumber="93">R. Snelling</collectorName>
, pers. obs.); no loc,
<date id="FF84102BFFB9FF937FC5FC075C80FC09" box="[151,254,907,933]" pageId="92" pageNumber="93" value="1963-06">
<collectingDate id="EFC0E9C3FFB9FF937FC5FC075C80FC09" box="[151,254,907,933]" pageId="92" pageNumber="93" value="1963-06">Jun 1963</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937E5CFC075D1BFC09" box="[270,357,907,933]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="92" pageNumber="93" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B523CB6FFB9FF937E2AFC075EE3FC65" ID-GBIF-Occurrence="3965991491" collectingDate="1984-05-07" collectionCode="LACM" country="United States of America" county="Riverside Co." location="Forest Service Rd" municipality="Cleveland National Forest" pageId="92" pageNumber="93" specimenCount="1" stateProvince="California">
<collectingCounty id="62E44E67FFB9FF937E2AFC075E6CFC09" box="[376,530,907,933]" pageId="92" pageNumber="93">
<emphasis id="B94EEAF9FFB9FF937E2AFC075E75FC09" box="[376,523,907,933]" italics="true" pageId="92" pageNumber="93">Riverside Co</emphasis>
.
</collectingCounty>
:
<collectingMunicipality id="6BE1AC91FFB9FF937D73FC075F32FC09" box="[545,844,907,933]" pageId="92" pageNumber="93">Cleveland National Forest</collectingMunicipality>
,
<location id="8EE56030FFB9FF937C0AFC07585DFC09" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937C0AFC07585DFC09" box="[856,1059,907,933]" country="United States of America" county="Riverside Co." municipality="Cleveland National Forest" name="Forest Service Rd" pageId="92" pageNumber="93" stateProvince="California">Forest Service Rd</location>
8S01 at
<locationDeviation id="9A372FC9FFB9FF937BDAFC075885FC09" box="[1160,1275,907,933]" location="Jct" pageId="92" pageNumber="93">
<quantity id="4CC29B0EFFB9FF937BDAFC0758C2FC09" box="[1160,1212,907,933]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="92" pageNumber="93" unit="mi" value="3.0">3 mi</quantity>
E Jct
</locationDeviation>
8S01 &amp; 8S04 (W of
<location id="8EE56030FFB9FF937FB6FC235DF9FC65" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937FB6FC235DF9FC65" box="[228,391,943,969]" country="United States of America" county="Riverside Co." municipality="Cleveland National Forest" name="Lake Elsinore" pageId="92" pageNumber="93" stateProvince="California">Lake Elsinore</location>
),
<date id="FF84102BFFB9FF937EC9FC235E53FC65" box="[411,557,943,969]" pageId="92" pageNumber="93" value="1984-05-07">
<collectingDate id="EFC0E9C3FFB9FF937EC9FC235E53FC65" box="[411,557,943,969]" pageId="92" pageNumber="93" value="1984-05-07">May 7, 1984</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937D6CFC235EEBFC65" box="[574,661,943,969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="92" pageNumber="93" type="Museum">LACM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937DF8FC2358DBFC65" ID-GBIF-Occurrence="3965991402" box="[682,1189,943,969]" collectingDate="1996-07-29" collectionCode="UCDC" country="United States of America" elevation="462" location="Lake Skinner" pageId="92" pageNumber="93" specimenCount="1" stateProvince="California">
<location id="8EE56030FFB9FF937DF8FC235F3AFC65" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937DF8FC235F3AFC65" box="[682,836,943,969]" country="United States of America" name="Lake Skinner" pageId="92" pageNumber="93" stateProvince="California">Lake Skinner</location>
,
<quantity id="4CC29B0EFFB9FF937C03FC235FE2FC65" box="[849,924,943,969]" metricMagnitude="2" metricUnit="m" metricValue="4.62" pageId="92" pageNumber="93" unit="m" value="462.0">
<elevation id="0017D1D8FFB9FF937C03FC235FE2FC65" box="[849,924,943,969]" metricMagnitude="2" metricUnit="m" metricValue="4.62" pageId="92" pageNumber="93" unit="m" value="462.0">462 m</elevation>
</quantity>
,
<date id="FF84102BFFB9FF937CF8FC235849FC65" box="[938,1079,943,969]" pageId="92" pageNumber="93" value="1996-07-29">
<collectingDate id="EFC0E9C3FFB9FF937CF8FC235849FC65" box="[938,1079,943,969]" pageId="92" pageNumber="93" value="1996-07-29">Jul 29, 1996</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937B1AFC2358E2FC65" box="[1096,1180,943,969]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="92" pageNumber="93" type="Museum">UCDC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B523CB6FFB9FF937BE3FC235916FC41" ID-GBIF-Occurrence="3965991330" collectingDate="1994-06-13" collectorName="M. Bennett" country="United States of America" county="San Bernardino Co." elevation="250" location="Chino Hills State Park" pageId="92" pageNumber="93" specimenCount="1" stateProvince="California">
<collectingCounty id="62E44E67FFB9FF937BE3FC2359EAFC65" box="[1201,1428,943,969]" pageId="92" pageNumber="93">
<emphasis id="B94EEAF9FFB9FF937BE3FC2359F0FC65" box="[1201,1422,943,969]" italics="true" pageId="92" pageNumber="93">San Bernardino Co</emphasis>
.
</collectingCounty>
:
<location id="8EE56030FFB9FF937FC5FC5F5DECFC41" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937FC5FC5F5DECFC41" box="[151,402,979,1005]" country="United States of America" county="San Bernardino Co." name="Chino Hills State Park" pageId="92" pageNumber="93" stateProvince="California">Chino Hills State Park</location>
at
<locationDeviation id="9A372FC9FFB9FF937EE6FC5F5EDEFC41" box="[436,672,979,1005]" location="Los Serranos" pageId="92" pageNumber="93">
<quantity id="4CC29B0EFFB9FF937EE6FC5F5D93FC41" box="[436,493,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="92" pageNumber="93" unit="km" value="6.0">6 km</quantity>
S Los Serranos
</locationDeviation>
,
<quantity id="4CC29B0EFFB9FF937DF9FC5E5E8AFC41" box="[683,756,978,1005]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="92" pageNumber="93" unit="m" value="250.0">
<elevation id="0017D1D8FFB9FF937DF9FC5E5E8AFC41" box="[683,756,978,1005]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="92" pageNumber="93" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<date id="FF84102BFFB9FF937C52FC5F5FEEFC41" box="[768,912,979,1005]" pageId="92" pageNumber="93" value="1994-06-13">
<collectingDate id="EFC0E9C3FFB9FF937C52FC5F5FEEFC41" box="[768,912,979,1005]" pageId="92" pageNumber="93" value="1994-06-13">Jun 13, 1994</collectingDate>
</date>
(not examined,
<collectorName id="26CF533DFFB9FF937B14FC5F58B8FC41" box="[1094,1222,979,1005]" pageId="92" pageNumber="93">M. Bennett</collectorName>
, pers. comm.)
</materialsCitation>
.
<materialsCitation id="3B523CB6FFB9FF937A20FC5F585FFBBD" ID-GBIF-Occurrence="3965991446" collectingDate="1952-05-13" collectingDateMax="1952-07-09" collectingDateMin="1952-05-13" collectionCode="LACM, USNM" country="United States of America" county="San Diego Co." location="San" pageId="92" pageNumber="93" specimenCount="1" stateProvince="California">
<collectingCounty id="62E44E67FFB9FF937A20FC5F5D72FBBD" pageId="92" pageNumber="93">
<emphasis id="B94EEAF9FFB9FF937A20FC5F5D7BFBBD" italics="true" pageId="92" pageNumber="93">San Diego Co</emphasis>
.
</collectingCounty>
:
<location id="8EE56030FFB9FF937E4EFC7B5D38FBBD" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937E4EFC7B5D38FBBD" box="[284,326,1015,1041]" country="United States of America" county="San Diego Co." name="San" pageId="92" pageNumber="93" stateProvince="California">San</location>
Isidro, 200,
<collectingDate id="EFC0E9C3FFB9FF937EB0FC7B5F2EFBBD" box="[482,848,1014,1041]" isEnumeration="true" pageId="92" pageNumber="93">
<date id="FF84102BFFB9FF937EB0FC7B5ED5FBBD" box="[482,683,1014,1041]" pageId="92" pageNumber="93" value="1952-05-13" valueMax="1952-05-15" valueMin="1952-05-13">May 1315, 1952</date>
&amp;
<date id="FF84102BFFB9FF937D83FC7B5F2EFBBD" box="[721,848,1014,1041]" pageId="92" pageNumber="93" value="1952-07-09">Jul 9, 1952</date>
</collectingDate>
(
<collectionCode id="ED2BAE2EFFB9FF937C32FC7B5FC8FBBD" box="[864,950,1015,1041]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="92" pageNumber="93" type="Museum">LACM</collectionCode>
;
<collectionCode id="ED2BAE2EFFB9FF937C91FC7B5867FBBD" box="[963,1049,1015,1041]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="92" pageNumber="93" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937B7FFC7B59FCFBBD" ID-GBIF-Occurrence="3965991351" box="[1069,1410,1015,1041]" collectingDate="1898-02" collectionCode="USNM" country="United States of America" location="Descanso" pageId="92" pageNumber="93" specimenCount="1" stateProvince="California">
<location id="8EE56030FFB9FF937B7FFC7B58E5FBBD" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937B7FFC7B58E5FBBD" box="[1069,1179,1015,1041]" country="United States of America" name="Descanso" pageId="92" pageNumber="93" stateProvince="California">Descanso</location>
,
<date id="FF84102BFFB9FF937BFAFC7B596CFBBD" box="[1192,1298,1015,1041]" pageId="92" pageNumber="93" value="1898-02">
<collectingDate id="EFC0E9C3FFB9FF937BFAFC7B596CFBBD" box="[1192,1298,1015,1041]" pageId="92" pageNumber="93" value="1898-02">Feb 1898</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937A70FC7B5907FBBD" box="[1314,1401,1015,1041]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="92" pageNumber="93" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937ADCFC7A5C85FBF5" ID-GBIF-Occurrence="3965991407" collectingDate="2003-06" collectionCode="UCDC" collectorName="R. Snelling" country="United States of America" elevation="957" location="Ecological Reserve" pageId="92" pageNumber="93" specimenCount="1" stateProvince="California">
<locationDeviation id="9A372FC9FFB9FF937ADCFC7A5D3AFB99" location="Descanso" pageId="92" pageNumber="93">
<quantity id="4CC29B0EFFB9FF937ADCFC7A5CCBFB99" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="92" pageNumber="93" unit="mi" value="5.0">5 mi</quantity>
E Descanso
</locationDeviation>
, 3000 (not examined,
<collectorName id="26CF533DFFB9FF937D1AFB975EB4FB99" box="[584,714,1051,1077]" pageId="92" pageNumber="93">R. Snelling</collectorName>
, pers. obs.); Santa Ysabel
<location id="8EE56030FFB9FF937CA4FB9758ACFB99" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937CA4FB9758ACFB99" box="[1014,1234,1051,1077]" country="United States of America" name="Ecological Reserve" pageId="92" pageNumber="93" stateProvince="California">Ecological Reserve</location>
,
<quantity id="4CC29B0EFFB9FF937B8DFB965957FB99" box="[1247,1321,1050,1077]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="92" pageNumber="93" unit="m" value="957.0">
<elevation id="0017D1D8FFB9FF937B8DFB965957FB99" box="[1247,1321,1050,1077]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="92" pageNumber="93" unit="m" value="957.0">957 m</elevation>
</quantity>
,
<date id="FF84102BFFB9FF937A67FB9759E2FB99" box="[1333,1436,1051,1077]" pageId="92" pageNumber="93" value="2003-06">
<collectingDate id="EFC0E9C3FFB9FF937A67FB9759E2FB99" box="[1333,1436,1051,1077]" pageId="92" pageNumber="93" value="2003-06">Jun 2003</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937FCDFBB35C8DFBF5" box="[159,243,1087,1113]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="92" pageNumber="93" type="Museum">UCDC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937E54FBB35F05FBF5" ID-GBIF-Occurrence="3965991411" box="[262,891,1087,1113]" collectingDate="2003-05-29" collectingDateMax="2003-06-03" collectingDateMin="2003-05-29" collectionCode="UCDC" country="United States of America" elevation="100" location="Sweetwater River" pageId="92" pageNumber="93" specimenCount="1" stateProvince="California">
<location id="8EE56030FFB9FF937E54FBB35DB1FBF5" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937E54FBB35DB1FBF5" box="[262,463,1087,1113]" country="United States of America" name="Sweetwater River" pageId="92" pageNumber="93" stateProvince="California">Sweetwater River</location>
,
<quantity id="4CC29B0EFFB9FF937E8BFBB35E5FFBF5" box="[473,545,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="92" pageNumber="93" unit="m" value="100.0">
<elevation id="0017D1D8FFB9FF937E8BFBB35E5FFBF5" box="[473,545,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="92" pageNumber="93" unit="m" value="100.0">100 m</elevation>
</quantity>
,
<date id="FF84102BFFB9FF937D7EFBB35F6FFBF5" box="[556,785,1087,1113]" pageId="92" pageNumber="93" value="2003-05-29" valueMax="2003-06-03" valueMin="2003-05-29">
<collectingDate id="EFC0E9C3FFB9FF937D7EFBB35F6FFBF5" box="[556,785,1087,1113]" pageId="92" pageNumber="93" value="2003-05-29" valueMax="2003-06-03" valueMin="2003-05-29">May 29Jun 3, 2003</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937C4DFBB35F0DFBF5" box="[799,883,1087,1113]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="92" pageNumber="93" type="Museum">UCDC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523CB6FFB9FF937CD7FBB35DF0FBD0" ID-GBIF-Occurrence="3965991454" collectingDate="2004-05-01" collectionCode="UCDC" country="United States of America" elevation="25" location="Penasquitos Canyon" municipality="Sycamore Crossing" pageId="92" pageNumber="93" specimenCount="1" stateProvince="California">
<collectingMunicipality id="6BE1AC91FFB9FF937CD7FBB3581CFBF5" box="[901,1122,1087,1113]" pageId="92" pageNumber="93">Sycamore Crossing</collectingMunicipality>
,
<location id="8EE56030FFB9FF937B3FFBB3592DFBF5" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937B3FFBB3592DFBF5" box="[1133,1363,1087,1113]" country="United States of America" municipality="Sycamore Crossing" name="Penasquitos Canyon" pageId="92" pageNumber="93" stateProvince="California">Peñasquitos Canyon</location>
,
<quantity id="4CC29B0EFFB9FF937A0FFBB259E9FBF5" box="[1373,1431,1086,1113]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="92" pageNumber="93" unit="m" value="25.0">
<elevation id="0017D1D8FFB9FF937A0FFBB259E9FBF5" box="[1373,1431,1086,1113]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="92" pageNumber="93" unit="m" value="25.0">25 m</elevation>
</quantity>
,
<date id="FF84102BFFB9FF937FC5FBEF5D5AFBD0" box="[151,292,1122,1149]" pageId="92" pageNumber="93" value="2004-05-01">
<collectingDate id="EFC0E9C3FFB9FF937FC5FBEF5D5AFBD0" box="[151,292,1122,1149]" pageId="92" pageNumber="93" value="2004-05-01">May 1, 2004</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937E63FBEE5DFBFBD0" box="[305,389,1122,1148]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="92" pageNumber="93" type="Museum">UCDC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B523CB6FFB9FF937EC5FBEF58AEFBD0" ID-GBIF-Occurrence="3965991359" box="[407,1232,1122,1149]" collectingDate="1986-05-05" collectionCode="JTLC" country="United States of America" elevation="980" location="San Marcos Pass" pageId="92" pageNumber="93" specimenCount="1" stateProvince="California">
<emphasis id="B94EEAF9FFB9FF937EC5FBEF5E1DFBD1" box="[407,611,1122,1149]" italics="true" pageId="92" pageNumber="93">Santa Barbara Co</emphasis>
.:
<quantity id="4CC29B0EFFB9FF937D25FBEE5ED1FBD0" box="[631,687,1122,1148]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="92" pageNumber="93" unit="km" value="4.0">4 km</quantity>
E
<location id="8EE56030FFB9FF937D9EFBEE5FF4FBD1" LSID="urn:lsid:plazi:treatment:039387FDFFBEFFAD7FC5FF1B5FD7FEED:8EE56030FFB9FF937D9EFBEE5FF4FBD1" box="[716,906,1122,1149]" country="United States of America" name="San Marcos Pass" pageId="92" pageNumber="93" stateProvince="California">San Marcos Pass</location>
,
<quantity id="4CC29B0EFFB9FF937CC7FBEE5FA3FBD1" box="[917,989,1122,1149]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="92" pageNumber="93" unit="m" value="980.0">
<elevation id="0017D1D8FFB9FF937CC7FBEE5FA3FBD1" box="[917,989,1122,1149]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="92" pageNumber="93" unit="m" value="980.0">980 m</elevation>
</quantity>
,
<date id="FF84102BFFB9FF937CBAFBEF580AFBD0" box="[1000,1140,1122,1149]" pageId="92" pageNumber="93" value="1986-05-05">
<collectingDate id="EFC0E9C3FFB9FF937CBAFBEF580AFBD0" box="[1000,1140,1122,1149]" pageId="92" pageNumber="93" value="1986-05-05">May 5, 1986</collectingDate>
</date>
(
<collectionCode id="ED2BAE2EFFB9FF937BD0FBEE58B6FBD0" box="[1154,1224,1122,1148]" pageId="92" pageNumber="93">JTLC</collectionCode>
)
</materialsCitation>
;
<quantity id="4CC29B0EFFB9FF937B88FBEE595EFBD0" box="[1242,1312,1122,1148]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="92" pageNumber="93" unit="km" value="21.0">21 km</quantity>
ESE Santa Ynez,
<quantity id="4CC29B0EFFB9FF937FB2FB0A5D57FB0D" box="[224,297,1158,1185]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="92" pageNumber="93" unit="m" value="980.0">980 m</quantity>
,
<date id="FF84102BFFB9FF937E67FB0B5DBDFB0C" box="[309,451,1158,1185]" pageId="92" pageNumber="93" value="1986-05-05">May 5, 1986</date>
(not examined, M. Bennett, pers. comm.) (
<figureCitation id="13012A6EFFB9FF937CF5FB0A5867FB0D" box="[935,1049,1158,1185]" captionStart="FIGURE 52" captionStartId="97.[151,250,1652,1676]" captionTargetBox="[151,1434,633,1628]" captionTargetId="figure-213@97.[151,1435,633,1629]" captionTargetPageId="97" captionText="FIGURE 52. Geographic distribution of Veromessor stoddardi. The larger black circle denotes the type locality." figureDoi="http://doi.org/10.5281/zenodo.7318946" httpUri="https://zenodo.org/record/7318946/files/figure.png" pageId="92" pageNumber="93">Figure 52</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3206560FFB9FF937F95FB265E1BFB4B" pageId="92" pageNumber="93" type="etymology">
<paragraph id="8B8536EBFFB9FF937F95FB265E1BFB4B" blockId="92.[151,1437,151,2043]" pageId="92" pageNumber="93">
<emphasis id="B94EEAF9FFB9FF937F95FB265D30FB68" bold="true" box="[199,334,1194,1220]" pageId="92" pageNumber="93">Etymology.</emphasis>
The specific epithet,
<taxonomicName id="4C3A4D68FFB9FF937D15FB265ECFFB68" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[583,689,1194,1220]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFB9FF937D15FB265ECFFB68" box="[583,689,1194,1220]" italics="true" pageId="92" pageNumber="93">stoddardi</emphasis>
</taxonomicName>
, was presumably derived from someone named Stoddard, but no reference to this person has been located.
</paragraph>
</subSubSection>
<subSubSection id="C3206560FFB9FF937F95FB7D58BBF857" pageId="92" pageNumber="93" type="discussion">
<paragraph id="8B8536EBFFB9FF937F95FB7D5D62F914" blockId="92.[151,1437,151,2043]" pageId="92" pageNumber="93">
<emphasis id="B94EEAF9FFB9FF937F95FB7D5D32FAA7" bold="true" box="[199,332,1265,1291]" pageId="92" pageNumber="93">Discussion.</emphasis>
The poorly developed psammophore distinguishes
<taxonomicName id="4C3A4D68FFB9FF937CDBFB7E5870FAA7" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[905,1038,1265,1291]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFB9FF937CDBFB7E5870FAA7" box="[905,1038,1265,1291]" italics="true" pageId="92" pageNumber="93">V. stoddardi</emphasis>
</taxonomicName>
from all congeners except
<taxonomicName id="4C3A4D68FFB9FF937A68FB7E59E2FAA7" box="[1338,1436,1265,1291]" class="Insecta" family="Formicidae" genus="Novomessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="andrei">
<emphasis id="B94EEAF9FFB9FF937A68FB7E59E2FAA7" box="[1338,1436,1265,1291]" italics="true" pageId="92" pageNumber="93">V. andrei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D68FFB9FF937F97FA9A5D2AFA83" authorityName="Smith" authorityYear="1956" box="[197,340,1301,1327]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="chicoensis">
<emphasis id="B94EEAF9FFB9FF937F97FA9A5D2AFA83" box="[197,340,1301,1327]" italics="true" pageId="92" pageNumber="93">V. chicoensis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C3A4D68FFB9FF937E33FA9A5E4EFA83" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[353,560,1301,1327]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="andrei">
<emphasis id="B94EEAF9FFB9FF937E33FA9A5E4EFA83" box="[353,560,1301,1327]" italics="true" pageId="92" pageNumber="93">Veromessor andrei</emphasis>
</taxonomicName>
is the only congener with a poorly developed psammophore that is likely to cooccur with
<taxonomicName id="4C3A4D68FFB9FF937E47FAB65DE4FAFF" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[277,410,1337,1363]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFB9FF937E47FAB65DE4FAFF" box="[277,410,1337,1363]" italics="true" pageId="92" pageNumber="93">V. stoddardi</emphasis>
</taxonomicName>
. The two species can be distinguished based on: (1)
<taxonomicName id="4C3A4D68FFB9FF937CBAFAB65813FAFF" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[1000,1133,1337,1363]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFB9FF937CBAFAB65813FAFF" box="[1000,1133,1337,1363]" italics="true" pageId="92" pageNumber="93">V. stoddardi</emphasis>
</taxonomicName>
has short propodeal spines (length less than the distance between their bases), (2) strongly polymorphic workers, and (3) weak, regular rugae on cephalic dorsum posterior to eyes and dorsum of pronotum. In
<taxonomicName id="4C3A4D68FFB9FF937C01FA0D5FC7FA36" baseAuthorityName="Mayr" baseAuthorityYear="1886" box="[851,953,1408,1435]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="andrei">
<emphasis id="B94EEAF9FFB9FF937C01FA0D5F1AFA36" box="[851,868,1409,1434]" italics="true" pageId="92" pageNumber="93">V</emphasis>
.
<emphasis id="B94EEAF9FFB9FF937C23FA0C5FC7FA36" box="[881,953,1408,1434]" italics="true" pageId="92" pageNumber="93">andrei</emphasis>
</taxonomicName>
: (1) propodeal spines long (length&gt; 3× the distance between their bases), (2) workers mostly monomorphic, and (3) notably coarse rugae on cephalic dorsum posterior to eyes and dorsum of pronotum.
<taxonomicName id="4C3A4D68FFB9FF937DDAFA445F02FA4E" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[648,892,1480,1506]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFB9FF937DDAFA445F02FA4E" box="[648,892,1480,1506]" italics="true" pageId="92" pageNumber="93">Veromessor stoddardi</emphasis>
</taxonomicName>
is most similar to the closely related species
<taxonomicName id="4C3A4D68FFB9FF937ADAFA445D72F9A9" authorityName="Smith" authorityYear="1956" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="chicoensis">
<emphasis id="B94EEAF9FFB9FF937ADAFA445D72F9A9" italics="true" pageId="92" pageNumber="93">V. chicoensis</emphasis>
</taxonomicName>
, from which it can be distinguished by: (1) antennal scape narrowest immediately distad of basal bend; maximum basal width of scape about equal to maximum preapical width, and (2) in dorsal view, the mesonotum usually smooth and shining, rugae distinctly weaker than those on pronotum. In
<taxonomicName id="4C3A4D68FFB9FF937B7EF9B858C0F9E1" authorityName="Smith" authorityYear="1956" box="[1068,1214,1587,1613]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="chicoensis">
<emphasis id="B94EEAF9FFB9FF937B7EF9B858C0F9E1" box="[1068,1214,1587,1613]" italics="true" pageId="92" pageNumber="93">V. chicoensis</emphasis>
</taxonomicName>
: (1) antennal scape narrowest near midlength; maximum basal width of scape greater than maximum preapical width, and (2) in dorsal view, rugae on mesonotum similar to those on pronotum, interrugae weakly dull (also see discussion under
<taxonomicName id="4C3A4D68FFB9FF937ADAF9F75D72F914" authorityName="Smith" authorityYear="1956" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="chicoensis">
<emphasis id="B94EEAF9FFB9FF937ADAF9F75D72F914" italics="true" pageId="92" pageNumber="93">P. chicoensis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B8536EBFFB9FF937F95F94E588BF8EB" blockId="92.[151,1437,151,2043]" pageId="92" pageNumber="93">
Workers of these two species display few consistent morphological differences, but queens and males differ in both morphology and size. Worker allometry is similar for both species (see above), but queens and males of
<taxonomicName id="4C3A4D68FFB9FF937ADAF96A5D7FF88F" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFB9FF937ADAF96A5D7FF88F" italics="true" pageId="92" pageNumber="93">V. stoddardi</emphasis>
</taxonomicName>
are distinctly larger than those of
<taxonomicName id="4C3A4D68FFB9FF937D29F8865F74F88F" authorityName="Smith" authorityYear="1956" box="[635,778,1801,1827]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="chicoensis">
<emphasis id="B94EEAF9FFB9FF937D29F8865F74F88F" box="[635,778,1801,1827]" italics="true" pageId="92" pageNumber="93">V. chicoensis</emphasis>
</taxonomicName>
(
<figureCitation id="13012A6EFFB9FF937C4AF8855FF6F88F" box="[792,904,1801,1827]" captionStart="FIGURE 19" captionStartId="38.[151,250,1949,1973]" captionTargetBox="[217,1373,196,1910]" captionTargetId="figure-26@38.[203,1384,181,1925]" captionTargetPageId="38" captionText="FIGURE 19. Bivariate plots for workers, queens, and males of Veromessor chicoensis and V. stoddardi: (A) head width versus head length and (B) head width versus mesosoma length. Sample size for each caste is in parentheses." figureDoi="http://doi.org/10.5281/zenodo.7318868" httpUri="https://zenodo.org/record/7318868/files/figure.png" pageId="92" pageNumber="93">Figure 19</figureCitation>
). A molecular phylogeny based on UCEs shows that both species form divergent, reciprocally monophyletic lineages (M.L. Borowiec, unpub. data).
</paragraph>
<paragraph id="8B8536EBFFB9FF937F95F8DD58BBF857" blockId="92.[151,1437,151,2043]" pageId="92" pageNumber="93">
<bibRefCitation id="EFAB4B1AFFB9FF937F95F8DD5E53F8C7" author="Wheeler, W. M. &amp; Creighton, W. S." box="[199,557,1873,1899]" pageId="92" pageNumber="93" pagination="341 - 387" refId="ref73843" refString="Wheeler, W. M. &amp; Creighton, W. S. (1934) A study of the ant genera Novomessor and Veromessor. Proceedings of the American Academy of Arts and Sciences, 69 (9), 341 - 387. https: // doi. org / 10.2307 / 20023057" type="journal article" year="1934">Wheeler and Creighton (1934)</bibRefCitation>
wrongly restricted the
<typeStatus id="54818849FFB9FF937C17F8DD5F0BF8C7" box="[837,885,1873,1899]" pageId="92" pageNumber="93">type</typeStatus>
locality of
<taxonomicName id="4C3A4D68FFB9FF937B57F8DE58EEF8C7" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[1029,1168,1873,1899]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFB9FF937B57F8DE58EEF8C7" box="[1029,1168,1873,1899]" italics="true" pageId="92" pageNumber="93">V. stoddardi</emphasis>
</taxonomicName>
to San Diego County, indicating, “the
<typeStatus id="54818849FFB9FF937E1FF8F95D03F823" box="[333,381,1909,1935]" pageId="92" pageNumber="93">type</typeStatus>
series comes from several localities all in San Diego County,
<collectingRegion id="49FEF809FFB9FF937B6CF8F858CDF822" box="[1086,1203,1908,1934]" country="United States of America" name="California" pageId="92" pageNumber="93">California</collectingRegion>
. It seems preferable, for this reason to regard San Diego County as the
<typeStatus id="54818849FFB9FF937D9EF8155E82F81F" box="[716,764,1945,1971]" pageId="92" pageNumber="93">type</typeStatus>
locality.” This is a remarkable statement given that none of the
<typeStatus id="54818849FFB9FF937FEDF8315C91F87B" box="[191,239,1981,2007]" pageId="92" pageNumber="93">type</typeStatus>
material came from San Diego County. The locality cited in the description by
<bibRefCitation id="EFAB4B1AFFB9FF937B0EF8315889F87B" author="Emery, C." box="[1116,1271,1980,2007]" pageId="92" pageNumber="93" pagination="257 - 360" refId="ref67769" refString="Emery, C. (1895) Beitrage zur Kenntniss der nordamerikanischen Ameisenfauna. Zoologische Jahrbucher. Abtheilung fur Systematik, Geographie und Biologie der Tiere, 8, 257 - 360." type="journal article" year="1895">Emery (1895)</bibRefCitation>
is San Jacinto, Riverside County,
<collectingRegion id="49FEF809FFB9FF937E3AF86C5DA2F856" box="[360,476,2016,2042]" country="United States of America" name="California" pageId="92" pageNumber="93">California</collectingRegion>
, and this is the correct
<typeStatus id="54818849FFB9FF937D8CF86D5F70F857" box="[734,782,2017,2043]" pageId="92" pageNumber="93">type</typeStatus>
locality, as noted by
<bibRefCitation id="EFAB4B1AFFB9FF937CAEF86C58BFF857" author="Creighton, W. S." box="[1020,1217,2016,2043]" pageId="92" pageNumber="93" pagination="1 - 585" refId="ref67372" refString="Creighton, W. S. (1950) The ants of North America. Bulletin of the Museum of Comparative Zoology, 104, 1 - 585." type="journal article" year="1950">Creighton (1950)</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF456663FFB8FF927FC5F81B58ECF878" ID-DOI="http://doi.org/10.5281/zenodo.7318938" ID-Zenodo-Dep="7318938" httpUri="https://zenodo.org/record/7318938/files/figure.png" pageId="93" pageNumber="94" startId="93.[151,250,1943,1967]" targetBox="[404,1184,181,1920]" targetPageId="93">
<paragraph id="8B8536EBFFB8FF927FC5F81B58ECF878" blockId="93.[151,1437,1943,2004]" pageId="93" pageNumber="94">
<emphasis id="B94EEAF9FFB8FF927FC5F81B5D5FF803" bold="true" box="[151,289,1943,1967]" pageId="93" pageNumber="94">FIGURE 48.</emphasis>
Photograph of
<taxonomicName id="4C3A4D68FFB8FF927E90F8145EE0F81C" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[450,670,1944,1968]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFB8FF927E90F8145EE0F81C" box="[450,670,1944,1968]" italics="true" pageId="93" pageNumber="94">Veromessor stoddardi</emphasis>
</taxonomicName>
minor worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT0922825). Photographs by Michele Esposito from www.AntWeb.org.
</paragraph>
</caption>
<caption id="DF456663FFBBFF917FC5F81B58ECF87F" ID-DOI="http://doi.org/10.5281/zenodo.7318940" ID-Zenodo-Dep="7318940" httpUri="https://zenodo.org/record/7318940/files/figure.png" pageId="94" pageNumber="95" startId="94.[151,250,1943,1967]" targetBox="[420,1168,181,1919]" targetPageId="94">
<paragraph id="8B8536EBFFBBFF917FC5F81B58ECF87F" blockId="94.[151,1437,1943,2003]" pageId="94" pageNumber="95">
<emphasis id="B94EEAF9FFBBFF917FC5F81B5D5FF803" bold="true" box="[151,289,1943,1967]" pageId="94" pageNumber="95">FIGURE 49.</emphasis>
Photograph of
<taxonomicName id="4C3A4D68FFBBFF917E90F8145EE1F803" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[450,671,1943,1967]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFBBFF917E90F8145EE1F803" box="[450,671,1943,1967]" italics="true" pageId="94" pageNumber="95">Veromessor stoddardi</emphasis>
</taxonomicName>
major worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT0922824). Photographs by Michele Esposito from www.AntWeb.org.
</paragraph>
</caption>
<caption id="DF456663FFBAFF907FC5F8145873F878" ID-DOI="http://doi.org/10.5281/zenodo.7318942" ID-Zenodo-Dep="7318942" httpUri="https://zenodo.org/record/7318942/files/figure.png" pageId="95" pageNumber="96" startId="95.[151,250,1944,1968]" targetBox="[404,1184,181,1920]" targetPageId="95">
<paragraph id="8B8536EBFFBAFF907FC5F8145873F878" blockId="95.[151,1436,1944,2004]" pageId="95" pageNumber="96">
<emphasis id="B94EEAF9FFBAFF907FC5F8145D61F81C" bold="true" box="[151,287,1944,1968]" pageId="95" pageNumber="96">FIGURE 50.</emphasis>
Photograph of
<taxonomicName id="4C3A4D68FFBAFF907EEAF8155EEDF81C" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[440,659,1944,1968]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FFBAFF907EEAF8155EEDF81C" box="[440,659,1944,1968]" italics="true" pageId="95" pageNumber="96">Veromessor stoddardi</emphasis>
</taxonomicName>
alate queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (LACMENT370303). Photographs by Wade Lee from www.AntWeb.org.
</paragraph>
</caption>
<caption id="DF456663FF85FFAF7FC5F81B585FF878" ID-DOI="http://doi.org/10.5281/zenodo.7318944" ID-Zenodo-Dep="7318944" httpUri="https://zenodo.org/record/7318944/files/figure.png" pageId="96" pageNumber="97" startId="96.[151,250,1943,1967]" targetBox="[405,1180,181,1920]" targetPageId="96">
<paragraph id="8B8536EBFF85FFAF7FC5F81B585FF878" blockId="96.[151,1436,1943,2004]" pageId="96" pageNumber="97">
<emphasis id="B94EEAF9FF85FFAF7FC5F81B5D61F803" bold="true" box="[151,287,1943,1967]" pageId="96" pageNumber="97">FIGURE 51.</emphasis>
Photograph of
<taxonomicName id="4C3A4D68FF85FFAF7EE8F8145EE8F81C" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[442,662,1944,1968]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FF85FFAF7EE8F8145EE8F81C" box="[442,662,1944,1968]" italics="true" pageId="96" pageNumber="97">Veromessor stoddardi</emphasis>
</taxonomicName>
male: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (LACMENT364104). Photographs by Michele Esposito from www.AntWeb.org.
</paragraph>
</caption>
<subSubSection id="C3206560FF84FFAD7F95FF1A5FD7FEED" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98" type="biology_ecology">
<paragraph id="8B8536EBFF84FFAE7F95FF1A58FBFEED" blockId="97.[151,1437,150,609]" pageId="97" pageNumber="98">
<emphasis id="B94EEAF9FF84FFAE7F95FF1A5D59FF1C" bold="true" box="[199,295,150,176]" pageId="97" pageNumber="98">Biology.</emphasis>
Workers of
<taxonomicName id="4C3A4D68FF84FFAE7EFFFF1B5E4FFF1D" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[429,561,151,177]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FF84FFAE7EFFFF1B5E4FFF1D" box="[429,561,151,177]" italics="true" pageId="97" pageNumber="98">V. stoddardi</emphasis>
</taxonomicName>
are solitary foragers that commence activities near dusk and forage through the night (M. Bennett, pers. comm.). Another observer indicated that colonies have long foraging columns (K.L. Jorda, pers. comm.), such that more information is needed to understand foraging biology of this species. Colonies probably contain
<geoCoordinate id="EE0E502CFF84FFAE7FA2FE8F5D50FEB1" box="[240,302,259,285]" degrees="1,000" direction="north" orientation="latitude" pageId="97" pageNumber="98" precision="55" value="1.0">1,000</geoCoordinate>
<geoCoordinate id="EE0E502CFF84FFAE7E7FFE8F5D02FEB1" box="[301,380,259,285]" degrees="2,000" direction="west" orientation="longitude" pageId="97" pageNumber="98" precision="55" value="-2.0">2,000</geoCoordinate>
workers (
<bibRefCitation id="EFAB4B1AFF84FFAE7EB8FE8F5EDAFEB1" author="Creighton, W. S." box="[490,676,258,285]" pageId="97" pageNumber="98" pagination="1 - 18" refId="ref67401" refString="Creighton, W. S. (1953) New data on the habits of the ants of the genus Veromessor. American Museum Novitates, 1612, 1 - 18." type="journal article" year="1953">Creighton, 1953</bibRefCitation>
; R.A. Johnson, pers. obs.). Workers are strongly polymorphic. Nests are often placed in hard clay soil and often have two to three entrances (
<bibRefCitation id="EFAB4B1AFF84FFAE7CEFFEAB5806FEED" author="Creighton, W. S." box="[957,1144,294,321]" pageId="97" pageNumber="98" pagination="1 - 18" refId="ref67401" refString="Creighton, W. S. (1953) New data on the habits of the ants of the genus Veromessor. American Museum Novitates, 1612, 1 - 18." type="journal article" year="1953">Creighton, 1953</bibRefCitation>
).
</paragraph>
<paragraph id="8B8536EBFF84FFAE7F95FEC75947FE25" blockId="97.[151,1437,150,609]" pageId="97" pageNumber="98">
Gland chemistry has not been examined in
<emphasis id="B94EEAF9FF84FFAE7D96FEC75F28FEC9" box="[708,854,331,357]" italics="true" pageId="97" pageNumber="98">
<taxonomicName id="4C3A4D68FF84FFAE7D96FEC75F2CFEC9" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[708,850,331,357]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="stoddardi">V. stoddardi</taxonomicName>
.
</emphasis>
Like other small-colony congeners, workers of
<taxonomicName id="4C3A4D68FF84FFAE7ADAFEC75D7FFE25" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FF84FFAE7ADAFEC75D7FFE25" italics="true" pageId="97" pageNumber="98">V. stoddardi</emphasis>
</taxonomicName>
have a small pygidial gland reservoir and lack a textured tergal cuticle (
<bibRefCitation id="EFAB4B1AFF84FFAE7B7AFEE35950FE25" author="Holldobler, B. &amp; Plowes, N. J. R. &amp; Johnson, R. A. &amp; Nishshanka, U. &amp; Liu, C. &amp; Attygalle, A. B." box="[1064,1326,367,393]" pageId="97" pageNumber="98" pagination="863 - 869" refId="ref69008" refString="Holldobler, B., Plowes, N. J. R., Johnson, R. A., Nishshanka, U., Liu, C. &amp; Attygalle, A. B. (2013) Pygidial gland chemistry and potential alarm-recruitment function in column foraging, but not solitary, Nearctic Messor harvesting ants (Hymenoptera: Formicidae: Myrmicinae). Journal of Insect Physiology, 59 (9), 863 - 869. https: // doi. org / 10.1016 / j. jinsphys. 2013.06.006" type="journal article" year="2013">
Hölldobler
<emphasis id="B94EEAF9FF84FFAE7BF9FEE3589DFE25" box="[1195,1251,367,393]" italics="true" pageId="97" pageNumber="98">et al.</emphasis>
, 2013
</bibRefCitation>
).
</paragraph>
<paragraph id="8B8536EBFF84FFAE7F95FE1F5FC5FE7D" blockId="97.[151,1437,150,609]" pageId="97" pageNumber="98">
Mating flights occur near dusk and post-dawn hours in June and July (
<bibRefCitation id="EFAB4B1AFF84FFAE7B54FE1F58BAFE01" author="Creighton, W. S." box="[1030,1220,402,429]" pageId="97" pageNumber="98" pagination="1 - 18" refId="ref67401" refString="Creighton, W. S. (1953) New data on the habits of the ants of the genus Veromessor. American Museum Novitates, 1612, 1 - 18." type="journal article" year="1953">Creighton, 1953</bibRefCitation>
; M. Bennett, pers. comm.). Nothing is known about mating frequency or colony founding.
</paragraph>
<paragraph id="8B8536EBFF84FFAE7F95FE575D63FDCD" blockId="97.[151,1437,150,609]" pageId="97" pageNumber="98">
<taxonomicName id="4C3A4D68FF84FFAE7F95FE575DC5FE59" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[199,443,475,501]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FF84FFAE7F95FE575DC5FE59" box="[199,443,475,501]" italics="true" pageId="97" pageNumber="98">Veromessor stoddardi</emphasis>
</taxonomicName>
occurs at elevations from
<quantity id="4CC29B0EFF84FFAE7DBAFE575F22FE59" box="[744,860,475,501]" metricMagnitude="2" metricUnit="m" metricValue="5.2" metricValueMax="9.8" metricValueMin="0.6" pageId="97" pageNumber="98" unit="m" value="520.0" valueMax="980.0" valueMin="60.0">60980 m</quantity>
, and it appears to be mostly restricted to soils with higher clay content (M. Bennett, pers. comm.). This species occurs in the
<collectingRegion id="49FEF809FF84FFAE7C80FE735800FDB5" box="[978,1150,511,537]" country="Mexico" name="Baja California" pageId="97" pageNumber="98">Baja California</collectingRegion>
desert,
<collectingRegion id="49FEF809FF84FFAE7B84FE735936FDB5" box="[1238,1352,511,537]" country="United States of America" name="California" pageId="97" pageNumber="98">California</collectingRegion>
coastal sage and chaparral, and
<collectingRegion id="49FEF809FF84FFAE7EF7FDAF5E69FD91" box="[421,535,547,573]" country="United States of America" name="California" pageId="97" pageNumber="98">California</collectingRegion>
montane chaparral and woodlands ecoregions, as defined by
<bibRefCitation id="EFAB4B1AFF84FFAE7B9AFDAF59E2FD91" author="Olson, D. M. &amp; Dinerstein, E. &amp; Wikramanayake, E. D. &amp; Burgess, N. D. &amp; Powell, G. V. N. &amp; Underwood, E. C. &amp; D'Amico, J. A. &amp; Itoua, I. &amp; Strand, H. E. &amp; Morrison, J. C. &amp; Loucks, C. J. &amp; Allnutt, T. F. &amp; Ricketts, T. H. &amp; Kura, Y. &amp; Lamoreux, J. F. &amp; Wettengel, W. W. &amp; Hedao, P. &amp; Kassem, K. R." box="[1224,1436,547,573]" pageId="97" pageNumber="98" pagination="933 - 938" refId="ref70788" refString="Olson, D. M., Dinerstein, E., Wikramanayake, E. D., Burgess, N. D., Powell, G. V. N., Underwood, E. C., D'Amico, J. A., Itoua, I., Strand, H. E., Morrison, J. C., Loucks, C. J., Allnutt, T. F., Ricketts, T. H., Kura, Y., Lamoreux, J. F., Wettengel, W. W., Hedao, P. &amp; Kassem, K. R. (2001) Terrestrial ecoregions of the world: a new map of life on Earth. Bioscience, 51 (11), 933 - 938. https: // doi. org / 10.1641 / 0006 - 3568 (2001) 051 [0933: TEOTWA] 2.0. CO; 2" type="journal article" year="2001">
Olson
<emphasis id="B94EEAF9FF84FFAE7A40FDAF5935FD91" box="[1298,1355,547,573]" italics="true" pageId="97" pageNumber="98">et al.</emphasis>
(2001)
</bibRefCitation>
(
<figureCitation id="13012A6EFF84FFAE7FCDFDCB5D6FFDCD" box="[159,273,582,609]" captionStart="FIGURE 52" captionStartId="97.[151,250,1652,1676]" captionTargetBox="[151,1434,633,1628]" captionTargetId="figure-213@97.[151,1435,633,1629]" captionTargetPageId="97" captionText="FIGURE 52. Geographic distribution of Veromessor stoddardi. The larger black circle denotes the type locality." figureDoi="http://doi.org/10.5281/zenodo.7318946" httpUri="https://zenodo.org/record/7318946/files/figure.png" pageId="97" pageNumber="98">Figure 52</figureCitation>
).
</paragraph>
<caption id="DF456663FF84FFAE7FC5F9F85976F921" ID-DOI="http://doi.org/10.5281/zenodo.7318946" ID-Zenodo-Dep="7318946" box="[151,1288,1652,1677]" httpUri="https://zenodo.org/record/7318946/files/figure.png" pageId="97" pageNumber="98" startId="97.[151,250,1652,1676]" targetBox="[151,1434,633,1628]" targetPageId="97">
<paragraph id="8B8536EBFF84FFAE7FC5F9F85976F921" blockId="97.[151,1288,1652,1677]" box="[151,1288,1652,1677]" pageId="97" pageNumber="98">
<emphasis id="B94EEAF9FF84FFAE7FC5F9F85D5EF921" bold="true" box="[151,288,1652,1677]" pageId="97" pageNumber="98">FIGURE 52.</emphasis>
Geographic distribution of
<taxonomicName id="4C3A4D68FF84FFAE7D6BF9F95F6BF921" authorityName="Ward, Brady, Fisher, and Schultz" authorityYear="2015" box="[569,789,1653,1677]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="stoddardi">
<emphasis id="B94EEAF9FF84FFAE7D6BF9F95F6BF921" box="[569,789,1653,1677]" italics="true" pageId="97" pageNumber="98">Veromessor stoddardi</emphasis>
</taxonomicName>
. The larger black circle denotes the type locality.
</paragraph>
</caption>
<paragraph id="8B8536EBFF84FFAE7FC5F9535D1BF955" blockId="97.[151,357,1759,1785]" box="[151,357,1759,1785]" pageId="97" pageNumber="98">
<emphasis id="B94EEAF9FF84FFAE7FC5F9535D1BF955" bold="true" box="[151,357,1759,1785]" pageId="97" pageNumber="98">Additional notes</emphasis>
</paragraph>
<paragraph id="8B8536EBFF84FFAD7FC5F8AB5FD7FEED" blockId="97.[151,1437,1831,2037]" lastBlockId="98.[151,1437,151,321]" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98">
We also note a putative new species of
<taxonomicName id="4C3A4D68FF84FFAE7D20F8A45E8AF8ED" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[626,756,1832,1857]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAF9FF84FFAE7D20F8A45E8AF8ED" box="[626,756,1832,1857]" italics="true" pageId="97" pageNumber="98">Veromessor</emphasis>
</taxonomicName>
that is unavailable to describe until additional material is collected. Two workers of this species were collected from the northern end of the Mohawk Dunes,
<taxonomicName id="4C3A4D68FF84FFAE7BAEF8C759E6F8C9" authority="County" authorityName="County" box="[1276,1432,1867,1893]" class="Insecta" genus="Yuma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">Yuma County</taxonomicName>
, Arizona, during summer (year unknown). These specimens were examined by Roy Snelling, who stated that they were a new species closely related to
<taxonomicName id="4C3A4D68FF84FFAE7D15F8185EABF801" box="[583,725,1939,1965]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis id="B94EEAF9FF84FFAE7D15F8185EABF801" box="[583,725,1939,1965]" italics="true" pageId="97" pageNumber="98">V. pergandei</emphasis>
</taxonomicName>
. These
<specimenCount id="9D3CFD62FF84FFAE7C62F81F5FA9F801" box="[816,983,1939,1965]" pageId="97" pageNumber="98" type="generic">two specimens</specimenCount>
were undoubtedly deposited at LACM, but the senior author could not find these workers during a visit to LACM in 2018, and they are presumed lost. The senior author and others (including RRS) have revisited Mohawk Dunes multiple times through the years without finding additional specimens, and we write this addendum so that other collectors can be on the lookout for this species. We presume that this is a small-colony, nocturnal species given the difficulty finding additional specimens. As a note on geographic range, there are numerous sand dunes in the area, and we presume this species would also occur south into the sand dunes of the Gran Desierto Altar in northwestern
<collectingRegion id="49FEF809FF87FFAD7CB1FE8F584BFEB1" box="[995,1077,259,285]" country="Mexico" name="Sonora" pageId="98" pageNumber="99">Sonora</collectingRegion>
,
<collectingCountry id="F32D767BFF87FFAD7B13FE8F58E4FEB1" box="[1089,1178,259,285]" name="Mexico" pageId="98" pageNumber="99">Mexico</collectingCountry>
. We predict ecology of this species (
<tableCitation id="C6B80350FF87FFAD7E74FEAB5D05FEED" box="[294,379,295,321]" captionStart="TABLE 1" captionStartId="99.[151,244,643,669]" captionTargetPageId="99" captionText="TABLE 1. Summary of morphological and ecological traits for species of Veromessor." httpUri="http://table.plazi.org/id/DF456663FF86FFAC7FC5FD0F5847FD31" pageId="98" pageNumber="99" tableUuid="DF456663FF86FFAC7FC5FD0F5847FD31">Table 1</tableCitation>
and below), which we list as
<emphasis id="B94EEAF9FF87FFAD7D95FEAB5EA6FEEC" box="[711,728,295,320]" italics="true" pageId="98" pageNumber="99">V</emphasis>
. sp.
<taxonomicName id="4C3A4D68FF87FFAD7C5FFEAB5FDFFEED" box="[781,929,295,321]" class="Insecta" family="Formicidae" genus="Veromessor" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="pergandei">
cf.
<emphasis id="B94EEAF9FF87FFAD7C63FEAB5FDFFEED" box="[817,929,295,321]" italics="true" pageId="98" pageNumber="99">pergandei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>