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<document id="C6EB04ED4D84AA7631BE68C4A8F3E7BA" ID-CLB-Dataset="5815" ID-DOI="10.11646/zootaxa.3779.3.2" ID-GBIF-Dataset="2055f347-d478-4d50-aad0-7e78c4b3b9c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4910403" ID-ZooBank="23584B50-EFAC-4BCE-A20E-9B09C22529B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623118476769" checkinUser="felipe" docAuthor="Deein, Gridsada, Tangjitjaroen, Weerapongse &amp; Page, Lawrence M." docDate="2014" docId="038987E7FFCDDB2BC0EF59BE5C63A221" docLanguage="en" docName="zt03779p352.pdf" docOrigin="Zootaxa 3779 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidocephalus pallens" docType="treatment" docVersion="8" lastPageNumber="348" masterDocId="FFB0FF9FFFCBDB2CC0785C1C5D08A042" masterDocTitle="A revision of the spirit loaches, genus Lepidocephalus (Cypriniformes, Cobitidae)" masterLastPageNumber="352" masterPageNumber="341" pageNumber="347" updateTime="1699003940845" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6E204D84AD07463C8D4BA24A20C62E42">A revision of the spirit loaches, genus Lepidocephalus (Cypriniformes, Cobitidae)</mods:title>
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<mods:namePart id="6C873F319B23CEFD39741AE6CB0F0273">Deein, Gridsada</mods:namePart>
<mods:affiliation id="4C55D468A7984EBB6DE074967EE3CDE0">Inland Fisheries Research and Development Bureau, Department of Fisheries, NIFI Building, Kasetklang, Chatuchak, Bangkok 10900 Thailand. E-mail: dgridsada @ hotmail. com</mods:affiliation>
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<mods:namePart id="50832990B1981F419D8E2314ED279C91">Tangjitjaroen, Weerapongse</mods:namePart>
<mods:affiliation id="BF1360735370299CBDEADE6076CDF80C">Faculty of Veterinary Medicine, Chiang Mai University, Chiang Mai, 50100, Thailand. E-mail: weerapongse @ yahoo. com</mods:affiliation>
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<mods:namePart id="6E925C899BEB5449D96E893C72F51E1A">Page, Lawrence M.</mods:namePart>
<mods:affiliation id="31F71891C461132A562DE02441A1FE44">Florida Museum of Natural History, University of Florida, Gainesville, FL 32611 - 7800 USA. E-mail: lpage 1 @ ufl. edu</mods:affiliation>
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<mods:title id="5E162A71617F3CBEA27FBCD95C72BBD7">Zootaxa</mods:title>
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<mods:date id="F3CAF4377CD4D710848E3AB2584E9C3C">2014</mods:date>
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<treatment id="038987E7FFCDDB2BC0EF59BE5C63A221" ID-DOI="http://doi.org/10.5281/zenodo.4910386" ID-GBIF-Taxon="183237224" ID-Zenodo-Dep="4910386" LSID="urn:lsid:plazi:treatment:038987E7FFCDDB2BC0EF59BE5C63A221" httpUri="http://treatment.plazi.org/id/038987E7FFCDDB2BC0EF59BE5C63A221" lastPageId="7" lastPageNumber="348" pageId="6" pageNumber="347">
<subSubSection id="C33A657AFFCDDB2AC0EF59BE5C63A59C" pageId="6" pageNumber="347" type="nomenclature">
<paragraph id="8B9F36F1FFCDDB2AC0EF59BE5F71A5FF" blockId="6.[151,633,1442,1538]" box="[151,633,1442,1469]" pageId="6" pageNumber="347">
<heading id="D0D7819DFFCDDB2AC0EF59BE5F71A5FF" bold="true" box="[151,633,1442,1469]" fontSize="11" level="1" pageId="6" pageNumber="347" reason="1">
<taxonomicName id="4C204D72FFCDDB2AC0EF59BE5F71A5FF" ID-CoL="6PB87" authority="(Vaillant, 1902)" baseAuthorityName="Vaillant" baseAuthorityYear="1902" box="[151,633,1442,1469]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="347" phylum="Chordata" rank="species" species="pallens">
<emphasis id="B954EAE3FFCDDB2AC0EF59BE5F71A5FF" bold="true" box="[151,633,1442,1469]" pageId="6" pageNumber="347">
<emphasis id="B954EAE3FFCDDB2AC0EF59BE5CBAA5FE" bold="true" box="[151,434,1442,1468]" italics="true" pageId="6" pageNumber="347">Lepidocephalus pallens</emphasis>
(
<bibRefCitation id="EFB14B00FFCDDB2AC1BA59BF5F79A5FF" author="Vaillant, L. L." box="[450,625,1443,1469]" pageId="6" pageNumber="347" pagination="1 - 166" refId="ref8282" refString="Vaillant, L. L. (1902) Resultats zoologiques de l'expedition scientifique neerlandaise au Borneo central, poissons. Notes from the Leyden Museum, 24 (1), 1 - 166." type="journal article" year="1902">Vaillant, 1902</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9F36F1FFCDDB2AC0EF59D95C63A59C" blockId="6.[151,633,1442,1538]" box="[151,363,1477,1502]" pageId="6" pageNumber="347">Pallid Spirit Loach</paragraph>
</subSubSection>
<subSubSection id="C33A657AFFCDDB2AC0EF59F55DF5A640" box="[151,253,1513,1538]" pageId="6" pageNumber="347" type="description">
<paragraph id="8B9F36F1FFCDDB2AC0EF59F55DF5A640" blockId="6.[151,633,1442,1538]" box="[151,253,1513,1538]" pageId="6" pageNumber="347">
(
<figureCitation id="131B2A74FFCDDB2AC0E759F55DFCA640" box="[159,244,1513,1538]" captionStart="FIGURE 1" captionStartId="2.[151,249,1690,1712]" captionTargetBox="[153,1434,151,1673]" captionTargetId="figure-188@2.[153,1438,141,1673]" captionTargetPageId="2" captionText="FIGURE 1. Species of Lepidocephalus. (A) L. nanensis, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B) L. macrochir, syntype, BMNH 1866.5.2.55, 77.7 mm SL, Sumatra or Java; (C) L. pallens, holotype, RMNH 7783, 41.9 mm SL; (D) L. pahangensis, holotype, ZRC 490, 35.8 mm SL; (E) L. spectrum, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos A, C, D and E by Z. Randall; B by M. Sabaj Perez." figureDoi="http://doi.org/10.5281/zenodo.4910405" httpUri="https://zenodo.org/record/4910405/files/figure.png" pageId="6" pageNumber="347">Fig. 1C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33A657AFFCDDB2AC0EF5A2D5CB7A624" pageId="6" pageNumber="347" type="reference_group">
<paragraph id="8B9F36F1FFCDDB2AC0EF5A2D5CB7A624" blockId="6.[151,1436,1585,1638]" pageId="6" pageNumber="347">
<treatmentCitationGroup id="AB3011DFFFCDDB2AC0EF5A2D5CB7A624" pageId="6" pageNumber="347">
<treatmentCitation id="0A8110E0FFCDDB2AC0EF5A2D5F8BA605" author="Vaillant, L. L." box="[151,643,1585,1607]" page="153" pageId="6" pageNumber="347" year="1902">
<taxonomicName id="4C204D72FFCDDB2AC0EF5A2D5F8BA605" ID-CoL="6PB87" authority="Vaillant, 1902: 153" authorityName="Vaillant" authorityPageNumber="153" authorityYear="1902" box="[151,643,1585,1607]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalichthys" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="347" phylum="Chordata" rank="species" species="pallens">
<emphasis id="B954EAE3FFCDDB2AC0EF5A2D5CB2A605" box="[151,442,1585,1607]" italics="true" pageId="6" pageNumber="347">Lepidocephalichthys pallens</emphasis>
<materialsCitation id="3B483CACFFCDDB2AC1BB5A2D5F8BA605" ID-GBIF-Occurrence="3313289307" box="[451,643,1585,1607]" collectionCode="RMNH" location="Kapuas River" pageId="6" pageNumber="347" specimenCode="RMNH 7783" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EFB14B00FFCDDB2AC1BB5A2D5F8BA605" author="Vaillant, L. L." box="[451,643,1585,1607]" pageId="6" pageNumber="347" pagination="1 - 166" refId="ref8282" refString="Vaillant, L. L. (1902) Resultats zoologiques de l'expedition scientifique neerlandaise au Borneo central, poissons. Notes from the Leyden Museum, 24 (1), 1 - 166." type="journal article" year="1902">Vaillant, 1902:153</bibRefCitation>
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</treatmentCitation>
. Type locality &quot;Bords du Kapoeas (Sintang?)&quot; [
<location id="8EFF602AFFCDDB2AC4095A2E5808A605" LSID="urn:lsid:plazi:treatment:038987E7FFCDDB2BC0EF59BE5C63A221:8EFF602AFFCDDB2AC4095A2E5808A605" box="[1137,1280,1585,1607]" name="Kapuas River" pageId="6" pageNumber="347">Kapuas River</location>
(at Sintang?)].
<typeStatus id="549B8853FFCDDB2AC0BF5A4C5C21A624" box="[199,297,1616,1638]" pageId="6" pageNumber="347" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DB869E8AFFCDDB2AC14A5A4D5CB2A624" box="[306,442,1616,1638]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="347" type="Museum">RMNH 7783</specimenCode>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33A657AFFCDDB2AC0EF5A8F5EC2A77E" pageId="6" pageNumber="347" type="diagnosis">
<paragraph id="8B9F36F1FFCDDB2AC0EF5A8F5EC2A77E" blockId="6.[151,1436,1683,2031]" pageId="6" pageNumber="347">
<emphasis id="B954EAE3FFCDDB2AC0EF5A8F5C19A6E8" box="[151,273,1683,1708]" inLineHeading="true" pageId="6" pageNumber="347" reason="1">
<emphasis id="B954EAE3FFCDDB2AC0EF5A8F5C02A6EE" bold="true" box="[151,266,1683,1708]" pageId="6" pageNumber="347">Diagnosis</emphasis>
.
</emphasis>
A species of
<taxonomicName id="4C204D72FFCDDB2AC1C85A8F5F69A6E9" authorityName="Bleeker" authorityYear="1858" box="[432,609,1683,1707]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="347" phylum="Chordata" rank="genus">
<emphasis id="B954EAE3FFCDDB2AC1C85A8F5F69A6E9" box="[432,609,1683,1707]" italics="true" pageId="6" pageNumber="347">Lepidocephalus</emphasis>
</taxonomicName>
that is readily distinguishable (
<figureCitation id="131B2A74FFCDDB2AC3B25A8F5906A6EE" box="[970,1038,1683,1708]" captionStart="FIGURE 1" captionStartId="2.[151,249,1690,1712]" captionTargetBox="[153,1434,151,1673]" captionTargetId="figure-188@2.[153,1438,141,1673]" captionTargetPageId="2" captionText="FIGURE 1. Species of Lepidocephalus. (A) L. nanensis, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B) L. macrochir, syntype, BMNH 1866.5.2.55, 77.7 mm SL, Sumatra or Java; (C) L. pallens, holotype, RMNH 7783, 41.9 mm SL; (D) L. pahangensis, holotype, ZRC 490, 35.8 mm SL; (E) L. spectrum, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos A, C, D and E by Z. Randall; B by M. Sabaj Perez." figureDoi="http://doi.org/10.5281/zenodo.4910405" httpUri="https://zenodo.org/record/4910405/files/figure.png" pageId="6" pageNumber="347">Fig. 1</figureCitation>
,
<tableCitation id="C6A2034AFFCDDB2AC4665A8F5979A6EE" box="[1054,1137,1683,1708]" captionStart="TABLE 1" captionStartId="9.[151,241,476,499]" captionTargetPageId="9" captionText="TABLE 1. Characteristics that vary among species of Lepidocephalus. Measurements are in mm or percentages of standard length (SL). Range is followed by average in parentheses." httpUri="http://table.plazi.org/id/DF5F6679FFC2DB25C0EF5DC05E6AA251" pageId="6" pageNumber="347" tableUuid="DF5F6679FFC2DB25C0EF5DC05E6AA251">Table 1</tableCitation>
) from all other species of
<taxonomicName id="4C204D72FFCDDB2AC0EF5AAB5C40A68D" authorityName="Bleeker" authorityYear="1858" box="[151,328,1719,1743]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="347" phylum="Chordata" rank="genus">
<emphasis id="B954EAE3FFCDDB2AC0EF5AAB5C40A68D" box="[151,328,1719,1743]" italics="true" pageId="6" pageNumber="347">Lepidocephalus</emphasis>
</taxonomicName>
by having the dorsal-fin origin over the pelvic-fin origin (vs. dorsal-fin origin behind the pelvic-fin origin), and a concomitantly shorter predorsal length (59.9 vs. 62.471.3% SL).
<taxonomicName id="4C204D72FFCDDB2AC4425AC7584EA6B1" baseAuthorityName="Vaillant" baseAuthorityYear="1902" box="[1082,1350,1755,1779]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="347" phylum="Chordata" rank="species" species="pallens">
<emphasis id="B954EAE3FFCDDB2AC4425AC7584EA6B1" box="[1082,1350,1755,1779]" italics="true" pageId="6" pageNumber="347">Lepidocephalus pallens</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C204D72FFCDDB2AC15C5B1C5CC7A755" authorityName=", de Beaufort" authorityYear="1933" box="[292,463,1791,1815]" family="Cobitidae" genus="Acanthophthalmus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="347" phylum="Chordata" rank="species" species="pahangensis">
<emphasis id="B954EAE3FFCDDB2AC15C5B1C5CC7A755" box="[292,463,1791,1815]" italics="true" pageId="6" pageNumber="347">L. pahangensis</emphasis>
</taxonomicName>
in having scales on top of the head, and from
<taxonomicName id="4C204D72FFCDDB2AC3A15B1C5956A755" authorityName="Roberts" authorityYear="1989" box="[985,1118,1792,1815]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="347" phylum="Chordata" rank="species" species="spectrum">
<emphasis id="B954EAE3FFCDDB2AC3A15B1C5956A755" box="[985,1118,1792,1815]" italics="true" pageId="6" pageNumber="347">L. spectrum</emphasis>
</taxonomicName>
by the presence of eyes and darker pigment on the body, and absence of tubules along the lateral line.
</paragraph>
</subSubSection>
<subSubSection id="C33A657AFFCDDB2BC0BF5B5A5E59A1ED" lastPageId="7" lastPageNumber="348" pageId="6" pageNumber="347" type="description">
<paragraph id="8B9F36F1FFCDDB2AC0BF5B5A5ECDA7E5" blockId="6.[151,1436,1683,2031]" pageId="6" pageNumber="347">
<emphasis id="B954EAE3FFCDDB2AC0BF5B5A5C50A71C" box="[199,344,1862,1887]" inLineHeading="true" pageId="6" pageNumber="347" reason="1">
<emphasis id="B954EAE3FFCDDB2AC0BF5B5A5C59A71D" bold="true" box="[199,337,1862,1887]" pageId="6" pageNumber="347">Description</emphasis>
.
</emphasis>
Body deep, slab-sided; greatest depth 17.4% SL. Head narrow, length 23.2% SL. Eye small, in dorsal half of head in shallow depression above bifid suborbital spine. Origin of dorsal fin over origin of pelvic fin. No axillary lobe on pelvic fin. Small fleshy lobe at origin of pectoral fin.
</paragraph>
<paragraph id="8B9F36F1FFCDDB2BC0BF5BAF5828A102" blockId="6.[151,1436,1683,2031]" lastBlockId="7.[151,1436,151,611]" lastPageId="7" lastPageNumber="348" pageId="6" pageNumber="347">
Scales on top of head, cheek and opercle; body completely covered with minute, partially embedded scales. Lateral line complete, ~200 scales along lateral line; no tubules along lateral line. Mouth horseshoe-shaped; upper lip without median indentation; lower lip with median indentation on lower edge. Two pairs of rostral barbels; inner pair reaching to corner of mouth, outer pair slightly longer and reaching slightly past corner of mouth; one pair of maxillary barbels, reaching approximately to vertical at posterior end of groove containing suborbital spine. Large flap on anterior nostril. Dorsal rays iii,8; pectoral rays 11; pelvic rays 7; anal rays ii,5; upper branched caudal rays 7; lower branched caudal rays 7; vertebrae 28 abdominal + 15 caudal = 43 total (
<bibRefCitation id="EFB14B00FFCCDB2BC42B5D3B581BA102" author="Roberts, T. R." box="[1107,1299,295,320]" pageId="7" pageNumber="348" pagination="1 - 210" refId="ref8105" refString="Roberts, T. R. (1989) The freshwater fishes of western Borneo (Kalimantan Barat, Indonesia). Memoirs of the California Academy of Sciences, 14, 1 - 210." type="journal article" year="1989">Roberts 1989:96</bibRefCitation>
).
</paragraph>
<paragraph id="8B9F36F1FFCCDB2BC0BF5D525874A1C9" blockId="7.[151,1436,151,611]" pageId="7" pageNumber="348">
Only specimen known is the
<typeStatus id="549B8853FFCCDB2BC2745D525F79A125" box="[524,625,334,359]" pageId="7" pageNumber="348" type="holotype">holotype</typeStatus>
, which appears to be a female. The pectoral fin is falcate with the 2
<superScript id="7C559BB9FFCCDB2BC51A5D575879A11B" attach="left" box="[1378,1393,331,345]" fontSize="6" pageId="7" pageNumber="348">nd</superScript>
ray longer but not thicker than other rays (20.0% SL). Pelvic fin small, but larger than in other species at 11.5% SL.
</paragraph>
<paragraph id="8B9F36F1FFCCDB2BC0BF5D8A5E59A1ED" blockId="7.[151,1436,151,611]" box="[199,849,406,431]" pageId="7" pageNumber="348">
<emphasis id="B954EAE3FFCCDB2BC0BF5D8A5C92A1ED" bold="true" box="[199,410,406,431]" inLineHeading="true" pageId="7" pageNumber="348" reason="1">Coloration in life.</emphasis>
Described as pallid by
<bibRefCitation id="EFB14B00FFCCDB2BC2D95D8A5E45A1ED" author="Vaillant, L. L." box="[673,845,406,431]" pageId="7" pageNumber="348" pagination="1 - 166" refId="ref8282" refString="Vaillant, L. L. (1902) Resultats zoologiques de l'expedition scientifique neerlandaise au Borneo central, poissons. Notes from the Leyden Museum, 24 (1), 1 - 166." type="journal article" year="1902">Vaillant (1902)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A657AFFCCDB2BC0BF5DA65815A191" box="[199,1309,442,467]" pageId="7" pageNumber="348" type="distribution">
<paragraph id="8B9F36F1FFCCDB2BC0BF5DA65815A191" blockId="7.[151,1436,151,611]" box="[199,1309,442,467]" pageId="7" pageNumber="348">
<emphasis id="B954EAE3FFCCDB2BC0BF5DA65C68A191" bold="true" box="[199,352,442,467]" inLineHeading="true" pageId="7" pageNumber="348" reason="1">Distribution.</emphasis>
Known only from the
<typeStatus id="549B8853FFCCDB2BC2185DA75F98A191" box="[608,656,443,467]" pageId="7" pageNumber="348">type</typeStatus>
locality, the Kapuas River presumably at Sintang (
<figureCitation id="131B2A74FFCCDB2BC4B25DA65818A191" box="[1226,1296,442,467]" captionStart="FIGURE 2" captionStartId="4.[151,249,1699,1721]" captionTargetBox="[153,1429,349,1673]" captionTargetId="figure-196@4.[151,1436,339,1677]" captionTargetPageId="4" captionText="FIGURE 2. Distribution of specimens of Lepidocephalus examined." figureDoi="http://doi.org/10.5281/zenodo.4910407" httpUri="https://zenodo.org/record/4910407/files/figure.png" pageId="7" pageNumber="348">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A657AFFCCDB2BC0BF5DC25C63A221" pageId="7" pageNumber="348" type="discussion">
<paragraph id="8B9F36F1FFCCDB2BC0BF5DC25C63A221" blockId="7.[151,1436,151,611]" pageId="7" pageNumber="348">
<emphasis id="B954EAE3FFCCDB2BC0BF5DC25C33A1B4" box="[199,315,478,503]" inLineHeading="true" pageId="7" pageNumber="348" reason="1">
<emphasis id="B954EAE3FFCCDB2BC0BF5DC25C3CA1B5" bold="true" box="[199,308,478,503]" pageId="7" pageNumber="348">Remarks</emphasis>
.
</emphasis>
<bibRefCitation id="EFB14B00FFCCDB2BC13A5DC25CE4A1B5" author="Roberts, T. R." box="[322,492,478,503]" pageId="7" pageNumber="348" pagination="1 - 210" refId="ref8105" refString="Roberts, T. R. (1989) The freshwater fishes of western Borneo (Kalimantan Barat, Indonesia). Memoirs of the California Academy of Sciences, 14, 1 - 210." type="journal article" year="1989">Roberts (1989)</bibRefCitation>
treated
<taxonomicName id="4C204D72FFCCDB2BC23F5DFC5FBEA1B5" baseAuthorityName="Vaillant" baseAuthorityYear="1902" box="[583,694,479,503]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="pallens">
<emphasis id="B954EAE3FFCCDB2BC23F5DFC5FBEA1B5" box="[583,694,479,503]" italics="true" pageId="7" pageNumber="348">L. pallens</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="4C204D72FFCCDB2BC3065DFC5907A1B5" baseAuthorityName="Bleeker" baseAuthorityYear="1854" box="[894,1039,479,503]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="macrochir">
<emphasis id="B954EAE3FFCCDB2BC3065DFC5907A1B5" box="[894,1039,479,503]" italics="true" pageId="7" pageNumber="348">L. macrochir</emphasis>
</taxonomicName>
. However, as noted above, the two species are easily distinguished by the position of the dorsal-fin origin relative to that of the pelvic-fin origin (
<figureCitation id="131B2A74FFCCDB2BC5175E1E5DACA27D" captionStart="FIGURE 1" captionStartId="2.[151,249,1690,1712]" captionTargetBox="[153,1434,151,1673]" captionTargetId="figure-188@2.[153,1438,141,1673]" captionTargetPageId="2" captionText="FIGURE 1. Species of Lepidocephalus. (A) L. nanensis, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B) L. macrochir, syntype, BMNH 1866.5.2.55, 77.7 mm SL, Sumatra or Java; (C) L. pallens, holotype, RMNH 7783, 41.9 mm SL; (D) L. pahangensis, holotype, ZRC 490, 35.8 mm SL; (E) L. spectrum, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos A, C, D and E by Z. Randall; B by M. Sabaj Perez." figureDoi="http://doi.org/10.5281/zenodo.4910405" httpUri="https://zenodo.org/record/4910405/files/figure.png" pageId="7" pageNumber="348">Fig. 1</figureCitation>
).
<taxonomicName id="4C204D72FFCCDB2BC0C65E3B5CC1A27D" baseAuthorityName="Vaillant" baseAuthorityYear="1902" box="[190,457,551,575]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="pallens">
<emphasis id="B954EAE3FFCCDB2BC0C65E3B5CC1A27D" box="[190,457,551,575]" italics="true" pageId="7" pageNumber="348">Lepidocephalus pallens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C204D72FFCCDB2BC27B5E345F81A27D" authorityName="Roberts" authorityYear="1989" box="[515,649,552,575]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="spectrum">
<emphasis id="B954EAE3FFCCDB2BC27B5E345F81A27D" box="[515,649,552,575]" italics="true" pageId="7" pageNumber="348">L. spectrum</emphasis>
</taxonomicName>
were both described from the Kapuas River in Borneo, and possibly occur syntopically.
</paragraph>
</subSubSection>
</treatment>
</document>