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<document id="DD36C91E0B113B4C6D2C60BB91099B92" ID-CLB-Dataset="37322" ID-DOI="10.11646/zootaxa.4211.1.1" ID-GBIF-Dataset="7af458af-d56d-44e3-a6f7-af97868175d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="272750" ID-ZooBank="6B86C6BA-6AFE-4AAD-870D-04794C138D47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484386885843" checkinUser="plazi" docAuthor="Antić, Dragan Ž. &amp; Makarov, Slobodan E." docDate="2016" docId="03B7878CFFA7CD4DFF4CE8E2B352EBCD" docLanguage="en" docName="zootaxa.4211.1.1.pdf.imf" docOrigin="Zootaxa 4211 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Enghoffiella insolita" docType="treatment" docVersion="8" lastPageNumber="116" masterDocId="FF8EFFF4FFD7CD3EFFDBE875B173E949" masterDocTitle="The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)" masterLastPageNumber="205" masterPageNumber="1" pageNumber="113" updateTime="1698435542289" updateUser="ExternalLinkService">
<mods:mods id="0EBB3837A873E84BAE7947DEE0EBC79F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DC0013EC4F30AD5B5DF19FCB3A8CA6F2">
<mods:title id="AF0DE36508EF5DBE370357472A9B98F5">The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)</mods:title>
</mods:titleInfo>
<mods:name id="C5E80C66ADD954B107BF30583E8CA424" type="personal">
<mods:role id="6A1F5D7EBA8403C7E93563E66CC4CA8D">
<mods:roleTerm id="CF5981E00DA5EFAB0B652EC116061341">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0A267D33EEFD6B94272BC0FA57BD049B">Antić, Dragan Ž.</mods:namePart>
</mods:name>
<mods:name id="C47E0CDAAF525510E09F7BB0F8FC1099" type="personal">
<mods:role id="D19E3386A76AEAF164C904C5D664A1D0">
<mods:roleTerm id="873743185A4D9B8B55BC7B21534C0CA1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CE22701D05584F87A85C016A00C6A5B5">Makarov, Slobodan E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="310D596DF6D847F46F7AE8605D70A3FB">text</mods:typeOfResource>
<mods:relatedItem id="CBC8F5BF9A311F6CAFF9FD454B2B4198" type="host">
<mods:titleInfo id="E233083146DCDF67422BE3FA9A19C48F">
<mods:title id="11F9941085B5640E74A5EFBCD28AB512">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8681C777AE597A3B9D21A0A739B38CB9">
<mods:date id="52D97B7774A23C6A0442C776E75B971F">2016</mods:date>
<mods:detail id="197AB7EE3C7934C5A153541AA4CF918D" type="volume">
<mods:number id="7629A03ED07FB39EEB3BFF1637D3D4C9">4211</mods:number>
</mods:detail>
<mods:detail id="3253CD7D66539AF09C5750D7AE677927" type="issue">
<mods:number id="28AF61E5B18098C9A6F6055D3A7EF268">1</mods:number>
</mods:detail>
<mods:extent id="E0FD1089243E6BA9C08F6DF02F14161B" unit="page">
<mods:start id="5928EBA5639F5544DA0B3DC7BA415C78">1</mods:start>
<mods:end id="25370AFC62DB512493A621BD0A2070F1">205</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0BB7032F8FC9FE2B3B439E9BA4C2D4FB">journal article</mods:classification>
<mods:identifier id="88A0B5F5F8B86AEA2C49F480DD786CB5" type="CLB-Dataset">37322</mods:identifier>
<mods:identifier id="2CD93A995A3B1E3F784E316E7C2B4A7C" type="DOI">10.11646/zootaxa.4211.1.1</mods:identifier>
<mods:identifier id="A4D07D33919CC6C9982C300ADD8EF39D" type="GBIF-Dataset">7af458af-d56d-44e3-a6f7-af97868175d9</mods:identifier>
<mods:identifier id="4CE1BA7B06FCF513854EFB190D5FA599" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9E712D2D406F258B034BF1459E13D8C5" type="Zenodo-Dep">272750</mods:identifier>
<mods:identifier id="6CF7194FF57C289A121810274B845A2C" type="ZooBank">6B86C6BA-6AFE-4AAD-870D-04794C138D47</mods:identifier>
</mods:mods>
<treatment id="03B7878CFFA7CD4DFF4CE8E2B352EBCD" ID-DOI="http://doi.org/10.5281/zenodo.6074439" ID-GBIF-Taxon="127419147" ID-Zenodo-Dep="6074439" LSID="urn:lsid:plazi:treatment:03B7878CFFA7CD4DFF4CE8E2B352EBCD" httpUri="http://treatment.plazi.org/id/03B7878CFFA7CD4DFF4CE8E2B352EBCD" lastPageId="115" lastPageNumber="116" pageId="112" pageNumber="113">
<subSubSection id="C3046511FFA7CD4EFF4CE8E2B066E99A" pageId="112" pageNumber="113" type="nomenclature">
<paragraph id="8BA1369AFFA7CD4EFF4CE8E2B33EE9FB" blockId="112.[151,589,151,211]" box="[151,589,151,178]" pageId="112" pageNumber="113">
<heading id="D0E981F6FFA7CD4EFF4CE8E2B33EE9FB" bold="true" box="[151,589,151,178]" fontSize="11" level="1" pageId="112" pageNumber="113" reason="1">
<emphasis id="B96AEA88FFA7CD4EFF4CE8E2B33EE9FB" bold="true" box="[151,589,151,178]" pageId="112" pageNumber="113">
<taxonomicName id="4C1E4D19FFA7CD4EFF4CE8E2B0E3E9F8" ID-CoL="848MB" box="[151,400,151,177]" class="Diplopoda" family="Anthroleucosomatidae" genus="Enghoffiella" kingdom="Animalia" order="Chordeumatida" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="B96AEA88FFA7CD4EFF4CE8E2B0E3E9F8" bold="true" box="[151,400,151,177]" italics="true" pageId="112" pageNumber="113">Enghoffiella insolita</emphasis>
</taxonomicName>
gen. et sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1369AFFA7CD4EFF4CE8CFB066E99A" blockId="112.[151,589,151,211]" box="[151,277,186,211]" pageId="112" pageNumber="113">
<figureCitation id="13252A1FFFA7CD4EFF4CE8CFB066E99A" box="[151,277,186,211]" captionStart-0="FIGURE 92" captionStart-1="FIGURE 93" captionStart-2="FIGURE 94" captionStartId-0="112.[151,250,1948,1970]" captionStartId-1="113.[151,250,1828,1850]" captionStartId-2="114.[151,250,1701,1723]" captionTargetBox-0="[188,1398,1020,1927]" captionTargetBox-1="[188,1398,207,1798]" captionTargetBox-2="[212,1381,188,1627]" captionTargetId-0="figure@112.[188,1398,1020,1927]" captionTargetId-1="figure@113.[188,1398,207,1798]" captionTargetId-2="figure@114.[188,1398,177,1679]" captionTargetPageId-0="112" captionTargetPageId-1="113" captionTargetPageId-2="114" captionText-0="FIGURE 92. Enghoffiella insolita gen. et sp. nov., paratype female habitus. Scale line = 1 mm." captionText-1="FIGURE 93. Enghoffiella insolita gen. et sp. nov., holotype male. A. Leg 3. B. Leg 4. C. Leg 5. D. Leg 7. E. Leg 10. Scale lines = 0.2 mm." captionText-2="FIGURE 94. Enghoffiella insolita gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, lateral view. E. Posterior gonopods, posterior view. c = coxite; l = lamella; p 1, p 2, p 3 and p 4 = processes on anterior gonopods; t = telopodite; ap = anterior part of posterior gonopods; cp = coxal process; cv = coxal vesicle; hl = hairy lever; pp = posterior part of posterior gonopods; sl = sternal lamella; pmp = posteromedial part. Scale line = 0.3 mm." httpUri-0="https://zenodo.org/record/272841/files/figure.png" httpUri-1="https://zenodo.org/record/272842/files/figure.png" httpUri-2="https://zenodo.org/record/272843/files/figure.png" pageId="112" pageNumber="113">Figs 92–94</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FFA7CD4EFF4CE977B0ABE852" box="[151,472,258,283]" pageId="112" pageNumber="113" type="diagnosis">
<paragraph id="8BA1369AFFA7CD4EFF4CE977B0ABE852" blockId="112.[151,1437,258,966]" box="[151,472,258,283]" pageId="112" pageNumber="113">
<emphasis id="B96AEA88FFA7CD4EFF4CE977B062E852" bold="true" box="[151,273,258,283]" pageId="112" pageNumber="113">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FFA7CD4EFF1CE953B296E876" box="[199,997,294,319]" pageId="112" pageNumber="113" type="etymology">
<paragraph id="8BA1369AFFA7CD4EFF1CE953B296E876" blockId="112.[151,1437,258,966]" box="[199,997,294,319]" pageId="112" pageNumber="113">
<emphasis id="B96AEA88FFA7CD4EFF1CE953B03DE876" bold="true" box="[199,334,294,319]" pageId="112" pageNumber="113">Etymology.</emphasis>
An adjective; to emphasize the unusual anterior gonopods.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FFA7CD4EFF1CE93FB38BEB5E" pageId="112" pageNumber="113" type="materials_examined">
<paragraph id="8BA1369AFFA7CD4EFF1CE93FB36CE8E2" blockId="112.[151,1437,258,966]" pageId="112" pageNumber="113">
<materialsCitation id="3B763CC7FFA7CD4EFF1CE93FB36CE8E2" ID-GBIF-Occurrence="1424567114" collectingDate="1986-07-18" collectionCode="ZMUM" collectorName="S. Golovatch" country="Russia" elevation="750" location="Argun Valley" municipality="Material" pageId="112" pageNumber="113" specimenCount="26" specimenCount-female="3" specimenCount-juvenile="22" specimenCount-male="1" stateProvince="Chechnya" typeStatus="holotype">
<emphasis id="B96AEA88FFA7CD4EFF1CE93FB0FEE82A" bold="true" box="[199,397,330,355]" pageId="112" pageNumber="113">
<collectingMunicipality id="6BC5ACE0FFA7CD4EFF1CE93FB05CE82A" box="[199,303,330,355]" pageId="112" pageNumber="113">Material</collectingMunicipality>
studied
</emphasis>
(total:
<specimenCount id="9D18FD13FFA7CD4EFE05E93EB35FE82A" box="[478,556,330,355]" pageId="112" pageNumber="113" type="male">1 male</specimenCount>
,
<specimenCount id="9D18FD13FFA7CD4EFDE3E93EB3D4E82A" box="[568,679,330,355]" pageId="112" pageNumber="113" type="female">3 females</specimenCount>
,
<specimenCount id="9D18FD13FFA7CD4EFD69E93EB24CE82A" box="[690,831,330,355]" pageId="112" pageNumber="113" type="juvenile">22 juveniles</specimenCount>
).
<emphasis id="B96AEA88FFA7CD4EFC8AE93FB2B6E82A" bold="true" box="[849,965,330,355]" pageId="112" pageNumber="113">
<typeStatus id="54A58838FFA7CD4EFC8AE93FB2B3E82A" box="[849,960,330,355]" pageId="112" pageNumber="113" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F309760AFFA7CD4EFC17E93EB547E82A" box="[972,1076,331,355]" name="Russia" pageId="112" pageNumber="113">RUSSIA</collectingCountry>
: male,
<collectingRegion id="49DAF878FFA7CD4EFB58E93FB585E82A" box="[1155,1270,330,355]" country="Russia" name="Chechnya" pageId="112" pageNumber="113">Chechnya</collectingRegion>
,
<location id="8EC16041FFA7CD4EFADAE93EB4EBE82A" LSID="urn:lsid:plazi:treatment:03B7878CFFA7CD4DFF4CE8E2B352EBCD:8EC16041FFA7CD4EFADAE93EB4EBE82A" box="[1281,1432,330,355]" country="Russia" municipality="Material" name="Argun Valley" pageId="112" pageNumber="113" stateProvince="Chechnya">Argun Valley</location>
,
<locationDeviation id="9A132FB8FFA7CD4EFF4CE91BB0E7E8CE" box="[151,404,366,391]" pageId="112" pageNumber="113">
<quantity id="4CE69B7FFFA7CD4EFF4CE91BB1A3E8CE" box="[151,208,366,391]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="112" pageNumber="113" unit="km" value="5.0">5 km</quantity>
N of Sovetskoye
</locationDeviation>
,
<taxonomicName id="4C1E4D19FFA7CD4EFE44E91BB08BE8CE" box="[415,504,366,391]" class="Magnoliopsida" family="Betulaceae" genus="Corylus" kingdom="Plantae" order="Fagales" pageId="112" pageNumber="113" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FFA7CD4EFE44E91BB08BE8CE" box="[415,504,366,391]" italics="true" pageId="112" pageNumber="113">Corylus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D19FFA7CD4EFDDEE91AB338E8CF" box="[517,587,367,390]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="112" pageNumber="113" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FFA7CD4EFDDEE91AB338E8CF" box="[517,587,367,390]" italics="true" pageId="112" pageNumber="113">Fagus</emphasis>
</taxonomicName>
,
<emphasis id="B96AEA88FFA7CD4EFD81E91AB3BAE8CF" box="[602,713,367,390]" italics="true" pageId="112" pageNumber="113">
<taxonomicName id="4C1E4D19FFA7CD4EFD81E91AB3B6E8CF" box="[602,709,367,390]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="112" pageNumber="113" phylum="Tracheophyta" rank="genus">Carpinus</taxonomicName>
,
</emphasis>
etc. forest,
<elevation id="0033D1A9FFA7CD4EFC97E91BB2E0E8CF" box="[844,915,366,391]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="112" pageNumber="113" unit="m" value="750.0">
<quantity id="4CE69B7FFFA7CD4EFC97E91BB2E0E8CF" box="[844,915,366,391]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="112" pageNumber="113" unit="m" value="750.0">750 m</quantity>
</elevation>
asl, litter, under stones and bark,
<collectingDate id="EFE4E9B2FFA7CD4EFAD0E91AB4EBE8CE" box="[1291,1432,366,391]" pageId="112" pageNumber="113" value="1986-07-18">
<date id="FFA0105AFFA7CD4EFAD0E91AB4EBE8CE" box="[1291,1432,366,391]" pageId="112" pageNumber="113" value="1986-07-18">18 Jul. 1986</date>
</collectingDate>
,
<collectorName id="26EB534CFFA7CD4EFF4CE9E6B05AE8E2" box="[151,297,402,427]" pageId="112" pageNumber="113">S. Golovatch</collectorName>
leg. (
<collectionCode id="ED0FAE5FFFA7CD4EFEB3E9E6B0B2E8E2" box="[360,449,403,427]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="112" pageNumber="113">ZMUM</collectionCode>
ρ3342).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFFA7CD4EFF1CE9C3B210E8BA" blockId="112.[151,1437,258,966]" pageId="112" pageNumber="113">
<emphasis id="B96AEA88FFA7CD4EFF1CE9C3B04CE886" bold="true" box="[199,319,438,463]" pageId="112" pageNumber="113">Paratypes</emphasis>
(total: 3 females, 22 juveniles).
<materialsCitation id="3B763CC7FFA7CD4EFD7DE9C3B210E8BA" ID-GBIF-Occurrence="1424567106" collectingDate="1986-07-18" collectionCode="ZMUM" collectorName="S. Golovatch" country="Russia" elevation="750" location="Argun Valley" municipality="All" pageId="112" pageNumber="113" specimenCount="25" specimenCount-female="3" specimenCount-juvenile="22" stateProvince="Chechnya">
<collectingMunicipality id="6BC5ACE0FFA7CD4EFD7DE9C3B3B9E886" box="[678,714,438,463]" pageId="112" pageNumber="113">All</collectingMunicipality>
from
<collectingCountry id="F309760AFFA7CD4EFCD6E9C2B200E886" box="[781,883,439,463]" name="Russia" pageId="112" pageNumber="113">RUSSIA</collectingCountry>
,
<collectingRegion id="49DAF878FFA7CD4EFCA5E9C3B282E886" box="[894,1009,438,463]" country="Russia" name="Chechnya" pageId="112" pageNumber="113">Chechnya</collectingRegion>
,
<location id="8EC16041FFA7CD4EFC20E9C2B5E1E886" LSID="urn:lsid:plazi:treatment:03B7878CFFA7CD4DFF4CE8E2B352EBCD:8EC16041FFA7CD4EFC20E9C2B5E1E886" box="[1019,1170,438,463]" country="Russia" municipality="All" name="Argun Valley" pageId="112" pageNumber="113" stateProvince="Chechnya">Argun Valley</location>
,
<locationDeviation id="9A132FB8FFA7CD4EFB47E9C3B4EAE886" box="[1180,1433,438,463]" pageId="112" pageNumber="113">
<quantity id="4CE69B7FFFA7CD4EFB47E9C3B5A6E886" box="[1180,1237,438,463]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="112" pageNumber="113" unit="km" value="5.0">5 km</quantity>
N of Sovetskoye
</locationDeviation>
:
<specimenCount id="9D18FD13FFA7CD4EFF4CE9AEB075E8BA" box="[151,262,474,499]" pageId="112" pageNumber="113" type="female">3 females</specimenCount>
,
<specimenCount id="9D18FD13FFA7CD4EFECAE9AEB0EFE8BA" box="[273,412,474,499]" pageId="112" pageNumber="113" type="juvenile">22 juveniles</specimenCount>
, same data as holotype (
<collectionCode id="ED0FAE5FFFA7CD4EFD77E9AEB276E8BA" box="[684,773,475,499]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="112" pageNumber="113">ZMUM</collectionCode>
ρ3349).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFFA7CD4EFF1CE98BB38BEB5E" blockId="112.[151,1437,258,966]" box="[199,760,510,535]" pageId="112" pageNumber="113">
<materialsCitation id="3B763CC7FFA7CD4EFF1CE98BB38BEB5E" ID-GBIF-Occurrence="1424567123" box="[199,760,510,535]" country="Russia" location="Argun Valley" pageId="112" pageNumber="113" specimenCount="1" stateProvince="Chechnya" typeStatus="holotype">
<emphasis id="B96AEA88FFA7CD4EFF1CE98BB016EB5E" bold="true" box="[199,357,510,535]" pageId="112" pageNumber="113">
<typeStatus id="54A58838FFA7CD4EFF1CE98BB071EB5E" box="[199,258,510,535]" pageId="112" pageNumber="113">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F309760AFFA7CD4EFEB7E98AB0A7EB5E" box="[364,468,511,535]" name="Russia" pageId="112" pageNumber="113">RUSSIA</collectingCountry>
:
<collectingRegion id="49DAF878FFA7CD4EFE05E98BB322EB5E" box="[478,593,510,535]" country="Russia" name="Chechnya" pageId="112" pageNumber="113">Chechnya</collectingRegion>
,
<location id="8EC16041FFA7CD4EFD87E98AB387EB5E" LSID="urn:lsid:plazi:treatment:03B7878CFFA7CD4DFF4CE8E2B352EBCD:8EC16041FFA7CD4EFD87E98AB387EB5E" box="[604,756,510,535]" country="Russia" name="Argun Valley" pageId="112" pageNumber="113" stateProvince="Chechnya">Argun Valley</location>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FFA7CD4DFF1CEA57B352EBCD" lastPageId="115" lastPageNumber="116" pageId="112" pageNumber="113" type="description">
<paragraph id="8BA1369AFFA7CD4EFF1CEA57B2D1EB72" blockId="112.[151,1437,258,966]" box="[199,930,546,571]" pageId="112" pageNumber="113">
<emphasis id="B96AEA88FFA7CD4EFF1CEA57B02AEB72" bold="true" box="[199,345,546,571]" pageId="112" pageNumber="113">Description.</emphasis>
Body with 31 segments (including telson) in adults.
</paragraph>
<paragraph id="8BA1369AFFA7CD4EFF1CEA32B2B3EBCB" blockId="112.[151,1437,258,966]" pageId="112" pageNumber="113">
MEASUREMENTS.
<typeStatus id="54A58838FFA7CD4EFE49EA33B089EB16" box="[402,506,582,607]" pageId="112" pageNumber="113" type="holotype">Holotype</typeStatus>
male
<quantity id="4CE69B7FFFA7CD4EFD99EA32B3E0EB17" box="[578,659,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="113" unit="mm" value="10.0">10 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFFA7CD4EFB03EA32B440EB17" box="[1240,1331,583,607]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="112" pageNumber="113" unit="mm" value="0.8">0.8 mm</quantity>
. Females
<quantity id="4CE69B7FFFA7CD4EFF4CEA1EB07CEBCB" box="[151,271,619,643]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="112" pageNumber="113" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFFA7CD4EFC9CEA1EB2CFEBCB" box="[839,956,619,643]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="112" pageNumber="113" unit="mm" value="0.95" valueMax="1.0" valueMin="0.9">0.9–1 mm</quantity>
.
</paragraph>
<paragraph id="8BA1369AFFA7CD4EFF1CEAFAB42FEBEE" blockId="112.[151,1437,258,966]" box="[199,1372,654,679]" pageId="112" pageNumber="113">
COLORATION (
<figureCitation id="13252A1FFFA7CD4EFEB7EAFBB0B3EBEE" box="[364,448,654,679]" captionStart="FIGURE 92" captionStartId="112.[151,250,1948,1970]" captionTargetBox="[188,1398,1020,1927]" captionTargetId="figure@112.[188,1398,1020,1927]" captionTargetPageId="112" captionText="FIGURE 92. Enghoffiella insolita gen. et sp. nov., paratype female habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272841/files/figure.png" pageId="112" pageNumber="113">Fig. 92</figureCitation>
). Prozonites greyish, metazonites pale brown. Head and first few segments darker.
</paragraph>
<paragraph id="8BA1369AFFA7CD4EFF1CEAC6B42FEA36" blockId="112.[151,1437,258,966]" pageId="112" pageNumber="113">
HEAD. Flattened in male. Labrum with three medial teeth and 5+5 labral and 2+2 supralabral setae. Promentum triangular, without setae. Lingual plates with 5+5 setae, on each plate arranged in two rows. Stipites with
<emphasis id="B96AEA88FFA7CD4EFAE8EAA2B43EEBA7" box="[1331,1357,727,750]" italics="true" pageId="112" pageNumber="113">ca</emphasis>
20+20 setae. Antennae
<quantity id="4CE69B7FFFA7CD4EFE95EA8EB0D7EA5B" box="[334,420,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="112" pageNumber="113" unit="mm" value="1.3">1.3 mm</quantity>
long in
<typeStatus id="54A58838FFA7CD4EFE24EA8FB316EA5A" box="[511,613,762,787]" pageId="112" pageNumber="113" type="holotype">holotype</typeStatus>
. Length of antennomeres (in mm): I (0.09), II (0.15), III (0.33), IV (0.17), V (0.3), VI (0.13), VII (0.11) and VIII (0.02). Length/breadth ratios of antennomeres I–VII: I (1), II (1.4), III (3.6), IV (2), V (2.7), VI (1.2) and VII (1.2). Antennomeres II, IV, V, VI and VII with one, three, one, four and one sensillum, respectively. Number of ocelli 15–18, arranged in 4 rows in
<typeStatus id="54A58838FFA7CD4EFC68EB13B56BEA36" box="[947,1048,870,895]" pageId="112" pageNumber="113" type="holotype">holotype</typeStatus>
;
<quantity id="4CE69B7FFFA7CD4EFBFFEB12B5F4EA36" box="[1060,1159,870,895]" metricMagnitude="-1" metricUnit="m" metricValue="4.699" metricValueMax="5.08" metricValueMin="4.318" pageId="112" pageNumber="113" unit="in" value="18.5" valueMax="20.0" valueMin="17.0">17–20 in</quantity>
5 rows in females.
</paragraph>
<paragraph id="8BA1369AFFA7CD4EFF1CEBFEB189EA8F" blockId="112.[151,1437,258,966]" pageId="112" pageNumber="113">COLLUM. Narrower than head, with six macrochaetae. Anterior edge semi-circular, posterior margin gently concave.</paragraph>
<caption id="DF616612FFA7CD4EFF4CEFE9B510EEFB" box="[151,1123,1948,1970]" httpUri="https://zenodo.org/record/272841/files/figure.png" pageId="112" pageNumber="113" targetBox="[188,1398,1020,1927]" targetPageId="112">
<paragraph id="8BA1369AFFA7CD4EFF4CEFE9B510EEFB" blockId="112.[151,1123,1948,1970]" box="[151,1123,1948,1970]" pageId="112" pageNumber="113">
<emphasis id="B96AEA88FFA7CD4EFF4CEFE9B053EEF8" bold="true" box="[151,288,1948,1970]" pageId="112" pageNumber="113">FIGURE 92.</emphasis>
<taxonomicName id="4C1E4D19FFA7CD4EFEFEEFE9B086EEFB" box="[293,501,1948,1970]" class="Diplopoda" family="Anthroleucosomatidae" genus="Enghoffiella" kingdom="Animalia" order="Chordeumatida" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="B96AEA88FFA7CD4EFEFEEFE9B086EEFB" box="[293,501,1948,1970]" italics="true" pageId="112" pageNumber="113">Enghoffiella insolita</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FFA7CD4EFE20EFE9B3EBEEF8" bold="true" box="[507,664,1948,1969]" pageId="112" pageNumber="113">gen. et sp. nov.</emphasis>
, paratype female habitus. Scale line = 1 mm.
</paragraph>
</caption>
<caption id="DF616612FFA6CD4FFF4CEF51B045EE11" httpUri="https://zenodo.org/record/272842/files/figure.png" pageId="113" pageNumber="114" targetBox="[188,1398,207,1798]" targetPageId="113">
<paragraph id="8BA1369AFFA6CD4FFF4CEF51B045EE11" blockId="113.[151,1436,1828,1880]" pageId="113" pageNumber="114">
<emphasis id="B96AEA88FFA6CD4FFF4CEF51B053EE70" bold="true" box="[151,288,1828,1850]" pageId="113" pageNumber="114">FIGURE 93.</emphasis>
<taxonomicName id="4C1E4D19FFA6CD4FFEF3EF51B08BEE73" box="[296,504,1828,1850]" class="Diplopoda" family="Anthroleucosomatidae" genus="Enghoffiella" kingdom="Animalia" order="Chordeumatida" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="B96AEA88FFA6CD4FFEF3EF51B08BEE73" box="[296,504,1828,1850]" italics="true" pageId="113" pageNumber="114">Enghoffiella insolita</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FFA6CD4FFDDBEF51B3D3EE70" bold="true" box="[512,672,1828,1849]" pageId="113" pageNumber="114">gen. et sp. nov.</emphasis>
, holotype male.
<emphasis id="B96AEA88FFA6CD4FFC96EF51B215EE70" bold="true" box="[845,870,1828,1849]" pageId="113" pageNumber="114">A.</emphasis>
Leg 3.
<emphasis id="B96AEA88FFA6CD4FFC6DEF51B2BDEE70" bold="true" box="[950,974,1828,1849]" pageId="113" pageNumber="114">B.</emphasis>
Leg 4.
<emphasis id="B96AEA88FFA6CD4FFBC5EF51B544EE73" bold="true" box="[1054,1079,1828,1850]" pageId="113" pageNumber="114">C.</emphasis>
Leg 5.
<emphasis id="B96AEA88FFA6CD4FFB5CEF51B5D3EE70" bold="true" box="[1159,1184,1828,1849]" pageId="113" pageNumber="114">D.</emphasis>
Leg 7.
<emphasis id="B96AEA88FFA6CD4FFB2AEF51B47BEE70" bold="true" box="[1265,1288,1828,1849]" pageId="113" pageNumber="114">E.</emphasis>
Leg 10. Scale lines = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFFA6CD4FFF1CEFF5B36BEEA9" blockId="113.[151,1437,1919,2016]" pageId="113" pageNumber="114">
BODY SEGMENTS (
<figureCitation id="13252A1FFFA6CD4FFE7CEF0AB377EED1" box="[423,516,1919,1944]" captionStart="FIGURE 92" captionStartId="112.[151,250,1948,1970]" captionTargetBox="[188,1398,1020,1927]" captionTargetId="figure@112.[188,1398,1020,1927]" captionTargetPageId="112" captionText="FIGURE 92. Enghoffiella insolita gen. et sp. nov., paratype female habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272841/files/figure.png" pageId="113" pageNumber="114">Fig. 92</figureCitation>
). Lateral keels not developed; only barely visible lateral swellings present. Macrochaetae short and trichoid. CIX (pleurotergite 15) ~ 0.8; MIX (pleurotergite 15) ~ 2; PIX (pleurotergite 15) ~ 0.1; MA (pleurotergite 15) ~ 145˚.
</paragraph>
<caption id="DF616612FFA5CD4CFF4CEED0B268EE7C" httpUri="https://zenodo.org/record/272843/files/figure.png" pageId="114" pageNumber="115" targetBox="[212,1381,188,1627]" targetPageId="114">
<paragraph id="8BA1369AFFA5CD4CFF4CEED0B268EE7C" blockId="114.[151,1436,1701,1846]" pageId="114" pageNumber="115">
<emphasis id="B96AEA88FFA5CD4CFF4CEED0B053EFF3" bold="true" box="[151,288,1701,1723]" pageId="114" pageNumber="115">FIGURE 94.</emphasis>
<taxonomicName id="4C1E4D19FFA5CD4CFEFDEED0B086EFF2" box="[294,501,1701,1723]" class="Diplopoda" family="Anthroleucosomatidae" genus="Enghoffiella" kingdom="Animalia" order="Chordeumatida" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="B96AEA88FFA5CD4CFEFDEED0B086EFF2" box="[294,501,1701,1723]" italics="true" pageId="114" pageNumber="115">Enghoffiella insolita</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FFA5CD4CFE27EED0B3EAEFF3" bold="true" box="[508,665,1701,1722]" pageId="114" pageNumber="115">gen. et sp. nov.</emphasis>
, holotype male gonopods.
<emphasis id="B96AEA88FFA5CD4CFC77EED0B2B6EFF3" bold="true" box="[940,965,1701,1722]" pageId="114" pageNumber="115">A.</emphasis>
Anterior gonopods, anterior view.
<emphasis id="B96AEA88FFA5CD4CFAF3EED0B433EFF3" bold="true" box="[1320,1344,1701,1722]" pageId="114" pageNumber="115">B.</emphasis>
Anterior gonopods, posterior view.
<emphasis id="B96AEA88FFA5CD4CFE76EEB1B0B6EF93" bold="true" box="[429,453,1732,1754]" pageId="114" pageNumber="115">C.</emphasis>
Anterior gonopods, lateral view.
<emphasis id="B96AEA88FFA5CD4CFCF1EEB1B230EF90" bold="true" box="[810,835,1732,1753]" pageId="114" pageNumber="115">D.</emphasis>
Posterior gonopods, lateral view.
<emphasis id="B96AEA88FFA5CD4CFB76EEB1B5B7EF90" bold="true" box="[1197,1220,1732,1753]" pageId="114" pageNumber="115">E.</emphasis>
Posterior gonopods, posterior view.
<emphasis id="B96AEA88FFA5CD4CFEEFEE96B04CEFB1" box="[308,319,1763,1784]" italics="true" pageId="114" pageNumber="115">c</emphasis>
= coxite;
<emphasis id="B96AEA88FFA5CD4CFE7DEE97B0DEEFB1" box="[422,429,1762,1784]" italics="true" pageId="114" pageNumber="115">l</emphasis>
= lamella;
<emphasis id="B96AEA88FFA5CD4CFDFAEE97B349EFB1" box="[545,570,1762,1784]" italics="true" pageId="114" pageNumber="115">p1</emphasis>
,
<emphasis id="B96AEA88FFA5CD4CFD9CEE97B313EFB1" box="[583,608,1762,1784]" italics="true" pageId="114" pageNumber="115">p2</emphasis>
,
<emphasis id="B96AEA88FFA5CD4CFDB6EE97B3F6EFB1" box="[621,645,1762,1784]" italics="true" pageId="114" pageNumber="115">p3</emphasis>
and
<emphasis id="B96AEA88FFA5CD4CFD63EE97B3A2EFB1" box="[696,721,1762,1784]" italics="true" pageId="114" pageNumber="115">p4</emphasis>
= processes on anterior gonopods;
<emphasis id="B96AEA88FFA5CD4CFBE1EE96B532EFB1" box="[1082,1089,1763,1784]" italics="true" pageId="114" pageNumber="115">t</emphasis>
= telopodite;
<emphasis id="B96AEA88FFA5CD4CFB14EE96B59BEFB1" box="[1231,1256,1763,1784]" italics="true" pageId="114" pageNumber="115">ap</emphasis>
= anterior part of posterior gonopods;
<emphasis id="B96AEA88FFA5CD4CFEB0EF77B0F7EE5E" box="[363,388,1794,1815]" italics="true" pageId="114" pageNumber="115">cp</emphasis>
= coxal process;
<emphasis id="B96AEA88FFA5CD4CFDE7EF77B321EE5E" box="[572,594,1794,1815]" italics="true" pageId="114" pageNumber="115">cv</emphasis>
= coxal vesicle;
<emphasis id="B96AEA88FFA5CD4CFCDCEF74B268EE5E" box="[775,795,1793,1815]" italics="true" pageId="114" pageNumber="115">hl</emphasis>
= hairy lever;
<emphasis id="B96AEA88FFA5CD4CFC63EF77B2A1EE5E" box="[952,978,1794,1815]" italics="true" pageId="114" pageNumber="115">pp</emphasis>
= posterior part of posterior gonopods;
<emphasis id="B96AEA88FFA5CD4CFAAFEF74B4F6EE5E" box="[1396,1413,1793,1815]" italics="true" pageId="114" pageNumber="115">sl</emphasis>
= sternal lamella;
<emphasis id="B96AEA88FFA5CD4CFEE3EF54B017EE7F" box="[312,356,1825,1846]" italics="true" pageId="114" pageNumber="115">pmp</emphasis>
= posteromedial part. Scale line = 0.3 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFFA5CD4CFF1CEF16B39FEED7" blockId="114.[151,1436,1890,2023]" pageId="114" pageNumber="115">TELSON. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 apical setae. Paraprocts with 3+3 marginal setae.</paragraph>
<paragraph id="8BA1369AFFA5CD4CFF1CEFDEB295EEAF" blockId="114.[151,1436,1890,2023]" pageId="114" pageNumber="115">WALKING LEGS. In both sexes, leg-pairs 1 and 2 with tarsal combs; prefemora with several long and robust setae; femora and postfemora with a group of several long and robust setae.</paragraph>
<paragraph id="8BA1369AFFA4CD4DFF1CE8EDB5BAE854" blockId="115.[151,1437,151,645]" pageId="115" pageNumber="116">
MALE SEXUAL CHARACTERS (
<figureCitation id="13252A1FFFA4CD4DFDCCE8E2B318E9F9" box="[535,619,151,176]" captionStart="FIGURE 93" captionStartId="113.[151,250,1828,1850]" captionTargetBox="[188,1398,207,1798]" captionTargetId="figure@113.[188,1398,207,1798]" captionTargetPageId="113" captionText="FIGURE 93. Enghoffiella insolita gen. et sp. nov., holotype male. A. Leg 3. B. Leg 4. C. Leg 5. D. Leg 7. E. Leg 10. Scale lines = 0.2 mm." httpUri="https://zenodo.org/record/272842/files/figure.png" pageId="115" pageNumber="116">Fig. 93</figureCitation>
). Leg-pairs 3–7 somewhat enlarged. Leg-pairs 3 and 4 each with a basal external protrusion on prefemur. Leg-pair 5 with a basal, oral, subtriangular protrusion on prefemur. Leg-pair 6 without peculiarities. Leg-pair 7 with a posterior, wide, squarish protrusion on coxae. Leg-pair 10 with coxal glands and coxal rounded protrusion. Leg-pair 11 with coxal glands, without other peculiarities.
</paragraph>
<paragraph id="8BA1369AFFA4CD4DFF1CE95DB2BCE8BD" blockId="115.[151,1437,151,645]" pageId="115" pageNumber="116">
ANTERIOR GONOPODS (
<figureCitation id="13252A1FFFA4CD4DFE0CE952B35CE809" box="[471,559,295,320]" captionStart="FIGURE 94" captionStartId="114.[151,250,1701,1723]" captionTargetBox="[212,1381,188,1627]" captionTargetId="figure@114.[188,1398,177,1679]" captionTargetPageId="114" captionText="FIGURE 94. Enghoffiella insolita gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, lateral view. E. Posterior gonopods, posterior view. c = coxite; l = lamella; p 1, p 2, p 3 and p 4 = processes on anterior gonopods; t = telopodite; ap = anterior part of posterior gonopods; cp = coxal process; cv = coxal vesicle; hl = hairy lever; pp = posterior part of posterior gonopods; sl = sternal lamella; pmp = posteromedial part. Scale line = 0.3 mm." httpUri="https://zenodo.org/record/272843/files/figure.png" pageId="115" pageNumber="116">Fig. 94</figureCitation>
A–C). Unusual. Barely visible, very thin, somewhat membranous and low anterior sternal lamella (
<emphasis id="B96AEA88FFA4CD4DFE75E939B0B2E82C" box="[430,449,332,357]" italics="true" pageId="115" pageNumber="116">sl</emphasis>
) present. Coxal processes (
<emphasis id="B96AEA88FFA4CD4DFD27E938B264E82D" box="[764,791,333,356]" italics="true" pageId="115" pageNumber="116">cp</emphasis>
) divided distally. Anteriorly, with four pairs of long and slender processes (
<emphasis id="B96AEA88FFA4CD4DFEA9E905B0FDE8C1" box="[370,398,368,392]" italics="true" pageId="115" pageNumber="116">p1</emphasis>
,
<emphasis id="B96AEA88FFA4CD4DFE44E905B0CFE8C1" box="[415,444,368,392]" italics="true" pageId="115" pageNumber="116">p2</emphasis>
,
<emphasis id="B96AEA88FFA4CD4DFE16E905B099E8C1" box="[461,490,368,392]" italics="true" pageId="115" pageNumber="116">p3</emphasis>
and
<emphasis id="B96AEA88FFA4CD4DFDFCE905B330E8C1" box="[551,579,368,392]" italics="true" pageId="115" pageNumber="116">p4</emphasis>
). Three pairs of these processes acuminate and bare, while one pair (
<emphasis id="B96AEA88FFA4CD4DFAADE905B4E0E8C1" box="[1398,1427,368,392]" italics="true" pageId="115" pageNumber="116">p3</emphasis>
) characterized by an extended and hairy apex. The lateralmost acuminate process (
<emphasis id="B96AEA88FFA4CD4DFBF7E9E1B53BE8E5" box="[1068,1096,404,428]" italics="true" pageId="115" pageNumber="116">p4</emphasis>
) connected to base of anterior gonopods by a lamella (
<emphasis id="B96AEA88FFA4CD4DFE76E9C2B0C6E899" box="[429,437,439,464]" italics="true" pageId="115" pageNumber="116">l</emphasis>
). Posteromedial part (
<emphasis id="B96AEA88FFA4CD4DFD6EE9CCB396E899" box="[693,741,441,464]" italics="true" pageId="115" pageNumber="116">pmp</emphasis>
) of coxites wing-like, highly hairy, with mesal edges folded inside. Posterior side of anterior gonopods with a pair of hairy levers (
<emphasis id="B96AEA88FFA4CD4DFC72E9A9B2CCE8BC" box="[937,959,476,501]" italics="true" pageId="115" pageNumber="116">hl</emphasis>
).
</paragraph>
<paragraph id="8BA1369AFFA4CD4DFF1CEA75B352EBCD" blockId="115.[151,1437,151,645]" pageId="115" pageNumber="116">
POSTERIOR GONOPODS (
<figureCitation id="13252A1FFFA4CD4DFE3BE98AB34AEB51" box="[480,569,511,536]" captionStart="FIGURE 94" captionStartId="114.[151,250,1701,1723]" captionTargetBox="[212,1381,188,1627]" captionTargetId="figure@114.[188,1398,177,1679]" captionTargetPageId="114" captionText="FIGURE 94. Enghoffiella insolita gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, lateral view. E. Posterior gonopods, posterior view. c = coxite; l = lamella; p 1, p 2, p 3 and p 4 = processes on anterior gonopods; t = telopodite; ap = anterior part of posterior gonopods; cp = coxal process; cv = coxal vesicle; hl = hairy lever; pp = posterior part of posterior gonopods; sl = sternal lamella; pmp = posteromedial part. Scale line = 0.3 mm." httpUri="https://zenodo.org/record/272843/files/figure.png" pageId="115" pageNumber="116">Fig. 94</figureCitation>
D, E). Coxites (
<emphasis id="B96AEA88FFA4CD4DFD21EA74B275EB51" box="[762,774,513,536]" italics="true" pageId="115" pageNumber="116">c</emphasis>
) divided. Telopodites (
<emphasis id="B96AEA88FFA4CD4DFBC6EA74B556EB51" box="[1053,1061,513,536]" italics="true" pageId="115" pageNumber="116">t</emphasis>
) present on posterolateral side, without setae (only one telopodite with a short seta on a small knob). Coxal processes (
<emphasis id="B96AEA88FFA4CD4DFBA2EA50B5E0EB75" box="[1145,1171,549,572]" italics="true" pageId="115" pageNumber="116">cp</emphasis>
) orientated posteriorly, consisting of an anterior part (
<emphasis id="B96AEA88FFA4CD4DFE2CEA3CB360EB29" box="[503,531,585,608]" italics="true" pageId="115" pageNumber="116">ap</emphasis>
) connected laterally to a posterior, spoon-shaped structure (
<emphasis id="B96AEA88FFA4CD4DFB12EA3CB596EB29" box="[1225,1253,585,608]" italics="true" pageId="115" pageNumber="116">pp</emphasis>
) by lamella (
<emphasis id="B96AEA88FFA4CD4DFA5FEA32B4FFEB29" box="[1412,1420,583,608]" italics="true" pageId="115" pageNumber="116">l</emphasis>
). Mesally, coxal vesicle (
<emphasis id="B96AEA88FFA4CD4DFE44EA18B0CBEBCD" box="[415,440,621,644]" italics="true" pageId="115" pageNumber="116">cv</emphasis>
) present.
</paragraph>
</subSubSection>
</treatment>
</document>
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