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<document id="FE36EB083DC8C0FE695D9BEC52127CA3" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A135F532CFDBCFBE9FEEBFBE1" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus rugitarsis Moret 2020, sp. nov." docType="treatment" docUuid="924335EA-EF3D-4630-8607-DE9A9749EEFA" docUuidSource="ZooBank" docVersion="8" lastPageNumber="26" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="24" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="0C6A7D5C6C6B753D1B5C88D20B1E48FC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BDCFBBC8198D3BE03012EE1BA49BD70B">
<mods:title id="A2FFDB550BA0B141EC963495C0D9BFBB">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
</mods:titleInfo>
<mods:name id="EA7CA134854D46F0D8C29CCDE4986A08" type="personal">
<mods:role id="42180045ADCD1604B60F3AF6359A8715">
<mods:roleTerm id="355C37A55B905BA18ECD2875F6A24531">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7881EC1864C724557BEBFF0A441C3596">Moret, Pierre</mods:namePart>
<mods:nameIdentifier id="36B96C4AB2A8EB129AAE8B9201E1E4C8" type="LSID">2E194645-D255-438B-819E-A2B6C39AD699</mods:nameIdentifier>
<mods:affiliation id="C9E58E225583F8DDAF0688F4BFDC71C8">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="38E759B6C10A22801A701DAA987B0489" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="7931049A296719D42280C8F0B134E063" type="personal">
<mods:role id="B7559688B2EBE876FD9466A51293F36C">
<mods:roleTerm id="D996EEB5C78051F4BF40F34202832FF2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="06D5C73B6E36CBC60970C392E8DDB17C">Murienne, Jérôme</mods:namePart>
<mods:nameIdentifier id="3B52C26EF27A71AF5A66F6070B68AB0E" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
<mods:affiliation id="DC18841B81DCDD41EAF7FDC81BC760A4">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
<mods:nameIdentifier id="C91D56E6A87B0FB153B949EDA718FA09" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A557E45DBD6FAD379B053571F432D18C">text</mods:typeOfResource>
<mods:relatedItem id="EB58147061CAA19CE1019ED2C2C1FD34" type="host">
<mods:titleInfo id="EC3A3F5C2E14E5BA9A10C076AAC33575">
<mods:title id="05E37B428B4A860E2B5A253B9106E890">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="34ECF4F77DCD3AA3C854DED2C2C93E11">
<mods:date id="A5B2F1C3D9CDD8EBA37675C1CF8B0A19">2020</mods:date>
<mods:detail id="0F664E17F212A85EDF78654425B13471" type="pubDate">
<mods:number id="73100B8253392AE631DC574FCB96E388">2020-05-15</mods:number>
</mods:detail>
<mods:detail id="4FEDC65686BF3A78D297BF6D4C17CFE3" type="volume">
<mods:number id="7D96B9D32A7D75572810E55212C98099">646</mods:number>
</mods:detail>
<mods:extent id="A3C74FCBE259D2F199D4EA46015FD5E4" unit="page">
<mods:start id="B6D8170ACB64E34917B8D6C7FCE732E1">1</mods:start>
<mods:end id="F71C61FB20A3AB552E4948098EEFCC0D">55</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B0C7BD5CB33148D8F4AFE7307BB51C17">journal article</mods:classification>
<mods:identifier id="6114E4ECB0898892BB7F6E6F5BF24509" type="CLB-Dataset">22092</mods:identifier>
<mods:identifier id="F4C2DA69F31661CB270FA265DDE92FF0" type="DOI">10.5852/ejt.2020.646</mods:identifier>
<mods:identifier id="8B25495CC261DA9C4B4E505D4EB99186" type="GBIF-Dataset">c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b</mods:identifier>
<mods:identifier id="88627065797C3076722F145672339673" type="Zenodo-Dep">3829682</mods:identifier>
<mods:identifier id="C8965F22037A40CA713363F07CA5DE39" type="ZooBank">4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B</mods:identifier>
</mods:mods>
<treatment id="03B7726A135F532CFDBCFBE9FEEBFBE1" ID-DOI="http://doi.org/10.5281/zenodo.3848377" ID-GBIF-Taxon="164160403" ID-Zenodo-Dep="3848377" LSID="urn:lsid:zoobank.org:act:924335EA-EF3D-4630-8607-DE9A9749EEFA" httpUri="http://treatment.plazi.org/id/03B7726A135F532CFDBCFBE9FEEBFBE1" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<subSubSection id="C30490F7135F5322FDBCFBE9FB42FBD2" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BA1C37C135F5322FDBCFBE9FC61FBAD" blockId="23.[369,1220,1043,1140]" box="[589,999,1043,1070]" pageId="23" pageNumber="24">
<heading id="D0E97410135F5322FDBCFBE9FC61FBAD" box="[589,999,1043,1070]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<taxonomicName id="4C1EB8FF135F5322FDBCFBE9FC03FBAD" ID-CoL="8MG5N" authority="Moret" authorityName="Moret" authorityYear="2020" box="[589,901,1043,1070]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="rugitarsis" status="sp. nov.">
<emphasis id="B96A1F6E135F5322FDBCFBE9FCB0FBAE" bold="true" box="[589,822,1043,1070]" italics="true" pageId="23" pageNumber="24">Dyscolus rugitarsis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215135F5322FC7DFBEEFC61FBAD" box="[908,999,1044,1070]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BA1C37C135F5322FE80FBCCFB42FBD2" blockId="23.[369,1220,1043,1140]" box="[369,1220,1078,1105]" pageId="23" pageNumber="24">
<uri id="FF8FCF7E135F5322FE80FBCCFB42FBD2" box="[369,1220,1078,1105]" pageId="23" pageNumber="24">
urn:lsid:zoobank.org:act:
<uuid id="FFB8F9A9135F5322FD78FBCCFB42FBD2" box="[649,1220,1078,1105]" pageId="23" pageNumber="24">924335EA-EF3D-4630-8607-DE9A9749EEFA</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30490F7135F5322FD29FBA0FCDDFBF7" box="[728,859,1114,1140]" pageId="23" pageNumber="24" type="description">
<paragraph id="8BA1C37C135F5322FD29FBA0FCDDFBF7" blockId="23.[369,1220,1043,1140]" box="[728,859,1114,1140]" pageId="23" pageNumber="24">
<figureCitation id="1325DFF9135F5322FD29FBA0FCDDFBF7" box="[728,859,1114,1140]" captionStart="Figs 19–24" captionStartId="24.[189,241,1879,1905]" captionTargetBox="[306,1275,267,1823]" captionTargetId="figure@24.[269,1319,262,1835]" captionTargetPageId="24" captionText="Figs 19–24. Dyscolus spp., habitus (19–21) and aedeagus, median lobe in lateral view (22–24). 19. D. rugitarsis Moret sp. nov., male holotype (QACZ). 20. D. sulcipedis Moret sp. nov., male holotype (QACZ). 21. D. marini Moret sp. nov., female paratype. 22. D. rugitarsis Moret sp. nov. 23. D. sulcipedis Moret sp. nov. 24. D. marini Moret sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3829694" httpUri="https://zenodo.org/record/3829694/files/figure.png" pageId="23" pageNumber="24">Figs 19, 22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30490F7135F5322FF4CFB66FCCEFB67" box="[189,840,1180,1252]" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BA1C37C135F5322FF4CFB66FEC2FB35" blockId="23.[189,324,1180,1206]" box="[189,324,1180,1206]" pageId="23" pageNumber="24">
<heading id="D0E97410135F5322FF4CFB66FEC2FB35" bold="true" box="[189,324,1180,1206]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B96A1F6E135F5322FF4CFB66FEC2FB35" bold="true" box="[189,324,1180,1206]" pageId="23" pageNumber="24">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C135F5322FF4CFB30FCCEFB67" blockId="23.[189,840,1226,1252]" box="[189,840,1226,1252]" pageId="23" pageNumber="24">Compound Latin adjective meaning ‘with rugose tarsi’.</paragraph>
</subSubSection>
<subSubSection id="C30490F7135F532CFF4CFAF1FEEBFBE1" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BA1C37C135F5322FF4CFAF1FEEAFAA6" blockId="23.[189,364,1291,1317]" box="[189,364,1291,1317]" pageId="23" pageNumber="24">
<heading id="D0E97410135F5322FF4CFAF1FEEAFAA6" bold="true" box="[189,364,1291,1317]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B96A1F6E135F5322FF4CFAF1FEEAFAA6" bold="true" box="[189,364,1291,1317]" pageId="23" pageNumber="24">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C135F5322FF1BFAC0FEDAFAD7" blockId="23.[189,1399,1338,1435]" box="[234,348,1338,1364]" pageId="23" pageNumber="24">
<emphasis id="B96A1F6E135F5322FF1BFAC0FEDAFAD7" bold="true" box="[234,348,1338,1364]" pageId="23" pageNumber="24">
<typeStatus id="54A57DDE135F5322FF1BFAC0FEDAFAD7" box="[234,348,1338,1364]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA1C37C135F5322FF4CFAA7FBB8FA19" blockId="23.[189,1399,1338,1435]" pageId="23" pageNumber="24">
<materialsCitation id="3B76C921135F5322FF4CFAA7FBB8FA19" ID-GBIF-Occurrence="2620303355" collectingDate="2016-08-04" collectionCode="QCAZ" collectorName="P. Moret &amp; S. Aguirre &amp; E. Moreno" country="Ecuador" elevation="3240" latitude="-4.711861" location="Parque Nacional Yacuri" longLatPrecision="1" longitude="-79.44035" municipality="Parque Nacional Yacuri" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Loja Province" typeStatus="holotype">
<collectingCountry id="F30983EC135F5322FF4CFAA7FECDFAF4" box="[189,331,1373,1399]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
•
<specimenCount id="9D1808F5135F5322FE96FAA1FEF8FAF4" box="[359,382,1371,1399]" pageId="23" pageNumber="24" type="male">♂</specimenCount>
;
<collectingRegion id="49DA0D9E135F5322FE7FFAA7FDBEFAF4" box="[398,568,1373,1399]" country="Ecuador" name="Loja" pageId="23" pageNumber="24">Loja Province</collectingRegion>
,
<collectingMunicipality id="6BC55906135F5322FDB4FAA4FCE4FAF4" box="[581,866,1373,1400]" pageId="23" pageNumber="24">Parque Nacional Yacuri</collectingMunicipality>
, Waypoint 167;
<geoCoordinate id="EE2AA5BB135F5322FBD7FAA7FB31FAF4" box="[1062,1207,1373,1399]" degrees="4.711861" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="-4.711861">4.711861º S</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB135F5322FB34FAA7FAF4FAFB" box="[1221,1394,1373,1400]" degrees="79.440355" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="-79.44035">79.440355º W</geoCoordinate>
;
<elevation id="0033244F135F5322FF4CFA7AFED0FA19" box="[189,342,1408,1435]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="23" pageNumber="24" unit="m" value="3240.0">
<quantity id="4CE66E99135F5322FF4CFA7AFE90FA18" box="[189,278,1408,1435]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="23" pageNumber="24" unit="m" value="3240.0">3240 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC135F5322FE92FA7AFE71FA19" box="[355,503,1408,1434]" pageId="23" pageNumber="24" value="2016-08-04">
<collectingDate id="EFE41C54135F5322FE92FA7AFE71FA19" box="[355,503,1408,1434]" pageId="23" pageNumber="24" value="2016-08-04">4 Aug. 2016</collectingDate>
</date>
;
<collectorName id="26EBA6AA135F5322FDF2FA7BFDECFA18" box="[515,618,1409,1435]" pageId="23" pageNumber="24">P. Moret</collectorName>
,
<collectorName id="26EBA6AA135F5322FD84FA7AFD6BFA19" box="[629,749,1408,1434]" pageId="23" pageNumber="24">S. Aguirre</collectorName>
and
<collectorName id="26EBA6AA135F5322FCD6FA7BFC22FA18" box="[807,932,1409,1435]" pageId="23" pageNumber="24">E. Moreno</collectorName>
leg.;
<collectionCode id="ED0F5BB9135F5322FC17FA7AFBBCFA19" box="[998,1082,1408,1434]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="23" pageNumber="24" type="Museum">QCAZ</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA1C37C135F5322FF1BFA38FD90FA5F" blockId="23.[189,1399,1472,1711]" box="[234,534,1472,1500]" pageId="23" pageNumber="24">
<emphasis id="B96A1F6E135F5322FF1BFA38FEE1FA5F" bold="true" box="[234,359,1474,1500]" pageId="23" pageNumber="24">
<typeStatus id="54A57DDE135F5322FF1BFA38FEE1FA5F" box="[234,359,1474,1500]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
</emphasis>
(9 ♂♂,
<specimenCount id="9D1808F5135F5322FE3BFA3BFD8CFA5F" box="[458,522,1473,1500]" pageId="23" pageNumber="24" type="female">5 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BA1C37C135F5322FF4CFA1FFE2EF92C" blockId="23.[189,1399,1472,1711]" pageId="23" pageNumber="24">
<materialsCitation id="3B76C921135F5322FF4CFA1FFBEEFA7C" ID-GBIF-Occurrence="2620303421" box="[189,1128,1509,1535]" collectingDate="2016-08-04" collectionCode="CPM" collectorName="P. Moret &amp; S. Aguirre &amp; E. Moreno" country="Ecuador" elevation="3240" latitude="-4.711861" location="Loja Province" longLatPrecision="1" longitude="-79.44035" municipality="Parque Nacional Yacuri" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Loja Province">
<collectingCountry id="F30983EC135F5322FF4CFA1FFECDFA7C" box="[189,331,1509,1535]" name="Ecuador" pageId="23" pageNumber="24">ECUADOR</collectingCountry>
–
<emphasis id="B96A1F6E135F5322FE98FA1FFD9CFA7C" bold="true" box="[361,538,1509,1535]" pageId="23" pageNumber="24">
<collectingRegion id="49DA0D9E135F5322FE98FA1FFD9CFA7C" box="[361,538,1509,1535]" country="Ecuador" name="Loja" pageId="23" pageNumber="24">Loja Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D1808F5135F5322FDC5FA1FFDE4FA7C" box="[564,610,1509,1535]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB9135F5322FBDBFA1FFBEEFA7C" box="[1066,1128,1509,1535]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="23" pageNumber="24">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921135F5322FB73FA1FFB36F9A1" ID-GBIF-Occurrence="2620303377" collectingDate="2016-08-04" collectionCode="CPM" collectorName="P. Moret &amp; S. Aguirre &amp; E. Moreno" country="Ecuador" elevation="3240" latitude="-4.711861" location="Waypoint" longLatPrecision="1" longitude="-79.44035" municipality="Parque Nacional Yacuri" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F5135F5322FB73FA1FFB36FA7C" box="[1154,1200,1507,1535]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; same collection data as for holotype; COI voucher PM167-05, BOLD sequence SUM060-18;
<collectionCode id="ED0F5BB9135F5322FB83F9F2FB36F9A1" box="[1138,1200,1544,1570]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="23" pageNumber="24">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921135F5322FB21F9F2FEB1F9EA" ID-GBIF-Occurrence="2620303411" collectingDate="2013-08-11" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" elevation="3240" latitude="-4.71198" location="Cordillera Lagunillas" longLatPrecision="1" longitude="-79.44045" municipality="Cordillera Lagunillas" pageId="23" pageNumber="24" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Loja">
<specimenCount id="9D1808F5135F5322FB21F9F2FA91F9A1" box="[1232,1303,1542,1570]" pageId="23" pageNumber="24" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="9D1808F5135F5322FAD8F9F2FAE9F9A0" box="[1321,1391,1544,1571]" pageId="23" pageNumber="24" type="female">3 ♀♀</specimenCount>
;
<collectingMunicipality id="6BC55906135F5322FF4CF9D1FE3BF9C6" box="[189,445,1579,1605]" pageId="23" pageNumber="24">Cordillera Lagunillas</collectingMunicipality>
, Waypoint 3;
<geoCoordinate id="EE2AA5BB135F5322FD99F9D1FD6AF9C6" box="[616,748,1579,1605]" degrees="4.71198" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="-4.71198">4.71198º S</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB135F5322FD0DF9D1FC1DF9C5" box="[764,923,1579,1606]" degrees="79.44045" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="-79.44045">79.44045º W</geoCoordinate>
;
<elevation id="0033244F135F5322FC5BF9D1FBCFF9C6" box="[938,1097,1579,1606]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="23" pageNumber="24" unit="m" value="3240.0">
<quantity id="4CE66E99135F5322FC5BF9D1FB80F9C5" box="[938,1030,1579,1606]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="23" pageNumber="24" unit="m" value="3240.0">3240 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC135F5322FBA8F9D1FA87F9C6" box="[1113,1281,1579,1605]" pageId="23" pageNumber="24" value="2013-08-11">
<collectingDate id="EFE41C54135F5322FBA8F9D1FA87F9C6" box="[1113,1281,1579,1605]" pageId="23" pageNumber="24" value="2013-08-11">11 Aug. 2013</collectingDate>
</date>
;
<collectorName id="26EBA6AA135F5322FAE1F9D6FAF1F9C5" box="[1296,1399,1580,1606]" pageId="23" pageNumber="24">P. Moret</collectorName>
leg.;
<collectionCode id="ED0F5BB9135F5322FF08F9B5FEB1F9EA" box="[249,311,1615,1641]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="23" pageNumber="24">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921135F5322FEA2F9B5FC30F9EA" ID-GBIF-Occurrence="2620303412" box="[339,950,1613,1641]" collectingDate="2013-08-11" collectionCode="MNHN" collectorName="P. Moret" country="Ecuador" elevation="3240" latitude="-4.71198" location="Waypoint" longLatPrecision="1" longitude="-79.44045" municipality="Cordillera Lagunillas" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F5135F5322FEA2F9B5FE07F9EA" box="[339,385,1613,1641]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED0F5BB9135F5322FCADF9B5FC30F9EA" box="[860,950,1615,1641]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921135F5322FC20F9B5FE38F90F" ID-GBIF-Occurrence="2620303335" collectingDate="2013-08-11" collectionCode="QCAZ" collectorName="P. Moret" country="Ecuador" elevation="3240" latitude="-4.71198" location="Waypoint" longLatPrecision="1" longitude="-79.44045" municipality="Cordillera Lagunillas" pageId="23" pageNumber="24" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Loja">
<specimenCount id="9D1808F5135F5322FC20F9B5FB90F9EA" box="[977,1046,1613,1641]" pageId="23" pageNumber="24" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D1808F5135F5322FBD4F9B5FBD5F9EA" box="[1061,1107,1615,1641]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED0F5BB9135F5322FE9FF988FE38F90F" box="[366,446,1650,1676]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="23" pageNumber="24" type="Museum">QCAZ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921135F5322FE2AF988FE22F92C" ID-GBIF-Occurrence="2620303314" collectingDate="2001-07-15" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" elevation="3200" location="Via Jimbura-Zumba" municipality="Cordillera Lagunillas" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F5135F5322FE2AF988FD8CF90F" box="[475,522,1648,1676]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6BC55906135F5322FDEAF988FC9CF90F" box="[539,794,1650,1676]" pageId="23" pageNumber="24">Cordillera Lagunillas</collectingMunicipality>
,
<location id="8EC195A7135F5322FCD6F988FB99F90F" LSID="urn:lsid:plazi:treatment:03B7726A135F532CFDBCFBE9FEEBFBE1:8EC195A7135F5322FCD6F988FB99F90F" box="[807,1055,1650,1676]" country="Ecuador" municipality="Cordillera Lagunillas" name="Via Jimbura-Zumba" pageId="23" pageNumber="24" stateProvince="Loja">Vía Jimbura–Zumba</location>
;
<elevation id="0033244F135F5322FBDCF988FB4CF90F" box="[1069,1226,1650,1676]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="23" pageNumber="24" unit="m" value="3200.0">
<quantity id="4CE66E99135F5322FBDCF988FB0EF90F" box="[1069,1160,1650,1676]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="23" pageNumber="24" unit="m" value="3200.0">3200 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC135F5322FB2BF98BFAF4F90F" box="[1242,1394,1649,1676]" pageId="23" pageNumber="24" value="2001-07-15">
<collectingDate id="EFE41C54135F5322FB2BF98BFAF4F90F" box="[1242,1394,1649,1676]" pageId="23" pageNumber="24" value="2001-07-15">15 Jul. 2001</collectingDate>
</date>
;
<collectorName id="26EBA6AA135F5322FF4CF96FFEA7F92C" box="[189,289,1685,1711]" pageId="23" pageNumber="24">P. Moret</collectorName>
leg.;
<collectionCode id="ED0F5BB9135F5322FE93F96FFE22F92C" box="[354,420,1685,1711]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="23" pageNumber="24">CPM</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA1C37C135F5322FF4CF92CFE52F973" blockId="23.[189,468,1750,1776]" box="[189,468,1750,1776]" pageId="23" pageNumber="24">
<heading id="D0E97410135F5322FF4CF92CFE52F973" bold="true" box="[189,468,1750,1776]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B96A1F6E135F5322FF4CF92CFE52F973" bold="true" box="[189,468,1750,1776]" pageId="23" pageNumber="24">Diagnostic description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C135F5322FF4CF8FFFAF0F84C" blockId="23.[189,1399,1797,1999]" pageId="23" pageNumber="24">
Habitus:
<figureCitation id="1325DFF9135F5322FEDDF8FFFE01F89C" box="[300,391,1797,1823]" captionStart="Figs 19–24" captionStartId="24.[189,241,1879,1905]" captionTargetBox="[306,1275,267,1823]" captionTargetId="figure@24.[269,1319,262,1835]" captionTargetPageId="24" captionText="Figs 19–24. Dyscolus spp., habitus (19–21) and aedeagus, median lobe in lateral view (22–24). 19. D. rugitarsis Moret sp. nov., male holotype (QACZ). 20. D. sulcipedis Moret sp. nov., male holotype (QACZ). 21. D. marini Moret sp. nov., female paratype. 22. D. rugitarsis Moret sp. nov. 23. D. sulcipedis Moret sp. nov. 24. D. marini Moret sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3829694" httpUri="https://zenodo.org/record/3829694/files/figure.png" pageId="23" pageNumber="24">Fig. 19</figureCitation>
. Wingless. Body length: 10.1–11.0 mm. Body colour variable, from brunneous to brownish black; femora brownish; rest of the legs, antennae and mouthparts reddish brown. Elytral microsculpture variable, mostly isodiametric in the middle part of the intervals, oblong near the striae. Head broad, weakly constricted basally, eyes moderately bulging, genae flat. Pronotum transverse, sides slightly arcuate anterad, weakly sinuate posterad; hind angles obtuse; two pairs of lateral setae. Elytra ovoid, slightly convex; striae entire, well impressed, relatively broad; intervals variable, flat to slightly
</paragraph>
<caption id="DF6193F41350532DFF4CF8ADFD3BF858" ID-DOI="http://doi.org/10.5281/zenodo.3829694" ID-Zenodo-Dep="3829694" httpUri="https://zenodo.org/record/3829694/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,241,1879,1905]" targetBox="[306,1275,267,1823]" targetPageId="24">
<paragraph id="8BA1C37C1350532DFF4CF8ADFD3BF858" blockId="24.[189,1399,1879,2011]" pageId="24" pageNumber="25">
<emphasis id="B96A1F6E1350532DFF4CF8ADFED6F8F2" bold="true" box="[189,336,1879,1905]" pageId="24" pageNumber="25">Figs 19–24.</emphasis>
<taxonomicName id="4C1EB8FF1350532DFEAEF8ADFE4DF8F2" authorityName="Dejean" authorityYear="1831" box="[351,459,1879,1905]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B96A1F6E1350532DFEAEF8ADFE4DF8F2" box="[351,459,1879,1905]" italics="true" pageId="24" pageNumber="25">Dyscolus</emphasis>
</taxonomicName>
spp., habitus (19–21) and aedeagus, median lobe in lateral view (22–24).
<emphasis id="B96A1F6E1350532DFF4CF880FF5CF817" bold="true" box="[189,218,1914,1940]" pageId="24" pageNumber="25">19</emphasis>
.
<taxonomicName id="4C1EB8FF1350532DFF19F881FE4DF816" authority="Moret" authorityName="Moret" authorityYear="2020" box="[232,459,1915,1941]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rugitarsis" status="sp. nov.">
<emphasis id="B96A1F6E1350532DFF19F881FEFAF817" box="[232,380,1915,1940]" italics="true" pageId="24" pageNumber="25">D. rugitarsis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151350532DFE21F881FDAAF816" box="[464,556,1915,1941]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (QACZ).
<emphasis id="B96A1F6E1350532DFCA5F880FCF7F817" bold="true" box="[852,881,1914,1940]" pageId="24" pageNumber="25">20</emphasis>
.
<taxonomicName id="4C1EB8FF1350532DFC8FF881FBE2F816" authority="Moret" authorityName="Moret" authorityYear="2020" box="[894,1124,1915,1941]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sulcipedis" status="sp. nov.">
<emphasis id="B96A1F6E1350532DFC8FF881FB90F816" box="[894,1046,1915,1941]" italics="true" pageId="24" pageNumber="25">D. sulcipedis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151350532DFB9BF881FB40F816" box="[1130,1222,1915,1941]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (QACZ).
<emphasis id="B96A1F6E1350532DFEDCF864FECCF83B" bold="true" box="[301,330,1950,1976]" pageId="24" pageNumber="25">21</emphasis>
.
<taxonomicName id="4C1EB8FF1350532DFEA6F865FD93F83B" authority="Moret" authorityName="Moret" authorityYear="2020" box="[343,533,1950,1976]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="marini" status="sp. nov.">
<emphasis id="B96A1F6E1350532DFEA6F865FE41F83B" box="[343,455,1951,1976]" italics="true" pageId="24" pageNumber="25">D. marini</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151350532DFDEBF864FDF3F83B" box="[538,629,1950,1976]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, female paratype.
<emphasis id="B96A1F6E1350532DFCB7F864FCE5F83B" bold="true" box="[838,867,1950,1976]" pageId="24" pageNumber="25">22</emphasis>
.
<taxonomicName id="4C1EB8FF1350532DFC80F865FBD4F83B" authority="Moret" authorityName="Moret" authorityYear="2020" box="[881,1106,1950,1976]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rugitarsis" status="sp. nov.">
<emphasis id="B96A1F6E1350532DFC80F865FB83F83B" box="[881,1029,1951,1976]" italics="true" pageId="24" pageNumber="25">D. rugitarsis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151350532DFBA6F864FB36F83B" box="[1111,1200,1950,1976]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B96A1F6E1350532DFB44F864FB54F83B" bold="true" box="[1205,1234,1950,1976]" pageId="24" pageNumber="25">23</emphasis>
.
<taxonomicName id="4C1EB8FF1350532DFB2EF865FE83F858" authority="Moret" authorityName="Moret" authorityYear="2020" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sulcipedis" status="sp. nov.">
<emphasis id="B96A1F6E1350532DFB2EF865FAF0F83B" box="[1247,1398,1950,1976]" italics="true" pageId="24" pageNumber="25">D. sulcipedis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151350532DFEFDF83BFEE1F858" box="[268,359,1985,2011]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
24.
<taxonomicName id="4C1EB8FF1350532DFE6BF838FDDDF858" authority="Moret" authorityName="Moret" authorityYear="2020" box="[410,603,1985,2011]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="marini" status="sp. nov.">
<emphasis id="B96A1F6E1350532DFE6BF838FD8DF858" box="[410,523,1986,2011]" italics="true" pageId="24" pageNumber="25">D. marini</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151350532DFD93F83BFD3BF858" box="[610,701,1985,2011]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BA1C37C1351532CFF4CFEF6FC06FE7B" blockId="25.[189,1406,267,505]" pageId="25" pageNumber="26">
convex. Third elytral interval with 2–3 setae (two specimens lack the anterior seta on one elytron only). Last visible abdominal ventrite with one pair (♂) or two pairs (
<specimenCount id="9D1808F51351532CFC68FED5FC2AFECA" box="[921,940,303,329]" pageId="25" pageNumber="26" type="female">♀</specimenCount>
) of setae along its apical margin. Legs: tarsi rugose dorsally, metatarsomeres 1–4 bisulcate, fourth metatarsomere with one pair of dorsolateral subapical setae, apical lobes moderately long, the external lobe two times longer than the inner lobe; fifth metatarsomere broadened and dorsally depressed in basal half, ventrally asetose.
<emphasis id="B96A1F6E1351532CFB30FE62FAE8FE31" box="[1217,1390,408,434]" italics="true" pageId="25" pageNumber="26">Male genitalia</emphasis>
:
<figureCitation id="1325DFF91351532CFF4CFE41FE9EFE56" box="[189,280,443,469]" captionStart="Figs 19–24" captionStartId="24.[189,241,1879,1905]" captionTargetBox="[306,1275,267,1823]" captionTargetId="figure@24.[269,1319,262,1835]" captionTargetPageId="24" captionText="Figs 19–24. Dyscolus spp., habitus (19–21) and aedeagus, median lobe in lateral view (22–24). 19. D. rugitarsis Moret sp. nov., male holotype (QACZ). 20. D. sulcipedis Moret sp. nov., male holotype (QACZ). 21. D. marini Moret sp. nov., female paratype. 22. D. rugitarsis Moret sp. nov. 23. D. sulcipedis Moret sp. nov. 24. D. marini Moret sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3829694" httpUri="https://zenodo.org/record/3829694/files/figure.png" pageId="25" pageNumber="26">Fig. 22</figureCitation>
. Median lobe arcuate, apex relatively large, very thin in lateral view, endophallus with one subapical sclerotized structure.
<emphasis id="B96A1F6E1351532CFDDEFE25FD70FE7A" box="[559,758,479,505]" italics="true" pageId="25" pageNumber="26">Female genitalia</emphasis>
: unstudied.
</paragraph>
<paragraph id="8BA1C37C1351532CFF4CFDDAFEE5FDB9" blockId="25.[189,355,544,570]" box="[189,355,544,570]" pageId="25" pageNumber="26">
<heading id="D0E974101351532CFF4CFDDAFEE5FDB9" bold="true" box="[189,355,544,570]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B96A1F6E1351532CFF4CFDDAFEE5FDB9" bold="true" box="[189,355,544,570]" pageId="25" pageNumber="26">Comparisons</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1351532CFF4CFDB5FD61FCBF" blockId="25.[189,1399,590,828]" pageId="25" pageNumber="26">
<taxonomicName id="4C1EB8FF1351532CFF4CFDB5FE7FFDEA" authority="Moret" authorityName="Moret" authorityYear="2020" box="[189,505,591,617]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rugitarsis" status="sp. nov.">
<emphasis id="B96A1F6E1351532CFF4CFDB5FE20FDEB" box="[189,422,591,617]" italics="true" pageId="25" pageNumber="26">Dyscolus rugitarsis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151351532CFDF5FDB5FDE4FDEA" box="[516,610,591,617]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
is closely related, within the
<taxonomicName id="4C1EB8FF1351532CFC23FDB5FB1DFDEA" authority="Moret" authorityName="Moret" authorityYear="2020" box="[978,1179,591,617]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="marini" status="sp. nov.">
<emphasis id="B96A1F6E1351532CFC23FDB5FBCEFDEB" box="[978,1096,591,616]" italics="true" pageId="25" pageNumber="26">D. marini</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151351532CFB57FDB5FA82FDEA" box="[1190,1284,591,617]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
clade, to
<taxonomicName id="4C1EB8FF1351532CFF4CFD89FEE3FD0F" authorityName="Perrault" authorityYear="1993" box="[189,357,627,652]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="moretianus">
<emphasis id="B96A1F6E1351532CFF4CFD89FEE3FD0F" box="[189,357,627,652]" italics="true" pageId="25" pageNumber="26">D. moretianus</emphasis>
</taxonomicName>
which inhabits the western part of the
<collectingRegion id="49DA0D9E1351532CFCCAFD88FC67FD0F" box="[827,993,626,652]" country="Ecuador" name="Loja" pageId="25" pageNumber="26">Loja Province</collectingRegion>
in the Parque Podocarpus,
<quantity id="4CE66E991351532CFADAFD88FAF0FD0F" box="[1323,1398,626,652]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="26" unit="km" value="70.0">70 km</quantity>
northwest from the
<typeStatus id="54A57DDE1351532CFE5AFD6FFE58FD2C" box="[427,478,661,687]" pageId="25" pageNumber="26">type</typeStatus>
locality of this new species.
<taxonomicName id="4C1EB8FF1351532CFCCEFD6FFBFEFD2C" authority="Moret" authorityName="Moret" authorityYear="2020" box="[831,1144,661,687]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rugitarsis" status="sp. nov.">
<emphasis id="B96A1F6E1351532CFCCEFD6FFBA0FD2C" box="[831,1062,661,687]" italics="true" pageId="25" pageNumber="26">Dyscolus rugitarsis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151351532CFB70FD6FFB58FD2C" box="[1153,1246,661,687]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
differs from the latter by the blunter hind angles of the pronotum, the presence of at least one seta in the basal half of the third elytral interval (third interval asetose in
<taxonomicName id="4C1EB8FF1351532CFCD4FD26FC4BFD76" authorityName="Perrault" authorityYear="1993" box="[805,973,732,757]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="moretianus">
<emphasis id="B96A1F6E1351532CFCD4FD26FC4BFD76" box="[805,973,732,757]" italics="true" pageId="25" pageNumber="26">D. moretianus</emphasis>
</taxonomicName>
), slightly longer antennae and tarsi, and a markedly longer external lobe at the apex of the fourth metatarsomere. For comparison with
<taxonomicName id="4C1EB8FF1351532CFF4CFCD9FE23FCBF" authority="Moret" authorityName="Moret" authorityYear="2020" box="[189,421,802,828]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sulcipedis" status="sp. nov.">
<emphasis id="B96A1F6E1351532CFF4CFCD9FED0FCBF" box="[189,342,802,828]" italics="true" pageId="25" pageNumber="26">D. sulcipedis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151351532CFE5CFCD8FD8CFCBF" box="[429,522,802,828]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
, see below (p. 27).
</paragraph>
<paragraph id="8BA1C37C1351532CFF4CFC99FE9BFCFE" blockId="25.[189,285,867,893]" box="[189,285,867,893]" pageId="25" pageNumber="26">
<heading id="D0E974101351532CFF4CFC99FE9BFCFE" bold="true" box="[189,285,867,893]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B96A1F6E1351532CFF4CFC99FE9BFCFE" bold="true" box="[189,285,867,893]" pageId="25" pageNumber="26">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1351532CFF4CFC68FEFFFC4C" blockId="25.[189,1399,913,975]" pageId="25" pageNumber="26">
Upper montane forest, at around
<quantity id="4CE66E991351532CFDB3FC6BFD62FC2F" box="[578,740,913,940]" metricMagnitude="3" metricUnit="m" metricValue="3.225" metricValueMax="3.25" metricValueMin="3.2" pageId="25" pageNumber="26" unit="m" value="3225.0" valueMax="3250.0" valueMin="3200.0">3200–3250 m</quantity>
a.s.l. Active at the beginning of the night on the surface of the leaf litter.
</paragraph>
<paragraph id="8BA1C37C1351532CFF4CFC0CFE6DFB93" blockId="25.[189,491,1014,1040]" box="[189,491,1014,1040]" pageId="25" pageNumber="26">
<heading id="D0E974101351532CFF4CFC0CFE6DFB93" bold="true" box="[189,491,1014,1040]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B96A1F6E1351532CFF4CFC0CFE6DFB93" bold="true" box="[189,491,1014,1040]" pageId="25" pageNumber="26">Geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1351532CFF4CFBDFFEEBFBE1" blockId="25.[189,1399,1061,1122]" pageId="25" pageNumber="26">
Only known from the
<typeStatus id="54A57DDE1351532CFE15FBDFFD91FBBC" box="[484,535,1061,1087]" pageId="25" pageNumber="26">type</typeStatus>
locality in Southern
<collectingCountry id="F30983EC1351532CFCDEFBDFFC15FBBC" box="[815,915,1061,1087]" name="Ecuador" pageId="25" pageNumber="26">Ecuador</collectingCountry>
, in Parque Nacional Yacuri. Probably microendemic.
</paragraph>
</subSubSection>
</treatment>
</document>
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