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<document id="B2ABB50EFE107F380239C740EB3DBAC9" ID-DOI="10.5852/ejt.2024.957.2651" ID-ISSN="2118-9773" ID-Zenodo-Dep="13773385" ID-ZooBank="9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726582870537" checkinUser="plazi" docAuthor="Selnekovič, Dávid, Jäch, Manfred A. &amp; Kodada, Ján" docDate="2024" docId="038B5001DA58E71CFDDDFA03FEAF25F0" docLanguage="en" docName="EJT.2024.957.1-229.pdf" docOrigin="European Journal of Taxonomy 957" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Elmomorphus bryanti Hinton 1935" docType="treatment" docVersion="2" lastPageNumber="84" masterDocId="FFB22879DA08E74FFF9EFFBAFFE32440" masterDocTitle="Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)" masterLastPageNumber="229" masterPageNumber="1" pageNumber="81" updateTime="1726584008522" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9BBD0E95AB29A12CFBF115EFC1ADFFCF">Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)</mods:title>
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<mods:namePart id="DB0A79CD8AE73A618EA28C353AF3ADDE">Selnekovič, Dávid</mods:namePart>
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<mods:affiliation id="BD006A4F789E9F00A4E306120AB5035E">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="A1CB32538DB131EE97CE88C4D7D06722" type="email">david.selnekovic@uniba.sk</mods:nameIdentifier>
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<mods:namePart id="930A24719F0FAE232988C45C7AC04181">Jäch, Manfred A.</mods:namePart>
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<mods:affiliation id="4209306F8A5D06AAFC54EB0B55BA5061">Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria.</mods:affiliation>
<mods:nameIdentifier id="E284FAED599D07448937BB8E8D589D4F" type="email">manfred.jaech@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:namePart id="0FDD5874789A06FC544012D0FE41D33A">Kodada, Ján</mods:namePart>
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<mods:affiliation id="33E29B09B9E9FEB98CC6D15FA322CE6D">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="AEA98E143E9014FB4B0896EBC71B0522" type="email">jan.kodada@uniba.sk</mods:nameIdentifier>
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<mods:title id="C99107C0E1CFAEF8AE7A320FBD3784AD">European Journal of Taxonomy</mods:title>
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<mods:date id="5A515C4E54011CCD2B1904774C90A63E">2024</mods:date>
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<subSubSection id="C338B29CDA58E71FFDDDFA03FC132195" box="[579,1008,1465,1493]" pageId="80" pageNumber="81" type="nomenclature">
<paragraph id="8B9DE117DA58E71FFDDDFA03FC132195" blockId="80.[579,1008,1465,1528]" box="[579,1008,1465,1493]" pageId="80" pageNumber="81">
<heading id="D0D5567BDA58E71FFDDDFA03FC132195" box="[579,1008,1465,1493]" centered="true" fontSize="11" level="2" pageId="80" pageNumber="81" reason="2">
<taxonomicName id="4C229A94DA58E71FFDDDFA03FC132195" authority="Hinton, 1935" authorityName="Hinton" authorityYear="1935" box="[579,1008,1465,1493]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B9563D05DA58E71FFDDDFA03FCAD2194" bold="true" box="[579,846,1465,1492]" italics="true" pageId="80" pageNumber="81">Elmomorphus bryanti</emphasis>
<bibRefCitation id="EFB39CE6DA58E71FFCCBFA00FC132195" author="Hinton H. E." box="[853,1008,1466,1493]" pageId="80" pageNumber="81" pagination="169 - 179" refId="ref99677" refString="Hinton H. E. 1935. Notes on the Dryopoidea (Col.). Stylops 4 (8): 169 - 179. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00660. x" type="journal article" year="1935">Hinton, 1935</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C338B29CDA58E71FFD59FA64FC8E21B8" box="[711,877,1502,1528]" pageId="80" pageNumber="81" type="description">
<paragraph id="8B9DE117DA58E71FFD59FA64FC8E21B8" blockId="80.[579,1008,1465,1528]" box="[711,877,1502,1528]" pageId="80" pageNumber="81">
<figureCitation id="1319FD92DA58E71FFD59FA64FCC221B8" box="[711,801,1502,1528]" captionStart="Fig" captionStartId="81.[189,232,1909,1935]" captionTargetBox="[389,1148,269,1844]" captionTargetId="figure-25@81.[374,1161,265,1861]" captionTargetPageId="81" captionText="Fig. 38. Elmomorphus bryanti Hinton, 1935, male from Gombak River, Malaysia (BMNH), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773488" httpUri="https://zenodo.org/record/13773488/files/figure.png" pageId="80" pageNumber="81">Figs 38</figureCitation>
,
<figureCitation id="1319FD92DA58E71FFCB3FA64FC8E21B8" box="[813,877,1502,1528]" captionStart="Fig" captionStartId="218.[189,232,1871,1897]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-40@218.[189,1399,265,1830]" captionTargetPageId="218" captionText="Fig. 110. Distribution maps.A. Elmomorphus corpulentus sp. nov.B. E. ellipticus sp. nov.C. E. mazzoldii sp. nov. D. E. bryanti Hinton, 1935. E. E. prosternalis Hinton, 1935. F. E. montanus (Grouvelle, 1913)." figureDoi="http://doi.org/10.5281/zenodo.13773714" httpUri="https://zenodo.org/record/13773714/files/figure.png" pageId="80" pageNumber="81">110D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA58E71FFF23F99AFCF3223C" pageId="80" pageNumber="81" type="reference_group">
<paragraph id="8B9DE117DA58E71FFF23F99AFC3D227A" blockId="80.[189,990,1568,1595]" box="[189,990,1568,1595]" pageId="80" pageNumber="81">
<treatmentCitationGroup id="AB32C639DA58E71FFF23F99AFC3D227A" box="[189,990,1568,1595]" pageId="80" pageNumber="81">
<treatmentCitation id="0A83C706DA58E71FFF23F99AFD36227A" author="Hinton H. E." box="[189,725,1568,1595]" page="172" pageId="80" pageNumber="81" year="1935">
<taxonomicName id="4C229A94DA58E71FFF23F99AFD36227A" authority="Hinton, 1935: 171 - 172" authorityName="Hinton" authorityPageNumber="171 - 172" authorityYear="1935" box="[189,725,1568,1595]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B9563D05DA58E71FFF23F99AFE5D227A" box="[189,446,1568,1594]" italics="true" pageId="80" pageNumber="81">Elmomorphus bryanti</emphasis>
<bibRefCitation id="EFB39CE6DA58E71FFE5BF99AFD36227A" author="Hinton H. E." box="[453,725,1568,1595]" pageId="80" pageNumber="81" pagination="169 - 179" refId="ref99677" refString="Hinton H. E. 1935. Notes on the Dryopoidea (Col.). Stylops 4 (8): 169 - 179. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00660. x" type="journal article" year="1935">Hinton, 1935: 171172</bibRefCitation>
</taxonomicName>
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(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9DE117DA58E71FFF23F9D8FCF3223C" blockId="80.[189,784,1634,1660]" box="[189,784,1634,1660]" pageId="80" pageNumber="81">
<treatmentCitationGroup id="AB32C639DA58E71FFF23F9D8FCF3223C" box="[189,784,1634,1660]" pageId="80" pageNumber="81">
<taxonomicName id="4C229A94DA58E71FFF23F9D8FE5D223C" authorityName="Hinton" authorityYear="1935" box="[189,446,1634,1660]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B9563D05DA58E71FFF23F9D8FE5D223C" box="[189,446,1634,1660]" italics="true" pageId="80" pageNumber="81">Elmomorphus bryanti</emphasis>
</taxonomicName>
<treatmentCitation id="0A83C706DA58E71FFE45F9D8FCEF223C" author="Shepard W. D. &amp; Sites R. W." box="[475,780,1634,1660]" page="93" pageId="80" pageNumber="81" year="2016">
<bibRefCitation id="EFB39CE6DA58E71FFE45F9D8FCEF223C" author="Shepard W. D. &amp; Sites R. W." box="[475,780,1634,1660]" pageId="80" pageNumber="81" pagination="89 - 126" refId="ref100782" refString="Shepard W. D. &amp; Sites R. W. 2016. Aquatic beetles of the families Dryopidae and Elmidae (Insecta: Coleoptera: Byrrhoidea) of Thailand: Annotated list and illustrated key to genera. Natural History Bulletin of the Siam Society 61 (2): 89 - 126." type="journal article" year="2016">Shepard &amp; Sites 2016: 93</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA58E71FFF23F91EFE0A23A8" pageId="80" pageNumber="81" type="diagnosis">
<paragraph id="8B9DE117DA58E71FFF23F91EFE2A22FE" blockId="80.[189,457,1700,1726]" box="[189,457,1700,1726]" pageId="80" pageNumber="81">
<heading id="D0D5567BDA58E71FFF23F91EFE2A22FE" bold="true" box="[189,457,1700,1726]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
<emphasis id="B9563D05DA58E71FFF23F91EFE2A22FE" bold="true" box="[189,457,1700,1726]" pageId="80" pageNumber="81">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA58E71FFF23F969FE0A23A8" blockId="80.[189,1400,1747,2024]" pageId="80" pageNumber="81">
<taxonomicName id="4C229A94DA58E71FFF23F969FE2722AD" authorityName="Hinton" authorityYear="1935" box="[189,452,1747,1773]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B9563D05DA58E71FFF23F969FE2722AD" box="[189,452,1747,1773]" italics="true" pageId="80" pageNumber="81">Elmomorphus bryanti</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C229A94DA58E71FFDCBF96EFD0D22AD" baseAuthorityName="Grouvelle" baseAuthorityYear="1913" box="[597,750,1748,1773]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B9563D05DA58E71FFDCBF96EFD0D22AD" box="[597,750,1748,1773]" italics="true" pageId="80" pageNumber="81">E. montanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C229A94DA58E71FFC9DF96EFC3022AD" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[771,979,1748,1773]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="paramontanus" status="sp. nov.">
<emphasis id="B9563D05DA58E71FFC9DF96EFC3022AD" box="[771,979,1748,1773]" italics="true" pageId="80" pageNumber="81">E. paramontanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA58E71FFC7CF96EFBA622AE" box="[994,1093,1748,1774]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C229A94DA58E71FFB11F96EFAA022AD" authorityName="Hinton" authorityYear="1935" box="[1167,1347,1747,1773]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="prosternalis">
<emphasis id="B9563D05DA58E71FFB11F96EFAA022AD" box="[1167,1347,1747,1773]" italics="true" pageId="80" pageNumber="81">E. prosternalis</emphasis>
</taxonomicName>
but differs in the smaller body size, TL:
<quantity id="4CDA4CF2DA58E71FFDE5F94DFCE82352" box="[635,779,1783,1810]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="80" pageNumber="81" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.62.8 mm</quantity>
versus TL:
<quantity id="4CDA4CF2DA58E71FFC01F94DFC102351" box="[927,1011,1783,1809]" metricMagnitude="-2" metricUnit="m" metricValue="8.128" pageId="80" pageNumber="81" unit="in" value="3.2">3.20 in</quantity>
<taxonomicName id="4C229A94DA58E71FFC63F942FB022352" authority=", TL" authorityName="TL" box="[1021,1249,1783,1810]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="prosternalis">
<emphasis id="B9563D05DA58E71FFC63F942FB4E2351" box="[1021,1197,1783,1809]" italics="true" pageId="80" pageNumber="81">E. prosternalis</emphasis>
, TL
</taxonomicName>
<quantity id="4CDA4CF2DA58E71FFB75F94DFAB52352" box="[1259,1366,1783,1810]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" pageId="80" pageNumber="81" unit="mm" value="3.35">3.35 mm</quantity>
in
<taxonomicName id="4C229A94DA58E71FFF23F8A6FEB12375" baseAuthorityName="Grouvelle" baseAuthorityYear="1913" box="[189,338,1820,1845]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B9563D05DA58E71FFF23F8A6FEB12375" box="[189,338,1820,1845]" italics="true" pageId="80" pageNumber="81">E. montanus</emphasis>
</taxonomicName>
, and TL:
<specimenCount id="9D242A9EDA58E71FFE53F8A6FE1A2376" box="[461,505,1820,1846]" count="1" pageId="80" pageNumber="81" type="male">♂♂</specimenCount>
<quantity id="4CDA4CF2DA58E71FFD9DF8A1FD532376" box="[515,688,1819,1846]" metricMagnitude="-3" metricUnit="m" metricValue="3.115" metricValueMax="3.33" metricValueMin="2.9" pageId="80" pageNumber="81" unit="mm" value="3.115" valueMax="3.33" valueMin="2.9">2.903.33 mm</quantity>
(3.07± 0.12, n=26),
<specimenCount id="9D242A9EDA58E71FFC2CF8A6FC3D2376" box="[946,990,1820,1846]" count="1" pageId="80" pageNumber="81" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DA58E71FFC76F8A1FB762376" box="[1000,1173,1819,1846]" metricMagnitude="-3" metricUnit="m" metricValue="3.115" metricValueMax="3.35" metricValueMin="2.88" pageId="80" pageNumber="81" unit="mm" value="3.115" valueMax="3.35" valueMin="2.88">2.883.35 mm</quantity>
(3.20 ± 0.14, n =9) in
<taxonomicName id="4C229A94DA58E71FFF42F8FAFE452319" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[220,422,1856,1881]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="paramontanus">
<emphasis id="B9563D05DA58E71FFF42F8FAFE452319" box="[220,422,1856,1881]" italics="true" pageId="80" pageNumber="81">E. paramontanus</emphasis>
</taxonomicName>
. In
<taxonomicName id="4C229A94DA58E71FFE46F8FAFDAD2319" authorityName="Hinton" authorityYear="1935" box="[472,590,1855,1881]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B9563D05DA58E71FFE46F8FAFDAD2319" box="[472,590,1855,1881]" italics="true" pageId="80" pageNumber="81">E. bryanti</emphasis>
</taxonomicName>
, the elytral plastron bands are wider, occupying approximately two-thirds of the elytral width, while in the other mentioned species, they are narrower (up to one-third of the elytral width, measured at elytral midlength).
<taxonomicName id="4C229A94DA58E71FFC92F83CFBED23E0" authorityName="Hinton" authorityYear="1935" box="[780,1038,1926,1952]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B9563D05DA58E71FFC92F83CFBED23E0" box="[780,1038,1926,1952]" italics="true" pageId="80" pageNumber="81">Elmomorphus bryanti</emphasis>
</taxonomicName>
further differs in the presence of long setae on the labrum and in two clusters on the male prosternal process, and in the unique shape of the aedeagus (
<figureCitation id="1319FD92DA58E71FFE1DF874FE3823A8" box="[387,475,1998,2024]" captionStart="Fig" captionStartId="81.[189,232,1909,1935]" captionTargetBox="[389,1148,269,1844]" captionTargetId="figure-25@81.[374,1161,265,1861]" captionTargetPageId="81" captionText="Fig. 38. Elmomorphus bryanti Hinton, 1935, male from Gombak River, Malaysia (BMNH), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773488" httpUri="https://zenodo.org/record/13773488/files/figure.png" pageId="80" pageNumber="81">Fig. 38</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF5DB19FDA59E71EFF23F8CFFCA523F3" ID-DOI="http://doi.org/10.5281/zenodo.13773488" ID-Zenodo-Dep="13773488" httpUri="https://zenodo.org/record/13773488/files/figure.png" pageId="81" pageNumber="82" startId="81.[189,232,1909,1935]" targetBox="[389,1148,269,1844]" targetPageId="81">
<paragraph id="8B9DE117DA59E71EFF23F8CFFCA523F3" blockId="81.[189,1399,1909,1971]" pageId="81" pageNumber="82">
<emphasis id="B9563D05DA59E71EFF23F8CFFEF923CF" bold="true" box="[189,282,1909,1935]" pageId="81" pageNumber="82">Fig. 38.</emphasis>
<taxonomicName id="4C229A94DA59E71EFEBDF8CFFD2A23D0" authority="Hinton, 1935" authorityName="Hinton" authorityYear="1935" box="[291,713,1909,1936]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B9563D05DA59E71EFEBDF8CFFDC723CF" box="[291,548,1909,1935]" italics="true" pageId="81" pageNumber="82">Elmomorphus bryanti</emphasis>
<bibRefCitation id="EFB39CE6DA59E71EFDB5F8CFFD2A23D0" author="Hinton H. E." box="[555,713,1909,1936]" pageId="81" pageNumber="82" pagination="169 - 179" refId="ref99677" refString="Hinton H. E. 1935. Notes on the Dryopoidea (Col.). Stylops 4 (8): 169 - 179. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00660. x" type="journal article" year="1935">Hinton, 1935</bibRefCitation>
</taxonomicName>
, male from Gombak River, Malaysia (BMNH), aedeagus.
<emphasis id="B9563D05DA59E71EFF23F822FF3123F2" bold="true" box="[189,210,1944,1970]" pageId="81" pageNumber="82">A</emphasis>
. Lateral aspect.
<emphasis id="B9563D05DA59E71EFE0DF822FE4423F2" bold="true" box="[403,423,1944,1970]" pageId="81" pageNumber="82">B</emphasis>
. Ventral aspect. Scale bar: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C338B29CDA5AE71DFF23FEB3FE0A27CD" pageId="82" pageNumber="83" type="materials_examined">
<paragraph id="8B9DE117DA5AE71DFF23FEB3FE8F2562" blockId="82.[189,364,264,291]" box="[189,364,264,291]" pageId="82" pageNumber="83">
<heading id="D0D5567BDA5AE71DFF23FEB3FE8F2562" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
<emphasis id="B9563D05DA5AE71DFF23FEB3FE8F2562" bold="true" box="[189,364,264,291]" pageId="82" pageNumber="83">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF74FE8DFEBF2511" blockId="82.[189,1398,311,408]" box="[234,348,311,337]" pageId="82" pageNumber="83">
<emphasis id="B9563D05DA5AE71DFF74FE8DFEBF2511" bold="true" box="[234,348,311,337]" pageId="82" pageNumber="83">
<typeStatus id="54995FB5DA5AE71DFF74FE8DFEBF2511" box="[234,348,311,337]" pageId="82" pageNumber="83" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23FEE0FED725D8" blockId="82.[189,1398,311,408]" pageId="82" pageNumber="83">
<materialsCitation id="3B4AEB4ADA5AE71DFF23FEE0FED725D8" collectingDate="1932-04-08" collectionCode="NHMUK" collectorName="H. M. Pendlebury" country="Malaysia" location="Selangor State" pageId="82" pageNumber="83" specimenCount="1" specimenCount-female="1" stateProvince="Selangor" typeStatus="holotype">
<collectingCountry id="F335A187DA5AE71DFF23FEE0FEB72534" box="[189,340,346,372]" name="Malaysia" pageId="82" pageNumber="83">MALAYSIA</collectingCountry>
<emphasis id="B9563D05DA5AE71DFEF5FEE0FDFC2534" bold="true" box="[363,543,346,372]" pageId="82" pageNumber="83">
<collectingRegion id="49E62FF5DA5AE71DFEF5FEE0FDFC2534" box="[363,543,346,372]" country="Malaysia" name="Selangor" pageId="82" pageNumber="83">Selangor State</collectingRegion>
</emphasis>
<specimenCount id="9D242A9EDA5AE71DFDA8FEE1FDAE2535" box="[566,589,347,373]" count="1" pageId="82" pageNumber="83" type="female"></specimenCount>
; “MALAYA
<collectingRegion id="49E62FF5DA5AE71DFD7BFEE0FC732535" box="[741,912,346,373]" country="Malaysia" name="Kuala Lumpur" pageId="82" pageNumber="83">Kuala Lumpur</collectingRegion>
at light
<date id="FF9CC7D7DA5AE71DFC77FEE0FB702534" box="[1001,1171,346,372]" pageId="82" pageNumber="83" value="1932-04-08">
<collectingDate id="EFD83E3FDA5AE71DFC77FEE0FB702534" box="[1001,1171,346,372]" pageId="82" pageNumber="83" value="1932-04-08">April 8th 1932</collectingDate>
</date>
<collectorName id="26D784C1DA5AE71DFB06FEE1FA8B2534" box="[1176,1384,346,373]" pageId="82" pageNumber="83">H.M. Pendlebury</collectorName>
”;
<collectionCode id="ED3379D2DA5AE71DFF23FEC4FED325D8" box="[189,304,382,408]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="82" pageNumber="83" type="Museum">NHMUK</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF74FE05FE842599" blockId="82.[188,1399,447,684]" box="[234,359,447,473]" pageId="82" pageNumber="83">
<emphasis id="B9563D05DA5AE71DFF74FE05FE842599" bold="true" box="[234,359,447,473]" pageId="82" pageNumber="83">
<typeStatus id="54995FB5DA5AE71DFF74FE05FE842599" box="[234,359,447,473]" pageId="82" pageNumber="83" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23FE58FEDA26EC" blockId="82.[188,1399,447,684]" pageId="82" pageNumber="83">
According to the original description: “Three with same data as above [
<typeStatus id="54995FB5DA5AE71DFBB3FE58FB7F25BC" box="[1069,1180,482,508]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
], but collected on different dates, as follows: one on 1 Aug., 1929; two in Feb., 1931” (
<bibRefCitation id="EFB39CE6DA5AE71DFC7FFDBFFB982660" author="Hinton H. E." box="[993,1147,517,544]" pageId="82" pageNumber="83" pagination="169 - 179" refId="ref99677" refString="Hinton H. E. 1935. Notes on the Dryopoidea (Col.). Stylops 4 (8): 169 - 179. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00660. x" type="journal article" year="1935">Hinton 1935</bibRefCitation>
). There is
<specimenCount id="9D242A9EDA5AE71DFB6BFDBCFA94265F" box="[1269,1399,517,544]" count="1" pageId="82" pageNumber="83" type="female">one female</specimenCount>
in the Hinton collection in
<collectionCode id="ED3379D2DA5AE71DFE6CFD93FD872603" box="[498,612,553,579]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="82" pageNumber="83" type="Museum">NHMUK</collectionCode>
, with labels corresponding to the data mentioned above and the date “
<date id="FF9CC7D7DA5AE71DFF56FDF6FEA72626" box="[200,324,588,614]" pageId="82" pageNumber="83" value="1931-02">Feb. 1931</date>
”. However, this specimen is labelled “
<typeStatus id="54995FB5DA5AE71DFC97FDF6FC962626" box="[777,885,588,614]" pageId="82" pageNumber="83" type="paratype">Paratype</typeStatus>
, pusillus, H.E. Hinton”. It is quite possible, that this specimen is a
<typeStatus id="54995FB5DA5AE71DFE5DFDD5FDCB26C9" box="[451,552,623,649]" pageId="82" pageNumber="83" type="paratype">paratype</typeStatus>
of
<emphasis id="B9563D05DA5AE71DFDD3FDCAFD2926C9" box="[589,714,623,649]" italics="true" pageId="82" pageNumber="83">
<taxonomicName id="4C229A94DA5AE71DFDD3FDCAFD2526C9" authorityName="Hinton" authorityYear="1935" box="[589,710,623,649]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="bryanti">E. bryanti</taxonomicName>
.
</emphasis>
Maybe Hinton had originally planned to name this species “pusillus”.
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23FD69FDD726AD" blockId="82.[189,564,723,749]" box="[189,564,723,749]" pageId="82" pageNumber="83">
<heading id="D0D5567BDA5AE71DFF23FD69FDD726AD" bold="true" box="[189,564,723,749]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
<emphasis id="B9563D05DA5AE71DFF23FD69FDD726AD" bold="true" box="[189,564,723,749]" pageId="82" pageNumber="83">Additional material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23FCB8FB32275D" blockId="82.[189,1233,770,797]" box="[189,1233,770,797]" pageId="82" pageNumber="83">
<materialsCitation id="3B4AEB4ADA5AE71DFF23FCB8FB2E275D" box="[189,1229,770,797]" collectionCode="NHMUK" collectorName="Gombak R. Malaya &amp; J. E. Bishop" country="Malaysia" location="Selangor State" pageId="82" pageNumber="83" specimenCount="1" specimenCount-male="1" stateProvince="Selangor">
<collectingCountry id="F335A187DA5AE71DFF23FCB8FEB7275C" box="[189,340,770,796]" name="Malaysia" pageId="82" pageNumber="83">MALAYSIA</collectingCountry>
<emphasis id="B9563D05DA5AE71DFEEFFCB8FDD4275C" bold="true" box="[369,567,770,796]" pageId="82" pageNumber="83">
<collectingRegion id="49E62FF5DA5AE71DFEEFFCB8FDC5275C" box="[369,550,770,796]" country="Malaysia" name="Selangor" pageId="82" pageNumber="83">Selangor State</collectingRegion>
</emphasis>
<specimenCount id="9D242A9EDA5AE71DFDA1FCB8FD8D275D" box="[575,622,770,797]" count="1" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
; “
<collectorName id="26D784C1DA5AE71DFD1BFCB8FC8D275C" box="[645,878,770,797]" pageId="82" pageNumber="83">Gombak R. Malaya</collectorName>
1969
<collectorName id="26D784C1DA5AE71DFC29FCB9FBA7275C" box="[951,1092,770,797]" pageId="82" pageNumber="83">J.E. Bishop</collectorName>
”;
<collectionCode id="ED3379D2DA5AE71DFBC4FCB9FB2E275D" box="[1114,1229,771,797]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="82" pageNumber="83" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23FCFEFEBE271D" blockId="82.[189,349,835,862]" box="[189,349,835,862]" pageId="82" pageNumber="83">
<heading id="D0D5567BDA5AE71DFF23FCFEFEBE271D" bold="true" box="[189,349,835,862]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
<emphasis id="B9563D05DA5AE71DFF23FCFEFEBE271D" bold="true" box="[189,349,835,862]" pageId="82" pageNumber="83">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23FCC8FE0A27CD" blockId="82.[189,489,882,909]" box="[189,489,882,909]" pageId="82" pageNumber="83">
<collectingCountry id="F335A187DA5AE71DFF23FCC8FECF27CC" box="[189,300,882,908]" name="Malaysia" pageId="82" pageNumber="83">Malaysia</collectingCountry>
,
<collectingRegion id="49E62FF5DA5AE71DFEA9FCC8FE0527CD" box="[311,486,882,909]" country="Malaysia" name="Kuala Lumpur" pageId="82" pageNumber="83">Kuala Lumpur</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA5AE71DFF23FC09FE4E238D" pageId="82" pageNumber="83" type="description">
<paragraph id="8B9DE117DA5AE71DFF23FC09FE88278D" blockId="82.[189,363,947,973]" box="[189,363,947,973]" pageId="82" pageNumber="83">
<heading id="D0D5567BDA5AE71DFF23FC09FE88278D" bold="true" box="[189,363,947,973]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
<emphasis id="B9563D05DA5AE71DFF23FC09FE88278D" bold="true" box="[189,363,947,973]" pageId="82" pageNumber="83">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23FC58FB72205F" blockId="82.[189,1398,994,1056]" pageId="82" pageNumber="83">
Body elongate oval, moderately convex dorsally, TL:
<quantity id="4CDA4CF2DA5AE71DFCFDFC58FC1927BD" box="[867,1018,994,1021]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="82" pageNumber="83" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.62.8 mm</quantity>
. Colouration brown; head and pronotum darkened; legs, anterior pronotal margin, antennae, and mouthparts paler.
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23FBFDFE3B20AD" blockId="82.[189,1399,1095,1261]" pageId="82" pageNumber="83">Cranial surface covered with plastron, except for narrow strip along anterior clypeal margin and wedge-shaped area on vertex and frons reaching to approximately midlength of eyes; round setiferous punctures separated by 12 × puncture diameters, each smaller than eye facet. Labrum transverse, anterior margin slightly emarginate, exposed portion with small setiferous punctures; males with long setae extending beyond anterior margin.</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23FAAFFC482208" blockId="82.[189,1399,1301,1609]" pageId="82" pageNumber="83">Pronotum transverse, weakly convex, disc nearly flat, PW/PL 1.661.91; plastron present on triangular areas in antero-lateral angles; punctures approximately as wide as on vertex; lateral sides weakly curved before anterior angles. Prosternal process without plastron; lateral sides rounded; lateral portions slightly raised, each with cluster of long setae in male; median keel flat. Metaventrite flat; narrow lateral sides of metaventral process raised. Elytra widest in middle, moderately convex, EL/EW: 1.641.70; punctures scattered; plastron forming posteriorly expanding lateral band on each elytron extending to two-thirds of elytron width in dorsal aspect. Tibiae approximately as long as lateral pronotal side, weakly bent. Each tarsus approximately two-thirds as long as corresponding tibia; terminal tarsomere as long as preceding segments combined; claws strongly bent, not expanded in male.</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23F9CAFEFB2357" blockId="82.[189,1399,1648,1815]" pageId="82" pageNumber="83">
Ventrites covered with plastron except for middle of ventrites 12. Intercoxal process of ventrite 1 very feebly punctate in
<typeStatus id="54995FB5DA5AE71DFE42F929FDA522ED" box="[476,582,1683,1709]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
, without admedian keels. Ventrite 5 rounded at apex and with short longitudinal keel in female, truncate and without keel in male.Aedeagus (
<figureCitation id="1319FD92DA5AE71DFB99F90CFBBD2290" box="[1031,1118,1718,1744]" captionStart="Fig" captionStartId="81.[189,232,1909,1935]" captionTargetBox="[389,1148,269,1844]" captionTargetId="figure-25@81.[374,1161,265,1861]" captionTargetPageId="81" captionText="Fig. 38. Elmomorphus bryanti Hinton, 1935, male from Gombak River, Malaysia (BMNH), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773488" httpUri="https://zenodo.org/record/13773488/files/figure.png" pageId="82" pageNumber="83">Fig. 38</figureCitation>
): phallobase moderately long and slender, PhL/PrL: 1.36; parameres slightly bent ventrad, apices rounded; sclerotised fibula slender.
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23F884FDD42318" blockId="82.[189,567,1854,1880]" box="[189,567,1854,1880]" pageId="82" pageNumber="83">
<heading id="D0D5567BDA5AE71DFF23F884FDD42318" bold="true" box="[189,567,1854,1880]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
<emphasis id="B9563D05DA5AE71DFF23F884FDD42318" bold="true" box="[189,567,1854,1880]" pageId="82" pageNumber="83">Secondary sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA5AE71DFF23F8D7FE4E238D" blockId="82.[189,1399,1901,1997]" pageId="82" pageNumber="83">The single male specimen known possesses clusters of long setae on labrum and prosternal process. Ventrite 5 rounded at apex; with short subapical keel in females, truncate and without keel in the male specimen examined.</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA5BE71CFF23FEB2FEAF25F0" pageId="83" pageNumber="84" type="distribution">
<paragraph id="8B9DE117DA5BE71CFF23FEB2FEB52562" blockId="83.[189,342,264,290]" box="[189,342,264,290]" pageId="83" pageNumber="84">
<heading id="D0D5567BDA5BE71CFF23FEB2FEB52562" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="B9563D05DA5BE71CFF23FEB2FEB52562" bold="true" box="[189,342,264,290]" pageId="83" pageNumber="84">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA5BE71CFF23FE8EFEAF25F0" blockId="83.[187,1399,308,432]" pageId="83" pageNumber="84">
<collectingCountry id="F335A187DA5BE71CFF23FE8EFECA250E" box="[189,297,308,334]" name="Malaysia" pageId="83" pageNumber="84">Malaysia</collectingCountry>
(
<collectingRegion id="49E62FF5DA5BE71CFEA6FE8EFE04250F" box="[312,487,308,335]" country="Malaysia" name="Kuala Lumpur" pageId="83" pageNumber="84">Kuala Lumpur</collectingRegion>
,
<collectingRegion id="49E62FF5DA5BE71CFE6DFE8EFD83250E" box="[499,608,308,334]" country="Malaysia" name="Selangor" pageId="83" pageNumber="84">Selangor</collectingRegion>
),?
<collectingCountry id="F335A187DA5BE71CFD19FE8EFD0C250E" box="[647,751,308,334]" name="Thailand" pageId="83" pageNumber="84">Thailand</collectingCountry>
(checklist published by
<bibRefCitation id="EFB39CE6DA5BE71CFB90FE8EFAF1250E" author="Shepard W. D. &amp; Sites R. W." box="[1038,1298,308,334]" pageId="83" pageNumber="84" pagination="89 - 126" refId="ref100782" refString="Shepard W. D. &amp; Sites R. W. 2016. Aquatic beetles of the families Dryopidae and Elmidae (Insecta: Coleoptera: Byrrhoidea) of Thailand: Annotated list and illustrated key to genera. Natural History Bulletin of the Siam Society 61 (2): 89 - 126." type="journal article" year="2016">Shepard &amp; Sites 2016</bibRefCitation>
, without any further locality details; identification needs confirmation; it cannot be excluded that the specimens recorded from
<collectingCountry id="F335A187DA5BE71CFEF9FECFFE2C25CF" box="[359,463,373,399]" name="Thailand" pageId="83" pageNumber="84">Thailand</collectingCountry>
actually belong to the closely related
<taxonomicName id="4C229A94DA5BE71CFC1EFECCFBAA25CF" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[896,1097,374,400]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="paramontanus" status="sp. nov.">
<emphasis id="B9563D05DA5BE71CFC1EFECCFC7125CF" box="[896,914,374,399]" italics="true" pageId="83" pageNumber="84">E</emphasis>
.
<emphasis id="B9563D05DA5BE71CFC3EFECCFBAA25CF" box="[928,1097,374,399]" italics="true" pageId="83" pageNumber="84">paramontanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA5BE71CFBD1FECCFB4B25D0" box="[1103,1192,374,400]" pageId="83" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
described herein) (
<figureCitation id="1319FD92DA5BE71CFF5AFE2CFEA325F0" box="[196,320,406,432]" captionStart="Fig" captionStartId="218.[189,232,1871,1897]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-40@218.[189,1399,265,1830]" captionTargetPageId="218" captionText="Fig. 110. Distribution maps.A. Elmomorphus corpulentus sp. nov.B. E. ellipticus sp. nov.C. E. mazzoldii sp. nov. D. E. bryanti Hinton, 1935. E. E. prosternalis Hinton, 1935. F. E. montanus (Grouvelle, 1913)." figureDoi="http://doi.org/10.5281/zenodo.13773714" httpUri="https://zenodo.org/record/13773714/files/figure.png" pageId="83" pageNumber="84">Fig. 110D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>